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<document id="4EC8016CED8C34C897D403AEB86DF1A0" ID-CLB-Dataset="28120" ID-DOI="10.11646/zootaxa.4568.3.5" ID-GBIF-Dataset="2d4a9aef-6b65-4af8-9d92-32924bf6827f" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601710" ID-ZooBank="AD20F4C5-9E81-4B0C-A7F3-294B0FD06C04" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553173036048" checkinUser="plazi" docAuthor="Silva, Alessandra C. G. &amp; Grossi, Paschoal C." docDate="2019" docId="AB2F87FAFFA8FFCBFF57148E85C9F9C2" docLanguage="en" docName="zootaxa.4568.3.5.pdf" docOrigin="Zootaxa 4568 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Casignetus humboldti" docType="treatment" docVersion="7" lastPageNumber="504" masterDocId="5716FF82FFABFFC8FFC01601817BFFC5" masterDocTitle="Immature stages of the genus Casignetus MacLeay, 1819 (Coleoptera: Lucanidae: Lucaninae)" masterLastPageNumber="519" masterPageNumber="501" pageNumber="504" updateTime="1698703561375" updateUser="ExternalLinkService">
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<mods:title id="167FB9A91E74FC82CF8D8C36E1260E70">Immature stages of the genus Casignetus MacLeay, 1819 (Coleoptera: Lucanidae: Lucaninae)</mods:title>
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<mods:namePart id="D759145EF48F59A48C43CEFF07A93127">Silva, Alessandra C. G.</mods:namePart>
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<treatment id="AB2F87FAFFA8FFCBFF57148E85C9F9C2" ID-DOI="http://doi.org/10.5281/zenodo.5941846" ID-GBIF-Taxon="154475028" ID-Zenodo-Dep="5941846" LSID="urn:lsid:plazi:treatment:AB2F87FAFFA8FFCBFF57148E85C9F9C2" httpUri="http://treatment.plazi.org/id/AB2F87FAFFA8FFCBFF57148E85C9F9C2" lastPageNumber="504" pageId="3" pageNumber="504">
<subSubSection id="6B9C6567FFA8FFCBFF57148E8050FB06" pageId="3" pageNumber="504" type="nomenclature">
<paragraph id="233936ECFFA8FFCBFF57148E8306FD6F" blockId="3.[151,637,655,682]" box="[151,637,655,682]" pageId="3" pageNumber="504">
<heading id="78718180FFA8FFCBFF57148E8306FD6F" bold="true" box="[151,637,655,682]" fontSize="11" level="1" pageId="3" pageNumber="504" reason="1">
<taxonomicName id="E4864D6FFFA8FFCBFF57148E8306FD6F" ID-CoL="5XD42" authority="(Gyllenhal, 1817)" baseAuthorityName="Gyllenhal" baseAuthorityYear="1817" box="[151,637,655,682]" class="Insecta" family="Lucanidae" genus="Casignetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="humboldti">
<emphasis id="11F2EAFEFFA8FFCBFF57148E8306FD6F" bold="true" box="[151,637,655,682]" pageId="3" pageNumber="504">
<emphasis id="11F2EAFEFFA8FFCBFF57148E80DBFD6C" bold="true" box="[151,416,655,681]" italics="true" pageId="3" pageNumber="504">Casignetus humboldti</emphasis>
(Gyllenhal, 1817)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="233936ECFFA8FFCBFF5714D78050FB06" blockId="3.[151,1437,726,1543]" pageId="3" pageNumber="504">
<emphasis id="11F2EAFEFFA8FFCBFF5714D78311FD2A" bold="true" box="[151,618,726,751]" pageId="3" pageNumber="504">
Larvae description (
<figureCitation id="BBBD2A69FFA8FFCBFE4814D78369FD2A" box="[392,530,726,751]" captionStart="FIGURES 6473" captionStartId="13.[151,264,1933,1956]" captionTargetBox="[201,1385,232,1890]" captionTargetId="figure@13.[181,1405,193,1921]" captionTargetPageId="13" captionText="FIGURES 6473. Casignetus humboldti larvae. 64, Labrum; 65, epipharynx; 6667, right mandible (ventral, dorsal); 6869, left mandible (dorsal, ventral); 70, maxillae and labium (ventral); 71, maxillae and hypopharynx (dorsal); 72, mesothoracic leg (external); 73, metathoracic leg (internal). Scale bars = 1 mm. Photographs: M. Cherman." httpUri="https://zenodo.org/record/2601728/files/figure.png" pageId="3" pageNumber="504">Figs. 6473</figureCitation>
). Body.
