treatments-xml/data/73/26/E1/7326E116DA81573B8B86E79970E71B52.xml
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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.227.100850" ID-Pensoft-Pub="1314-2003-227-123" ID-Pensoft-UUID="5E6B1E50FAF35B7DAC1BB68271C262E7" ModsDocID="1314-2003-227-123" checkinTime="1686000155700" checkinUser="pensoft" docAuthor="Selvi, Federico, Gonnelli, Cristina &amp; Bianchi, Elisabetta" docDate="2023" docId="7326E116DA81573B8B86E79970E71B52" docLanguage="en" docName="PhytoKeys 227: 123-134" docOrigin="PhytoKeys 227" docPubDate="2023-06-05" docSource="http://dx.doi.org/10.3897/phytokeys.227.100850" docTitle="Silene isabellae Selvi &amp; Bianchi 2023, sp. nov." docType="treatment" docVersion="2" id="5E6B1E50FAF35B7DAC1BB68271C262E7" lastPageNumber="123" masterDocId="5E6B1E50FAF35B7DAC1BB68271C262E7" masterDocTitle="Silene isabellae (Caryophyllaceae), a new campion species from serpentine soils of Albania" masterLastPageNumber="134" masterPageNumber="123" pageNumber="123" updateTime="1686000456488" updateUser="ExternalLinkService">
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<mods:title id="B73C63CBA4D4C7C5195BDB678453A435">Silene isabellae (Caryophyllaceae), a new campion species from serpentine soils of Albania</mods:title>
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<mods:namePart id="12D9917731534C39E6D8CCC3E3ABD2D1">Selvi, Federico</mods:namePart>
<mods:nameIdentifier id="67A197D4E7380AE87A8ED94E3ABA3D4F" type="ORCID">https://orcid.org/0000-0002-3820-125X</mods:nameIdentifier>
<mods:affiliation id="A7BA8A7A699BDA716F2EB3BFE99724B1">Department of Agriculture, Food, Environment and Forestry, Laboratories of Botany, University of Florence, Piazzale delle Cascine 28, I- 50144, Florence, Italy &amp; NBFC, National Biodiversity Future Center, I- 90133, Palermo, Italy</mods:affiliation>
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<mods:namePart id="B1A93E58090740099E5FC74C0DAD251E">Gonnelli, Cristina</mods:namePart>
<mods:nameIdentifier id="47054766E1146A5970EB8C4FB317D6FF" type="ORCID">https://orcid.org/0000-0001-5201-586X</mods:nameIdentifier>
<mods:affiliation id="656C844985125292F1FC9D64D1237BB0">NBFC, National Biodiversity Future Center, I- 90133, Palermo, Italy &amp; Department of Agriculture, Food, Environment and Forestry, Laboratories of Botany, University of Florence, Piazzale delle Cascine 28, I- 50144, Florence, Italy</mods:affiliation>
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<mods:namePart id="C4D844A61DC6A4E1EEADF42CD3FB9A65">Bianchi, Elisabetta</mods:namePart>
<mods:nameIdentifier id="EA6C95D6E313CAC3074277D21E888BE6" type="ORCID">https://orcid.org/0000-0003-1197-4081</mods:nameIdentifier>
<mods:affiliation id="FB65D2BCFD214904946FE868425567C6">Department of Agriculture, Food, Environment and Forestry, Laboratories of Botany, University of Florence, Piazzale delle Cascine 28, I- 50144, Florence, Italy &amp; NBFC, National Biodiversity Future Center, I- 90133, Palermo, Italy</mods:affiliation>
<mods:nameIdentifier id="B65CC51AAF3949DFBC3CCDD62D2666C6" type="email">e.bianchi@unifi.it</mods:nameIdentifier>
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<mods:date id="824456F75E80DC674F34F8B0C1D56BD8">2023</mods:date>
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<mods:identifier id="A804B301BE4DDEF21E47210351C723BA" type="DOI">http://dx.doi.org/10.3897/phytokeys.227.100850</mods:identifier>
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<treatment id="7326E116DA81573B8B86E79970E71B52" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52" httpUri="http://treatment.plazi.org/id/7326E116DA81573B8B86E79970E71B52" lastPageNumber="123" pageId="0" pageNumber="123">
<subSubSection id="2519B3633EAC79548C5E8B86F3004D4E" pageId="0" pageNumber="123" type="nomenclature">
<paragraph id="7A8586CC815948A47AE573F597EF2172" pageId="0" pageNumber="123">
<taxonomicName id="575B732B15A564CC044326BCA5864E6C" LSID="7326E116-DA81-573B-8B86-E79970E71B52" authority="Selvi &amp; Bianchi" authorityName="Selvi &amp; Bianchi" authorityYear="2023" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Silene isabellae" order="Caryophyllales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="species" species="isabellae" status="sp. nov.">Silene isabellae Selvi &amp; Bianchi</taxonomicName>
<taxonomicNameLabel id="0B07E2BA211B59AC38A6CD48C85BEB2F" pageId="0" pageNumber="123">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="6B7D5961F47BCFB511AA8BC1B6BB35AC" pageId="0" pageNumber="123" type="description">
<paragraph id="E31296DACBE7EEBCCEF95EB19EE5298C" pageId="0" pageNumber="123">
<figureCitation id="9AA2A0A0AE6BA69834E1FDD6013A30B2" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Silene isabellae A whole plant B flower with calyx and corolla C fruiting calyx D capsule and carpophore E seed. Original drawing by Laura Vivona." figureDoi="10.3897/phytokeys.227.100850.figure1" httpUri="https://binary.pensoft.net/fig/859764" pageId="0" pageNumber="123">Figs 1</figureCitation>
<figureCitation id="A8DC49CEA4D492B23297DD7F8B2C1FAA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Field photos of Silene isabellae A habitat B, C whole plants in natural habitat D inflorescence with flowers and fruiting calyces E flower F basal leaves G flowers at late stage H seed." figureDoi="10.3897/phytokeys.227.100850.figure2" httpUri="https://binary.pensoft.net/fig/859765" pageId="0" pageNumber="123">, 2</figureCitation>
<figureCitation id="8588EB93B6747999613DDB0D7BD3C7C7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Calyx teeth, capsule, and margin of basal leaves of Silene isabellae (respectively A-C), S. noctiflora (respectively D-F) and S. vallesia subsp. graminea (respectively G-I)." figureDoi="10.3897/phytokeys.227.100850.figure3" httpUri="https://binary.pensoft.net/fig/859766" pageId="0" pageNumber="123">, 3A-C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4CD8C7718A1AB2A23BFFDE9E86C5079D" pageId="0" pageNumber="123" type="materials_examined">
<paragraph id="55D16507054A5D2CA5ADAE342DDD9F22" pageId="0" pageNumber="123">
<typeStatus id="202C6A3C3D5E96CE2E7AD4619104C30D">Type</typeStatus>
.
</paragraph>
<paragraph id="BAE7F1A1E709E740BF8DE588B38F1248" pageId="0" pageNumber="123">
<materialsCitation id="E90A69C215A3CA372CB9B7C16AA769EA" accessionNumber="FI068201" collectingDate="2022-07-12" collectorName="F. Selvi, E. Bianchi, I. Colzi, A. Coppi" country="Albania" elevation="1100" latitude="41.522167" location="Kruje" longLatPrecision="11" longitude="19.887833" municipality="Qafe Shtame" specimenCount="1" typeStatus="holotype">
<emphasis id="33D8B9CB89E6D867A3E1E80AAEC1DDB1" bold="true" pageId="0" pageNumber="123">
<collectingCountry id="914650E6F7BEC304E1419E3B194751E2" name="Albania">Albania</collectingCountry>
</emphasis>
,
<normalizedToken id="8F7900F522A68C4257229767C6A955A4" originalValue="Krujë">
<location id="DC3622A802E99F81B9FC1695096D5BB5" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:DC3622A802E99F81B9FC1695096D5BB5" country="Albania" latitude="41.522167" longLatPrecision="11" longitude="19.887833" municipality="Qafe Shtame" name="Kruje">Kruje</location>
</normalizedToken>
, verso
<collectingMunicipality id="EFEABC0C54A1AE1D7349182DBC6209E1">
<normalizedToken id="77330611C45757F0EDE4F319E36CBAFD" originalValue="Qafë">Qafe</normalizedToken>
<normalizedToken id="A57C9BE828024B90E814C5626DE1F125" originalValue="Shtamë">Shtame</normalizedToken>
</collectingMunicipality>
, pendice ripida del monte sopra la fonte lungo la strada prima del passo, sottobosco della foresta rada di
<taxonomicName id="2ABA5AE2554D9E059C0CAEB4CEAC4944" lsidName="P. nigra" pageId="0" pageNumber="123" rank="species" species="nigra">
<emphasis id="7BC542086E60D5CCBC4258B87393EA39" italics="true" pageId="0" pageNumber="123">P. nigra</emphasis>
</taxonomicName>
, suolo roccioso ultramafico (serpentino),
<quantity id="AA44FC662E44F8E928E67430A58C1F8F" metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">
<elevation id="CFB200A86FFEB1A9286B91B6190773CA" metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">1100 m</elevation>
</quantity>
elev.,
<geoCoordinate id="1B75054614680ECEBBCDEB65300CF15F" degrees="41" direction="north" minutes="31.33" orientation="latitude" precision="9" value="41.522167">41°31.33'N</geoCoordinate>
,
<geoCoordinate id="B67C296C0767FAA855C9578CBCF116C9" degrees="19" direction="east" minutes="53.27" orientation="longitude" precision="9" value="19.887833">19°53.27'E</geoCoordinate>
;
<emphasis id="BACF547EF36BAFEF20A598D1403EE1E9" italics="true" pageId="0" pageNumber="123">
<collectorName id="1FB1F02A13BD07E5253403EE3A7F8D4F">F. Selvi</collectorName>
,
<collectorName id="8F05C4F3EE1D43591D303A28EA822AAA">E. Bianchi</collectorName>
,
<collectorName id="38DDEBFAE7AC0FA672AE2B23079F8F70">I. Colzi</collectorName>
&amp;
<collectorName id="242C1825DFBA43C3C0B5310487D624C5">A. Coppi</collectorName>
</emphasis>
,
<collectingDate id="DC89DB1E510B19940BACE8718A3EFF52" value="2022-07-12">12 Jul 2022</collectingDate>
(
<typeStatus id="FC160BB4B098D1010F8BA638DC670C7A">holotype</typeStatus>
:
<accessionNumber id="693A01222E0A62C9BFB6D7722B057B81" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/FI068201">FI068201</accessionNumber>
; isotypes: K, G, TIR)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BC821B5DFE70FA7D7D17D0FB1958BC1C" pageId="0" pageNumber="123" type="diagnosis">
<paragraph id="69A2C066A26AE7B0D5BB632E3D2306CE" pageId="0" pageNumber="123">Diagnosis.</paragraph>
<paragraph id="71216149110C6C52A714BC4D44FFC3DC" pageId="0" pageNumber="123">
<taxonomicName id="5B648E0D68E47D63E0D8B77BDEEEAAEF" authorityName="Selvi &amp; Bianchi" authorityYear="2023" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Silene isabellae" order="Caryophyllales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="species" species="isabellae">
<emphasis id="9025DD7D60FFAF17D800CB3713695EAF" italics="true" pageId="0" pageNumber="123">Silene isabellae</emphasis>
</taxonomicName>
differs from
<taxonomicName id="E065D66C072A0078A011507DAC827F50" lsidName="S. noctiflora" pageId="0" pageNumber="123" rank="species" species="noctiflora">
<emphasis id="E6E16C615F8742A3B116E9EA5B222D3A" italics="true" pageId="0" pageNumber="123">S. noctiflora</emphasis>
</taxonomicName>
by the perennial habit with stoloniferous stems forming thick mats (instead of annual), the sparsely and shortly glandular-pubescent stem (vs. densely hairy), the basal leaves of the sterile stems present and widely obovate-spathulate (vs. absent), the cauline leaves linear-lanceolate (3-5 mm vs. ovate-lanceolate 25-35 mm wide), with shortly ciliate margins (vs. densely pubescent on both surfaces), the longer calyx teeth (ca. 8.5 vs. 7.0 mm), the corolla unscented and open during daytime (instead of scented and opening at evening), the petal lobes dentate (vs. entire), the fruiting calyx with prominent longitudinal ribs and the longer carpophore (6-7 vs. 2-3 mm).
</paragraph>
</subSubSection>
<subSubSection id="17AE21140D6B274C44E7BB7593638D70" pageId="0" pageNumber="123" type="description">
<paragraph id="187FB0FA35A133ECBC9D9B900C80F496" pageId="0" pageNumber="123">Description.</paragraph>
<paragraph id="58F060EA32B4F625B564417FC74F8BFF" pageId="0" pageNumber="123">
Perennial herb forming thick mats, with branched stoloniferous stems thickened at the nodes. Flowering stems up to 40 cm, rigid, usually 1-4 branched in the upper half, rarely simple, shortly glandular-pubescent. Basal leaves mostly appressed to the ground, 30-80
<normalizedToken id="56D985207949B2C5F302753E8246E388" originalValue="×">x</normalizedToken>
8-18 mm, tapering into the petiole, widely obovate-spathulate, apiculate at tip; leaf blade thick, glabrous and finely papillose on both surfaces, shortly ciliate at the margins (hairs ca. 0.15 mm). Cauline leaves in pairs of 4-5, the uppermost narrowly lanceolate to almost linear, 30-50
<normalizedToken id="23B61D033DBF6ECC6C8AF0DAA65EAFDE" originalValue="×">x</normalizedToken>
3-5 mm, acuminate, the mid- and upper ones much shorter than the internodes, finely ciliate along margins with glandular and simple hairs on both surfaces. Inflorescence with 1-4 branches each bearing a single flower, rarely simple; branches finely glandular-pubescent, with two linear-lanceolate leaf-like bracts 6-8 mm long. Flowering calyx narrowly cylindrical, 25-30 mm, including teeth 8.2-8.6 mm, linear-lanceolate, surface glandular-pubescent, veins tinged with purplish, reticulate especially in the upper half; fruiting calyx obconical, truncate at the base and swollen at the middle, up to 13 mm wide, with 5-6 prominent longitudinal ribs. Corolla spreading 19-21 mm wide, petals 25-30 mm long (claw 15-20 mm long and slightly exserted), divided to ca. 1/3 in two lobes dentate at the outer margin, pale pink above, tinged with mauve red beneath; coronal scales exserted, truncate, nearly white; styles 3. Capsule smooth and glossy, ovoid-conical, 13-15
<normalizedToken id="709882A9B60739ED9E7743BEF0BBB96D" originalValue="×">x</normalizedToken>
4-5 mm, dehiscing with 6 teeth curved outwards, carpophore 6-7 mm. Seeds ca. 1.1
<normalizedToken id="75CF7ADB09A900084074E648258F05CA" originalValue="×">x</normalizedToken>
0.83 mm, reniform, with surface finely papillose-tuberculate in concentric crests around the hilum, dark brown to blackish.
</paragraph>
</subSubSection>
<subSubSection id="10810E9F8504ED3AE7111E6AE3ADBE21" pageId="0" pageNumber="123" type="etymology">
<paragraph id="232C90738BBE0EFC6706E311A10790B5" pageId="0" pageNumber="123">Etymology.</paragraph>
<paragraph id="3FA817142129E54755BCC103F5158428" pageId="0" pageNumber="123">
This species is dedicated to the first
<normalizedToken id="9ABB7AAFC7DF765A89840F24757A54D0" originalValue="authors">author's</normalizedToken>
wife, for her continuous support and advice during many botanical trips across the Mediterranean and the Middle East.
