144 lines
16 KiB
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144 lines
16 KiB
XML
<document id="A6A71088CFD7BA87202196CC107394CA" ID-DOI="10.1111/j.1096-3642.2011.00771.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5406728" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630346404524" checkinUser="marcus" docAuthor="Archibald, J. David & Averianov, Alexander" docDate="2012" docId="AB4D878FFF98612F51E5FF18C376FA35" docLanguage="en" docName="j.1096-3642.2011.00771.x.pdf" docOrigin="Zoological Journal of the Linnean Society 164 (2)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2011.00771.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Gallolestes pachymandibularis : Lillegraven 1976" docType="treatment" docVersion="6" lastPageNumber="405" masterDocId="5774FFF7FFB461035112FFDEC247FFF9" masterDocTitle="Phylogenetic analysis, taxonomic revision, and dental ontogeny of the Cretaceous Zhelestidae (Mammalia: Eutheria)" masterLastPageNumber="426" masterPageNumber="361" pageNumber="405" updateTime="1699238079856" updateUser="plazi">
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<mods:title id="B99B7F9D674D26EAE010BC70D4C7DC92">Phylogenetic analysis, taxonomic revision, and dental ontogeny of the Cretaceous Zhelestidae (Mammalia: Eutheria)</mods:title>
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<mods:namePart id="6FCD72558B9D5A2AC728CDD8EDCF490C">Archibald, J. David</mods:namePart>
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<mods:namePart id="B7502AE658FCF517787FA852D11FB06B">Averianov, Alexander</mods:namePart>
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<mods:title id="D070FB4B2D7E822B160948E13E3E7F27">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="EA4391CFA25EB2D3D3A72D4631E3B19E">2012</mods:date>
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<mods:number id="2EB79D2B23D1A77AF5FFCB8DE3AEA021">164</mods:number>
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<mods:identifier id="0EAFA26A713713BF412F34DE02FBB8CD" type="DOI">10.1111/j.1096-3642.2011.00771.x</mods:identifier>
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<mods:identifier id="1ECC543D7C4EAA57180E178248BA5AB9" type="ISSN">0024-4082</mods:identifier>
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<treatment id="AB4D878FFF98612F51E5FF18C376FA35" ID-DOI="http://doi.org/10.5281/zenodo.5479736" ID-Zenodo-Dep="5479736" LSID="urn:lsid:plazi:treatment:AB4D878FFF98612F51E5FF18C376FA35" httpUri="http://treatment.plazi.org/id/AB4D878FFF98612F51E5FF18C376FA35" lastPageNumber="405" pageId="44" pageNumber="405">
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<subSubSection id="6BFE6512FF98612F51E5FF18C3B0FEB8" pageId="44" pageNumber="405" type="nomenclature">
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<paragraph id="235B3699FF98612F51E5FF18C07EFF05" blockId="44.[247,655,198,252]" pageId="44" pageNumber="405">
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<heading id="781381F5FF98612F51E5FF18C07EFF05" centered="true" fontSize="8" level="2" pageId="44" pageNumber="405" reason="8">
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<taxonomicName id="E4E44D1AFF98612F51E5FF18C071FF05" authority="LILLEGRAVEN, 1976" authorityName=": Lillegraven" authorityYear="1976" class="Mammalia" family="Didymoconidae" genus="Gallolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pilosa" pageId="44" pageNumber="404" phylum="Chordata" rank="species" species="pachymandibularis">
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<emphasis id="1190EA8BFF98612F51E5FF18C0C8FF23" box="[247,655,198,219]" italics="true" pageId="44" pageNumber="405">GALLOLESTES PACHYMANDIBULARIS</emphasis>
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<bibRefCitation id="47754B68FF98612F505FFF38C071FF05" author="Lillegraven JA" box="[333,566,230,252]" pageId="44" pageNumber="405" pagination="437 - 443" refId="ref32245" refString="Lillegraven JA. 1976. A new genus of therian mammal from the Late Cretaceous ' El Gallo Formation, ' Baja California, Mexico. Journal of Paleontology 50: 437 - 443." type="journal article" year="1976">LILLEGRAVEN, 1976</bibRefCitation>
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</taxonomicName>
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.