</emphasis>
Larvae
<specimenCount id="3580FD65FFA8FFCBFD0614D683A0FD2B" box="[710,731,727,750]" pageId="3" pageNumber="504" type="female"></specimenCount>
, total length 21.98 mm.
<emphasis id="11F2EAFEFFA8FFCBFC2E14D78556FD2A" bold="true" box="[1006,1069,726,751]" pageId="3" pageNumber="504">Head</emphasis>
. Cephalic capsule (
<figureCitation id="BBBD2A69FFA8FFCBFACA14D78420FD2A" box="[1290,1371,726,751]" captionStart="FIGURES 1018" captionStartId="7.[151,264,1943,1966]" captionTargetBox="[191,1384,249,1841]" captionTargetId="figure@7.[181,1409,193,1926]" captionTargetPageId="7" captionText="FIGURES 1018. Head capsule (frontal), labrum and epipharynx of Casignetus larvae. 10, Casignetus humboldti; 11, C. spixi; 12, C. lindei; 13, labrum of C. humboldti; 14, labrum of C. spixi; 15, labrum of C. lindei; 16, epipharynx of C. humboldti; 17, epipharynx of C. spixi; 18, epipharynx of C. lindei. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601716/files/figure.png" pageId="3" pageNumber="504">Fig. 10</figureCitation>
) with 6.5 mm from apex of mandibles until posterior margin,
<quantity id="E47E9B09FFA8FFCBFCD514FB822CFCD7" box="[789,855,762,787]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="504" unit="mm" value="5.0">5 mm</quantity>
of wide; 1 lateroepicranial seta on each side. Each side of frons with 2 externofrontal setae. Base of clypeus (
<figureCitation id="BBBD2A69FFA8FFCBFC8A151F82D7FCF2" box="[842,940,798,823]" captionStart="FIGURES 1018" captionStartId="7.[151,264,1943,1966]" captionTargetBox="[191,1384,249,1841]" captionTargetId="figure@7.[181,1409,193,1926]" captionTargetPageId="7" captionText="FIGURES 1018. Head capsule (frontal), labrum and epipharynx of Casignetus larvae. 10, Casignetus humboldti; 11, C. spixi; 12, C. lindei; 13, labrum of C. humboldti; 14, labrum of C. spixi; 15, labrum of C. lindei; 16, epipharynx of C. humboldti; 17, epipharynx of C. spixi; 18, epipharynx of C. lindei. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601716/files/figure.png" pageId="3" pageNumber="504">Figs. 13</figureCitation>
,
<figureCitation id="BBBD2A69FFA8FFCBFC7C151F82A2FCF2" box="[956,985,798,823]" captionStart="FIGURES 6473" captionStartId="13.[151,264,1933,1956]" captionTargetBox="[201,1385,232,1890]" captionTargetId="figure@13.[181,1405,193,1921]" captionTargetPageId="13" captionText="FIGURES 6473. Casignetus humboldti larvae. 64, Labrum; 65, epipharynx; 6667, right mandible (ventral, dorsal); 6869, left mandible (dorsal, ventral); 70, maxillae and labium (ventral); 71, maxillae and hypopharynx (dorsal); 72, mesothoracic leg (external); 73, metathoracic leg (internal). Scale bars = 1 mm. Photographs: M. Cherman." httpUri="https://zenodo.org/record/2601728/files/figure.png" pageId="3" pageNumber="504">64</figureCitation>
) measuring about 2.5 times its length. Posterolabral seta located on left side.
