197 lines
31 KiB
XML
197 lines
31 KiB
XML
<document ID-CLB-Dataset="6710" ID-DOI="http://dx.doi.org/10.3897/dez.63.10241" ID-GBIF-Dataset="0a6965a7-b124-4871-95d4-9e35c49073de" ID-Pensoft-Pub="1860-1324-2-211" ID-Pensoft-UUID="51243BBBE9F158F58BBC9BF3EA964017" ID-Zenodo-Dep="167458" ID-ZooBank="45552C4EC6AE4F9499980C2D492333B3" ModsDocID="1860-1324-63-2-211" checkinTime="1621372567443" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2016" docId="2C7D280DE6B4BC2F982E21646ECA9D82" docLanguage="en" docName="DeutEntomolZeit 63(2): 211-270" docOrigin="Deutsche Entomologische Zeitschrift 63 (2)" docPubDate="2016-11-18" docSource="http://dx.doi.org/10.3897/dez.63.10241" docTitle="Cyphocoleus burwelli Liebherr 2016, sp. n." docType="treatment" docUuid="B5B3D4B3-E303-4330-B69C-7EB910795E93" docUuidSource="ZooBank" docVersion="5" id="51243BBBE9F158F58BBC9BF3EA964017" lastPageNumber="233" masterDocId="51243BBBE9F158F58BBC9BF3EA964017" masterDocTitle="Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia" masterLastPageNumber="270" masterPageNumber="211" pageNumber="233" updateTime="1691866435949" updateUser="ExternalLinkService">
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<mods:title id="3A90D4F28C141752884D9B1EB33987A2">Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia</mods:title>
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<mods:namePart id="7357ED5B410D9B76F50F11A053E1A3E0">Liebherr, James K.</mods:namePart>
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<mods:affiliation id="9BE35953C5C21C23A6ED1A06CD5BFA00">Cornell University Insect Collection, John H. and Anna B. Comstock Hall, Cornell University, Ithaca, NY 14853 - 2601, USA</mods:affiliation>
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<mods:nameIdentifier id="813938D502D2FBB61B3C9C5DBC504BEF" type="email">jkl5@cornell.edu</mods:nameIdentifier>
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<mods:title id="DCA890B7FB3A52F8BD62D3640564FA78">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date id="C8ED158CB9C1F6D07348A809121358F9">2016</mods:date>
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<mods:number id="C815542A94B3DE20F36A4CE35D11E9DD">2016-11-18</mods:number>
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<mods:number id="34556E8F14DB12ACCCC6FFFB374B109E">63</mods:number>
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<treatment id="2C7D280DE6B4BC2F982E21646ECA9D82" ID-GBIF-Taxon="182223725" LSID="urn:lsid:zoobank.org:act:B5B3D4B3-E303-4330-B69C-7EB910795E93" httpUri="http://treatment.plazi.org/id/2C7D280DE6B4BC2F982E21646ECA9D82" lastPageNumber="233" pageId="22" pageNumber="233">
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<subSubSection id="C5FEFBB7BFDD74C82FB8CC09FFFF17BF" pageId="22" pageNumber="233" type="nomenclature">
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<paragraph id="DD0E2349385C93A3D5B82ED1CAB8BE10" pageId="22" pageNumber="233">
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6.
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<taxonomicName id="4060A936A238A20EDCF1CED085797A6A" ID-CoL="6BWYX" LSID="http://zoobank.org/B5B3D4B3-E303-4330-B69C-7EB910795E93" authority="Liebherr, 2016" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus burwelli" order="Coleoptera" pageId="22" pageNumber="233" phylum="Arthropoda" rank="species" species="burwelli" status="sp. n.">Cyphocoleus burwelli</taxonomicName>
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<taxonomicNameLabel id="EF4AA154FECD363B3BED973A260D5689" pageId="22" pageNumber="233">sp. n.</taxonomicNameLabel>
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<figureCitation id="4AC4A1246E49261E7D5D556E7EB30974" captionStart="Figures 66–75" captionStartId="F13" captionText="Figures 66 - 75. Female right gonocoxae, ventral view, for Cyphocoleus spp. illustrating apical fringe setae of basal gonocoxite 1, and two or three lateral ensiform setae, one dorsal ensiform seta, and two apical nematiform setae on apical gonocoxite 2; 66, Cyphocoleus lissus, Mt. Humboldt; 67, Cyphocoleus prolixus, Mt. Humboldt; 68, Cyphocoleus heterogenus, Yahoue; 69, Cyphocoleus parovicollis, Mandjelia; 70, Cyphocoleus ovicollis, Ningua Res.; 71, Cyphocoleus burwelli, Mt. Humboldt; 72, Cyphocoleus monteithi, Mt. Mou; 73, Cyphocoleus miricollis, Foret Nord; 74, Cyphocoleus mirabilis, Mt. Koghi; 75, Cyphocoleus lescheni, apical gonocoxite damaged, Aoupinie. Scale bar, 0.10 mm." figureDoi="10.3897/dez.63.10241.figures66-75" httpUri="https://binary.pensoft.net/fig/111550" pageId="22" pageNumber="233">Figures 71</figureCitation>