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</heading>
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</paragraph>
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<paragraph id="235B3699FF98612F5182FED2C3B0FEB8" blockId="44.[144,758,268,321]" pageId="44" pageNumber="405">(See Appendix 4 for synonymies, referred illustrations, and referred specimens.)</paragraph>
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</subSubSection>
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<subSubSection id="6BFE6512FF98612F5182FEB6C376FA35" pageId="44" pageNumber="405" type="materials_examined">
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<paragraph id="235B3699FF98612F5182FEB6C397FE65" blockId="44.[144,758,360,413]" pageId="44" pageNumber="405">
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<materialsCitation id="938C3CC4FF98612F5182FEB6C397FE65" collectionCode="LACM" pageId="44" pageNumber="405" specimenCode="LACM 42633" specimenCount="1" typeStatus="holotype">
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<emphasis id="1190EA8BFF98612F5182FEB6C2BDFE84" box="[144,250,360,381]" italics="true" pageId="44" pageNumber="405">
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<typeStatus id="FC5F883BFF98612F5182FEB6C2B3FE84" box="[144,244,360,381]" pageId="44" pageNumber="405" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<specimenCode id="73429EE2FF98612F501EFEB6C3F7FE87" box="[268,432,360,382]" collectionCode="LACM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="44" pageNumber="405" type="Museum">LACM 42633</specimenCode>
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, right dentary with fragmented p4, dp5, and m1-3.
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</materialsCitation>
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</paragraph>
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<paragraph id="235B3699FF98612F5182FE1BC060FDEE" blockId="44.[144,758,452,535]" pageId="44" pageNumber="405">
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<emphasis id="1190EA8BFF98612F5182FE1BC3FAFE20" box="[144,445,452,474]" italics="true" pageId="44" pageNumber="405">Type locality and horizon:</emphasis>
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<materialsCitation id="938C3CC4FF98612F50DAFE1AC064FDEE" collectionCode="LACM" collectorName="El Gallo Formation'" country="Mexico" location="Baja California" pageId="44" pageNumber="405" specimenCount="1" stateProvince="Baja California">
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<collectionCode id="45F5AE5CFF98612F50DAFE1AC054FE20" box="[456,531,452,473]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" lsid="urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="44" pageNumber="405" type="Museum">LACM</collectionCode>
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locality 3302,
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<collectingRegion id="E120F87BFF98612F53D0FE1AC344FE00" country="Mexico" name="Baja California" pageId="44" pageNumber="405">Baja California</collectingRegion>
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del Norte,
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<collectingCountry id="5BF37609FF98612F5085FE3DC3ADFE00" box="[407,490,483,505]" name="Mexico" pageId="44" pageNumber="405">Mexico</collectingCountry>
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. ‘
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<collectorName id="8E11534FFF98612F5312FE3DC0B4FE00" box="[512,755,483,505]" pageId="44" pageNumber="405">El Gallo Formation’</collectorName>
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, Late Cretaceous (late Campanian)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="235B3699FF98612F5182FDE0C39DFD8B" blockId="44.[144,758,574,627]" pageId="44" pageNumber="405">
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<emphasis id="1190EA8BFF98612F5182FDE0C327FDAA" box="[144,352,574,595]" italics="true" pageId="44" pageNumber="405">Revised diagnosis:</emphasis>
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About 40% larger than
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<taxonomicName id="E4E44D1AFF98612F5361FDE1C2E9FD8A" authorityName=": Cifelli" authorityYear="1994" class="Mammalia" family="Didymoconidae" genus="Gallolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pilosa" pageId="44" pageNumber="405" phylum="Chordata" rank="species" species="agujaensis">
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<emphasis id="1190EA8BFF98612F5361FDE1C2E9FD8A" italics="true" pageId="44" pageNumber="405">G. agujaensis</emphasis>
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</taxonomicName>
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in dental measurements.