<emphasis id="11F2EAFEFFA8FFCBFD84154283DDFC9E" box="[580,678,835,859]" italics="true" pageId="3" pageNumber="504">Corypha</emphasis>
(
<figureCitation id="BBBD2A69FFA8FFCBFD7615438268FC9E" box="[694,787,834,859]" captionStart="FIGURES 1018" captionStartId="7.[151,264,1943,1966]" captionTargetBox="[191,1384,249,1841]" captionTargetId="figure@7.[181,1409,193,1926]" captionTargetPageId="7" captionText="FIGURES 1018. Head capsule (frontal), labrum and epipharynx of Casignetus larvae. 10, Casignetus humboldti; 11, C. spixi; 12, C. lindei; 13, labrum of C. humboldti; 14, labrum of C. spixi; 15, labrum of C. lindei; 16, epipharynx of C. humboldti; 17, epipharynx of C. spixi; 18, epipharynx of C. lindei. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601716/files/figure.png" pageId="3" pageNumber="504">Figs. 16</figureCitation>
,
<figureCitation id="BBBD2A69FFA8FFCBFCDE15438240FC9E" box="[798,827,834,859]" captionStart="FIGURES 6473" captionStartId="13.[151,264,1933,1956]" captionTargetBox="[201,1385,232,1890]" captionTargetId="figure@13.[181,1405,193,1921]" captionTargetPageId="13" captionText="FIGURES 6473. Casignetus humboldti larvae. 64, Labrum; 65, epipharynx; 6667, right mandible (ventral, dorsal); 6869, left mandible (dorsal, ventral); 70, maxillae and labium (ventral); 71, maxillae and hypopharynx (dorsal); 72, mesothoracic leg (external); 73, metathoracic leg (internal). Scale bars = 1 mm. Photographs: M. Cherman." httpUri="https://zenodo.org/record/2601728/files/figure.png" pageId="3" pageNumber="504">65</figureCitation>
) with 9 setae,
<emphasis id="11F2EAFEFFA8FFCBFC1D15428510FC9E" box="[989,1131,835,859]" italics="true" pageId="3" pageNumber="504">haptomerum</emphasis>
with 8 sensillae,
<emphasis id="11F2EAFEFFA8FFCBFAED154284E7FC9F" box="[1325,1436,835,858]" italics="true" pageId="3" pageNumber="504">acroparia</emphasis>
with 6 setae,
<emphasis id="11F2EAFEFFA8FFCBFEF3156780B1FCBB" box="[307,458,870,894]" italics="true" pageId="3" pageNumber="504">achantoparia</emphasis>
with 6 setae on each side, right
<emphasis id="11F2EAFEFFA8FFCBFC89156782ABFCBB" box="[841,976,870,894]" italics="true" pageId="3" pageNumber="504">chaetoparia</emphasis>
with 48 setae, left
<emphasis id="11F2EAFEFFA8FFCBFB7315678441FCBB" box="[1203,1338,870,894]" italics="true" pageId="3" pageNumber="504">chaetoparia</emphasis>
with 39 setae,
<emphasis id="11F2EAFEFFA8FFCBFF1F158A801AFC66" box="[223,353,907,931]" italics="true" pageId="3" pageNumber="504">protophoba</emphasis>
with 4 setae and 10 sensillae. Uncus of galea (