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<figureCitation id="460A1BEB62971195D1F53F5AE526C8C5" captionStart="Figures 76–81" captionStartId="F14" captionText="Figures 76 - 81. Distribution maps of Cyphocoleus spp.: 76, Cyphocoleus lissus; 77, Cyphocoleus heterogenus, Cyphocoleus prolixus; 78, Cyphocoleus parovicollis, Cyphocoleus ovicollis; 79, Cyphocoleus burwelli, Cyphocoleus angustatus, Cyphocoleus monteithi; 80, Cyphocoleus miricollis, Cyphocoleus fasciatus; 81, Cyphocoleus mirabilis, Cyphocoleus moorei, Cyphocoleus lescheni." figureDoi="10.3897/dez.63.10241.figures76-81" httpUri="https://binary.pensoft.net/fig/111551" pageId="22" pageNumber="233">, 79</figureCitation>
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<figureCitation id="3EA90451EC250412DC63824641232BBA" captionStart="Figures 82–85" captionStartId="F15" captionText="Figures 82 - 85. Dorsal habitus photographs of Cyphocoleus spp.: 82, Cyphocoleus ovicollis female, Ningua Res.; 83, Cyphocoleus burwelli female, Mt. Canala; 84, Cyphocoleus angustatus male, Mt. Panie; 85, Cyphocoleus monteithi male, Dzumac Rd. jct." figureDoi="10.3897/dez.63.10241.figures82-85" httpUri="https://binary.pensoft.net/fig/111552" pageId="22" pageNumber="233">, 83</figureCitation>
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<figureCitation id="83443A645E3C9B5CBC9F74B97C8984E3" captionStart="Figures 86–98" captionStartId="F16" captionText="Figures 86 - 98. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 86, Cyphocoleus burwelli, internal sac everted, Ningua Res.; 87, Cyphocoleus angustatus, dextroventral view, internal sac everted, Mt. Panie; 88, Cyphocoleus monteithi, Parque Prov. Riviere Bleue, Haute Pourina; 89, Cyphocoleus miricollis, Mt. Mou; 90, Cyphocoleus fasciatus, dextral view, sac partially everted, Mt. Panie; 91 - 98, Cyphocoleus cychroides; 91, internal sac everted, Mandjelia; 92 - 93, Mandjelia; 94, internal sac everted, Hienghene; 95, Touho; 96, Tionaka Tal; 97, Ateou; 98, Pic d'Amoa. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation." figureDoi="10.3897/dez.63.10241.figures86-98" httpUri="https://binary.pensoft.net/fig/111553" pageId="22" pageNumber="233">, 86</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="D79E76F4EE3007CD595F92FBFE40A70F" pageId="22" pageNumber="233" type="diagnosis">
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<paragraph id="40F5AEDE4BCCBB487F814DF426020B0F" pageId="22" pageNumber="233">Diagnosis.</paragraph>
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<paragraph id="BFF8CBDB82CD087F21E9987512DFD798" pageId="22" pageNumber="233">
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This species and the following,
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<taxonomicName id="988204DDEDCA04F9E6ECB4AEEA731163" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus angustatus" order="Diptera" pageId="22" pageNumber="233" phylum="Arthropoda" rank="species" species="angustatus">
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<emphasis id="1A27FF8CEB59CD6904BBBA6D98A179B0" italics="true" pageId="22" pageNumber="233">Cyphocoleus angustatus</emphasis>
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</taxonomicName>
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, comprise a second sibling species pair, these two characterized by the narrow body: i.e. pronotum and elytra. The pronotum of
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<taxonomicName id="0661C2F33CE5CC411311E08F72A1C5A8" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus burwelli" order="Diptera" pageId="22" pageNumber="233" phylum="Arthropoda" rank="species" species="burwelli">
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<emphasis id="41DC5CB5444CDF3CC7DCFB18344E95D4" italics="true" pageId="22" pageNumber="233">Cyphocoleus burwelli</emphasis>
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</taxonomicName>
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is more evenly ovate, with MPW/BPW = 1.72-1.83 versus values of 1.84-1.91 for
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<taxonomicName id="00C17B75EEBED3686247A46284C3A217" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus angustatus" order="Diptera" pageId="22" pageNumber="233" phylum="Arthropoda" rank="species" species="angustatus">
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<emphasis id="87F8F5A429F3F93EB059CCC2246AEF44" italics="true" pageId="22" pageNumber="233">Cyphocoleus angustatus</emphasis>
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</taxonomicName>
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. The pronotum is also relatively narrower overall, with MPW/PL = 0.68-0.72 compared to values of 0.72-0.77 recorded for
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<taxonomicName id="EDE9EC11828762A8A1C22191BC190CE8" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus angustatus" order="Diptera" pageId="22" pageNumber="233" phylum="Arthropoda" rank="species" species="angustatus">
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<emphasis id="CDB0D0086228FB86C05B4969DD15BABB" italics="true" pageId="22" pageNumber="233">Cyphocoleus angustatus</emphasis>
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</taxonomicName>
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. The elytra are narrower basally in
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<taxonomicName id="DC16D43474E7550FA9C2761648C9204E" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus burwelli" order="Diptera" pageId="22" pageNumber="233" phylum="Arthropoda" rank="species" species="burwelli">
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<emphasis id="7CC94A81E30534EFA3E82CA5CF181A77" italics="true" pageId="22" pageNumber="233">Cyphocoleus burwelli</emphasis>
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</taxonomicName>
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, with MEW/HuW = 2.78-3.19 versus 2.60-2.79 for
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<taxonomicName id="23B0F04FBA7C0DD9B0EBE26350835B6D" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus angustatus" order="Diptera" pageId="22" pageNumber="233" phylum="Arthropoda" rank="species" species="angustatus">
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<emphasis id="1D8C60AAC71DAF3273FE0E581E43D1F2" italics="true" pageId="22" pageNumber="233">Cyphocoleus angustatus</emphasis>
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</taxonomicName>
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, and the humeral angle is obtuse versus very distinctly right angled to slightly acute in
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<taxonomicName id="0A756227A35E22C9B08F2C5148235857" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus angustatus" order="Diptera" pageId="22" pageNumber="233" phylum="Arthropoda" rank="species" species="angustatus">
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<emphasis id="CDB7BC268ACA90036A9E3A5430D05E9D" italics="true" pageId="22" pageNumber="233">Cyphocoleus angustatus</emphasis>
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</taxonomicName>
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(Figs
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<figureCitation id="187112A712640F2807E1A50797CAF59A" captionStart="Figures 82–85" captionStartId="F15" captionText="Figures 82 - 85. Dorsal habitus photographs of Cyphocoleus spp.: 82, Cyphocoleus ovicollis female, Ningua Res.; 83, Cyphocoleus burwelli female, Mt. Canala; 84, Cyphocoleus angustatus male, Mt. Panie; 85, Cyphocoleus monteithi male, Dzumac Rd. jct." figureDoi="10.3897/dez.63.10241.figures82-85" httpUri="https://binary.pensoft.net/fig/111552" pageId="22" pageNumber="233">83</figureCitation>
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,
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<figureCitation id="E7AF0619A79F3B855A07BF03C6520704" captionStart="Figures 82–85" captionStartId="F15" captionText="Figures 82 - 85. Dorsal habitus photographs of Cyphocoleus spp.: 82, Cyphocoleus ovicollis female, Ningua Res.; 83, Cyphocoleus burwelli female, Mt. Canala; 84, Cyphocoleus angustatus male, Mt. Panie; 85, Cyphocoleus monteithi male, Dzumac Rd. jct." figureDoi="10.3897/dez.63.10241.figures82-85" httpUri="https://binary.pensoft.net/fig/111552" pageId="22" pageNumber="233">84</figureCitation>
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). A male dissection can be used to confirm this diagnosis, with the aedeagal median lobe apex narrowly rounded in this species, and the internal sac bearing a stout apical tooth (Fig.
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<figureCitation id="41FFDD70426965B7155FA7F2E94AAFF5" captionStart="Figures 86–98" captionStartId="F16" captionText="Figures 86 - 98. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 86, Cyphocoleus burwelli, internal sac everted, Ningua Res.; 87, Cyphocoleus angustatus, dextroventral view, internal sac everted, Mt. Panie; 88, Cyphocoleus monteithi, Parque Prov. Riviere Bleue, Haute Pourina; 89, Cyphocoleus miricollis, Mt. Mou; 90, Cyphocoleus fasciatus, dextral view, sac partially everted, Mt. Panie; 91 - 98, Cyphocoleus cychroides; 91, internal sac everted, Mandjelia; 92 - 93, Mandjelia; 94, internal sac everted, Hienghene; 95, Touho; 96, Tionaka Tal; 97, Ateou; 98, Pic d'Amoa. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation." figureDoi="10.3897/dez.63.10241.figures86-98" httpUri="https://binary.pensoft.net/fig/111553" pageId="22" pageNumber="233">86</figureCitation>
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). Standardized body length 11.1-12.2 mm.