</paragraph>
</subSubSection>
<subSubSection id="CAA07339F963231DF6C81EF8E23DBF8B" pageId="0" pageNumber="123" type="phenology">
<paragraph id="97D7125E9AB2A480114979E7571953CA" pageId="0" pageNumber="123">Phenology.</paragraph>
<paragraph id="DE641CD9ACFC55636731F7CC6AB03E1E" pageId="0" pageNumber="123">Flowering is from June to early July; fruiting is from end of June to end of July, depending on slope aspect and altitude.</paragraph>
</subSubSection>
<subSubSection id="89845EC0CC27ED4AC9E32452DB5716FA" pageId="0" pageNumber="123" type="distribution">
<paragraph id="039F8C83FF58C67DD30B6AB6D04907DB" pageId="0" pageNumber="123">Distribution and ecology.</paragraph>
<paragraph id="DD8E032D756058D690AF239B0064B7D4" pageId="0" pageNumber="123">
<taxonomicName id="EB120774BC8712C7298DF38DFF65FC04" authorityName="Selvi &amp; Bianchi" authorityYear="2023" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Silene isabellae" order="Caryophyllales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="species" species="isabellae">
<emphasis id="40395BA62CD2099383E178BA8AAA5BB3" italics="true" pageId="0" pageNumber="123">Silene isabellae</emphasis>
</taxonomicName>
is likely endemic to central Albania, precisely to the ultramafic sectors of the
<normalizedToken id="ACB8C4388DE085D16A65FAB1F4DC22DA" originalValue="Skënderbëut">Skenderbeut</normalizedToken>
mountain range running along the border between the districts of
<normalizedToken id="8E3A9DC571656C86A26BEF6F54ACE957" originalValue="Krujë">Kruje</normalizedToken>
and Mat (Fig.
<figureCitation id="4FD907CCF0DACC9B0DDA1E1999C85CA2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution of Silene isabellae, S. noctiflora (partial) and the taxa of the S. vallesia L. group." figureDoi="10.3897/phytokeys.227.100850.figure4" httpUri="https://binary.pensoft.net/fig/859767" pageId="0" pageNumber="123">4</figureCitation>
). We could not observe it in other parts of this mountain range with calcareous soil. Additionally, we could not find it during our botanical trips across other ultramafic mountain areas of the country (amongst which were
<normalizedToken id="56B97381246BB43C409FC1F1DFDD7F57" originalValue="Tropojë">Tropoje</normalizedToken>
, Pashtrik,
<normalizedToken id="806C227DA5783CFECB42BDF042C170B4" originalValue="Lurë">Lure</normalizedToken>
,
<normalizedToken id="ACFEC368B5E4A434B7F025722C017FD4" originalValue="Bulqizë">Bulqize</normalizedToken>
, Shebenik, Shpat,
<normalizedToken id="BEB20A8A3DE041C5170FEA2369204B47" originalValue="Moravë">Morave</normalizedToken>
and
<normalizedToken id="7FA6B9BF3C4DCE2A5547E05CBA8E3DB5" originalValue="Vallamarë">Vallamare</normalizedToken>
), suggesting a restricted range and, possibly, the reason why this species has remained unknown until present. The plant is found in patches in the understorey of the open
<taxonomicName id="C0192FE1AAB00155D70F446D31A3DEFD" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" lsidName="Pinus nigra" order="Pinales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="species" species="nigra">
<emphasis id="A566B3E1D2BB62787B0FB3C17186A3C0" italics="true" pageId="0" pageNumber="123">Pinus nigra</emphasis>
</taxonomicName>
forests and in the rocky grasslands on the mountain slopes around the pass
<normalizedToken id="D52AB08DDE938584BE216E723DC583CF" originalValue="Qafë">Qafe</normalizedToken>
<normalizedToken id="895ADD88E30A4511FB18A63CF60CC14F" originalValue="Shtamë">Shtame</normalizedToken>
, between 1100 and 1600 m a.s.l., but possibly even higher (Fig.
<figureCitation id="1155D0BFC9C27BDF4F684A65504FA0A1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Field photos of Silene isabellae A habitat B, C whole plants in natural habitat D inflorescence with flowers and fruiting calyces E flower F basal leaves G flowers at late stage H seed." figureDoi="10.3897/phytokeys.227.100850.figure2" httpUri="https://binary.pensoft.net/fig/859765" pageId="0" pageNumber="123">2A, B</figureCitation>
). Slope aspect was prevalently south and south-west. This area is geologically characterised by vast outcrops of ultramafic rocks hosting a rich serpentine flora including remarkable Balkan endemics, such as
<taxonomicName id="B0826AB2E27618EFBA943A4830A1308C" class="Magnoliopsida" family="Oleaceae" genus="Forsythia" kingdom="Plantae" lsidName="Forsythia europaea" order="Lamiales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="species" species="europaea">
<emphasis id="EB8D7331155EEE3CFD7B6A5A58AB2904" italics="true" pageId="0" pageNumber="123">Forsythia europaea</emphasis>
</taxonomicName>
Degen &amp; Bald.,
<taxonomicName id="35FE15A45E46453EA846B225469B6AE3" class="Liliopsida" family="Poaceae" genus="Festucopsis" kingdom="Plantae" lsidName="Festucopsis serpentini" order="Poales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="species" species="serpentini">
<emphasis id="AF22E90A0F6BAD9FE73FE4C8D4090EAC" italics="true" pageId="0" pageNumber="123">Festucopsis serpentini</emphasis>
</taxonomicName>
(C.E. Hubb.) Melderis and the Ni-hyperaccumulator
<taxonomicName id="3DE2EBC5F3053AB31F032D1C8DD83C62" class="Magnoliopsida" family="Brassicaceae" genus="Odontarrhena" kingdom="Plantae" lsidName="Odontarrhena smolikana" order="Brassicales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="species" species="smolikana">
<emphasis id="B84400884C22AAAC5BB8C8EC2119FA70" italics="true" pageId="0" pageNumber="123">Odontarrhena smolikana</emphasis>
</taxonomicName>
(Bald.)
<normalizedToken id="096DB3A776A5B19A22B4C9FCAEB6A32C" originalValue="Španiel">Spaniel</normalizedToken>
, Al-Shehbaz, D.A.German &amp; Marhold, subsp.
<taxonomicName id="C60E0AA4038720214B0279415A1A65D5" authorityName="L.Cecchi &amp; Selvi" authorityYear="2018" baseAuthorityName="Nyar. ex Markgr." class="Magnoliopsida" family="Brassicaceae" genus="Odontarrhena" kingdom="Plantae" lsidName="Odontarrhena smolikana subsp. glabra" order="Brassicales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="subSpecies" species="smolikana" subSpecies="glabra">Odontarrhena smolikana glabra</taxonomicName>
(
<normalizedToken id="136ACD3B54FBEEB40B7B79162C133A92" originalValue="Nyár">Nyar</normalizedToken>
. ex Markgr.) L.Cecchi &amp; Selvi. We assume that
<taxonomicName id="A3CB9ECAA0C847E05D441CDE0ED1480F" lsidName="S. isabellae" pageId="0" pageNumber="123" rank="species" species="isabellae">
<emphasis id="FD376441D497BA692F07263ABA90432C" italics="true" pageId="0" pageNumber="123">S. isabellae</emphasis>
</taxonomicName>
is an obligate serpentine endemic.
</paragraph>
</subSubSection>
<subSubSection id="1D096D33F08F4EB15D95EBE3FCA7CC47" pageId="0" pageNumber="123" type="conservation status">
<paragraph id="285CF6320AEE828894C73CF475BB54CF" pageId="0" pageNumber="123">Conservation status.</paragraph>
<paragraph id="E7557C07DA6EE0F7586020B27E9D9156" pageId="0" pageNumber="123">
<taxonomicName id="1550CC4A0F292709CB6C0C910F933635" authorityName="Selvi &amp; Bianchi" authorityYear="2023" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Silene isabellae" order="Caryophyllales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="species" species="isabellae">
<emphasis id="1C3752DE7CDB8D90A9E5E96434B04A1E" italics="true" pageId="0" pageNumber="123">Silene isabellae</emphasis>
</taxonomicName>
grows in a mountain area which has been subjected for a long time to grazing mainly by sheep and goats. This ancient land use form has implied the creation and conservation of large grassland areas at the expenses of the forest. Fire is also a recurrent disturbance especially to the pine forest on the mountain slopes. Over the area that we could visit (ca. 5 km2), the species was relatively frequent, though discontinuously distributed. Based on the present state of knowledge, its range and extent of occurrence seem quite restricted, suggesting categorising it as &quot;nearly threatened&quot; (NT), mainly according to IUCN criterion B (
<bibRefCitation id="4C92B860C27AF649B071676EEB71AFA0" author="Greuter, W" journalOrPublisher="Lubrecht &amp; Cramer" pageId="0" pageNumber="123" refId="B13" refString="IUCN [International Union for Conservation of Nature] (2012) IUCN red list categories and criteria. version 3.1, 2nd edn. IUCN, Gland and Cambridge." year="1997">IUCN 2012</bibRefCitation>
). However, more field studies are needed to define the correct conservation category of this species.
</paragraph>
</subSubSection>
<subSubSection id="7D50805CABDF6377C052C88BA6F4C9F9" pageId="0" pageNumber="123" type="discussion">
<paragraph id="D5FB8BD8C48E4CA5FB608B7949F07E38" pageId="0" pageNumber="123">Discussion.</paragraph>
<paragraph id="685B41F7883D87ED2C091C5892B49941" pageId="0" pageNumber="123">
<taxonomicName id="6BB23E4E0C30F015FC2252B22336E24A" authorityName="Selvi &amp; Bianchi" authorityYear="2023" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Silene isabellae" order="Caryophyllales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="species" species="isabellae">
<emphasis id="92018D815742A08D6E4D020BA03AF030" italics="true" pageId="0" pageNumber="123">Silene isabellae</emphasis>
</taxonomicName>
shows morphological affinities with taxa of
<taxonomicName id="4E7DABAD9268401A3265BB001B4EDE7C" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Silene (Behenantha) subsp. subg." order="Caryophyllales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="subGenus" subGenus="Behenantha">Silene subg. Behenantha</taxonomicName>
(Otth) Torr. &amp; A. Gray sect.
<taxonomicName id="E7B46537F4C0BEF30D852DA2DF8D46EF" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Silene" order="Caryophyllales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="section" section="Elisanthe">Silene Elisanthe</taxonomicName>
(Fenzl ex Endl.) Ledeb. and is assumed to belong here. This is a well-defined group supported by molecular evidence (
<bibRefCitation id="EE6D539A03A03853FE4A9F73F98C3858" DOI="https://doi.org/10.1002/tax.12230" author="Jafari, F" journalOrPublisher="Taxon" pageId="0" pageNumber="123" pagination="337 - 368" refId="B14" refString="Jafari, F, Zarre, S, Gholipour, A, Eggens, F, Rabeler, RK, Oxelman, B, 2020. A new taxonomic backbone for the infrageneric classification of the species-rich genus Silene (Caryophyllaceae). Taxon 69 (2): 337 - 368, DOI: https://doi.org/10.1002/tax.12230" title="A new taxonomic backbone for the infrageneric classification of the species-rich genus Silene (Caryophyllaceae)." url="https://doi.org/10.1002/tax.12230" volume="69" year="2020">Jafari et al. 2020</bibRefCitation>
) that includes annual, biennial and perennial species with large flowers, solitary or in dichasia, often dioecious, with bifid petal limb, prominent coronal scales, often five styles, capsule without basal septa dehiscing with 6 or 10 teeth and a very short carpophore (
<bibRefCitation id="5DBE9B26EBBE113EE228F73BDB9DE72B" author="Chater, AO" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="123" refId="B7" refString="Chater, AO, Walters, SM, Akeroyd, JR, Wrigley, F, 1993. Silene L. in Flora Europaea 2°edn., vol. 1 Cambridge." title="Silene L. in Flora Europaea 2 ° edn., vol. 1 Cambridge." year="1993">Chater et al. 1993</bibRefCitation>
). In Europe, the section includes widespread species, such as
<taxonomicName id="3CC147AF65B1AAD7E5C469EBE140B3D5" lsidName="S. latifolia" pageId="0" pageNumber="123" rank="species" species="latifolia">
<emphasis id="09A79226CF4521564798380BA594B72F" italics="true" pageId="0" pageNumber="123">S. latifolia</emphasis>
</taxonomicName>
Poir.,
<taxonomicName id="E3A56C9E97C69236B81F06E5DE4B7D2B" lsidName="S. dioica" pageId="0" pageNumber="123" rank="species" species="dioica">
<emphasis id="A6557532902D56C64907B711370996F8" italics="true" pageId="0" pageNumber="123">S. dioica</emphasis>
</taxonomicName>
(L.) Clairv. and
<taxonomicName id="F3179561BC946D430CD4CA4F0C79D8E5" lsidName="S. noctiflora" pageId="0" pageNumber="123" rank="species" species="noctiflora">
<emphasis id="BDD87EEB1757D195518A9B00CAEA1899" italics="true" pageId="0" pageNumber="123">S. noctiflora</emphasis>
</taxonomicName>
L. (
<bibRefCitation id="521E3766E1BC0E71B405304F567DE92C" DOI="https://doi.org/10.1111/j.1095-8339.1978.tb01399.x" author="Prentice, HC" journalOrPublisher="Botanical Journal of the Linnean Society" pageId="0" pageNumber="123" pagination="203 - 2016" refId="B22" refString="Prentice, HC, 1978. Experimental taxonomy of Silene section Elisanthe (Caryophyllaceae): Crossing experiment. Botanical Journal of the Linnean Society 77 (3): 203 - 2016, DOI: https://doi.org/10.1111/j.1095-8339.1978.tb01399.x" title="Experimental taxonomy of Silene section Elisanthe (Caryophyllaceae): Crossing experiment." url="https://doi.org/10.1111/j.1095-8339.1978.tb01399.x" volume="77" year="1978">Prentice 1978</bibRefCitation>
). The latter species, described by
<bibRefCitation id="3FC682BBA86280245D524A45F07408FB" author="Linnaeus, C" journalOrPublisher="Nordic Journal of Botany" pageId="0" pageNumber="123" refId="B17" refString="Linnaeus, C, 1753. Species Plantarum 1. Holmiae." title="Species Plantarum 1. Holmiae." year="1753">Linnaeus (1753)</bibRefCitation>
from Sweden and Germany (&quot;Habitat in Suecia, Germania&quot;; see
<bibRefCitation id="1793E9B0DC5E31C0EF3450C6EFDC9245" DOI="https://doi.org/10.1111/j.1756-1051.1994.tb00581.x" author="Jonsell, B" journalOrPublisher="Nordic Journal of Botany" pageId="0" pageNumber="123" pagination="145 - 164" refId="B16" refString="Jonsell, B, Jarvis, CE, 1994. Lectotypifications of Linnaean names for Flora Nordica Vol. 1 (Lycopodiaceae - Papaveraceae). Nordic Journal of Botany 14 (2): 145 - 164, DOI: https://doi.org/10.1111/j.1756-1051.1994.tb00581.x" title="Lectotypifications of Linnaean names for Flora Nordica Vol. 1 (Lycopodiaceae - Papaveraceae)." url="https://doi.org/10.1111/j.1756-1051.1994.tb00581.x" volume="14" year="1994">Jonsell and Jarvis (1994)</bibRefCitation>
), is apparently the more closely related one to
<taxonomicName id="681CDB125E285CD39AF292B79283C390" lsidName="S. isabellae" pageId="0" pageNumber="123" rank="species" species="isabellae">
<emphasis id="3F2FEE2999AACE535AC3ED0ECDD8280C" italics="true" pageId="0" pageNumber="123">S. isabellae</emphasis>
</taxonomicName>
, with which it shares the thick stem, the few-flowered inflorescence, the texture and shape of the calyx, the usually hermaphrodite flowers with three styles and the reniform seeds with a nearly identical ornamentation pattern. On the other hand,
<taxonomicName id="E7BD20BC3B767D14B57389D9AB999625" lsidName="S. isabellae" pageId="0" pageNumber="123" rank="species" species="isabellae">
<emphasis id="55B05F47BC32767530EAF895157E97C5" italics="true" pageId="0" pageNumber="123">S. isabellae</emphasis>
</taxonomicName>
differs from
<taxonomicName id="5CD6CC9BFFD9538ED56202EECB9E0D84" lsidName="S. noctiflora" pageId="0" pageNumber="123" rank="species" species="noctiflora">
<emphasis id="E91831DF43B36D77D6F69140D135935E" italics="true" pageId="0" pageNumber="123">S. noctiflora</emphasis>
</taxonomicName>
in several striking characters (see Table
<tableCitation id="01D253F9DE3A7E4B3A34D1B1F5AA265A" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Main diagnostic morphological characters of S. isabellae, S. noctiflora and S. vallesia subsp. graminea." httpUri="http://table.plazi.org/id/41CAFFFD02350AC5C1EAAFBE123462F1" pageId="0" pageNumber="123" tableUuid="41CAFFFD02350AC5C1EAAFBE123462F1">1</tableCitation>
). Most importantly, it is a perennial plant forming thick mats with sterile stoloniferous stems and basal leaves (Fig.