<figureCitation id="BBBD2A69FFA8FFCBFCBF158B82A4FC66" box="[895,991,906,931]" captionStart="FIGURES 3139" captionStartId="9.[151,264,1936,1959]" captionTargetBox="[226,1390,219,1898]" captionTargetId="figure@9.[182,1411,193,1920]" captionTargetPageId="9" captionText="FIGURES 3139. Mouthparts of Casignetus larvae. 3133, Casignetus humboldti labium, hypopharynx, and left maxilla (dorsal), respectively; 3436, C. spixi labium, hypopharynx, and left maxilla (dorsal), respectively; 3739, C. lindei labium, hypopharynx, and left maxilla (dorsal), respectively. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601720/files/figure.png" pageId="3" pageNumber="504">Figs. 33</figureCitation>
,
<figureCitation id="BBBD2A69FFA8FFCBFC2C158A854FFC66" box="[1004,1076,907,931]" captionStart="FIGURES 6473" captionStartId="13.[151,264,1933,1956]" captionTargetBox="[201,1385,232,1890]" captionTargetId="figure@13.[181,1405,193,1921]" captionTargetPageId="13" captionText="FIGURES 6473. Casignetus humboldti larvae. 64, Labrum; 65, epipharynx; 6667, right mandible (ventral, dorsal); 6869, left mandible (dorsal, ventral); 70, maxillae and labium (ventral); 71, maxillae and hypopharynx (dorsal); 72, mesothoracic leg (external); 73, metathoracic leg (internal). Scale bars = 1 mm. Photographs: M. Cherman." httpUri="https://zenodo.org/record/2601728/files/figure.png" pageId="3" pageNumber="504">7071</figureCitation>
) surrounded dorsally by 6 long setae; galea with a row of 11 long setae on inner ventral margin. Lacinia with 32 setae on inner lateral margin. Hypopharynx (
<figureCitation id="BBBD2A69FFA8FFCBFE8015D380E6FC2E" box="[320,413,978,1003]" captionStart="FIGURES 3139" captionStartId="9.[151,264,1936,1959]" captionTargetBox="[226,1390,219,1898]" captionTargetId="figure@9.[182,1411,193,1920]" captionTargetPageId="9" captionText="FIGURES 3139. Mouthparts of Casignetus larvae. 3133, Casignetus humboldti labium, hypopharynx, and left maxilla (dorsal), respectively; 3436, C. spixi labium, hypopharynx, and left maxilla (dorsal), respectively; 3739, C. lindei labium, hypopharynx, and left maxilla (dorsal), respectively. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601720/files/figure.png" pageId="3" pageNumber="504">Figs. 32</figureCitation>
,
<figureCitation id="BBBD2A69FFA8FFCBFE6815D280BEFC2E" box="[424,453,979,1003]" captionStart="FIGURES 6473" captionStartId="13.[151,264,1933,1956]" captionTargetBox="[201,1385,232,1890]" captionTargetId="figure@13.[181,1405,193,1921]" captionTargetPageId="13" captionText="FIGURES 6473. Casignetus humboldti larvae. 64, Labrum; 65, epipharynx; 6667, right mandible (ventral, dorsal); 6869, left mandible (dorsal, ventral); 70, maxillae and labium (ventral); 71, maxillae and hypopharynx (dorsal); 72, mesothoracic leg (external); 73, metathoracic leg (internal). Scale bars = 1 mm. Photographs: M. Cherman." httpUri="https://zenodo.org/record/2601728/files/figure.png" pageId="3" pageNumber="504">71</figureCitation>
) with 55 setae. Labium Figs. (31, 70) with 1 posterior seta on each side.