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</paragraph>
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<paragraph id="235B3699FF98612F5182FD45C3C3FD36" blockId="44.[144,759,666,719]" pageId="44" pageNumber="405">
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<emphasis id="1190EA8BFF98612F5182FD45C35FFD49" box="[144,280,667,688]" italics="true" pageId="44" pageNumber="405">Description:</emphasis>
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See
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<bibRefCitation id="47754B68FF98612F5077FD44C0FEFD49" author="Lillegraven JA" box="[357,697,666,688]" pageId="44" pageNumber="405" pagination="437 - 443" refId="ref32245" refString="Lillegraven JA. 1976. A new genus of therian mammal from the Late Cretaceous ' El Gallo Formation, ' Baja California, Mexico. Journal of Paleontology 50: 437 - 443." type="journal article" year="1976">Lillegraven (1976: 438–440)</bibRefCitation>
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and
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<bibRefCitation id="47754B68FF98612F5182FD67C3C7FD36" author="Clemens WA" box="[144,384,697,719]" pageId="44" pageNumber="405" pagination="1 - 10" refId="ref31771" refString="Clemens WA. 1980. Gallolestes pachymandibularis (Mammalia: Theria, incertae sedis) from Late Cretaceous deposits in Baja California del Norte, Mexico. PaleoBios 33: 1 - 10." type="journal article" year="1980">Clemens (1980: 2–3)</bibRefCitation>
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.
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</paragraph>
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<paragraph id="235B3699FF98612F5182FD29C0E6FCF5" blockId="44.[144,673,758,780]" box="[144,673,758,780]" pageId="44" pageNumber="405">
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<emphasis id="1190EA8BFF98612F5182FD29C37DFCF5" box="[144,314,759,780]" italics="true" pageId="44" pageNumber="405">Measurements:</emphasis>
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See
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<bibRefCitation id="47754B68FF98612F5064FD28C003FCF5" author="Lillegraven JA" box="[374,580,758,780]" pageId="44" pageNumber="405" pagination="437 - 443" refId="ref32245" refString="Lillegraven JA. 1976. A new genus of therian mammal from the Late Cretaceous ' El Gallo Formation, ' Baja California, Mexico. Journal of Paleontology 50: 437 - 443." type="journal article" year="1976">Lillegraven (1976</bibRefCitation>
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: tab. 1).
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</paragraph>
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<paragraph id="235B3699FF98612F5182FCEDC376FA35" blockId="44.[144,759,819,1484]" pageId="44" pageNumber="405">
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<emphasis id="1190EA8BFF98612F5182FCEDC349FCB1" box="[144,270,819,840]" italics="true" pageId="44" pageNumber="405">Comments:</emphasis>
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We follow
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<bibRefCitation id="47754B68FF98612F5098FCEDC05CFCB1" author="Butler PM" box="[394,539,819,841]" pageId="44" pageNumber="405" pagination="241 - 269" refId="ref31561" refString="Butler PM. 1977. Evolutionary radiation of the cheek teeth of Cretaceous placentals. Acta Palaeontologica Polonica 22: 241 - 269." type="journal article" year="1977">Butler (1977</bibRefCitation>
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,
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<bibRefCitation id="47754B68FF98612F5335FCEDC019FCB0" author="Butler PM" box="[551,606,819,841]" pageId="44" pageNumber="405" pagination="529 - 552" refId="ref31613" refString="Butler PM. 1990. Early trends in the evolution of tribosphenic molars. Biological Reviews 65: 529 - 552." type="journal article" year="1990">1990</bibRefCitation>
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) in the interpretation of the last premolar on the
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<typeStatus id="FC5F883BFF98612F5360FC8CC094FC91" box="[626,723,850,872]" pageId="44" pageNumber="405" type="holotype">holotype</typeStatus>
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of
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<taxonomicName id="E4E44D1AFF98612F5182FCAFC3C9FC7C" authorityName=": Lillegraven" authorityYear="1976" box="[144,398,880,902]" class="Mammalia" family="Didymoconidae" genus="Gallolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pilosa" pageId="44" pageNumber="405" phylum="Chordata" rank="species" species="pachymandibularis">
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<emphasis id="1190EA8BFF98612F5182FCAFC3C9FC7C" box="[144,398,880,902]" italics="true" pageId="44" pageNumber="405">G. pachymandibularis</emphasis>
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</taxonomicName>
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as a milk tooth, dp5, contra our previous claim that this tooth is a molariform p5 (Archibald & Averianov, 2001; see also discussion in