<figureCitation id="1917905C691A30AEFE66A76DFC0E043C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Silene isabellae A whole plant B flower with calyx and corolla C fruiting calyx D capsule and carpophore E seed. Original drawing by Laura Vivona." figureDoi="10.3897/phytokeys.227.100850.figure1" httpUri="https://binary.pensoft.net/fig/859764" pageId="0" pageNumber="123">1A</figureCitation>
), while
<taxonomicName id="BB549BB403E835992294C0929F26052F" lsidName="S. noctiflora" pageId="0" pageNumber="123" rank="species" species="noctiflora">
<emphasis id="AFAA5C330A711F20CA76B99DE4025F86" italics="true" pageId="0" pageNumber="123">S. noctiflora</emphasis>
</taxonomicName>
is annual with usually a single stem and lacking basal leaves. The latter is also different in the dense and long pubescence of the lower leaves and stems which are lacking in
<taxonomicName id="69CA187F7C48F59EF8573BE3CCA0BEFA" lsidName="S. isabellae" pageId="0" pageNumber="123" rank="species" species="isabellae">
<emphasis id="58A022ED7DDC668CC48DD0911A3165E0" italics="true" pageId="0" pageNumber="123">S. isabellae</emphasis>
</taxonomicName>
, where the lower leaves are nearly glabrous and the lower stem is very shortly glandular-pubescent. The flowers of
<taxonomicName id="5ECA28C11E18056DC63567C6FEDB30A3" lsidName="S. isabellae" pageId="0" pageNumber="123" rank="species" species="isabellae">
<emphasis id="27F4511386F13523E49F657B602F8B0B" italics="true" pageId="0" pageNumber="123">S. isabellae</emphasis>
</taxonomicName>
are open during daytime and unscented, while
<taxonomicName id="A745303D40A8CDC3173A2CBC04BD9E96" lsidName="S. noctiflora" pageId="0" pageNumber="123" rank="species" species="noctiflora">
<emphasis id="9FD4DCDA9262CFD1DDA5A693A48F0BAC" italics="true" pageId="0" pageNumber="123">S. noctiflora</emphasis>
</taxonomicName>
is characterised by scented flowers with petals enrolled during daytime, opening in evening and night. Moreover, the petal lobes of the latter are entire or nearly so, while these are distinctly dentate in the Albanian plant (Figs
<figureCitation id="89B7402296F77B4F7013352BDF90DAF7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Silene isabellae A whole plant B flower with calyx and corolla C fruiting calyx D capsule and carpophore E seed. Original drawing by Laura Vivona." figureDoi="10.3897/phytokeys.227.100850.figure1" httpUri="https://binary.pensoft.net/fig/859764" pageId="0" pageNumber="123">1B</figureCitation>
,
<figureCitation id="672AB1310369C3C64A6F445FFCE7ABDC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Field photos of Silene isabellae A habitat B, C whole plants in natural habitat D inflorescence with flowers and fruiting calyces E flower F basal leaves G flowers at late stage H seed." figureDoi="10.3897/phytokeys.227.100850.figure2" httpUri="https://binary.pensoft.net/fig/859765" pageId="0" pageNumber="123">2D, E</figureCitation>
). The fruiting calyx is more distinctly truncate at the base and obconical, provided with prominent veins forming longitudinal ribs (Figs
<figureCitation id="351F213989067A9DC55E912280188598" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Silene isabellae A whole plant B flower with calyx and corolla C fruiting calyx D capsule and carpophore E seed. Original drawing by Laura Vivona." figureDoi="10.3897/phytokeys.227.100850.figure1" httpUri="https://binary.pensoft.net/fig/859764" pageId="0" pageNumber="123">1C</figureCitation>
,
<figureCitation id="7DDF705B88972A588DEDFF6CC51F95DF" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Field photos of Silene isabellae A habitat B, C whole plants in natural habitat D inflorescence with flowers and fruiting calyces E flower F basal leaves G flowers at late stage H seed." figureDoi="10.3897/phytokeys.227.100850.figure2" httpUri="https://binary.pensoft.net/fig/859765" pageId="0" pageNumber="123">2D</figureCitation>
). The capsule is borne by a longer carpophore (Fig.
<figureCitation id="A98BB0B98913767DDE32589BFCBCA8B0" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Silene isabellae A whole plant B flower with calyx and corolla C fruiting calyx D capsule and carpophore E seed. Original drawing by Laura Vivona." figureDoi="10.3897/phytokeys.227.100850.figure1" httpUri="https://binary.pensoft.net/fig/859764" pageId="0" pageNumber="123">1D</figureCitation>
). Additional evidence comes from the comparison of the chorological and ecological traits of the two taxa. In fact,
<taxonomicName id="DFCBEAF7F41794C4DE9FF9EEC6BE619A" lsidName="S. noctiflora" pageId="0" pageNumber="123" rank="species" species="noctiflora">
<emphasis id="00D383EAEFAD94A77F6C1F921194654F" italics="true" pageId="0" pageNumber="123">S. noctiflora</emphasis>
</taxonomicName>
is widespread in Central and Eastern Europe (
<bibRefCitation id="DD7E9B4AA36C263333E929A6CD8071D7" author="Jalas, J" journalOrPublisher="Taxon" pageId="0" pageNumber="123" refId="B15" refString="Jalas, J, Suominen, J, 1988. Atlas Florae Europaeae, volume 3: distribution of vascular plants in Europe (Vol. 3), Caryophyllaceae. Cambridge." title="Atlas Florae Europaeae, volume 3: distribution of vascular plants in Europe (Vol. 3), Caryophyllaceae. Cambridge." year="1988">Jalas and Suominen 1988</bibRefCitation>
), including Serbia, Kosovo and Methojia, where it occurs south of
<normalizedToken id="B441EBC6BD6A6BDFF9182089224FDC8A" originalValue="Priština">Pristina</normalizedToken>
, on the eastern rim of the Kosovo Basin (
<bibRefCitation id="AEE70CFBBE532D88FD30A5D7EB659B16" DOI="https://doi.org/10.5281/zenodo.200403" author="Prodanovic, D" journalOrPublisher="Biologica Nyssana" pageId="0" pageNumber="123" pagination="83 - 86" refId="B23" refString="Prodanovic, D, Stanojevic, M, Krivosej, Z, 2016. Silene noctiflora L., present in the flora of Kosovo and Metohija (Serbia). Biologica Nyssana 7: 83 - 86, DOI: https://doi.org/10.5281/zenodo.200403" title="Silene noctiflora L., present in the flora of Kosovo and Metohija (Serbia)." url="https://doi.org/10.5281/zenodo.200403" volume="7" year="2016">
<normalizedToken id="34E8813DAB4A986B61E5216D1FF82C99" originalValue="Prodanović">Prodanovic</normalizedToken>
et al. 2016
</bibRefCitation>
; Fig.
<figureCitation id="81ADDAA064F8F10935C2805B736147D2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution of Silene isabellae, S. noctiflora (partial) and the taxa of the S. vallesia L. group." figureDoi="10.3897/phytokeys.227.100850.figure4" httpUri="https://binary.pensoft.net/fig/859767" pageId="0" pageNumber="123">4</figureCitation>
). The plant is not known from Albania (according to
<bibRefCitation id="B95389798F710B4E10A895E79503F411" author="Vangjeli, J" journalOrPublisher="Koeltz Botanical Books, Oberreifenberg" pageId="0" pageNumber="123" refId="B26" refString="Vangjeli, J, 2015. Excursion Flora of Albania. Koeltz Botanical Books, Oberreifenberg" title="Excursion Flora of Albania." year="2015">Vangjeli (2015)</bibRefCitation>
,
<bibRefCitation id="56B7D6500500EC74124FE88889C3E7A4" author="Pils, G" journalOrPublisher="Eigenverlag, St. Stefan" pageId="0" pageNumber="123" refId="B21" refString="Pils, G, 2016. Illustrated Flora of Albania. Eigenverlag, St. Stefan" title="Illustrated Flora of Albania." year="2016">Pils (2016)</bibRefCitation>
;
<bibRefCitation id="FA1F83818384E748DF5E494C9D075B25" DOI="https://doi.org/10.11646/phytotaxa.378.1.1" author="Barina, Z" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="123" pagination="1 - 339" refId="B1" refString="Barina, Z, Somogyi, G, Pifko, D, Rakaj, M, 2018. Checklist of vascular plants of Albania. Phytotaxa 378 (1): 1 - 339, DOI: https://doi.org/10.11646/phytotaxa.378.1.1" title="Checklist of vascular plants of Albania." url="https://doi.org/10.11646/phytotaxa.378.1.1" volume="378" year="2018">Barina et al. (2018)</bibRefCitation>
) and there is, thus, no range overlap with
<taxonomicName id="704AF98B4E08B8A2E78C9CBC7C5EFBC8" lsidName="S. isabellae" pageId="0" pageNumber="123" rank="species" species="isabellae">
<emphasis id="8CB2ACC1A138E74C876881BB7ACCFDA3" italics="true" pageId="0" pageNumber="123">S. isabellae</emphasis>
</taxonomicName>
. Moreover,
<taxonomicName id="589AF25214D3C4BD1EDD232FC0A944FF" lsidName="S. noctiflora" pageId="0" pageNumber="123" rank="species" species="noctiflora">
<emphasis id="75955C9D2577CBED795C6965B78F43A3" italics="true" pageId="0" pageNumber="123">S. noctiflora</emphasis>
</taxonomicName>
is a weed usually growing in ruderal places, arable fields, cultivated ground and other synanthropic habitats, usually in lowland areas (
<bibRefCitation id="6F95B08EC59031D7DA1505D08E830CE2" author="Chater, AO" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="123" refId="B7" refString="Chater, AO, Walters, SM, Akeroyd, JR, Wrigley, F, 1993. Silene L. in Flora Europaea 2°edn., vol. 1 Cambridge." title="Silene L. in Flora Europaea 2 ° edn., vol. 1 Cambridge." year="1993">Chater et al. 1993</bibRefCitation>
).
</paragraph>
<caption id="C47C8564B542AAD13D96FBB212D7AD77" doi="10.3897/phytokeys.227.100850.figure1" httpUri="https://binary.pensoft.net/fig/859764" pageId="0" pageNumber="123" start="Figure 1" startId="F1">
<paragraph id="BD8388A2EBD0BB8AA5CC015DC96536E0" pageId="0" pageNumber="123">
<emphasis id="AB47BC0C430FC89696B063A003C06A39" bold="true" pageId="0" pageNumber="123">Figure 1.</emphasis>
<taxonomicName id="2CE7D0002AA5DA3B0C46E7E5D82C81FB" authorityName="Selvi &amp; Bianchi" authorityYear="2023" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Silene isabellae" order="Caryophyllales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="species" species="isabellae">
<emphasis id="EB34D6CE81CB9B2A719CD4FDF0575161" italics="true" pageId="0" pageNumber="123">Silene isabellae</emphasis>
</taxonomicName>
<emphasis id="4423E1845321BD575C9EF72767F139D0" bold="true" pageId="0" pageNumber="123">A</emphasis>
whole plant
<emphasis id="31850DE5CB1DFC9D6E4A3DD0C1524946" bold="true" pageId="0" pageNumber="123">B</emphasis>
flower with calyx and corolla
<emphasis id="CE51ED80426F7F29423EEF1A706B2242" bold="true" pageId="0" pageNumber="123">C</emphasis>
fruiting calyx
<emphasis id="328214CF878A746E0BDBCB6D53CD7E16" bold="true" pageId="0" pageNumber="123">D</emphasis>
capsule and carpophore
<emphasis id="503B0FCE407ADA37587CBE6801D5B042" bold="true" pageId="0" pageNumber="123">E</emphasis>
seed. Original drawing by Laura Vivona.
</paragraph>
</caption>
<caption id="3FB0DE16C863F0DCC5C51993543297BB" doi="10.3897/phytokeys.227.100850.figure2" httpUri="https://binary.pensoft.net/fig/859765" pageId="0" pageNumber="123" start="Figure 2" startId="F2">
<paragraph id="0E205A0B24AC3F7570393B799D865011" pageId="0" pageNumber="123">
<emphasis id="1FF96CB6E0B21BE8AF0E9D40D4DFF52A" bold="true" pageId="0" pageNumber="123">Figure 2.</emphasis>
Field photos of
<taxonomicName id="F9BB2BB590F4D7DACC25C279F474AD04" authorityName="Selvi &amp; Bianchi" authorityYear="2023" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Silene isabellae" order="Caryophyllales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="species" species="isabellae">
<emphasis id="4F6D63C73CF1AE2580615B678C84D5EC" italics="true" pageId="0" pageNumber="123">Silene isabellae</emphasis>
</taxonomicName>
<emphasis id="BCB105964F17731AFB5026FFB1DCDDDE" bold="true" pageId="0" pageNumber="123">A</emphasis>
habitat
<emphasis id="5CC553FE79419F8EF02B7293E06A8219" bold="true" pageId="0" pageNumber="123">B, C</emphasis>
whole plants in natural habitat
<emphasis id="56290090469CDB3FF21D43E3CBDCEB75" bold="true" pageId="0" pageNumber="123">D</emphasis>
inflorescence with flowers and fruiting calyces
<emphasis id="C354DDC4A7EF84A858B4E66D61174C1D" bold="true" pageId="0" pageNumber="123">E</emphasis>
flower
<emphasis id="7FFCF377813378C81B3608EB697DCCBC" bold="true" pageId="0" pageNumber="123">F</emphasis>
basal leaves
<emphasis id="8F4DE51B4D12D6592DBB7B99AAF43993" bold="true" pageId="0" pageNumber="123">G</emphasis>
flowers at late stage
<emphasis id="BDE9E42B3FBFE49550D0CA1ADFD9022C" bold="true" pageId="0" pageNumber="123">H</emphasis>
seed.
</paragraph>
</caption>
<caption id="012F7EC11A85D1AE6D007B6F442D9D77" doi="10.3897/phytokeys.227.100850.figure3" httpUri="https://binary.pensoft.net/fig/859766" pageId="0" pageNumber="123" start="Figure 3" startId="F3">
<paragraph id="0723466BEADE2915A0207727A2C17702" pageId="0" pageNumber="123">
<emphasis id="983486ADE1959863F9F2AC5A6285C7FA" bold="true" pageId="0" pageNumber="123">Figure 3.</emphasis>
Calyx teeth, capsule, and margin of basal leaves of
<taxonomicName id="BBD5E303AB471F736260CB78FCD9DE6E" authorityName="Selvi &amp; Bianchi" authorityYear="2023" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Silene isabellae" order="Caryophyllales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="species" species="isabellae">
<emphasis id="AC6DBDA891F130CA7C68DA9107175D2F" italics="true" pageId="0" pageNumber="123">Silene isabellae</emphasis>
</taxonomicName>
(respectively
<emphasis id="5C309CF125F1D8188A9FD0A93B16A9B2" bold="true" pageId="0" pageNumber="123">A-C</emphasis>
),
<taxonomicName id="F524DBC8273236DBF85138F556BD3F31" lsidName="S. noctiflora" pageId="0" pageNumber="123" rank="species" species="noctiflora">
<emphasis id="7A344EC3EAA797518ADAF91019658BF8" italics="true" pageId="0" pageNumber="123">S. noctiflora</emphasis>
</taxonomicName>
(respectively
<emphasis id="543871E3B251E4601BC9ACC2E6C57648" bold="true" pageId="0" pageNumber="123">D-F</emphasis>
) and
<taxonomicName id="AFCE8B255B3D722531450C283248A625" lsidName="S. vallesia subsp. graminea" pageId="0" pageNumber="123" rank="subSpecies" species="vallesia" subSpecies="graminea">S. vallesia subsp. graminea</taxonomicName>
(respectively
<emphasis id="B2AF6B74A1E8ACD869BC2867FE7D7BA8" bold="true" pageId="0" pageNumber="123">G-I</emphasis>
).
</paragraph>
</caption>
<caption id="7223B7EB4A08B58ECA1944F76B05D730" doi="10.3897/phytokeys.227.100850.figure4" httpUri="https://binary.pensoft.net/fig/859767" pageId="0" pageNumber="123" start="Figure 4" startId="F4">
<paragraph id="EE6B7DB2C6441F3CA3CCD102C9707297" pageId="0" pageNumber="123">
<emphasis id="BBA7FEA29F91A825F2C691575DF78322" bold="true" pageId="0" pageNumber="123">Figure 4.</emphasis>
Distribution of
<taxonomicName id="3BF7AB1918403B3568244A417C14647C" authorityName="Selvi &amp; Bianchi" authorityYear="2023" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Silene isabellae" order="Caryophyllales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="species" species="isabellae">
<emphasis id="FEF6512C2A6C611BB089144F9344F87A" italics="true" pageId="0" pageNumber="123">Silene isabellae</emphasis>
</taxonomicName>
,
<taxonomicName id="2A0E187532592A28114392DB282C1D64" lsidName="S. noctiflora" pageId="0" pageNumber="123" rank="species" species="noctiflora">
<emphasis id="F3048CD85CB75711A18B54A0DBD843C3" italics="true" pageId="0" pageNumber="123">S. noctiflora</emphasis>
</taxonomicName>
(partial) and the taxa of the
<taxonomicName id="F5D5434DC9A13804AFB13314A84593E6" lsidName="S. vallesia" pageId="0" pageNumber="123" rank="species" species="vallesia">
<emphasis id="9AF1D80CBC0807574879B6A25A9CF545" italics="true" pageId="0" pageNumber="123">S. vallesia</emphasis>
</taxonomicName>
L. group.
</paragraph>
</caption>
<caption id="41CAFFFD02350AC5C1EAAFBE123462F1" ID-Table-UUID="41CAFFFD02350AC5C1EAAFBE123462F1" httpUri="http://table.plazi.org/id/41CAFFFD02350AC5C1EAAFBE123462F1" pageId="0" pageNumber="123" start="Table 1" startId="T1">
<paragraph id="CF327811F162EDB93B4AB1E907E8F2D8" pageId="0" pageNumber="123">
<emphasis id="8149AF7B1BEF9AE162C1B7CC30629C99" bold="true" pageId="0" pageNumber="123">Table 1.</emphasis>
Main diagnostic morphological characters of
<taxonomicName id="7CD2E7365C0BDC8DF37D1C4028A65F60" lsidName="S. isabellae" pageId="0" pageNumber="123" rank="species" species="isabellae">
<emphasis id="57B511452234470E4C0EAC58835F9170" italics="true" pageId="0" pageNumber="123">S. isabellae</emphasis>
</taxonomicName>
,
<taxonomicName id="EEA2EB160B0B050D38711A901DFD8ECE" lsidName="S. noctiflora" pageId="0" pageNumber="123" rank="species" species="noctiflora">
<emphasis id="81458F67C59E773C9D9D7D107D81B799" italics="true" pageId="0" pageNumber="123">S. noctiflora</emphasis>
</taxonomicName>
and
<taxonomicName id="4DF95DA680E5002331AE639486A47D04" lsidName="S. vallesia subsp. graminea" pageId="0" pageNumber="123" rank="subSpecies" species="vallesia" subSpecies="graminea">S. vallesia subsp. graminea</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="DD1DC0A79926C780005842DD44E2561D" pageId="0" pageNumber="123">
<table id="D4EB6549E8819C0BAF41E3D058812093" pageId="0" pageNumber="123">
<tr id="24F1254C5B79D92BE1624D27DF1B6D81" pageId="0" pageNumber="123">
<th id="650754A588C3CF82423F1086B08264D1" colspan="1" isEmpty="true" pageId="0" pageNumber="123" rowspan="1">-</th>
<th id="8113F91674A1EB143C764BE51B14D6D2" colspan="1" pageId="0" pageNumber="123" rowspan="1">
<taxonomicName id="15470C36BD0EA5432464025F8CB31A22" authorityName="Selvi &amp; Bianchi" authorityYear="2023" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Silene isabellae" order="Caryophyllales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="species" species="isabellae">
<emphasis id="38B78A8B2ADE800A77B7DB9D70A798DF" italics="true" pageId="0" pageNumber="123">Silene isabellae</emphasis>
</taxonomicName>
</th>
<th id="F097EE8E3AB775C727ED116BE0638FFA" colspan="1" pageId="0" pageNumber="123" rowspan="1">
<taxonomicName id="5DB1B2C685A073EEA37D0984D06FF87E" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Silene noctiflora" order="Caryophyllales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="species" species="noctiflora">
<emphasis id="97C7D155F295FD37E5C48207B197A396" italics="true" pageId="0" pageNumber="123">Silene noctiflora</emphasis>
</taxonomicName>
</th>
<th id="53B7D5312DEC066B1BD9D1482CCDDC83" colspan="1" pageId="0" pageNumber="123" rowspan="1">
<taxonomicName id="119825CFA7B90A39A8A274F3333056DE" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Silene vallesia subsp. graminea" order="Caryophyllales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="subSpecies" species="vallesia" subSpecies="graminea">Silene vallesia subsp. graminea</taxonomicName>
</th>
</tr>
<tr id="17125758098272C15085ABD68C049884" pageId="0" pageNumber="123">
<td id="F65250C5053C6683A5531038144AA147" colspan="1" pageId="0" pageNumber="123" rowspan="1">
<emphasis id="20E9BF32318BF1EB6B6D3DF8A9E1B867" bold="true" pageId="0" pageNumber="123">Habit and life cycle</emphasis>
</td>
<td id="2D333E097F0CE3C7E156607507D1513F" colspan="1" pageId="0" pageNumber="123" rowspan="1">perennial, mat-forming</td>
<td id="7D3F77B5247D10D71F5AE57BF5616654" colspan="1" pageId="0" pageNumber="123" rowspan="1">annual, not mat-forming</td>
<td id="5673CF00872640BC5E6E8BE9CEF9D749" colspan="1" pageId="0" pageNumber="123" rowspan="1">perennial, mat-forming</td>
</tr>
<tr id="1C7D8C766C5FD91B14CC767EF28C32E0" pageId="0" pageNumber="123">
<td id="2A7C54EC55987C14E0520E8182ED19CF" colspan="1" pageId="0" pageNumber="123" rowspan="1">
<emphasis id="719585620E469855670D8D2A3FB28467" bold="true" pageId="0" pageNumber="123">Flowering stems and branches</emphasis>
</td>
<td id="F8648FD5218D9A7C4E6035EC817FBA39" colspan="1" pageId="0" pageNumber="123" rowspan="1">sparsely and shortly glandular pubescent, 15-40 cm, usually branched, rarely simple, with 1-4 flowers</td>
<td id="93049BCB239D088829BEA71CC4E7B845" colspan="1" pageId="0" pageNumber="123" rowspan="1">densely glandular pubescent, 12-50 cm, usually simple with a single flower, rarely branched with more flowers</td>
<td id="AF8E7C9479055186C9EA299090351756" colspan="1" pageId="0" pageNumber="123" rowspan="1">sparsely pubescent, 5-15 cm, usually simple with a single flower</td>
</tr>
<tr id="DF5786A88C8A84CBA625932CEAD741A1" pageId="0" pageNumber="123">
<td id="D94FEC86322E755C30C923780F2FEBC9" colspan="1" pageId="0" pageNumber="123" rowspan="1">
<emphasis id="0E3513034C7E074EC93BBCBD7A0652CC" bold="true" pageId="0" pageNumber="123">Basal leaves of sterile stems</emphasis>
</td>
<td id="8B9DBBF9A75F8652A52BFF3AFCF6B598" colspan="1" pageId="0" pageNumber="123" rowspan="1">
widely obovate-spathulate, 8-18 mm wide, blade glabrous, but finely and densely ciliate along margins (Fig.
<figureCitation id="D6EBFDA661DEEDAC7B8A1777036FDE36" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Calyx teeth, capsule, and margin of basal leaves of Silene isabellae (respectively A-C), S. noctiflora (respectively D-F) and S. vallesia subsp. graminea (respectively G-I)." figureDoi="10.3897/phytokeys.227.100850.figure3" httpUri="https://binary.pensoft.net/fig/859766" pageId="0" pageNumber="123">3C</figureCitation>
)
</td>
<td id="9DBEBC512EC0FB43F3B583C7CC3653CC" colspan="1" pageId="0" pageNumber="123" rowspan="1">absent</td>
<td id="957FC2ED29DFC099DBC1B073F38BFBF4" colspan="1" pageId="0" pageNumber="123" rowspan="1">
linear 1.5-2 mm wide, blade with few simple hairs (Fig.
<figureCitation id="A7FBA6D6FE8B25C8F0D80D344C4C871B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Calyx teeth, capsule, and margin of basal leaves of Silene isabellae (respectively A-C), S. noctiflora (respectively D-F) and S. vallesia subsp. graminea (respectively G-I)." figureDoi="10.3897/phytokeys.227.100850.figure3" httpUri="https://binary.pensoft.net/fig/859766" pageId="0" pageNumber="123">3I</figureCitation>
)
</td>
</tr>
<tr id="974759DCA1797BE1951729E0B0B900DF" pageId="0" pageNumber="123">
<td id="9757AD40EEFACD4126737B7C25C642B6" colspan="1" pageId="0" pageNumber="123" rowspan="1">
<emphasis id="F7FE0F5A148613BBCA6CA61B660BD441" bold="true" pageId="0" pageNumber="123">Cauline leaves of flowering stems</emphasis>
</td>
<td id="DE4A7CF5CAD1DDAF6ABA8B2FABC3FA1D" colspan="1" pageId="0" pageNumber="123" rowspan="1">linear-lanceolate, 3-5 mm wide, shorter than internodes, shortly ciliate along the margins, blade with short glandular and simple hairs</td>
<td id="3867F6B00BFD346C4F2DE73C5B775119" colspan="1" pageId="0" pageNumber="123" rowspan="1">
ovate-lanceolate, up to 30 mm wide, blade and margins densely pubescent (Fig.
<figureCitation id="8B84C9F21F94C769395EFD1603325911" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Calyx teeth, capsule, and margin of basal leaves of Silene isabellae (respectively A-C), S. noctiflora (respectively D-F) and S. vallesia subsp. graminea (respectively G-I)." figureDoi="10.3897/phytokeys.227.100850.figure3" httpUri="https://binary.pensoft.net/fig/859766" pageId="0" pageNumber="123">3F</figureCitation>
)
</td>
<td id="F769A42A8C5F79462704A810FC6CBFF4" colspan="1" pageId="0" pageNumber="123" rowspan="1">linear-lanceolate, 1.5-2 mm wide, subequal or longer than internodes, marginally ciliate at the base, with simple and few glandular hairs</td>
</tr>
<tr id="CFACB678BE596C9CE7F731CDE0FD1294" pageId="0" pageNumber="123">
<td id="76BB26FCF53597F785417392EDCDB659" colspan="1" pageId="0" pageNumber="123" rowspan="1">
<emphasis id="BB2CAC5F5E46B9A80EB81E516012B3FC" bold="true" pageId="0" pageNumber="123">Flowers</emphasis>
</td>
<td id="D408775312C33D2AC5107FCDD44BE8C7" colspan="1" pageId="0" pageNumber="123" rowspan="1">
calyx 25-30 mm, teeth 8 mm, narrowly cylindrical in flower, truncate-obconical in fruit and prominently ribbed (Fig.
<figureCitation id="3CF97D2511AD454DBE60BE81B051A922" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Calyx teeth, capsule, and margin of basal leaves of Silene isabellae (respectively A-C), S. noctiflora (respectively D-F) and S. vallesia subsp. graminea (respectively G-I)." figureDoi="10.3897/phytokeys.227.100850.figure3" httpUri="https://binary.pensoft.net/fig/859766" pageId="0" pageNumber="123">3A</figureCitation>
); corolla unscented, open during daytime; petals not enrolled during daytime pale pink above, tinged with mauve red beneath, lobes dentate
</td>
<td id="1F9ECD9932D6977B1370513DAC27BC4C" colspan="1" pageId="0" pageNumber="123" rowspan="1">
calyx 20-30 mm, teeth ca. 7 mm (Fig.
<figureCitation id="AB64DDAB7ECE95A65C756A959F63DE9D" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Calyx teeth, capsule, and margin of basal leaves of Silene isabellae (respectively A-C), S. noctiflora (respectively D-F) and S. vallesia subsp. graminea (respectively G-I)." figureDoi="10.3897/phytokeys.227.100850.figure3" httpUri="https://binary.pensoft.net/fig/859766" pageId="0" pageNumber="123">3D</figureCitation>
); attenuate at the base in fruit, not ribbed; corolla opening in evening with petals enrolled during daytime, pink above and yellowish beneath, lobes entire
</td>
<td id="B9A297F04DB7CDC8C3EB2B9BD359A4A8" colspan="1" pageId="0" pageNumber="123" rowspan="1">
calyx 12-16 mm, teeth ca. 1.8 mm, narrowly obconical and not ribbed in fruit (Fig.
<figureCitation id="5FED92AD0D3FE7053EC2B4823C6CAF13" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Calyx teeth, capsule, and margin of basal leaves of Silene isabellae (respectively A-C), S. noctiflora (respectively D-F) and S. vallesia subsp. graminea (respectively G-I)." figureDoi="10.3897/phytokeys.227.100850.figure3" httpUri="https://binary.pensoft.net/fig/859766" pageId="0" pageNumber="123">3G</figureCitation>
); corolla unscented, open during daytime, petals usually white, lobes entire
</td>
</tr>
<tr id="9EC402E39445DDCD8916951E27189C0D" pageId="0" pageNumber="123">
<td id="2F1CE2E878F191985F13B25E17AC08A3" colspan="1" pageId="0" pageNumber="123" rowspan="1">
<emphasis id="8D754A45F430A01409D2713AD5C5416E" bold="true" pageId="0" pageNumber="123">Fruits</emphasis>
</td>
<td id="C55D23F77BE66AE5C03DDAAEDEF7F758" colspan="1" pageId="0" pageNumber="123" rowspan="1">
capsule conic-ovoid, 13-15
<normalizedToken id="B0C78B12978E4A73D5268C0751560736" originalValue="×">x</normalizedToken>
5 mm, surface smooth, glossy (Fig.
<figureCitation id="2CAC474A6357EE418C5677C1C63F6A61" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Calyx teeth, capsule, and margin of basal leaves of Silene isabellae (respectively A-C), S. noctiflora (respectively D-F) and S. vallesia subsp. graminea (respectively G-I)." figureDoi="10.3897/phytokeys.227.100850.figure3" httpUri="https://binary.pensoft.net/fig/859766" pageId="0" pageNumber="123">3B</figureCitation>
); carpophore 6-7 mm
</td>
<td id="31DA76E57F7CC01E5D66D7A9581F01BC" colspan="1" pageId="0" pageNumber="123" rowspan="1">
capsule ovoid 13-15
<normalizedToken id="429AA0994638A14BD3F32872D62E28AB" originalValue="×">x</normalizedToken>
8-10 mm, glossy (Fig.
<figureCitation id="9D21F0CF035128A9AFF60D8BB5953599" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Calyx teeth, capsule, and margin of basal leaves of Silene isabellae (respectively A-C), S. noctiflora (respectively D-F) and S. vallesia subsp. graminea (respectively G-I)." figureDoi="10.3897/phytokeys.227.100850.figure3" httpUri="https://binary.pensoft.net/fig/859766" pageId="0" pageNumber="123">3E</figureCitation>
); carpophore 2-3 mm
</td>
<td id="2866096D8665C644BC92F676FA36047A" colspan="1" pageId="0" pageNumber="123" rowspan="1">
capsule conic-ovoid 12-13
<normalizedToken id="332B4F2253C77BCD312CC62DABABA8B4" originalValue="×">x</normalizedToken>
ca. 4 mm, glabrous, but finely rugose (Fig.