<emphasis id="11F2EAFEFFA8FFCBFB3115D38452FC2E" bold="true" box="[1265,1321,978,1003]" pageId="3" pageNumber="504">Legs</emphasis>
. Posterior face of mesocoxa (
<figureCitation id="BBBD2A69FFA8FFCBFEAF15F780B5FBCA" box="[367,462,1014,1039]" captionStart="FIGURES 4051" captionStartId="10.[151,264,1882,1905]" captionTargetBox="[220,1373,203,1847]" captionTargetId="figure@10.[200,1392,193,1867]" captionTargetPageId="10" captionText="FIGURES 4051. FIGURES 4045. Stridulatory organs Casignetus larvae. 40, Pars stridens of C. humboldti; 41, pars stridens of C. spixi; 42, pars stridens of C. lindei; 43, plectrum of C. humboldti; 44, plectrum of C. spixi; 45, plectrum of C. lindei. Scale bars = 1 mm. FIGURES 4651. Last abdominal segment of Casignetus larvae. 46, Raster of C. humboldti; 47, raster of C. spixi; 48, raster of C. lindei; 49, anal lobes (posterior) of C. humboldti; 50, anal lobes (posterior) of C. spixi; 51, anal lobes (posterior) of C. lindei (dilated anal region due to the presence of excrement at death). Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601722/files/figure.png" pageId="3" pageNumber="504">Figs. 40</figureCitation>
,
<figureCitation id="BBBD2A69FFA8FFCBFE1C15F68081FBCA" box="[476,506,1015,1039]" captionStart="FIGURES 6473" captionStartId="13.[151,264,1933,1956]" captionTargetBox="[201,1385,232,1890]" captionTargetId="figure@13.[181,1405,193,1921]" captionTargetPageId="13" captionText="FIGURES 6473. Casignetus humboldti larvae. 64, Labrum; 65, epipharynx; 6667, right mandible (ventral, dorsal); 6869, left mandible (dorsal, ventral); 70, maxillae and labium (ventral); 71, maxillae and hypopharynx (dorsal); 72, mesothoracic leg (external); 73, metathoracic leg (internal). Scale bars = 1 mm. Photographs: M. Cherman." httpUri="https://zenodo.org/record/2601728/files/figure.png" pageId="3" pageNumber="504">72</figureCitation>
) strongly excavated. Pars stridens (
<figureCitation id="BBBD2A69FFA8FFCBFC4F15F78295FBCA" box="[911,1006,1014,1039]" captionStart="FIGURES 4051" captionStartId="10.[151,264,1882,1905]" captionTargetBox="[220,1373,203,1847]" captionTargetId="figure@10.[200,1392,193,1867]" captionTargetPageId="10" captionText="FIGURES 4051. FIGURES 4045. Stridulatory organs Casignetus larvae. 40, Pars stridens of C. humboldti; 41, pars stridens of C. spixi; 42, pars stridens of C. lindei; 43, plectrum of C. humboldti; 44, plectrum of C. spixi; 45, plectrum of C. lindei. Scale bars = 1 mm. FIGURES 4651. Last abdominal segment of Casignetus larvae. 46, Raster of C. humboldti; 47, raster of C. spixi; 48, raster of C. lindei; 49, anal lobes (posterior) of C. humboldti; 50, anal lobes (posterior) of C. spixi; 51, anal lobes (posterior) of C. lindei (dilated anal region due to the presence of excrement at death). Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601722/files/figure.png" pageId="3" pageNumber="504">Figs. 40</figureCitation>
,
<figureCitation id="BBBD2A69FFA8FFCBFC3C15F68562FBCA" box="[1020,1049,1015,1039]" captionStart="FIGURES 6473" captionStartId="13.[151,264,1933,1956]" captionTargetBox="[201,1385,232,1890]" captionTargetId="figure@13.[181,1405,193,1921]" captionTargetPageId="13" captionText="FIGURES 6473. Casignetus humboldti larvae. 64, Labrum; 65, epipharynx; 6667, right mandible (ventral, dorsal); 6869, left mandible (dorsal, ventral); 70, maxillae and labium (ventral); 71, maxillae and hypopharynx (dorsal); 72, mesothoracic leg (external); 73, metathoracic leg (internal). Scale bars = 1 mm. Photographs: M. Cherman." httpUri="https://zenodo.org/record/2601728/files/figure.png" pageId="3" pageNumber="504">72</figureCitation>
) with 74 teeth; main row with 36 teeth; one adjacent row comprising 4 slightly apparent teeth on anteroproximal extremity, and 2 rows with 11 and 5 similar teeth, respectively, and a grainy area on posteroproximal extremity; distal extremity with about 18 teeth arranged irregularly. Plectrum (