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<bibRefCitation id="47754B68FF98612F5182FC12C31CFC1B" author="Lillegraven JA" box="[144,347,972,994]" pageId="44" pageNumber="405" pagination="437 - 443" refId="ref32245" refString="Lillegraven JA. 1976. A new genus of therian mammal from the Late Cretaceous ' El Gallo Formation, ' Baja California, Mexico. Journal of Paleontology 50: 437 - 443." type="journal article" year="1976">Lillegraven, 1976</bibRefCitation>
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;
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<bibRefCitation id="47754B68FF98612F507BFC12C052FC18" author="Clemens WA" box="[361,533,972,994]" pageId="44" pageNumber="405" pagination="1 - 10" refId="ref31771" refString="Clemens WA. 1980. Gallolestes pachymandibularis (Mammalia: Theria, incertae sedis) from Late Cretaceous deposits in Baja California del Norte, Mexico. PaleoBios 33: 1 - 10." type="journal article" year="1980">Clemens, 1980</bibRefCitation>
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;
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<bibRefCitation id="47754B68FF98612F5330FC13C0B5FC18" author="Nesov LA & Archibald JD & Kielan-Jaworowska Z" box="[546,754,972,994]" pageId="44" pageNumber="405" pagination="40 - 88" refId="ref32857" refString="Nesov LA, Archibald JD, Kielan-Jaworowska Z. 1998. Ungulate-like mammals from the Late Cretaceous of Uzbekistan and a phylogenetic analysis of Ungulatomorpha. In: Beard KC, Dawson MR, eds. Dawn of the Age of Mammals in Asia. Bulletin of the Carnegie Museum of Natural History 34: 40 - 88." type="journal article" year="1998">
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Nesov
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<emphasis id="1190EA8BFF98612F5363FC13C0E6FC18" box="[625,673,972,994]" italics="true" pageId="44" pageNumber="405">et al</emphasis>
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., 1998
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</bibRefCitation>
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;
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<bibRefCitation id="47754B68FF98612F5182FC35C019FBF8" author="Kielan-Jaworowska Z & Cifelli RL & Luo Z-X" box="[144,606,1003,1025]" pageId="44" pageNumber="405" refId="ref32044" refString="Kielan-Jaworowska Z, Cifelli RL, Luo Z-X. 2004. Mammals from the age of dinosaurs: origins, evolution, and structure. New York: Columbia University Press." type="book" year="2004">Kielan-Jaworowska, Cifelli & Luo, 2004</bibRefCitation>
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). The deciduous nature of this tooth was rejected on the ground that ‘the roots are well formed, very unusual for a deciduous tooth that was soon destined to be shed’ (Archibald & Averianov, 2001: 543). The roots of this tooth, however, actually are formed as well as in some dp5s of zhelestids from Dzharakuduk having the same state of wear. The smaller size relative to m1 and the much heavier wear strongly argue that this is a milk tooth. As was pointed out by
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<bibRefCitation id="47754B68FF98612F532EFB20C0B5FAED" author="Clemens WA" box="[572,754,1278,1300]" pageId="44" pageNumber="405" pagination="1 - 10" refId="ref31771" refString="Clemens WA. 1980. Gallolestes pachymandibularis (Mammalia: Theria, incertae sedis) from Late Cretaceous deposits in Baja California del Norte, Mexico. PaleoBios 33: 1 - 10." type="journal article" year="1980">Clemens (1980)</bibRefCitation>
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, retention of the last deciduous premolar until the m3 has erupted is quite unusual. In the Asiatic Zhelestinae dp5 is replaced when m3 is not fully erupted (see the next section). A similar delay in replacement of dp5 was noted for the zalambdalestid
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<taxonomicName id="E4E44D1AFF98612F535FFA49C095FA55" authorityName="Kielan-Jaworowska" authorityYear="1975" box="[589,722,1431,1452]" class="Mammalia" family="Zalambdalestidae" genus="Barunlestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pilosa" pageId="44" pageNumber="405" phylum="Chordata" rank="genus">
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<emphasis id="1190EA8BFF98612F535FFA49C095FA55" box="[589,722,1431,1452]" italics="true" pageId="44" pageNumber="405">Barunlestes</emphasis>
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</taxonomicName>
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by
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<bibRefCitation id="47754B68FF98612F5182FA68C36AFA35" author="Butler PM" box="[144,301,1462,1484]" pageId="44" pageNumber="405" pagination="529 - 552" refId="ref31613" refString="Butler PM. 1990. Early trends in the evolution of tribosphenic molars. Biological Reviews 65: 529 - 552." type="journal article" year="1990">Butler (1990)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |