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<document id="9B0CD6463AA78494F6A213E3A857BECC" ID-DOI="10.11646/phytotaxa.193.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13642005" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725348533824" checkinUser="felipe" docAuthor="Sukkharak, Phiangphak" docDate="2015" docId="73083D48FF88BF07FF173D96FA3E944C" docLanguage="en" docName="phytotaxa.193.1.1.pdf" docOrigin="Phytotaxa 193 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.193.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.4:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="4" docTitle="Thysananthus montanus Gradstein, He &amp; Piippo" docType="treatment" docVersion="2" lastPageNumber="450" masterDocId="8F314530FFA4BF2BFF803C21FF9E9C03" masterDocTitle="A systematic monograph of the genus Thysananthus (Lejeuneaceae, Marchantiophyta)" masterLastPageNumber="450" masterPageNumber="448" pageNumber="448" updateTime="1725354303689" updateUser="ExternalLinkService">
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<mods:title id="2565023D709191AFE45361A5CC69540C">A systematic monograph of the genus Thysananthus (Lejeuneaceae, Marchantiophyta)</mods:title>
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<mods:title id="A80C7831B203B6BE4C341BCD2A06B48E">Phytotaxa</mods:title>
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<mods:date id="89A3BA3FE211F7E9F86A9E1E9560AB7D">2015</mods:date>
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<treatment id="73083D48FF88BF07FF173D96FA3E944C" LSID="urn:lsid:plazi:treatment:73083D48FF88BF07FF173D96FA3E944C" httpUri="http://treatment.plazi.org/id/73083D48FF88BF07FF173D96FA3E944C" pageId="44">
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<emphasis id="C9D5504CFF88BF07FF173D96FD699DD3" bold="true" box="[151,759,439,464]" pageId="44">
14.
<taxonomicName id="3CA1F7DDFF88BF07FF403D96FD699DD3" ID-CoL="56TGM" authority="Gradstein, He &amp; Piippo" authorityName="Gradstein, He &amp; Piippo" box="[192,759,439,464]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Thysananthus" kingdom="Plantae" order="Porellales" pageId="44" phylum="Marchantiophyta" rank="species" species="montanus">
<emphasis id="C9D5504CFF88BF07FF403D96FE449DCC" bold="true" box="[192,474,439,464]" italics="true" pageId="44">Thysananthus montanus</emphasis>
Gradstein, He &amp; Piippo
</taxonomicName>
</emphasis>
in
<bibRefCitation id="9F30F1AFFF88BF07FC9A3D96FA959DD3" author="Gradstein, S. R. &amp; He, X. - L. &amp; Piippo, S. &amp; Mizutani, M." box="[794,1291,439,464]" pageId="44" pagination="1 - 88" refId="ref47615" refString="Gradstein, S. R., He, X. - L., Piippo, S. &amp; Mizutani, M. (2002) Bryophyte flora of the Huon Peninsula, Papua New Guinea. LXVIII. Lejeuneaceae subfamily Ptychanthoideae (Hepaticae). Acta Botanica Fennica 174: 1 - 88." type="journal article" year="2002">Gradstein, He, Piippo &amp; Mizutani (2002: 77)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B3BBDFD5FF88BF07FF163E21FC3499D9" pageId="44" type="materials_examined">
<paragraph id="FB1E8C5EFF88BF07FF163E21FB7F9E37" blockId="44.[150,1435,512,564]" pageId="44">
<materialsCitation id="4BC98603FF88BF07FF163E21FB429E37" country="Papua New Guinea" county="Type" elevation="2625" latitude="-6.0416665" location="Teptep-Wantuat" longLatPrecision="925" longitude="146.55" municipality="Lake Wamba" pageId="44" specimenCount="1" stateProvince="Morobe" typeStatus="holotype">
<collectingCounty id="127FF4D2FF88BF07FF163E21FF529E16" box="[150,204,512,533]" pageId="44">Type</collectingCounty>
:
<collectingCountry id="83B6CCCEFF88BF07FF563E21FE4E9E15" box="[214,464,512,534]" name="Papua New Guinea" pageId="44">PAPUA NEW GUINEA</collectingCountry>
.
<collectingRegion id="396542BCFF88BF07FE593E21FDB39E15" box="[473,557,512,534]" country="Papua New Guinea" name="Morobe" pageId="44">Morobe</collectingRegion>
:
<collectingMunicipality id="1B7A1624FF88BF07FDB73E21FD209E15" box="[567,702,512,534]" pageId="44">Lake Wamba</collectingMunicipality>
,
<locationDeviation id="EAAC957CFF88BF07FD483E21FCEA9E16" box="[712,884,512,534]" location="Teptep" pageId="44">
<quantity id="3C5921BBFF88BF07FD483E21FD659E15" box="[712,763,512,534]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="44" unit="km" value="5.0">5 km</quantity>
S of Teptep
</locationDeviation>
airstrip,
<location id="FE7EDA85FF88BF07FC4F3E21FBEE9E16" LSID="urn:lsid:plazi:treatment:73083D48FF88BF07FF173D96FA3E944C:FE7EDA85FF88BF07FC4F3E21FBEE9E16" box="[975,1136,512,533]" country="Papua New Guinea" county="Type" latitude="-6.0416665" longLatPrecision="925" longitude="146.55" municipality="Lake Wamba" name="Teptep-Wantuat" pageId="44" stateProvince="Morobe">Teptep-Wantuat</location>
trail
<locationDeviation id="EAAC957CFF88BF07FB243E21FAFE9E16" box="[1188,1376,512,534]" location="Teptep" pageId="44">
<quantity id="3C5921BBFF88BF07FB243E21FB7D9E15" box="[1188,1251,512,534]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="44" unit="km" value="10.0">10 km</quantity>
S of Teptep
</locationDeviation>
, open montane forest on ridge sloping NE,
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<elevation id="708C6B6DFF88BF07FDBD3E3FFD529E37" box="[573,716,542,564]" metricMagnitude="3" metricUnit="m" metricValue="2.625" metricValueMax="2.7" metricValueMin="2.55" pageId="44" unit="m" value="2625.0" valueMax="2700.0" valueMin="2550.0">25502700 m</elevation>
</quantity>
,
<geoCoordinate id="9E95EA99FF88BF07FD563E3FFCBB9E37" box="[726,805,542,564]" degrees="6" direction="south" minutes="2.5" orientation="latitude" pageId="44" precision="92" value="-6.0416665">6°2.5' S</geoCoordinate>
<geoCoordinate id="9E95EA99FF88BF07FCAC3E3FFC0C9E37" box="[812,914,542,564]" degrees="146" direction="east" minutes="33" orientation="longitude" pageId="44" precision="925" value="146.55">146°33' E</geoCoordinate>
,
<emphasis id="C9D5504CFF88BF07FC1C3E3EFBD49E37" box="[924,1098,542,564]" italics="true" pageId="44">Koponen 33958b</emphasis>
(
<typeStatus id="241A32FCFF88BF07FBD63E3FFB2F9E37" box="[1110,1201,542,564]" pageId="44" type="holotype">holotype</typeStatus>
H!)
</materialsCitation>
.
</paragraph>
<paragraph id="FB1E8C5EFF88BF07FF163E40FEDF99B5" blockId="44.[150,1437,609,2002]" pageId="44">
<emphasis id="C9D5504CFF88BF07FF163E40FF7C9E79" box="[150,226,609,634]" italics="true" pageId="44">Plants</emphasis>
dioicous, with projecting growth, turning upwards and becoming ascending to erect, pale brown in herbarium specimens, up to
<quantity id="3C5921BBFF88BF07FE5A3EA4FD8A9E9E" box="[474,532,645,670]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="44" unit="cm" value="4.0">4 cm</quantity>
long ×
<quantity id="3C5921BBFF88BF07FDED3EA4FD689E9E" box="[621,758,645,670]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.6" metricValueMin="1.5" pageId="44" unit="mm" value="2.05" valueMax="2.6" valueMin="1.5">1.52.6 mm</quantity>
wide.
<emphasis id="C9D5504CFF88BF07FCC33EA7FC1A9E9E" box="[835,900,646,669]" italics="true" pageId="44">Stems</emphasis>
rather rigid; ventral merophyte 810 cell rows wide; stem in cross section orbicular, 215240 µm high × 303329 µm wide, 1417 cell layers high, composed of 4348 epidermal cells surrounding 141150 medullary cells, epidermal cells as large as medullary cells.
<emphasis id="C9D5504CFF88BF07FACD3EEFFA029EE6" box="[1357,1436,718,741]" italics="true" pageId="44">Leaves</emphasis>
imbricate, when dry suberect and convolute, when moist weakly convex, apical part plane, not recurved; dorsal lobe symmetrically oblong, 1.61.8 ×
<quantity id="3C5921BBFF88BF07FDD53F34FD7D9F2E" box="[597,739,789,814]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="44" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0.60.8 mm</quantity>
, apex acute-apiculate, margin entire, dorsal base auriculate, auricle 5796 × 5776 µm, ventral margin incurved 1/2 × leaf length; cells elongate-hexagonal with acute ends, vitta present in midportion of lobe, extending to base, 2/3 × lobe length, 817 cell rows wide, 1320 cells long, marginal cells 1012 × 57 µm, median vitta 2545 × 57 µm, basal vitta 5067 × 1015 µm, trigones cordate, often coalesced, intermediate thickenings 02 per cell; oil bodies unknown.
<emphasis id="C9D5504CFF88BF07FC7A3F84FBCD9FBD" box="[1018,1107,933,958]" italics="true" pageId="44">Lobules</emphasis>
rectangular, 0.81 ×
<quantity id="3C5921BBFF88BF07FAC73F84FF589FE2" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="44" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.30.4 mm</quantity>
, 1/41/3 × lobe length; appendages on surface of lobule base on both sides of stem; keel without appendage; lobule apex oblique, free margin continuing into the ventral lobe margin, apex with one linear tooth, the tooth consisting of 79 cells, being 23 cells wide at base and ending in a row of 67 cells.
<emphasis id="C9D5504CFF88BF07FB0C3830FA809829" box="[1164,1310,1041,1066]" italics="true" pageId="44">Underleaves</emphasis>
imbricate, slightly squarrose, spathulate, 1.21.3 ×
<quantity id="3C5921BBFF88BF07FDE93814FD68984E" box="[617,758,1077,1102]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="44" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.60.7 mm</quantity>
, 23 × stem width, apex round, plane, margin entire, bases cuneate, underleaf bases not adnate with leaves; cells 2737 × 25 µm.
<emphasis id="C9D5504CFF88BF07FC4C3878FBDF9871" box="[972,1089,1113,1138]" italics="true" pageId="44">Androecia</emphasis>
terminal-intercalary on lateral branches, bracts, and bracteoles in 613(20) pairs, bracts hypostatic, 0.70.9 ×
<quantity id="3C5921BBFF88BF07FBA5385CFB319896" box="[1061,1199,1149,1174]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="44" unit="mm" value="0.35" valueMax="0.5" valueMin="0.2">0.20.5 mm</quantity>
, apex acute, margins entire; antheridia 2 per bract.
<taxonomicName id="3CA1F7DDFF88BF07FE653883FDCE98BA" authorityName="Doubleday" authorityYear="1844" box="[485,592,1186,1209]" class="Insecta" genus="Gynoecia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="44" phylum="Arthropoda" rank="genus">
<emphasis id="C9D5504CFF88BF07FE653883FDCE98BA" box="[485,592,1186,1209]" italics="true" pageId="44">Gynoecia</emphasis>
</taxonomicName>
with 2 lejeuneoid innovations forming a dichasial pattern; lobe lanceolate, 2.72.8 ×
<quantity id="3C5921BBFF88BF07FE8A38E4FE0A98DE" box="[266,404,1221,1246]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="44" unit="mm" value="0.85" valueMax="0.9" valueMin="0.8">0.80.9 mm</quantity>
, apex acute, margins entire or in upper 2/3 with triangular teeth, the teeth consisting of 35 cells, being 23 cells wide at base and ending in a row of 12 cells; lobules broadly ovate, 1/2 × lobe length, apex bifid, margin with triangular teeth, the teeth consisting of 35 cells, being 23 cells wide at base and ending in a row of 12 cells; bracteoles spathulate, 2.42.5 ×
<quantity id="3C5921BBFF88BF07FD403910FC89994A" box="[704,791,1329,1354]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="44" unit="mm" value="0.7">0.7 mm</quantity>
, apex emarginate to shortly bifid, entire or 1/2 × bracteole length with teeth, the teeth consisting of 35 cells, being 23 cells wide at base and ending in a row of 12 cells, margins plane.
<emphasis id="C9D5504CFF88BF07FEC83958FE299991" box="[328,439,1401,1426]" italics="true" pageId="44">Perianths</emphasis>
oblong-cylindrical, 2.62.7 ×
<quantity id="3C5921BBFF88BF07FC993958FC389992" box="[793,934,1401,1426]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="44" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.70.8 mm</quantity>
, keels entire; beak 6265 µm (45 cells) in length.
<figureCitation id="639A90DBFF88BF07FF6B39BCFEA399B5" box="[235,317,1437,1462]" captionStart="FIGURE 27" captionStartId="78.[151,250,1678,1700]" captionTargetBox="[195,1399,212,1608]" captionTargetId="figure-90@78.[151,1436,193,1657]" captionTargetPageId="78" captionText="FIGURE 27. Thysananthus montanus Gradst., X.-L. He &amp; Piippo. A. Portion of shoot, ventral view. B. Leaves, ventral view. C. Underleaves, ventral view. D. Female bracts, ventral view. E. Female bracteole. F. Perianth, dorsal view. G. Perianth, ventral view. H. Leaf apices. I. Margin cells of leaf lobe. J. Median cells of leaf lobe. K. Basal cells of leaf lobe. L. Leaf lobule apex. M. Cross section of stem. (All from the holotype)." figureDoi="http://doi.org/10.5281/zenodo.13642063" httpUri="https://zenodo.org/record/13642063/files/figure.png" pageId="44">Fig. 27</figureCitation>
.
</paragraph>
<paragraph id="FB1E8C5EFF88BF07FF4739E0FC3499D9" blockId="44.[150,1437,609,2002]" box="[199,938,1473,1498]" pageId="44">
<emphasis id="C9D5504CFF88BF07FF4739E0FD9C99D9" bold="true" box="[199,514,1473,1498]" pageId="44">Additional illustrations:—</emphasis>
<bibRefCitation id="9F30F1AFFF88BF07FD8239E0FD6699D9" author="Gradstein, S. R. &amp; He, X. - L. &amp; Piippo, S. &amp; Mizutani, M." box="[514,760,1473,1498]" pageId="44" pagination="1 - 88" refId="ref47615" refString="Gradstein, S. R., He, X. - L., Piippo, S. &amp; Mizutani, M. (2002) Bryophyte flora of the Huon Peninsula, Papua New Guinea. LXVIII. Lejeuneaceae subfamily Ptychanthoideae (Hepaticae). Acta Botanica Fennica 174: 1 - 88." type="journal article" year="2002">
Gradstein
<emphasis id="C9D5504CFF88BF07FDF539E3FD3099D9" box="[629,686,1473,1498]" italics="true" pageId="44">et al.</emphasis>
(2002
</bibRefCitation>
, p. 78, Fig. 49).
</paragraph>
</subSubSection>
<subSubSection id="B3BBDFD5FF88BF07FF4739C4FA3E944C" pageId="44" type="distribution">
<paragraph id="FB1E8C5EFF88BF07FF4739C4FDE69A21" blockId="44.[150,1437,609,2002]" pageId="44">
<emphasis id="C9D5504CFF88BF07FF4739C4FDB999FD" bold="true" box="[199,551,1509,1534]" pageId="44">Distribution and ecology:—</emphasis>
Endemic to
<collectingCountry id="83B6CCCEFF88BF07FD3939C7FC3D99FD" box="[697,931,1509,1534]" name="Papua New Guinea" pageId="44">Papua New Guinea</collectingCountry>
;
<quantity id="3C5921BBFF88BF07FC3039C4FBC699FE" box="[944,1112,1509,1534]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.3" metricValueMin="1.9" pageId="44" unit="m" value="2100.0" valueMax="2300.0" valueMin="1900.0">19002300 m</quantity>
; on tree trunks, lianas, and branches in open montane forests.
<figureCitation id="639A90DBFF88BF07FD9C3A28FDEA9A21" box="[540,628,1545,1570]" captionStart="FIGURE 5" captionStartId="14.[153,256,1811,1833]" captionTargetBox="[275,1327,210,1744]" captionTargetId="figure-115@14.[255,1340,197,1771]" captionTargetPageId="14" captionText="FIGURE 5. Distribution of Thysananthus species. A. Thysananthus amazonicus (Spruce) Schiffn. B. Thysananthus spathulistipus (Reinwardt, Blume &amp; Nees) Lindenb. C. Thysananthus convolutus Lindenb. D. Thysananthus mollis Steph., E. Thysananthus pancheri (Steph.) Hürl. F. Thysananthus anguiformis (Hook.f. &amp; Taylor) Taylor ex Gottsche, Lindenb. &amp; Nees G. Thysananthus comosus Lindenb. H. Thysananthus fruticosus (Lindenb. &amp; Gottsche) Schiffn. I. Thysananthus aculeatus Herzog. J. Thysananthus appendiculatus Steph. K. Thysananthus montanus Gradst., X.-L. He &amp; Piippo. L. Thysananthus retusus (Reinw., Blume &amp; Nees) B.M.Thiers &amp; Gradst. M. Thysananthus gottschei (Jack &amp; Steph.) Steph. N. Thysananthus ciliaris comb. nov. O. Thysananthus discretus Sukkharak &amp; Gradst." figureDoi="http://doi.org/10.5281/zenodo.13642015" httpUri="https://zenodo.org/record/13642015/files/figure.png" pageId="44">Fig. 5K</figureCitation>
.
</paragraph>
<paragraph id="FB1E8C5EFF88BF07FF473A0CFBFA9AB1" blockId="44.[150,1437,609,2002]" pageId="44">
<emphasis id="C9D5504CFF88BF07FF473A0CFC919A45" bold="true" box="[199,783,1581,1606]" pageId="44">
Representative specimens:—
<collectingCountry id="83B6CCCEFF88BF07FD9E3A0CFC949A45" box="[542,778,1581,1606]" name="Papua New Guinea" pageId="44">Papua New Guinea</collectingCountry>
.
</emphasis>
<collectingRegion id="396542BCFF88BF07FC973A0FFB909A47" box="[791,1038,1582,1605]" country="Papua New Guinea" name="Eastern Highlands" pageId="44">
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<smallCapsWord id="FDF81A82FF88BF07FC073A0FFB909A47" baselines="1599,1599" box="[903,1038,1582,1605]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Highlands" pageId="44">HIGHLANDS</smallCapsWord>
</collectingRegion>
: Gahavisuka National Park,
<emphasis id="C9D5504CFF88BF07FAD43A0CFF4F9A69" italics="true" pageId="44">Thiers 3656</emphasis>
(G).—
<collectingRegion id="396542BCFF88BF07FEA33A70FE2C9A6B" box="[291,434,1617,1642]" country="Papua New Guinea" name="Milne Bay" pageId="44">
<smallCapsWord id="FDF81A82FF88BF07FEA33A70FEEB9A6B" baselines="1636,1635" box="[291,373,1617,1642]" lowerCaseFontSize="10" mainFontSize="10" normCase="title" normString="Milne" pageId="44">MILNE</smallCapsWord>
<smallCapsWord id="FDF81A82FF88BF07FEFD3A73FE2C9A6B" baselines="1635,1635" box="[381,434,1618,1641]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Bay" pageId="44">BAY</smallCapsWord>
</collectingRegion>
: Fergusson Island,
<emphasis id="C9D5504CFF88BF07FD1B3A73FCB09A69" box="[667,814,1617,1642]" italics="true" pageId="44">Brass 27072</emphasis>
(L).—
<smallCapsWord id="FDF81A82FF88BF07FCFE3A70FC769A6B" baselines="1636,1635" box="[894,1000,1617,1642]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Morobe" pageId="44">
<collectingRegion id="396542BCFF88BF07FCFE3A70FC769A6B" box="[894,1000,1617,1642]" country="Papua New Guinea" name="Morobe" pageId="44">MOROBE</collectingRegion>
</smallCapsWord>
: Gumi divide,
<emphasis id="C9D5504CFF88BF07FB1E3A73FAFB9A69" box="[1182,1381,1617,1642]" italics="true" pageId="44">Streimann 22707</emphasis>
(JE, LAE); Mt. Kaindi,
<emphasis id="C9D5504CFF88BF07FEF73A57FDA79A8D" box="[375,569,1653,1678]" italics="true" pageId="44">Streimann 22511</emphasis>
(LAE); Araulu logging area,
<emphasis id="C9D5504CFF88BF07FC143A57FBC69A8D" box="[916,1112,1653,1678]" italics="true" pageId="44">Streimann 13639</emphasis>
(LAE); Nauti logging area, upper Watut river,
<emphasis id="C9D5504CFF88BF07FEE83ABBFDB99AB1" box="[360,551,1689,1714]" italics="true" pageId="44">Streimann 17150</emphasis>
(LAE); Kaisenik logging area,
<emphasis id="C9D5504CFF88BF07FC093AB8FC609AB1" box="[905,1022,1689,1714]" italics="true" pageId="44">Shea 6359</emphasis>
(NICH).
</paragraph>
<paragraph id="FB1E8C5EFF88BF07FF473A9CFC059BAD" blockId="44.[150,1437,609,2002]" pageId="44">
<emphasis id="C9D5504CFF88BF07FF473A9CFE269AD5" bold="true" box="[199,440,1725,1750]" pageId="44">Taxonomic notes:—</emphasis>
The
<typeStatus id="241A32FCFF88BF07FE6C3A9FFD839AD5" box="[492,541,1726,1750]" pageId="44">type</typeStatus>
specimen was previously identified as
<taxonomicName id="3CA1F7DDFF88BF07FC623A9FFBD49AD5" authorityName="Sande Lacoste" authorityYear="1855" box="[994,1098,1725,1750]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Thysananthus" kingdom="Plantae" order="Porellales" pageId="44" phylum="Marchantiophyta" rank="species" species="planus">
<emphasis id="C9D5504CFF88BF07FC623A9FFBD49AD5" box="[994,1098,1725,1750]" italics="true" pageId="44">T. planus</emphasis>
</taxonomicName>
(=
<taxonomicName id="3CA1F7DDFF88BF07FBF43A9FFB7F9AD6" authorityName="B. M. Thiers &amp; Gradstein" authorityYear="1989" baseAuthorityName="Reinwardt, Blume &amp; Nees" baseAuthorityYear="1824" box="[1140,1249,1726,1749]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Thysananthus" kingdom="Plantae" order="Porellales" pageId="44" phylum="Marchantiophyta" rank="species" species="retusus">
<emphasis id="C9D5504CFF88BF07FBF43A9FFB7F9AD6" box="[1140,1249,1726,1749]" italics="true" pageId="44">T. retusus</emphasis>
</taxonomicName>
). By its oblong, entire leaves
<taxonomicName id="3CA1F7DDFF88BF07FEA93AC3FE2F9AFA" box="[297,433,1762,1785]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Thysananthus" kingdom="Plantae" order="Porellales" pageId="44" phylum="Marchantiophyta" rank="species" species="montanus">
<emphasis id="C9D5504CFF88BF07FEA93AC3FE2F9AFA" box="[297,433,1762,1785]" italics="true" pageId="44">T. montanus</emphasis>
</taxonomicName>
approaches
<taxonomicName id="3CA1F7DDFF88BF07FDBC3AC3FD399AFA" authorityName="B. M. Thiers &amp; Gradstein" authorityYear="1989" baseAuthorityName="Reinwardt, Blume &amp; Nees" baseAuthorityYear="1824" box="[572,679,1762,1785]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Thysananthus" kingdom="Plantae" order="Porellales" pageId="44" phylum="Marchantiophyta" rank="species" species="retusus">
<emphasis id="C9D5504CFF88BF07FDBC3AC3FD399AFA" box="[572,679,1762,1785]" italics="true" pageId="44">T. retusus</emphasis>
</taxonomicName>
but differs from the latter by (1) leaves when dry convolute (widely spreading, loosely reflexed or plane in
<taxonomicName id="3CA1F7DDFF88BF07FDD53B27FD5D9B1E" authorityName="B. M. Thiers &amp; Gradstein" authorityYear="1989" baseAuthorityName="Reinwardt, Blume &amp; Nees" baseAuthorityYear="1824" box="[597,707,1798,1821]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Thysananthus" kingdom="Plantae" order="Porellales" pageId="44" phylum="Marchantiophyta" rank="species" species="retusus">
<emphasis id="C9D5504CFF88BF07FDD53B27FD5D9B1E" box="[597,707,1798,1821]" italics="true" pageId="44">T. retusus</emphasis>
</taxonomicName>
), (2) dorsal base auriculate (cordate in
<taxonomicName id="3CA1F7DDFF88BF07FB063B27FB6A9B1E" authorityName="B. M. Thiers &amp; Gradstein" authorityYear="1989" baseAuthorityName="Reinwardt, Blume &amp; Nees" baseAuthorityYear="1824" box="[1158,1268,1798,1821]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Thysananthus" kingdom="Plantae" order="Porellales" pageId="44" phylum="Marchantiophyta" rank="species" species="retusus">
<emphasis id="C9D5504CFF88BF07FB063B27FB6A9B1E" box="[1158,1268,1798,1821]" italics="true" pageId="44">T. retusus</emphasis>
</taxonomicName>
), (3) leaf cells elongate-hexagonal, vitta 817 cell rows wide (subisodiametric,
<quantity id="3C5921BBFF88BF07FCE93B08FC2E9B41" box="[873,944,1833,1858]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="1.524" metricValueMin="1.016" pageId="44" unit="in" value="5.0" valueMax="6.0" valueMin="4.0">46 in</quantity>
<taxonomicName id="3CA1F7DDFF88BF07FC363B0BFBBC9B42" authorityName="B. M. Thiers &amp; Gradstein" authorityYear="1989" baseAuthorityName="Reinwardt, Blume &amp; Nees" baseAuthorityYear="1824" box="[950,1058,1834,1857]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Thysananthus" kingdom="Plantae" order="Porellales" pageId="44" phylum="Marchantiophyta" rank="species" species="retusus">
<emphasis id="C9D5504CFF88BF07FC363B0BFBBC9B42" box="[950,1058,1834,1857]" italics="true" pageId="44">T. retusus</emphasis>
</taxonomicName>
), (4) being dioicous (autoicous in
<taxonomicName id="3CA1F7DDFF88BF07FF173B6FFE9B9B66" authorityName="B. M. Thiers &amp; Gradstein" authorityYear="1989" baseAuthorityName="Reinwardt, Blume &amp; Nees" baseAuthorityYear="1824" box="[151,261,1870,1893]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Thysananthus" kingdom="Plantae" order="Porellales" pageId="44" phylum="Marchantiophyta" rank="species" species="retusus">
<emphasis id="C9D5504CFF88BF07FF173B6FFE9B9B66" box="[151,261,1870,1893]" italics="true" pageId="44">T. retusus</emphasis>
</taxonomicName>
), and (5) entire perianths (toothed perianths in
<taxonomicName id="3CA1F7DDFF88BF07FCA43B6FFC0C9B66" authorityName="B. M. Thiers &amp; Gradstein" authorityYear="1989" baseAuthorityName="Reinwardt, Blume &amp; Nees" baseAuthorityYear="1824" box="[804,914,1870,1893]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Thysananthus" kingdom="Plantae" order="Porellales" pageId="44" phylum="Marchantiophyta" rank="species" species="retusus">
<emphasis id="C9D5504CFF88BF07FCA43B6FFC0C9B66" box="[804,914,1870,1893]" italics="true" pageId="44">T. retusus</emphasis>
</taxonomicName>
).
<taxonomicName id="3CA1F7DDFF88BF07FC2A3B6CFB239B66" box="[938,1213,1869,1894]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Thysananthus" kingdom="Plantae" order="Porellales" pageId="44" phylum="Marchantiophyta" rank="species" species="montanus">
<emphasis id="C9D5504CFF88BF07FC2A3B6CFB239B66" box="[938,1213,1869,1894]" italics="true" pageId="44">Thysananthus montanus</emphasis>
</taxonomicName>
is morphologically most similar to
<taxonomicName id="3CA1F7DDFF88BF07FED33B53FE279B89" authorityName="Stephani" authorityYear="1912" box="[339,441,1905,1930]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Thysananthus" kingdom="Plantae" order="Porellales" pageId="44" phylum="Marchantiophyta" rank="species" species="mollis">
<emphasis id="C9D5504CFF88BF07FED33B53FE279B89" box="[339,441,1905,1930]" italics="true" pageId="44">T. mollis</emphasis>
</taxonomicName>
. The latter species differs essentially by its leaf, which is lanceolate in outline, the emarginate-lunulate apex of the underleaf, and the toothed perianths.
</paragraph>
<paragraph id="FB1E8C5EFF88BF07FF473B98FB3A9BD1" blockId="44.[150,1437,609,2002]" box="[199,1188,1977,2002]" pageId="44">
The female bracts and bracteoles in
<taxonomicName id="3CA1F7DDFF88BF07FDDB3B98FCF49BD2" box="[603,874,1977,2002]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Thysananthus" kingdom="Plantae" order="Porellales" pageId="44" phylum="Marchantiophyta" rank="species" species="montanus">
<emphasis id="C9D5504CFF88BF07FDDB3B98FCF49BD2" box="[603,874,1977,2002]" italics="true" pageId="44">Thysananthus montanus</emphasis>
</taxonomicName>
vary from entire to dentate.
</paragraph>
<paragraph id="FB1E8C5EFF88BF07FF153417FA3E944C" blockId="44.[149,610,2102,2123]" box="[149,1440,2098,2127]" lastBlockId="44.[956,1440,2098,2127]" pageId="44">
<taxonomicName id="3CA1F7DDFF88BF07FF153417FE859448" box="[149,283,2102,2123]" class="Jungermanniopsida" family="Lejeuneaceae" genus="Thysananthus" kingdom="Plantae" order="Porellales" pageId="44" phylum="Marchantiophyta" rank="genus">
<emphasis id="C9D5504CFF88BF07FF153417FE859448" box="[149,283,2102,2123]" italics="true" pageId="44">Thysananthus</emphasis>
</taxonomicName>
(
<taxonomicName id="3CA1F7DDFF88BF07FEA73417FE309448" baseAuthorityName="Gradstein" baseAuthorityYear="1994" box="[295,430,2102,2123]" class="Jungermanniopsida" family="Lejeuneaceae" kingdom="Plantae" order="Porellales" pageId="44" phylum="Marchantiophyta" rank="family">Lejeuneaceae</taxonomicName>
, Marchantiophyta)
<emphasis id="C9D5504CFF88BF07FC3C3416FB83944F" box="[956,1053,2103,2124]" italics="true" pageId="44">Phytotaxa</emphasis>
193 (1) © 2015 Magnolia Press •
<emphasis id="C9D5504CFF88BF07FA003413FA3E944C" bold="true" box="[1408,1440,2098,2127]" pageId="44">45</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>