<figureCitation id="BBBD2A69FFA8FFCBFDCB1263830BFBBE" box="[523,624,1122,1147]" captionStart="FIGURES 4051" captionStartId="10.[151,264,1882,1905]" captionTargetBox="[220,1373,203,1847]" captionTargetId="figure@10.[200,1392,193,1867]" captionTargetPageId="10" captionText="FIGURES 4051. FIGURES 4045. Stridulatory organs Casignetus larvae. 40, Pars stridens of C. humboldti; 41, pars stridens of C. spixi; 42, pars stridens of C. lindei; 43, plectrum of C. humboldti; 44, plectrum of C. spixi; 45, plectrum of C. lindei. Scale bars = 1 mm. FIGURES 4651. Last abdominal segment of Casignetus larvae. 46, Raster of C. humboldti; 47, raster of C. spixi; 48, raster of C. lindei; 49, anal lobes (posterior) of C. humboldti; 50, anal lobes (posterior) of C. spixi; 51, anal lobes (posterior) of C. lindei (dilated anal region due to the presence of excrement at death). Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601722/files/figure.png" pageId="3" pageNumber="504">Figs. 43</figureCitation>
,
<figureCitation id="BBBD2A69FFA8FFCBFD42126283E4FBBE" box="[642,671,1123,1147]" captionStart="FIGURES 6473" captionStartId="13.[151,264,1933,1956]" captionTargetBox="[201,1385,232,1890]" captionTargetId="figure@13.[181,1405,193,1921]" captionTargetPageId="13" captionText="FIGURES 6473. Casignetus humboldti larvae. 64, Labrum; 65, epipharynx; 6667, right mandible (ventral, dorsal); 6869, left mandible (dorsal, ventral); 70, maxillae and labium (ventral); 71, maxillae and hypopharynx (dorsal); 72, mesothoracic leg (external); 73, metathoracic leg (internal). Scale bars = 1 mm. Photographs: M. Cherman." httpUri="https://zenodo.org/record/2601728/files/figure.png" pageId="3" pageNumber="504">73</figureCitation>
) comprising 44 teeth. Posterodistal extremity of metatrochanter forming an angle of approximately 35°.
<emphasis id="11F2EAFEFFA8FFCBFD97128783B7FB5A" bold="true" box="[599,716,1158,1183]" pageId="3" pageNumber="504">Abdomen</emphasis>
. Raster (
<figureCitation id="BBBD2A69FFA8FFCBFCF0128782F9FB5A" box="[816,898,1158,1183]" captionStart="FIGURES 4051" captionStartId="10.[151,264,1882,1905]" captionTargetBox="[220,1373,203,1847]" captionTargetId="figure@10.[200,1392,193,1867]" captionTargetPageId="10" captionText="FIGURES 4051. FIGURES 4045. Stridulatory organs Casignetus larvae. 40, Pars stridens of C. humboldti; 41, pars stridens of C. spixi; 42, pars stridens of C. lindei; 43, plectrum of C. humboldti; 44, plectrum of C. spixi; 45, plectrum of C. lindei. Scale bars = 1 mm. FIGURES 4651. Last abdominal segment of Casignetus larvae. 46, Raster of C. humboldti; 47, raster of C. spixi; 48, raster of C. lindei; 49, anal lobes (posterior) of C. humboldti; 50, anal lobes (posterior) of C. spixi; 51, anal lobes (posterior) of C. lindei (dilated anal region due to the presence of excrement at death). Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601722/files/figure.png" pageId="3" pageNumber="504">Fig. 46</figureCitation>
) comprising of 270 short setae. Anal lobes (
<figureCitation id="BBBD2A69FFA8FFCBFAAF128781CFFB06" captionStart="FIGURES 4051" captionStartId="10.[151,264,1882,1905]" captionTargetBox="[220,1373,203,1847]" captionTargetId="figure@10.[200,1392,193,1867]" captionTargetPageId="10" captionText="FIGURES 4051. FIGURES 4045. Stridulatory organs Casignetus larvae. 40, Pars stridens of C. humboldti; 41, pars stridens of C. spixi; 42, pars stridens of C. lindei; 43, plectrum of C. humboldti; 44, plectrum of C. spixi; 45, plectrum of C. lindei. Scale bars = 1 mm. FIGURES 4651. Last abdominal segment of Casignetus larvae. 46, Raster of C. humboldti; 47, raster of C. spixi; 48, raster of C. lindei; 49, anal lobes (posterior) of C. humboldti; 50, anal lobes (posterior) of C. spixi; 51, anal lobes (posterior) of C. lindei (dilated anal region due to the presence of excrement at death). Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601722/files/figure.png" pageId="3" pageNumber="504">Fig. 49</figureCitation>
) glabrous.