<figureCitation id="1E15C2D86E1298426EB5C3866C24EAF1" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Calyx teeth, capsule, and margin of basal leaves of Silene isabellae (respectively A-C), S. noctiflora (respectively D-F) and S. vallesia subsp. graminea (respectively G-I)." figureDoi="10.3897/phytokeys.227.100850.figure3" httpUri="https://binary.pensoft.net/fig/859766" pageId="0" pageNumber="123">3H</figureCitation>
); carpophore 9 mm
</td>
</tr>
</table>
</paragraph>
<paragraph id="C36170F7C0449D1E7D701758A6BB7A11" pageId="0" pageNumber="123">
<taxonomicName id="CEA4F44277E8420B7D7543CC43A95B9E" authorityName="Selvi &amp; Bianchi" authorityYear="2023" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Silene isabellae" order="Caryophyllales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="species" species="isabellae">
<emphasis id="AE9E4247BCB469E6DF93013CAB0E3609" italics="true" pageId="0" pageNumber="123">Silene isabellae</emphasis>
</taxonomicName>
shows a weak morphological affinity also with the taxa of the
<taxonomicName id="E55D08BBEB89A710DAE02A0707B2DC75" lsidName="S. vallesia" pageId="0" pageNumber="123" rank="species" species="vallesia">
<emphasis id="F12E4CB4EDA597069FFB7A6F8614B0B7" italics="true" pageId="0" pageNumber="123">S. vallesia</emphasis>
</taxonomicName>
group [sect.
<taxonomicName id="4F4EC56FFF684C3B15AEDB64408C2C70" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Silene" order="Caryophyllales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="section" section="Auriculatae">Silene Auriculatae</taxonomicName>
(Boiss.) Schischk.]. This group includes taxa endemic to some mountain regions of southern Europe (Fig.
<figureCitation id="9FF4B35A9C3EA6D78896DAFFEE33BD26" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution of Silene isabellae, S. noctiflora (partial) and the taxa of the S. vallesia L. group." figureDoi="10.3897/phytokeys.227.100850.figure4" httpUri="https://binary.pensoft.net/fig/859767" pageId="0" pageNumber="123">4</figureCitation>
), in France (
<taxonomicName id="D5F217DD4540CCDB7F88200E8831D1C0" lsidName="S. petrarchae" pageId="0" pageNumber="123" rank="species" species="petrarchae">
<emphasis id="A1A9262E5078DBCAB2685E2669BE0D93" italics="true" pageId="0" pageNumber="123">S. petrarchae</emphasis>
</taxonomicName>
Ferrarini &amp; Cecchi), the western Alps (
<taxonomicName id="4307171156CF3021D635D533EAB63FC3" authority="L. subsp. vallesia" lsidName="S. vallesia subsp. vallesia" pageId="0" pageNumber="123" rank="subSpecies" species="vallesia" subSpecies="vallesia">S. vallesia L. subsp. vallesia</taxonomicName>
), the Apuan Alps in Italy (
<taxonomicName id="00853612A22CD1A4D59BAA79F775BEB1" lsidName="S. pichiana" pageId="0" pageNumber="123" rank="species" species="pichiana">
<emphasis id="22B170AD8E2A305DE5987E365C0620DF" italics="true" pageId="0" pageNumber="123">S. pichiana</emphasis>
</taxonomicName>
Ferrarini &amp; Cecchi), the central Apennines (
<taxonomicName id="FF256B6D5D383C63060FEF28A200080F" lsidName="S. cattariniana" pageId="0" pageNumber="123" rank="species" species="cattariniana">
<emphasis id="0F6B637162B2EE565509DB63DB175598" italics="true" pageId="0" pageNumber="123">S. cattariniana</emphasis>
</taxonomicName>
Ferrarini &amp; Cecchi) and the Balkan massifs [
<taxonomicName id="C5CCEA27A70836731772FC90CA5AA6F3" lsidName="S. vallesia subsp. graminifolia" pageId="0" pageNumber="123" rank="subSpecies" species="vallesia" subSpecies="graminifolia">S. vallesia subsp. graminifolia</taxonomicName>
(Vis ex. Rchb.) Nyman,
<bibRefCitation id="09353D2D75F306ED2408E204B9B2B413" DOI="https://doi.org/10.1080/00837792.2001.10670717" author="Ferrarini, E" journalOrPublisher="Webbia" pageId="0" pageNumber="123" pagination="241 - 263" refId="B11" refString="Ferrarini, E, Cecchi, O, 2001. Nuove specie del genere &quot; Silene &quot; (Caryophyllaceae) delle Alpi Apuane, dell'Appennino centrale (Italia) e della Francia meridionale. Webbia 56 (2): 241 - 263, DOI: https://doi.org/10.1080/00837792.2001.10670717" title="Nuove specie del genere &quot; Silene &quot; (Caryophyllaceae) delle Alpi Apuane, dell'Appennino centrale (Italia) e della Francia meridionale." url="https://doi.org/10.1080/00837792.2001.10670717" volume="56" year="2001">Ferrarini and Cecchi 2001</bibRefCitation>
]. Similarities are in the perennial habit with stoloniferous stems forming mats, the glandular-pubescent stems and leaves, the few-flowered inflorescence, the large flowers with narrowly cylindrical and reticulate-veined calyx, truncate at base, the corolla with bifid petals, the conical-ovoid capsule and the reniform seeds with a similar surface ornamentation. However, the only taxon of this group occurring in the Balkans and Albania,
<taxonomicName id="6C464531607E8949B79D8D20A72C9357" lsidName="S. vallesia subsp. graminea" pageId="0" pageNumber="123" rank="subSpecies" species="vallesia" subSpecies="graminea">S. vallesia subsp. graminea</taxonomicName>
, is strongly differentiated from
<taxonomicName id="777D61E7564DAE5B6980E6A279A20247" lsidName="S. isabellae" pageId="0" pageNumber="123" rank="species" species="isabellae">
<emphasis id="613AC2915B1CAD38AEFB3ED7D3937646" italics="true" pageId="0" pageNumber="123">S. isabellae</emphasis>
</taxonomicName>
in the much smaller size of the plant, the shape of the leaves (narrowly linear-lanceolate), the usually single flower per stem and other features of the calyx teeth and capsule (Table
<tableCitation id="4D2EFF11A2AE1C32D5D1728C19EACD6A" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Main diagnostic morphological characters of S. isabellae, S. noctiflora and S. vallesia subsp. graminea." httpUri="http://table.plazi.org/id/41CAFFFD02350AC5C1EAAFBE123462F1" pageId="0" pageNumber="123" tableUuid="41CAFFFD02350AC5C1EAAFBE123462F1">1</tableCitation>
, Fig.
<figureCitation id="B863917C812EA0AA806B5C61A246BA4A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Calyx teeth, capsule, and margin of basal leaves of Silene isabellae (respectively A-C), S. noctiflora (respectively D-F) and S. vallesia subsp. graminea (respectively G-I)." figureDoi="10.3897/phytokeys.227.100850.figure3" httpUri="https://binary.pensoft.net/fig/859766" pageId="0" pageNumber="123">3G</figureCitation>
). Moreover, this taxon is only found in the Alpine massifs of the north-east of the country in the district of
<normalizedToken id="3C5C732FA4B432AEFC04949101299EDD" originalValue="Kükes">Kuekes</normalizedToken>
and close to the border with Kosovo and Macedonia, such as Mt. Pashtrik, Mt. Gjallica and Mt. Koritnik, where it reaches the southern limit of its distribution. This taxon is, thus, likely to be allopatric and geographically isolated with respect to
<taxonomicName id="9DE3B6B2E7E4CEE9EF37895063F82B10" lsidName="S. isabellae" pageId="0" pageNumber="123" rank="species" species="isabellae">
<emphasis id="C1D8D2407ED0169D539305D598ED1E0D" italics="true" pageId="0" pageNumber="123">S. isabellae</emphasis>
</taxonomicName>
, which is found over 80 km to the southwest (Fig.
<figureCitation id="F31FB7A349DB6D0B42753EDCEC5BE274" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Distribution of Silene isabellae, S. noctiflora (partial) and the taxa of the S. vallesia L. group." figureDoi="10.3897/phytokeys.227.100850.figure4" httpUri="https://binary.pensoft.net/fig/859767" pageId="0" pageNumber="123">4</figureCitation>
). Based on
<bibRefCitation id="11DFD3A469001ACE56E2AD0076D0C5F3" author="Vangjeli, J" journalOrPublisher="Koeltz Botanical Books, Oberreifenberg" pageId="0" pageNumber="123" refId="B26" refString="Vangjeli, J, 2015. Excursion Flora of Albania. Koeltz Botanical Books, Oberreifenberg" title="Excursion Flora of Albania." year="2015">Vangjeli (2015)</bibRefCitation>
,
<taxonomicName id="6C6BE6CF44B80D73D0A5BBEE35FB6682" lsidName="S. vallesia subsp. graminea" pageId="0" pageNumber="123" rank="subSpecies" species="vallesia" subSpecies="graminea">S. vallesia subsp. graminea</taxonomicName>
is found in the alpine belt at 1700-2000 m a.s.l., thus, at higher elevation than
<taxonomicName id="2DE7FC29573F00114573F7AB2E3EF285" lsidName="S. isabellae" pageId="0" pageNumber="123" rank="species" species="isabellae">
<emphasis id="9032F57A4ABF4FAE74DCE89860248112" italics="true" pageId="0" pageNumber="123">S. isabellae</emphasis>
</taxonomicName>
and is typical of calcareous substrates especially limestone (
<bibRefCitation id="2F62BC1EAF9F60A2D71C6939F5AC7559" DOI="https://doi.org/10.1080/00837792.2001.10670717" author="Ferrarini, E" journalOrPublisher="Webbia" pageId="0" pageNumber="123" pagination="241 - 263" refId="B11" refString="Ferrarini, E, Cecchi, O, 2001. Nuove specie del genere &quot; Silene &quot; (Caryophyllaceae) delle Alpi Apuane, dell'Appennino centrale (Italia) e della Francia meridionale. Webbia 56 (2): 241 - 263, DOI: https://doi.org/10.1080/00837792.2001.10670717" title="Nuove specie del genere &quot; Silene &quot; (Caryophyllaceae) delle Alpi Apuane, dell'Appennino centrale (Italia) e della Francia meridionale." url="https://doi.org/10.1080/00837792.2001.10670717" volume="56" year="2001">Ferrarini and Cecchi 2001</bibRefCitation>
;
<bibRefCitation id="450A401EF58685C0D5C6D56905767F3F" author="Pils, G" journalOrPublisher="Eigenverlag, St. Stefan" pageId="0" pageNumber="123" refId="B21" refString="Pils, G, 2016. Illustrated Flora of Albania. Eigenverlag, St. Stefan" title="Illustrated Flora of Albania." year="2016">Pils 2016</bibRefCitation>
).
</paragraph>
<paragraph id="76C88BB26A4DE01406104DC91C105CAB" pageId="0" pageNumber="123">
<taxonomicName id="7773988944B40D42275F027C0BAD676C" authorityName="Selvi &amp; Bianchi" authorityYear="2023" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Silene isabellae" order="Caryophyllales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="species" species="isabellae">
<emphasis id="06F51055C49B78839FF3E4416F29CCDF" italics="true" pageId="0" pageNumber="123">Silene isabellae</emphasis>
</taxonomicName>
is apparently found only on serpentine soils. Thanks to this specialisation, the plant can be categorised as a
<normalizedToken id="39B8998750D72C8F28F1B21A5896C853" originalValue="“metallophyte”">&quot;metallophyte&quot;</normalizedToken>
, being able to cope with the typical chemical and physical anomalies of the serpentine soils and especially the poverty in nutrients, the high levels of Mg and in trace metals like Ni, Zn and Cr (
<bibRefCitation id="46722E999917340823D910E269A4431B" DOI="https://doi.org/10.1016/bs.abr.2016.12.002" author="Wojcik, M" journalOrPublisher="Advances in Botanical Research" pageId="0" pageNumber="123" pagination="1 - 42" refId="B27" refString="Wojcik, M, Gonnelli, C, Selvi, F, Dresler, S, Rostanski, A, Vangronsveld, J, 2017. Metallophytes of Serpentine and Calamine Soils - Their Unique Ecophysiology and Potential for Phytoremediation. Advances in Botanical Research 83: 1 - 42, DOI: https://doi.org/10.1016/bs.abr.2016.12.002" title="Metallophytes of Serpentine and Calamine Soils - Their Unique Ecophysiology and Potential for Phytoremediation." url="https://doi.org/10.1016/bs.abr.2016.12.002" volume="83" year="2017">
<normalizedToken id="844C98F74CB5328D1FC125F3929169A1" originalValue="Wójcik">Wojcik</normalizedToken>
et al. 2017
</bibRefCitation>
). We plan further studies to assess the concentration of these metals within the plant and, consequently, the strategies that it adopts to thrive on such hostile soils. Adaptation to serpentine soils is not uncommon in the genus
<taxonomicName id="7D759BCC7DE4E80266BA21756119435D" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Silene" order="Caryophyllales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="genus">
<emphasis id="3EAA50D83C1DBA65B553F9E07541C2AB" italics="true" pageId="0" pageNumber="123">Silene</emphasis>
</taxonomicName>
, being found in several other taxa of especially the Balkan flora. According to
<bibRefCitation id="9E4B1192F7639464452966F121FF00CE" DOI="https://doi.org/10.1007/s00606-003-0044-8" author="Stefanovic, V" journalOrPublisher="Plant Systematics and Evolution" pageId="0" pageNumber="123" pagination="149 - 170" refId="B24" refString="Stefanovic, V, Tan, K, Iatrou, G, 2003. Distribution of the endemic Balkan fora on serpentine I.-Obligate serpentine endemics. Plant Systematics and Evolution 242 (1-4): 149 - 170, DOI: https://doi.org/10.1007/s00606-003-0044-8" title="Distribution of the endemic Balkan fora on serpentine I. - Obligate serpentine endemics." url="https://doi.org/10.1007/s00606-003-0044-8" volume="242" year="2003">
<normalizedToken id="7BA824B890EFAD7E9DFEBA7A28B7BC1F" originalValue="Stefanović">Stefanovic</normalizedToken>
et al. (2003)
</bibRefCitation>
, the Balkan countries host seven taxa of this genus that can be classified as obligate serpentine endemics, while others are found on serpentine as well as non-serpentine soils (the so-called &quot;facultative serpentinophytes&quot;). The distant phylogenetic position of these species suggests that ecological speciation driven by the &quot;serpentine factor&quot; has occurred through multiple and independent events, as in other plant groups of the Mediterranean and Balkan flora (
<bibRefCitation id="DAFFB002B38FC364E0DD59D93836C4AB" DOI="https://doi.org/10.1002/tax.583002" author="Cecchi, L" journalOrPublisher="Taxon" pageId="0" pageNumber="123" pagination="700 - 714" refId="B2" refString="Cecchi, L, Selvi, F, 2009. Phylogenetic relationships of the monotypic genera Halacsya and Paramoltkia and the origins of serpentine adaptation in circum-mediterranean Lithospermeae (Boraginaceae): Insights from ITS and matK DNA sequences. Taxon 58 (3): 700 - 714, DOI: https://doi.org/10.1002/tax.583002" title="Phylogenetic relationships of the monotypic genera Halacsya and Paramoltkia and the origins of serpentine adaptation in circum-mediterranean Lithospermeae (Boraginaceae): Insights from ITS and matK DNA sequences." url="https://doi.org/10.1002/tax.583002" volume="58" year="2009">Cecchi and Selvi 2009</bibRefCitation>
;
<bibRefCitation id="9FF0225264FA70B496B5C1A7B106F6B6" DOI="https://doi.org/10.1007/s00606-011-0506-3" author="Cecchi, L" journalOrPublisher="Plant Systematics and Evolution" pageId="0" pageNumber="123" pagination="185 - 199" refId="B4" refString="Cecchi, L, Coppi, A, Selvi, F, 2011. Evolutionary dynamics of serpentine adaptation in Onosma (Boraginaceae) as revealed by ITS sequence data. Plant Systematics and Evolution 297 (3-4): 185 - 199, DOI: https://doi.org/10.1007/s00606-011-0506-3" title="Evolutionary dynamics of serpentine adaptation in Onosma (Boraginaceae) as revealed by ITS sequence data." url="https://doi.org/10.1007/s00606-011-0506-3" volume="297" year="2011">Cecchi et al. 2011</bibRefCitation>
,
<bibRefCitation id="A21AD0746694754AF48BA1ECED6F7D8F" DOI="https://doi.org/10.1111/boj.12084" author="Cecchi, L" journalOrPublisher="Botanical Journal of the Linnean Society" pageId="0" pageNumber="123" pagination="269 - 289" refId="B5" refString="Cecchi, L, Colzi, I, Coppi, A, Gonnelli, C, Selvi, F, 2013. Diversity and biogeography of Ni-hyperaccumulators of Alyssum section Odontarrhena (Brassicaceae) in the central western Mediterranean: Evidence from karyology, morphology and DNA sequence data. Botanical Journal of the Linnean Society 173 (2): 269 - 289, DOI: https://doi.org/10.1111/boj.12084" title="Diversity and biogeography of Ni-hyperaccumulators of Alyssum section Odontarrhena (Brassicaceae) in the central western Mediterranean: Evidence from karyology, morphology and DNA sequence data." url="https://doi.org/10.1111/boj.12084" volume="173" year="2013">2013</bibRefCitation>
). Remarkably, some of the serpentine-tolerant species were also found at the same locality of
<taxonomicName id="0593C960721B91B582A7A3F924EC3469" lsidName="S. isabellae" pageId="0" pageNumber="123" rank="species" species="isabellae">
<emphasis id="BA65E863932285193191575D0189CBD8" italics="true" pageId="0" pageNumber="123">S. isabellae</emphasis>
</taxonomicName>
, such as
<taxonomicName id="5F1EC937AEED898805B4E01907393E89" lsidName="S. paradoxa" pageId="0" pageNumber="123" rank="species" species="paradoxa">
<emphasis id="D70AAEC6BE871245631831DFDB7A0239" italics="true" pageId="0" pageNumber="123">S. paradoxa</emphasis>
</taxonomicName>
L.,
<taxonomicName id="1562D246610E4B5EB1F18887F04EF690" lsidName="S. italica" pageId="0" pageNumber="123" rank="species" species="italica">
<emphasis id="32194B4FCBDDB8F7B2779DDAA48E560A" italics="true" pageId="0" pageNumber="123">S. italica</emphasis>
</taxonomicName>
L.,
<taxonomicName id="299E68499828CAA6A7C8D32A1992FD19" lsidName="S. saxifraga" pageId="0" pageNumber="123" rank="species" species="saxifraga">
<emphasis id="FD399AB85805EC8FFA65D04FA3AE0E73" italics="true" pageId="0" pageNumber="123">S. saxifraga</emphasis>
</taxonomicName>
L.,
<taxonomicName id="510AE82DFDFB5111CBEAFEC09B8B4DAE" lsidName="S. vulgaris subsp. prostrata" pageId="0" pageNumber="123" rank="subSpecies" species="vulgaris" subSpecies="prostrata">S. vulgaris L. subsp. prostrata</taxonomicName>
(Gaud.) Schinz &amp; Thell. and
<taxonomicName id="F5BB4D2C7B1B9A90348720131BBFC823" lsidName="S. damboldtiana" pageId="0" pageNumber="123" rank="species" species="damboldtiana">
<emphasis id="9C0E6762C4F09407D6CBFDDFF46665C2" italics="true" pageId="0" pageNumber="123">S. damboldtiana</emphasis>
</taxonomicName>
Greuter.
</paragraph>
<paragraph id="0E3058FE065129D789B46DA18BC430B5" pageId="0" pageNumber="123">
<taxonomicName id="0EA6F72BB1A07B8179E8AA0CD9CDA9B6" authorityName="Selvi &amp; Bianchi" authorityYear="2023" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Silene isabellae" order="Caryophyllales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="species" species="isabellae">
<emphasis id="E9E1E3DBB3EA38F8DC15DEC51F188C2D" italics="true" pageId="0" pageNumber="123">Silene isabellae</emphasis>
</taxonomicName>
shows some variation in quantitative characters associated with site conditions. Plants from the coniferous forest at lower altitude (1000-1200 m a.s.l.) were characterised by taller flowering stems and larger leaves, in particular those of the sterile stems. In plants from open and rocky sites, we observed a general reduction in the size of the plants, also associated with altitude (above 1400 m a.s.l.), while qualitative characters remained constant.
</paragraph>
</subSubSection>
<subSubSection id="77118D656AAF7A25BD405C95096AC825" pageId="0" pageNumber="123" type="materials_examined">
<paragraph id="FEDC1766295A6B4BB9C4F7D924AF2091" pageId="0" pageNumber="123">Additional specimens examined.</paragraph>
<paragraph id="CDF53BD667AA2EFB6967C1B0B1306C4C" pageId="0" pageNumber="123">
<taxonomicName id="015E663AE60B123B5469B8E93DE28E42" authorityName="Selvi &amp; Bianchi" authorityYear="2023" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Silene isabellae" order="Caryophyllales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="species" species="isabellae">
<emphasis id="5E925E48B7A5712DFA5F43ABCE1492A3" bold="true" italics="true" pageId="0" pageNumber="123">Silene isabellae</emphasis>
</taxonomicName>
:
<materialsCitation id="43CD5895328196D3166569339C7CE1F4" accessionNumber="FI053966, FI068202" collectingDate="2018-06-30" collectingDateMax="2022-07-12" collectingDateMin="2018-06-30" collectorName="F. Selvi, I. Bettarini, E. Bianchi, I. Colzi, A. Coppi" country="Albania" county="District of Kruje" elevation="1100" latitude="41.5251" location="District of Kruje" longLatPrecision="6" longitude="19.5251" municipality="Qafe Shtame" specimenCount="1">
<emphasis id="4B8F19BD4A60EE4B4CDE01A310D51CE2" bold="true" pageId="0" pageNumber="123">
<collectingCountry id="F4C04CC857FC83448E723F2DC05E699F" name="Albania">Albania</collectingCountry>
.
</emphasis>
<location id="8C55F3980B28E5C56B4376DB2F7550FC" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:8C55F3980B28E5C56B4376DB2F7550FC" country="Albania" county="District of Kruje" latitude="41.5251" longLatPrecision="6" longitude="19.5251" municipality="Qafe Shtame" name="District of Kruje">
District of
<normalizedToken id="5FDBB3EBF4F9BF28616658FB9BC44257" originalValue="Krujë">Kruje</normalizedToken>
</location>
,
<collectingMunicipality id="EED2E9305D1369F9A921645B12BF634F">
<normalizedToken id="3789AD5917403C7625FA3DA4DCE4CAFD" originalValue="Qafë">Qafe</normalizedToken>
<normalizedToken id="92ABFFA276F575AA40855A6E8AA08A26" originalValue="Shtamë">Shtame</normalizedToken>
</collectingMunicipality>
, rocce di serpentino nella foresta di
<taxonomicName id="2092EB4AA60F797683E6E7C0C5ACBBF0" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" lsidName="Pinus nigra" order="Pinales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="species" species="nigra">
<emphasis id="C3C27B8628F099A6A387EB07B12A0CD0" italics="true" pageId="0" pageNumber="123">Pinus nigra</emphasis>
</taxonomicName>
, ca. 800 prima del passo,
<geoCoordinate id="DA5230596318B5C61BB7A01FD7814A75" degrees="41" direction="north" minutes="31.35" orientation="latitude" precision="9" value="41.5225">41°31.35'N</geoCoordinate>
,
<geoCoordinate id="560AED50ECBF003BA1F2DF3303031AF1" degrees="19" direction="east" minutes="53.32" orientation="longitude" precision="9" value="19.888666">19°53.32'E</geoCoordinate>
,
<quantity id="A0847AF5BED9325BEB03C2E6957EC73E" metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">
<elevation id="4E6E8FBAE7B00BF97218C12617E35CA1" metricMagnitude="3" metricUnit="m" metricValue="1.1" unit="m" value="1100.0">1100 m</elevation>
</quantity>
elev.,
<collectingDate id="25354872A1EB185B88E6BF26DD5101BD" value="2018-06-30">30 Jun 2018</collectingDate>
,
<emphasis id="5A8E760005D518D9B205D221ADC8AF36" italics="true" pageId="0" pageNumber="123">
<collectorName id="CE487A6BE46AB2D41FE49BADECB7F043">F. Selvi</collectorName>
&amp;
<collectorName id="6653F9E9ADA3922CC265D1723D7E4C46">I. Bettarini</collectorName>
</emphasis>
(
<accessionNumber id="7AB34E861532B1D12F723D379EDDACA1" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/FI053966">FI053966</accessionNumber>
);
<collectingCounty id="0D24ED3F850327AB9D991D9689ABD050">
District of
<normalizedToken id="C1634689E8072AABC449E156E015DE3C" originalValue="Krujë">Kruje</normalizedToken>
</collectingCounty>
, Qafe
<normalizedToken id="8E1D325A92FE22BEA1B99C0A1494D1CD" originalValue="Shtamë">Shtame</normalizedToken>
, pascoli rupestri e margini di boscaglia rocciosa lungo la pista che sale dal passo verso Nord, suolo ultramafico (serpentino),
<geoCoordinate id="660863B921CA1B62B1F4681578B241C7" degrees="41.5251" direction="north" orientation="latitude" precision="5" value="41.5251">41.5251°N</geoCoordinate>
,
<geoCoordinate id="72E59820EABE8771DD87F25B179F2758" degrees="19.5251" direction="east" orientation="longitude" precision="5" value="19.5251">19.5251°E</geoCoordinate>
,
<quantity id="2FA4AF4FDF3A5D04116A1D966013FC63" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.45" metricValueMin="1.15" unit="m" value="1300.0" valueMax="1450.0" valueMin="1150.0">
<elevation id="B88C43C645C12B8B89D2980BDCF0B4A6" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.45" metricValueMin="1.15" unit="m" value="1300.0" valueMax="1450.0" valueMin="1150.0">1150-1450 m</elevation>
</quantity>
elev.,
<collectingDate id="C6D95E65F6F1A409D9880C47DC90E827" value="2022-07-12">12 July 2022</collectingDate>
,
<emphasis id="C76E10B8BE7A90A1E461F8EA0867DE38" italics="true" pageId="0" pageNumber="123">
<collectorName id="49D2E730AB790204C51EE8768D48A65C">F. Selvi</collectorName>
,
<collectorName id="E130572D02C26F6A8DCDFB8994D58BB1">E. Bianchi</collectorName>
,
<collectorName id="CC089D436ED0025CE65E1B36F48A6CFB">I. Colzi</collectorName>
&amp;
<collectorName id="B39BE8A4A42B8892E5CC466B28D3CB78">A. Coppi</collectorName>
</emphasis>
(
<accessionNumber id="133422E42411C4D81C380784A2D96B02" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/FI068202">FI068202</accessionNumber>
)
</materialsCitation>
.
</paragraph>
<paragraph id="886E6FA32F12BB31850A09150EC3A704" pageId="0" pageNumber="123">
<emphasis id="61070A74E7B084473F74B96C7FA89254" bold="true" pageId="0" pageNumber="123">
<taxonomicName id="4BC4C68751EBCF7363F53CEE81CF628E" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Silene noctiflora" order="Caryophyllales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="species" species="noctiflora">
<emphasis id="9DB23ED6513B32F8DAF05B41F6E5A608" italics="true" pageId="0" pageNumber="123">Silene noctiflora</emphasis>
</taxonomicName>
:
<materialsCitation id="206D3C005A9D2DE8C70762FC3BA3C779" collectingDate="1989-11-07" collectingDateMax="2010-09-06" collectingDateMin="1989-11-07" collectorName="auf dem Autobahn-Rastplatz, Helmut M., Leonhartsberger S., Maurer W." country="Austria" county="Kaernten. Lavanttal" elevation="450" latitude="46.725" location="Bezirk Wolfsberg" longLatPrecision="55" longitude="15.45833" municipality="Autobahn-Rastplatz in St. Andrae" specimenCount="1" stateProvince="Steiermark">
<collectingCountry id="184FF68A5AED9220F588709DF88E5420" name="Austria">Austria</collectingCountry>
.
<collectingCounty id="A847789D0C3CDAB2C56C6C4F05AAB419">
<normalizedToken id="34886663AF3228CB5F5691366D42D993" originalValue="Kärnten">Kaernten</normalizedToken>
. Lavanttal
</collectingCounty>
:
<collectorName id="1B6844FBEB9D6D4266A7646E999061A2">
auf dem
<collectingMunicipality id="F3FC1BB322B399F991434FA17D0D95D0">
Autobahn-Rastplatz in St.
<normalizedToken id="23A82F107A85ABE2FD0EAA901B9835A6" originalValue="Andrä">Andrae</normalizedToken>
</collectingMunicipality>
</collectorName>
,
<location id="22DD191D89CFE72D27E846AE5DF4F9A2" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:22DD191D89CFE72D27E846AE5DF4F9A2" country="Austria" county="Kaernten. Lavanttal" latitude="46.725" longLatPrecision="55" longitude="15.45833" municipality="Autobahn-Rastplatz in St. Andrae" name="Bezirk Wolfsberg" stateProvince="Steiermark">Bezirk Wolfsberg</location>
,
<location id="27B541B1CD0D5F20AFD9C349493E2233" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:27B541B1CD0D5F20AFD9C349493E2233" country="Austria" county="Kaernten. Lavanttal" latitude="46.725" longLatPrecision="55" longitude="15.45833" municipality="Autobahn-Rastplatz in St. Andrae" name="Gemeinde Sankt Andrae" stateProvince="Steiermark">
Gemeinde Sankt
<normalizedToken id="2FF0B6A989D5EB2BC43D2941360CF48E" originalValue="Andrä">Andrae</normalizedToken>
</location>
,
<location id="4E509DBDFB84C3F5859D73F05DE894F5" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:4E509DBDFB84C3F5859D73F05DE894F5" country="Austria" county="Kaernten. Lavanttal" latitude="46.725" longLatPrecision="55" longitude="15.45833" municipality="Autobahn-Rastplatz in St. Andrae" name="Katastralgemeinde Kollegg" stateProvince="Steiermark">Katastralgemeinde Kollegg</location>
,
<geoCoordinate id="1831708EC8115AFEF82B02D7E7B9F3D9" degrees="46" direction="north" minutes="47" orientation="latitude" precision="15" seconds="08" value="46.785557">46°47'08&quot;N</geoCoordinate>
,
<geoCoordinate id="A3D54BFE6612EA43C087ACB2AB362704" degrees="14" direction="east" minutes="48" orientation="longitude" precision="15" seconds="46" value="14.8127775">14°48'46&quot;E</geoCoordinate>
,
<quantity id="55779C0FBE49F89B35FDD70ABE3FFDAC" metricMagnitude="2" metricUnit="m" metricValue="4.5" unit="m" value="450.0">
<elevation id="C798B810C4D62B501B95E522376D48D4" metricMagnitude="2" metricUnit="m" metricValue="4.5" unit="m" value="450.0">450 m</elevation>
</quantity>
elev.,
<collectingDate id="9B9B3D8560710533A35F39902607CC9C" value="2008-05-27">27 May 2008</collectingDate>
,
<emphasis id="B13EF0389A59D1556026DA2C30374A69" italics="true" pageId="0" pageNumber="123">
<collectorName id="0F5278B29769D4B6B77E06E510C67ED4">Helmut M.</collectorName>
</emphasis>
s.n. (GJO 0041410);
<normalizedToken id="C69D770DF3B59D7B369E01A9A2AAF432" originalValue="Österreich">Oesterreich</normalizedToken>
,
<collectingRegion id="F12522B2CB6053D4EA29EB2A15A1BA66" country="Austria" name="Steiermark">Steiermark</collectingRegion>
; Steirisches Randgebirge, Westliches Grazer Bergland; Bezirk Graz (Stadt);
<normalizedToken id="E7B865E0C52DF64686AEAA915D5B01E0" originalValue="Gösting">Goesting</normalizedToken>
, Ruine
<normalizedToken id="287DC004A67EC2BE0472874E1E1367E1" originalValue="Gösting">Goesting</normalizedToken>
,
<geoCoordinate id="E2F567FE630B9542ABF4E87C7BB0A526" degrees="47" direction="north" minutes="06" orientation="latitude" precision="1" seconds="13.44" value="47.103733">47°06'13.44&quot;N</geoCoordinate>
,
<geoCoordinate id="FD9742BC79249D1D36780A06FAA2858B" degrees="15" direction="east" minutes="22" orientation="longitude" precision="1" seconds="48.28" value="15.380078">15°22'48.28&quot;E</geoCoordinate>
,
<quantity id="982627CBCA44E8F1B0F67F5D64CE3CA6" metricMagnitude="2" metricUnit="m" metricValue="5.41" unit="m" value="541.0">
<elevation id="6016B82CBA7FCE32696D56C0C651DAE1" metricMagnitude="2" metricUnit="m" metricValue="5.41" unit="m" value="541.0">541 m</elevation>
</quantity>
elev.,
<collectingDate id="4636F777E1664F9EB120BE3FA1FBA334" value="2010-09-06">6 September 2010</collectingDate>
,
<collectorName id="659EAE0172FB6FA96DC0D7456FDCF0F3">
<emphasis id="3A301540BAF753C2B899114A5395FBF5" italics="true" pageId="0" pageNumber="123">Leonhartsberger S</emphasis>
.
</collectorName>
, (GJO 0062960); Windische
<normalizedToken id="C4136864942C1B2BB18ED765053ECC59" originalValue="Bühel">Buehel</normalizedToken>
, Kreuzberg bei Leutschach, Oberfahrenbach - Gamlitz - Karnerberg, Gasthof Eichbergho,
<geoCoordinate id="5674E7152BCBD24551B751342F20B381" degrees="46.725" direction="north" orientation="latitude" precision="55" value="46.725">46.725°N</geoCoordinate>
,
<geoCoordinate id="E0AA776D8EA9C64D8DC42D3AC4EFA065" degrees="15.45833" direction="east" orientation="longitude" precision="1" value="15.45833">15.45833°E</geoCoordinate>
,
<collectingDate id="E6AE9488CEBA15D70ABB1712CE9AC499" value="1989-11-07">07 November 1989</collectingDate>
,
<collectorName id="61F10D873077C342CFAC891030726E59">
<emphasis id="ACE122436804E6FA1804556B97C986C1" italics="true" pageId="0" pageNumber="123">Maurer W</emphasis>
.
</collectorName>
s.n., (GZU 000239138)
</materialsCitation>
</emphasis>
;
<materialsCitation id="D5382AEC0D193E3F058B23CC64377BB1" collectingDate="1863-07" collectingDateMax="1866-07" collectingDateMin="1863-07" country="Belgium" location="Hackerinver" municipality="Moissons" specimenCount="1">
<emphasis id="E64A3F47BA0A22623A6DE7575D96CA1E" bold="true" pageId="0" pageNumber="123">
<collectingCountry id="812BE1D117BF84D2E14E5D95EF49968D" name="Belgium">Belgium</collectingCountry>
.
</emphasis>
<collectingMunicipality id="106EAFB56DA2D46765B93E6497E0D308">Moissons</collectingMunicipality>
,
<location id="0C949A8908880179D7C55A237CBEA964" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:0C949A8908880179D7C55A237CBEA964" country="Belgium" municipality="Moissons" name="Hackerinver">Hackerinver</location>
,
<collectingDate id="9CEB97902BB3761554E203939D6557D0" value="1863-07">July 1863</collectingDate>
,
<emphasis id="79A739C9C9349A81447412587ECEE396" italics="true" pageId="0" pageNumber="123">Thieleus Armand</emphasis>
;
<normalizedToken id="CCC9855916D5CDF2C01E01B19F0F426D" originalValue="cultivè">cultive</normalizedToken>
de cinq pieds sauvages
<normalizedToken id="E4466733729B3AC817D32424409CD3E7" originalValue="trouvés">trouves</normalizedToken>
en 1865 entre Haekendover (Brabant) et Overhespen (
<normalizedToken id="7D62C9ADDD0CCE011D240C957BD896CD" originalValue="Liège">Liege</normalizedToken>
),
<collectingDate id="4CBDB2A27E03C9831640EEB818545117" value="1866-07">July 1866</collectingDate>
, (FI)
</materialsCitation>
.
<materialsCitation id="2FCCAC5E00E52E3886B372787C3044D9" collectingDate="1974-07-23" collectingDateMax="2017-07-14" collectingDateMin="1974-07-23" collectorName="Cesky Krumlov. VU Boletice, Grulich V., Dvorak F." country="Czech Republic" county="Nova Viska" elevation="280" latitude="48.86556" location="Brno" longLatPrecision="365" longitude="14.07" municipality="Kristanove" specimenCount="1" stateProvince="Bohemia">
<emphasis id="3FA9D02E1EDE7B77987D7F8F5F156964" bold="true" pageId="0" pageNumber="123">
<collectingCountry id="10B62B938D752515E032BA97D597ADA3" name="Czech Republic">Czech Republic</collectingCountry>
.
</emphasis>
<collectingRegion id="F67579B6741AAF697A8741AC252D2618" country="Czech Republic" name="Jihocesky">Bohemia</collectingRegion>
, distr.
<collectorName id="515E3CDCC82CD6CB27FD0E5349420446">
<normalizedToken id="8E9E3165E6EEAD08DEB0224829DC97B4" originalValue="Český">Cesky</normalizedToken>
Krumlov.
<normalizedToken id="C8617D8AED4F6E5E7D195312551C7EB6" originalValue="VÚ">VU</normalizedToken>
Boletice
</collectorName>
:
<collectingCounty id="6BBAEDCD36E145825A47385339003E94">
<normalizedToken id="D677C56193B4E8C019F5CA481F64DA52" originalValue="Nová">Nova</normalizedToken>
<normalizedToken id="F07D4057FF513E41DA9412DE99CD68B2" originalValue="Víska">Viska</normalizedToken>
</collectingCounty>
, okraj cesty
<normalizedToken id="013646BFC9AD497C93C3496D66FE8CFD" originalValue="přes">pres</normalizedToken>
louky
<quantity id="EF7CF5A882E79B7F3D7F0366B1CE399C" metricMagnitude="3" metricUnit="m" metricValue="6.0" unit="km" value="6.0">6 km</quantity>
JV od kostela v
<normalizedToken id="F8ED706F670C7C3683D52DF8EA5F5BCF" originalValue="Křišťanově">
<collectingMunicipality id="3944CE6D423047E8DF7188798411CA4C">Kristanove</collectingMunicipality>
</normalizedToken>
,
<geoCoordinate id="213D5B51BA57ED865EC48CD9035CDA86" degrees="48.86556" direction="north" orientation="latitude" precision="1" value="48.86556">48.86556°N</geoCoordinate>
,
<geoCoordinate id="1EA52B390962C5BD72D581A57C8059C0" degrees="14.07" direction="east" orientation="longitude" precision="555" value="14.07">14.07°E</geoCoordinate>
,
<collectingDate id="7D9FFB9259498C5F3383002F84C0958C" value="2017-07-14">14 July 2017</collectingDate>
,
<collectorName id="FE5590FFF8A49CBCC21013514D40A631">
<emphasis id="581696EBCE576BC175A97895E33AAC5C" italics="true" pageId="0" pageNumber="123">Grulich V</emphasis>
.
</collectorName>
(BRNU 636522);
<collectingRegion id="41C54E69B4B047CF8C0683FADDDD4827" country="Czech Republic" name="Jihomoravsky">Moravia</collectingRegion>
centr., distr
<location id="37D3620CE59CB49E14D103B14D6AC432" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:37D3620CE59CB49E14D103B14D6AC432" country="Czech Republic" county="Nova Viska" latitude="48.86556" longLatPrecision="365" longitude="14.07" municipality="Kristanove" name="Brno" stateProvince="Bohemia">Brno</location>
: inter segtes ad pedem collis
<location id="28B95F547626292998308A5ED02E92FF" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:28B95F547626292998308A5ED02E92FF" country="Czech Republic" county="Nova Viska" latitude="48.86556" longLatPrecision="365" longitude="14.07" municipality="Kristanove" name="Zlobica" stateProvince="Bohemia">Zlobica</location>
dicti situ septentr-orientali ad oppido
<normalizedToken id="735B75DF0B85100C187938A4B9274EEF" originalValue="Kuřim">
<location id="626AA4137CB1B7833499775C54D1AA00" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:626AA4137CB1B7833499775C54D1AA00" country="Czech Republic" county="Nova Viska" latitude="48.86556" longLatPrecision="365" longitude="14.07" municipality="Kristanove" name="Kurim" stateProvince="Bohemia">Kurim</location>
</normalizedToken>
,
<quantity id="80256FCC405F1308A34A78A710E4B6CA" metricMagnitude="2" metricUnit="m" metricValue="2.8" unit="m" value="280.0">
<elevation id="CA0CE9FEB63883969BA70D0B4CC2881C" metricMagnitude="2" metricUnit="m" metricValue="2.8" unit="m" value="280.0">280 m</elevation>
</quantity>
elev.,
<collectingDate id="85036F89A2007D81851C882A51028F6D" value="1974-07-23">23 July 1974</collectingDate>
,
<collectorName id="8F6DE75CAFC33A639D39A324A41A7CD6">
<emphasis id="250EDE89A5ABECE0300A58971CE4E6C5" italics="true" pageId="0" pageNumber="123">
<normalizedToken id="B5D3BF77EFAE22FE1E414FD4FC60BDDC" originalValue="Dvořàk">Dvorak</normalizedToken>
F
</emphasis>
.
</collectorName>
(FI)
</materialsCitation>
.
<materialsCitation id="00FDF2427120DF392291B6BEE84305F4" collectingDate="1853-08-17" collectingDateMax="1882-07-26" collectingDateMin="1853-08-17" collectorName="de Strasbourg, Nestler C. G., Caruel L." country="France" location="Douai" municipality="Environs de Strasbourg" specimenCount="1">
<emphasis id="CDB193703EACF1DAEC6BFED7C017D24B" bold="true" pageId="0" pageNumber="123">
<collectingCountry id="9C56FA3FC423734528E931199610DF76" name="France">France</collectingCountry>
.
</emphasis>
<collectingMunicipality id="007FEDCBF277205B17A9E6B9E1957C97">
Environs
<collectorName id="A2D204AE6634BA1C0AA5B9EC2EA5F08F">de Strasbourg</collectorName>
</collectingMunicipality>
, 1805,
<collectorName id="77A29CF61B89F4879E95FE0A2AE02CD8">
<emphasis id="BCCC076579B26113558DEAE35FA0E28B" italics="true" pageId="0" pageNumber="123">Nestler C. G</emphasis>
.
</collectorName>
sn, (G00215384);
<location id="77C1E6897BB99B16D7A5B53C31CC2A73" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:77C1E6897BB99B16D7A5B53C31CC2A73" country="France" municipality="Environs de Strasbourg" name="Douai">Douai</location>
(Nord), abondant dans un jardin
<normalizedToken id="2E847E38342FFAFCDFE6613C753DADCE" originalValue="où">ou</normalizedToken>
il a
<normalizedToken id="0A2ABFE489AF34C9AB90EAF5DF7C6940" originalValue="peut-être">peut-etre</normalizedToken>
<normalizedToken id="2981305730EE13CFECC7749C7F919426" originalValue="ètè">ete</normalizedToken>
introduit accidentallement,
<collectingDate id="1121C47D3700CDC43309309AEE1F4D3D" value="1882-07-26">26 July 1882</collectingDate>
,
<emphasis id="0763B74CB64407DBC0D8D63081049B25" italics="true" pageId="0" pageNumber="123">Maugin</emphasis>
(FI); Versailles,
<collectingDate id="B0931D06F713B20A6AD9B3E0303D75CD" value="1853-08-17">17 August 1853</collectingDate>
,
<collectorName id="A8196DA5209281137187CAF0EF269E1B">
<emphasis id="E331F7E5D058FC62D177CEB459227B4A" italics="true" pageId="0" pageNumber="123">Caruel L</emphasis>
.
</collectorName>
(FI)
</materialsCitation>
.
<materialsCitation id="CB980F70F3D1E492A2FF565A9C3A26EB" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="Wallroth K. F. W." country="Germany" location="Germany" specimenCount="1">
<emphasis id="C6EBD1AECFBA729C9A17E39BAEB9D80E" bold="true" pageId="0" pageNumber="123">
<collectingCountry id="16E64FFEFCDE4387D7CB7E1D00398765" name="Germany">Germany</collectingCountry>
.
</emphasis>
Thuring, 1822,
<collectorName id="C6588C48C8DAF3A26332F6C13EF49DA1">
<emphasis id="BA09E4AA139AD2EF2C12BF820AF147DF" italics="true" pageId="0" pageNumber="123">Wallroth K.F.W</emphasis>
.
</collectorName>
sn, (G00215386)
</materialsCitation>
.
<materialsCitation id="046A6EA698F2E921F70E39FA9A4D902F" collectingDate="1894-08-29" collectingDateMax="1971-09-10" collectingDateMin="1894-08-29" collectorName="di Tornimparte, Pelliccione G." country="Italy" county="Luoghi" elevation="1200" location="Valle del Lagaccio" municipality="Cretone di Tornimparte" specimenCount="1" stateProvince="Liguria">
<emphasis id="9DFABC16428122CACB21CEDB823D56EC" bold="true" pageId="0" pageNumber="123">
<collectingCountry id="D9563C8B212AC42A0B77012FD5165EB7" name="Italy">Italy</collectingCountry>
.
</emphasis>
<collectingCounty id="81349DAE14D9FA574CB2FC323C5238E3">Luoghi</collectingCounty>
erbosi presso
<collectingMunicipality id="CD473EA6184DBC31FB5A3F9F2FF67D39">
Cretone
<collectorName id="8D571D1B5DBB848C4DFE2321A08A215F">di Tornimparte</collectorName>
</collectingMunicipality>
(
<normalizedToken id="BC3920B4DCFF81BE89CD13FEC4E31F5D" originalValue="LAquila">L'Aquila</normalizedToken>
),
<quantity id="CF1425B8C765E1D2B6CA64600008C033" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">
<elevation id="6D05F4F9DC7F9E68A07678917D82A7F2" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</elevation>
</quantity>
elev.,
<collectingDate id="6E9C5882FEF901B0B55C930F7A92F91A" value="1971-09-10">10 September 1971</collectingDate>
,
<collectorName id="651613C7F16DE47C3C916BA80ADFEC6B">
<emphasis id="C70631594D31F8790CE1FC6F8A48B96D" italics="true" pageId="0" pageNumber="123">Pelliccione G</emphasis>
.
</collectorName>
(FI),
<location id="BD78E42D2D84607575C060EF393150CB" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:BD78E42D2D84607575C060EF393150CB" country="Italy" county="Luoghi" municipality="Cretone di Tornimparte" name="Valle del Lagaccio" stateProvince="Liguria">Valle del Lagaccio</location>
,
<collectingRegion id="84043E39099A380339CA883D21475684" country="Italy" name="Liguria">Liguria</collectingRegion>
, June,
<emphasis id="0F405CBE613244D003BEA11C83C5160C" italics="true" pageId="0" pageNumber="123">Baglietto</emphasis>
(FI); Cult.
<normalizedToken id="0B66FC937FF18E137B805BEE0A5E92FC" originalValue="nellOrto">nell'Orto</normalizedToken>
<location id="199E93DF1AE26964031E6DC71D67887F" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:199E93DF1AE26964031E6DC71D67887F" country="Italy" county="Luoghi" municipality="Cretone di Tornimparte" name="Botanico Sperimentale di Vallombrosa" stateProvince="Liguria">Botanico Sperimentale di Vallombrosa</location>
,
<collectingDate id="CF3B59987245405835614225AB551C16" value="1894-08-29">29 August 1894</collectingDate>
,
<emphasis id="AA3E19B7BE0C3691351EFF40D944CFE6" italics="true" pageId="0" pageNumber="123">Solla n.</emphasis>
1413 (FIAF); Giardino dei Semplici, 1830,
<emphasis id="648F53BAFE765C9582CC8580D675ECDB" italics="true" pageId="0" pageNumber="123">Ricasoli</emphasis>
(FI)
</materialsCitation>
.
</paragraph>
<paragraph id="A62128038CF0F52360091FC4A5725231" pageId="0" pageNumber="123">
<emphasis id="82F967C2A99A0A08A2C710A831FE16B5" bold="true" pageId="0" pageNumber="123">
<taxonomicName id="62052D7E9D3B924A01E828B80CC796E3" class="Magnoliopsida" family="Caryophyllaceae" genus="Silene" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Silene vallesia subsp. graminea" order="Caryophyllales" pageId="0" pageNumber="123" phylum="Tracheophyta" rank="subSpecies" species="vallesia" subSpecies="graminea">Silene vallesia subsp. graminea</taxonomicName>
:
<materialsCitation id="706DE46A215C78DFE8F48222C4CBD1C8" accessionNumber="BP591483, BP455289, BP455288" collectingDate="1877-07-24" collectingDateMax="1936-07-21" collectingDateMin="1877-07-24" collectorName="I. Dolfler, O. V. E. Behr, Distrikt Hasi, Pastrik, Doerfler, Ignaz" country="Albania" county="An Felsen in der Gipfelregion" elevation="1700" location="Velika Golia" municipality="Skrivanek" specimenCount="1">
<collectingCountry id="4751F638107299D9AEB8DEFA453A5E30" name="Albania">Albania</collectingCountry>
.