</paragraph>
</subSubSection>
<subSubSection id="6B9C6567FFA8FFCBFF0712CF85B9FB22" box="[199,1218,1230,1255]" pageId="3" pageNumber="504" type="materials_examined">
<paragraph id="233936ECFFA8FFCBFF0712CF85B9FB22" blockId="3.[151,1437,726,1543]" box="[199,1218,1230,1255]" pageId="3" pageNumber="504">
<emphasis id="11F2EAFEFFA8FFCBFF0712CF80CBFB22" bold="true" box="[199,432,1230,1255]" pageId="3" pageNumber="504">Material examined.</emphasis>
<materialsCitation id="93EE3CB1FFA8FFCBFE7712CF85C6FB22" ID-GBIF-Occurrence="2013965689" box="[439,1213,1230,1255]" location="Nova Friburgo" pageId="3" pageNumber="504" specimenCount="12" specimenCount-female="5" specimenCount-larva="6" specimenCount-male="1" stateProvince="Rio de Janeiro">
<specimenCount id="3580FD65FFA8FFCBFE7712CF836BFB22" box="[439,528,1230,1255]" pageId="3" pageNumber="504" type="larva">6 larvae</specimenCount>
(
<specimenCount id="3580FD65FFA8FFCBFDE112CE833FFB22" box="[545,580,1231,1255]" pageId="3" pageNumber="504" type="male">1♂</specimenCount>
e
<specimenCount id="3580FD65FFA8FFCBFD9E12CF83F3FB23" box="[606,648,1230,1255]" pageId="3" pageNumber="504" type="female">5 ♀</specimenCount>
,
<location id="26596037FFA8FFCBFD5612CE8245FB22" LSID="urn:lsid:plazi:treatment:AB2F87FAFFA8FFCBFF57148E85C9F9C2:26596037FFA8FFCBFD5612CE8245FB22" box="[662,830,1230,1255]" name="Nova Friburgo" pageId="3" pageNumber="504" stateProvince="Rio de Janeiro">Nova Friburgo</location>
,
<collectingRegion id="E142F80EFFA8FFCBFC8912CF8295FB22" box="[841,1006,1230,1255]" country="Brazil" name="Rio de Janeiro" pageId="3" pageNumber="504">Rio de Janeiro</collectingRegion>
, Brazil—CERPE)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6B9C6567FFA8FFCBFF0712F385C9F9C2" pageId="3" pageNumber="504" type="discussion">
<paragraph id="233936ECFFA8FFCBFF0712F385C9F9C2" blockId="3.[151,1437,726,1543]" pageId="3" pageNumber="504">
<emphasis id="11F2EAFEFFA8FFCBFF0712F38040FACE" bold="true" box="[199,315,1266,1291]" pageId="3" pageNumber="504">Remarks.</emphasis>
<taxonomicName id="E4864D6FFFA8FFCBFE8612F28344FACE" baseAuthorityName="Gyllenhal" baseAuthorityYear="1817" box="[326,575,1267,1291]" class="Insecta" family="Lucanidae" genus="Casignetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="humboldti">
<emphasis id="11F2EAFEFFA8FFCBFE8612F28344FACE" box="[326,575,1267,1291]" italics="true" pageId="3" pageNumber="504">Casignetus humboldti</emphasis>
</taxonomicName>
occurs in southeastern and southern regions of
<collectingCountry id="5B91767CFFA8FFCBFBB712F385C0FACE" box="[1143,1211,1266,1291]" name="Brazil" pageId="3" pageNumber="504">Brazil</collectingCountry>
(
<bibRefCitation id="47174B1DFFA8FFCBFB0E12F384ECFACE" author="Luderwaldt, H." box="[1230,1431,1266,1291]" pageId="3" pageNumber="504" pagination="447 - 574" refId="ref7541" refString="Luderwaldt, H. (1935) Monographia dos lucanideos brasileiros. Revista do Museu Paulista, 19, 447 - 574." type="journal article" year="1935">Lüderwaldt 1935</bibRefCitation>
;
<bibRefCitation id="47174B1DFFA8FFCBFF5713178035FAEA" author="Weinreich, E." box="[151,334,1302,1327]" pageId="3" pageNumber="504" pagination="183 - 212" refId="ref7989" refString="Weinreich, E. (1963) Revision sudamerikanischer Lucanidae (Ins. Col.). III. Die Gattungen Pholidotus, Aegognathus, Macrocrates, Auxicerus und Erganzungenzu Metadorcus, Scortizus, Beneshius und Sclerostomus. Senckenbergiana Biologica, 44, 183 - 212." type="journal article" year="1963">Weinreich 1963</bibRefCitation>
). However, among the material examined, and the collections studied by one of us (PCG), there were only specimens from southeastern region from the states of
<collectingRegion id="E142F80EFFA8FFCBFCB2133B856FFA96" box="[882,1044,1338,1363]" country="Brazil" name="Espirito Santo" pageId="3" pageNumber="504">Espírito Santo</collectingRegion>
,
<collectingRegion id="E142F80EFFA8FFCBFBDF133B85BFFA96" box="[1055,1220,1338,1363]" country="Brazil" name="Rio de Janeiro" pageId="3" pageNumber="504">Rio de Janeiro</collectingRegion>
,
<collectingRegion id="E142F80EFFA8FFCBFB0F133B8412FA96" box="[1231,1385,1338,1363]" country="Brazil" name="Minas Gerais" pageId="3" pageNumber="504">Minas Gerais</collectingRegion>
, and
<collectingRegion id="E142F80EFFA8FFCBFF57135E8070FAB2" box="[151,267,1374,1399]" country="Brazil" name="Sao Paulo" pageId="3" pageNumber="504">São Paulo</collectingRegion>
, and for the southern region there are only two known specimens (
<bibRefCitation id="47174B1DFFA8FFCBFC3A135F85C9FAB2" author="Weinreich, E." box="[1018,1202,1374,1399]" pageId="3" pageNumber="504" pagination="183 - 212" refId="ref7989" refString="Weinreich, E. (1963) Revision sudamerikanischer Lucanidae (Ins. Col.). III. Die Gattungen Pholidotus, Aegognathus, Macrocrates, Auxicerus und Erganzungenzu Metadorcus, Scortizus, Beneshius und Sclerostomus. Senckenbergiana Biologica, 44, 183 - 212." type="journal article" year="1963">Weinreich 1963</bibRefCitation>
). Considering this, it is necessary to confirm the records of the specimens from that region. Regarding the pupae of
<taxonomicName id="E4864D6FFFA8FFCBFB1D13828409FA5E" baseAuthorityName="Gyllenhal" baseAuthorityYear="1817" box="[1245,1394,1411,1435]" class="Insecta" family="Lucanidae" genus="Casignetus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="504" phylum="Arthropoda" rank="species" species="humboldti">
<emphasis id="11F2EAFEFFA8FFCBFB1D13828409FA5E" box="[1245,1394,1411,1435]" italics="true" pageId="3" pageNumber="504">C. humboldti</emphasis>
</taxonomicName>
we have on hand, both male and female were nearing emergence, especially the female, whose thecae were damaged on many parts. For this reason, we do not present here full descriptions for this species. Additionally, most of the description of the genus was based on these pupae, and we consider not necessary to repeat it.
</paragraph>
</subSubSection>
</treatment>
</document>