<location id="F1B56E3A600EE0891DAFB029CC138504" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:F1B56E3A600EE0891DAFB029CC138504" country="Albania" county="An Felsen in der Gipfelregion" municipality="Skrivanek" name="Velika Golia">Velika Golia</location>
,
<quantity id="DE24E676F70F9742E782F1951292CCC7" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<elevation id="F921BE56718905CEE34895C24969F6CB" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700 m</elevation>
</quantity>
elev.,
<collectingDate id="58A8F3D3A7CE142BA0DBA3E944253B4A" value="1904-07-17">17 July 1904</collectingDate>
(FI);
<location id="83426BC2FA5C52A4687BA2198B0B967F" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:83426BC2FA5C52A4687BA2198B0B967F" country="Albania" county="An Felsen in der Gipfelregion" municipality="Skrivanek" name="Hais Patrik An Felsen in der Gipfelregion">Hais Patrik An Felsen in der Gipfelregion</location>
,
<quantity id="66026A88736FC06FBA4BBE2EA1BE19B9" metricMagnitude="3" metricUnit="m" metricValue="1.8" unit="m" value="1800.0">
<elevation id="66AE90D45B6B766B030AB340E3DAA639" metricMagnitude="3" metricUnit="m" metricValue="1.8" unit="m" value="1800.0">1800 m</elevation>
</quantity>
elev.,
<collectingDate id="42419E5E8004DEA8D96E8A3577410213" value="1877-07-24">24 July 1877</collectingDate>
,
<emphasis id="533571F656C8639D19F1ABFE3E666429" italics="true" pageId="0" pageNumber="123">
<collectorName id="55A79BCBC7F4E97A827D79BCC1133534">I. Dolfler</collectorName>
</emphasis>
(W);
<location id="F4B8C16802C556696A84E0F3790DA320" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:F4B8C16802C556696A84E0F3790DA320" country="Albania" county="An Felsen in der Gipfelregion" municipality="Skrivanek" name="Aufdem Berge Patrik">Aufdem Berge Patrik</location>
bei
<location id="480A9A5528A02F3E0571211219CDE09E" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:480A9A5528A02F3E0571211219CDE09E" country="Albania" county="An Felsen in der Gipfelregion" municipality="Skrivanek" name="Prizzen">Prizzen</location>
,
<quantity id="64D995257ECBF76B2C2BF561767B9B3E" metricMagnitude="3" metricUnit="m" metricValue="1.9" unit="m" value="1900.0">
<elevation id="400D79385E8FB5E04656EFA4AD5ED93E" metricMagnitude="3" metricUnit="m" metricValue="1.9" unit="m" value="1900.0">1900 m</elevation>
</quantity>
elev.,
<collectingDate id="2CF8012D718E1774D9CF1A80AC2C117B" value="1936-07-20" valueMax="1936-07-21" valueMin="1936-07-20">20-21 July 1936</collectingDate>
,
<emphasis id="E72DB3CDEB48F1856118F054664E1B9C" italics="true" pageId="0" pageNumber="123">
<collectorName id="64E36784919168F477209D3668E780D5">O.V.E. Behr</collectorName>
</emphasis>
(FI);
<location id="B59CB0031A0DF7ECA864739E22272E50" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:B59CB0031A0DF7ECA864739E22272E50" country="Albania" county="An Felsen in der Gipfelregion" municipality="Skrivanek" name="Prizren">Prizren</location>
:
<location id="29C2A0A0BBF6DD166FDAFA411C09CBE5" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:29C2A0A0BBF6DD166FDAFA411C09CBE5" country="Albania" county="An Felsen in der Gipfelregion" municipality="Skrivanek" name="Mt. Pastrik">Mt. Pastrik</location>
in graminosis alpinis,
<collectingDate id="91BAEAD91D6E39429D430AC0F6CEED87" value="1936-07-21">21 July 1936</collectingDate>
,
<emphasis id="2ECF765B170E2A41984C4EE804B22CEE" italics="true" pageId="0" pageNumber="123">
<collectingMunicipality id="D65C3210908D98E3491D009ED8362036">Skrivanek</collectingMunicipality>
</emphasis>
(
<accessionNumber id="D80173D9664FB700AFF5192D7C207A8B" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/BP591483">BP591483</accessionNumber>
);
<collectorName id="EC5A0D48A8DCDC3681AF4D7FB9AFB763">Distrikt Hasi</collectorName>
,
<collectorName id="DDD4F9A124281250C075A586AA7C8CF5">Pastrik</collectorName>
,
<collectingCounty id="9D78C8978788B712FB46E9201B3D2F04">An Felsen in der Gipfelregion</collectingCounty>
,
<collectingDate id="61B3CBB822EF8F1E802F77A2BF6AEA3C" value="1918-07-24">24 July 1918</collectingDate>
,
<emphasis id="E6742917579D536F6BDB50AC55355AC0" italics="true" pageId="0" pageNumber="123">
<normalizedToken id="89848C5D105E66DAA7127F32AA0FA491" originalValue="Dörfler">
<collectorName id="690A1170160B4D7C55B4EB83ACCA3741">Doerfler</collectorName>
</normalizedToken>
&amp;
<collectorName id="EA73EA6B1CCE3518DFC81B362862929B">Ignaz</collectorName>
</emphasis>
(
<accessionNumber id="DDD2C4D03D7888EBD4F1D5A8319235AF" isEnumeration="true">BP455289, BP455288</accessionNumber>
)
</materialsCitation>
</emphasis>
.
<materialsCitation id="B56B4AA5A9447296BA3C2E5B73E1C15A" collectingDate="1897-07-27" collectorName="Gipfelregion der Golia, der Velica Golja, July, J, F. Faltis, E. Brandis" country="Bosnia and Herzegovina" county="Livno" elevation="1700" location="Sudskamn" municipality="KameKnica" specimenCount="1">
<emphasis id="48F3DEEC1C1B1AB64D7C9208CA643E21" bold="true" pageId="0" pageNumber="123">
<collectingCountry id="CD40586420748C60DE497906BF7039EC" name="Bosnia and Herzegovina">Bosnia and Herzegovina</collectingCountry>
.
</emphasis>
<collectorName id="06336F75D9A631BC6835A6247D58B990">Gipfelregion der Golia</collectorName>
:
<location id="625C88CF8A4A3F6D48379A7DAD66B531" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:625C88CF8A4A3F6D48379A7DAD66B531" country="Bosnia and Herzegovina" county="Livno" municipality="KameKnica" name="Sudskamn">Sudskamn</location>
(?)
<collectorName id="A14491047F2EE86E9003EB9B06F7CB47">der Velica Golja</collectorName>
,
<quantity id="F56AFB8AE96795F23A9248D192A14B27" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">
<elevation id="45BD9CCD79A4281D9D4BC14E36703B54" metricMagnitude="3" metricUnit="m" metricValue="1.7" unit="m" value="1700.0">1700 m</elevation>
</quantity>
elev., 18
<collectorName id="3EEDE31347844C69E1C4E089AE3BA100">
July,
<emphasis id="99520026B7E3D16120B4A12D75EA57EF" italics="true" pageId="0" pageNumber="123">
J. Stdlmann &amp;
<collectorName id="1C381B9E95F65E2E2CB4DB1A7BA86E86">F. Faltis</collectorName>
</emphasis>
</collectorName>
(FI);
<collectingMunicipality id="049A9D6E45C38373CBEE0CD8A315C2CF">KameKnica</collectingMunicipality>
bai
<collectingCounty id="E5A5163250C64971ACC3FE0EB7849965">Livno</collectingCounty>
,
<quantity id="668CB0723E2FA1C0D6025D082203731A" metricMagnitude="3" metricUnit="m" metricValue="1.8" unit="m" value="1800.0">
<elevation id="908B423D0E53A4A30DCD32EE85309740" metricMagnitude="3" metricUnit="m" metricValue="1.8" unit="m" value="1800.0">1800 m</elevation>
</quantity>
elev.,
<collectingDate id="0B9A6AB910BC3AA70C2C9A8A1FAAE095" value="1897-07-27">27 July 1897</collectingDate>
,
<emphasis id="0FA58CC622EED1714DC49C3EC4CCA7F6" italics="true" pageId="0" pageNumber="123">
<collectorName id="39BEB9144E5DC1A2150F9D2DC6C28365">E. Brandis</collectorName>
</emphasis>
(PAD)
</materialsCitation>
.
<materialsCitation id="634B74725F763FC0EB3086D4831A64D3" accessionNumber="PI042104" collectingDate="1868-07-18" collectorName="Dalmatien M. Prolog, Th. Pichler, G. Lengyel, R. Visiani" country="Croazia" county="Allau" elevation="1400" location="Ex" municipality="Risioica" specimenCount="1">
<emphasis id="73DEA9051F0924003EEFDBCB54C44541" bold="true" pageId="0" pageNumber="123">
<collectingCountry id="A2A209B0C7523B552C4BAF8C21E1AC49" name="Croatia">Croazia</collectingCountry>
.
</emphasis>
<location id="9B1BBF84DA4E6C596B818E55416D2760" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:9B1BBF84DA4E6C596B818E55416D2760" country="Croazia" county="Allau" municipality="Risioica" name="Ex">Ex</location>
rupibus
<location id="08139D598C995B565AB843647B2DF8DA" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:08139D598C995B565AB843647B2DF8DA" country="Croazia" county="Allau" municipality="Risioica" name="Dinaricis">Dinaricis</location>
ad
<normalizedToken id="12A34376E4A6D54DD91F04B8CAA5F375" originalValue="…">...</normalizedToken>
<location id="CCE5BBFF5ED27058C7DB3CD33A81364B" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:CCE5BBFF5ED27058C7DB3CD33A81364B" country="Croazia" county="Allau" municipality="Risioica" name="Dalmaticae">Dalmaticae</location>
,
<location id="498101C1C73E1B5AFDC1E62E51BC9AD7" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:498101C1C73E1B5AFDC1E62E51BC9AD7" country="Croazia" county="Allau" municipality="Risioica" name="Herb Levier">Herb Levier</location>
(FI);
<location id="89251F9DA7F1D458959D96A16E4AF787" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:89251F9DA7F1D458959D96A16E4AF787" country="Croazia" county="Allau" municipality="Risioica" name="In">In</location>
saxosis apricis,
<location id="32612E7460B0AD47ABF4BC800D2D8423" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:32612E7460B0AD47ABF4BC800D2D8423" country="Croazia" county="Allau" municipality="Risioica" name="Dinara">Dinara</location>
,
<location id="E58E272D33F6B0DEB200F95E047CFC4A" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:E58E272D33F6B0DEB200F95E047CFC4A" country="Croazia" county="Allau" municipality="Risioica" name="Ghnjat">Ghnjat</location>
,
<location id="B3C7FDC127607E44F2972D48CC40A06E" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:B3C7FDC127607E44F2972D48CC40A06E" country="Croazia" county="Allau" municipality="Risioica" name="Prologh">Prologh</location>
,
<emphasis id="9572DD0E7717E218342E59406A31A864" italics="true" pageId="0" pageNumber="123">
<location id="E4765E888E04BAF8F1EFDB1EDB5AAFB6" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:E4765E888E04BAF8F1EFDB1EDB5AAFB6" country="Croazia" county="Allau" municipality="Risioica" name="Visiani">Visiani</location>
</emphasis>
(PD);
<location id="1406EC41F97B059EA7BBE32A9F43D4FD" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:1406EC41F97B059EA7BBE32A9F43D4FD" country="Croazia" county="Allau" municipality="Risioica" name="Mont Ghnjat">Mont Ghnjat</location>
(PAD);
<location id="D25164B7648D0D71338E4C63CFE34E6B" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:D25164B7648D0D71338E4C63CFE34E6B" country="Croazia" county="Allau" municipality="Risioica" name="In">In</location>
dorso apricot et saxoso
<location id="4BCE40EF08DD538D1381B80B22C84C60" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:4BCE40EF08DD538D1381B80B22C84C60" country="Croazia" county="Allau" municipality="Risioica" name="Mts. Dinara">Mts. Dinara</location>
,
<location id="34AC3D4676F8E45A157C2686AB6621AA" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:34AC3D4676F8E45A157C2686AB6621AA" country="Croazia" county="Allau" municipality="Risioica" name="July">July</location>
, 51K (Jos. Kargl) (PAD);
<location id="2149B4F736A308BD33A6E79D88506443" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:2149B4F736A308BD33A6E79D88506443" country="Croazia" county="Allau" municipality="Risioica" name="Am Chemeschniza am Prolog">Am Chemeschniza am Prolog</location>
,
<collectingDate id="33D8FD768F70D025030CB5FDF4B2CD5F" value="1868-07-18">18 July 1868</collectingDate>
,
<emphasis id="FB0BC83BB3E9124D099D0C315825F5B0" italics="true" pageId="0" pageNumber="123">
<location id="B3B4E3D36B5F87E14CE1EB98D7740B0C" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:B3B4E3D36B5F87E14CE1EB98D7740B0C" country="Croazia" county="Allau" municipality="Risioica" name="Th Pichler">Th Pichler</location>
</emphasis>
(PAD);
<collectorName id="E1166F99792E7228FFF1FA7CFE130101">Dalmatien M. Prolog</collectorName>
,
<collectingDate id="5CD6A8D85387FB9A46103B1BB7D9AF99" value="1868-07-18">18 July 1868</collectingDate>
,
<emphasis id="99B60763D06AF6B505382CB8FE919A8C" italics="true" pageId="0" pageNumber="123">
<collectorName id="5CF914C50EA0ACA5A557D266521A3C1B">
<location id="8A3FF041B75F93B7324C2DD34DCFEA4B" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:8A3FF041B75F93B7324C2DD34DCFEA4B" country="Croazia" county="Allau" municipality="Risioica" name="Th. Pichler">Th. Pichler</location>
</collectorName>
</emphasis>
(TSB);
<location id="A73516C7DD7DE383D3399CB73938ADD1" LSID="urn:lsid:plazi:treatment:7326E116DA81573B8B86E79970E71B52:A73516C7DD7DE383D3399CB73938ADD1" country="Croazia" county="Allau" municipality="Risioica" name="Croatia. Velebit">Croatia. Velebit</location>
in lapidosis herbosis m.
<collectingMunicipality id="1E406FA270F1D119690BB65025169515">Risioica</collectingMunicipality>
supra
<collectingCounty id="30268E4937B7869064EA0BD1D538C2BB">Allau</collectingCounty>
, alt. ca.
<quantity id="D73948F218B071C95483694185F3CD5A" metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">
<elevation id="047FE3C6C13082728D54EA4218CB491C" metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">1400 m</elevation>
</quantity>
, 1908,
<emphasis id="1F47B4F62D24D82400BD243F5C8FFE72" italics="true" pageId="0" pageNumber="123">
<collectorName id="D5C3907B98997886764BAB7B8E229285">G. Lengyel</collectorName>
</emphasis>
(
<accessionNumber id="B901C6863EF7DB2E08C0BE5B959D6C33" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/PI042104">PI042104</accessionNumber>
); In montib. Dalmatia, s.d.,
<emphasis id="5FDA390341575CB54CBDF967174BDD40" italics="true" pageId="0" pageNumber="123">
<collectorName id="EA2D0BC8D4C847579663A7452E6B9569">R. Visiani</collectorName>
</emphasis>
(HAL0012278)
</materialsCitation>
.
<materialsCitation id="444FA03C60E9092093CD4D69AC47138C" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="M. Durmitor, J. Panic" country="Montenegro" location="Montenegro" specimenCount="1">
<emphasis id="C2A616644E3061097A302EFFC01B1DE4" bold="true" pageId="0" pageNumber="123">
<collectingCountry id="FD944D09A8F5085EAF1BC3643F9AABD8" name="Serbia">Montenegro</collectingCountry>
.
</emphasis>
In saxosis excelsioribus
<collectorName id="A1CDA9CCC3086BA2894801C9CC22E635">M. Durmitor</collectorName>
, 870,
<emphasis id="1E9E01A1A6DF1CC7D028612696A89D95" italics="true" pageId="0" pageNumber="123">
<collectorName id="07BB146B274A0AFB09996DE21C736386">J. Panic</collectorName>
</emphasis>
(PAD)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>