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</paragraph>
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</subSubSection>
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<subSubSection id="DADF3503986CC412EA94DB22CAE47D12" pageId="22" pageNumber="233" type="description">
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<paragraph id="AF3750F8F98A8195280A7D22D067E673" pageId="22" pageNumber="233">Description</paragraph>
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<paragraph id="496935669CD1A5EC1D0007D64F29ED4C" pageId="22" pageNumber="233">
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(n = 4). Head capsule moderately elongate, trapezoidal, with broad flange basad posterior mandibular articulation, neck distinctly constricted (Fig.
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<figureCitation id="772815463F7A29548C99AE51FFA5420C" captionStart="Figures 82–85" captionStartId="F15" captionText="Figures 82 - 85. Dorsal habitus photographs of Cyphocoleus spp.: 82, Cyphocoleus ovicollis female, Ningua Res.; 83, Cyphocoleus burwelli female, Mt. Canala; 84, Cyphocoleus angustatus male, Mt. Panie; 85, Cyphocoleus monteithi male, Dzumac Rd. jct." figureDoi="10.3897/dez.63.10241.figures82-85" httpUri="https://binary.pensoft.net/fig/111552" pageId="22" pageNumber="233">83</figureCitation>
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); frons broadly convex between hind margin of eyes, with shallow median dimple and progressively broadened frontal grooves anteriorly; eyes small and little convex, 23-28 ommatidia across horizontal diameter; supraorbital setae absent; mandibles elongate, length 2.4
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<normalizedToken id="86B44AB955B55DF8695D1DE3487CFE31" originalValue="×">x</normalizedToken>
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distance from antennal articulatory socket to lateroapical margin of labrum; antennae elongate, scape length 4.2
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<normalizedToken id="EF2E060289A454D76C81882BAEC4CD3A" originalValue="×">x</normalizedToken>
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maximal breadth; subgenal seta absent. Pronotum with front angles only slightly protruded, obtuse angulate; lateral marginal bead continuous to just mesad evenly rounded hind angles, basal margin not beaded medially; proepipleuron and proepisternum bulging outward in basal half of pronotal length, visible in dorsal view; median longitudinal impression shallowly and finely incised, wavering along length; anterior transverse impression very shallow, barely traceable as oblique impression that reaches lateral marginal bead behind front angle; prosternal process flat anteriorly, flat to slightly depressed medially on ventral surface, convex posteriorly between procoxae. Elytra narrowly ellipsoid, moderately convex; elytral striae deep, impunctate, associated intervals moderately convex; parascutellar seta present, articulatory socket not upraised above surface of elytra; humeral angles obtuse-angulate, the juncture of basal and marginal grooves tightly rounded; lateral margin evenly convex outside anterior series of lateral elytral setae; two to three dorsal elytral setae present (setae at anterior and middle position present or absent, seta at posterior position always present); lateral elytral setae arranged as 6 + (7)8; subapical and apical elytral setae present; subapical sinuation slightly, evenly concave, sinuation meeting lateral margin in broadly convex curve; elytral apices rounded, the apical margin invaginated to meet fused elytral suture. Mesepisternum impunctate; metepisternal dorsal length 0.9
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<normalizedToken id="738348B8D6428CF4D1130F2C3037168D" originalValue="×">x</normalizedToken>
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diagonal width. Abdomen of males with apical ventrite distinctly notched medially; two setae on apical margin of apical ventrite in males (one seta each side), four setae (two each side) present on apical margin of females. Legs elongate, gracile, mt1 length/tibial length = 0.27; metacoxae bisetose; tarsomeres with short, sparsely distributed setae on dorsal surface; mt4 length of outer lobe 1.3
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<normalizedToken id="D7997C790E36AEE528DAAA629B5E7F7D" originalValue="×">x</normalizedToken>
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median length, 6-8 ventrolateral setae each side. Microsculpture of frons isodiametric, longitudinally stretched near median dimple, head microsculpture transversely stretched on clypeus and further posterad on vertex, becoming dense transverse lines before neck constriction; pronotal and elytral disc with dense transverse-line microsculpture; pro-, meso-, and metasternum, metacoxae and abdominal ventrites with sparsely distributed pelage of moderately elongate microsetae. Coloration of head and pronotum piceous with metallic cyanotic iridescence due to microsculpture; elytra rufopiceous with similar iridescence; ventral body surface uniformly dark, piceous to rufopiceous on medial portions of apical three abdominal ventrites, apical ventrite with pale, rufoflavous apical margin; legs slightly paler than ventrites, femora and tibiae rufobrunneous with piceous case, tarsi brunneous.
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</paragraph>
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<paragraph id="BF81EAD0B6BCF2F1B474A98539A0C149" pageId="22" pageNumber="233">
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<emphasis id="83D31309AB63F9828A97DA485E72E2D9" italics="true" pageId="22" pageNumber="233">Male genitalia</emphasis>
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(n = 1). Male aedeagal median lobe robust, broadly parallel sided in basal half, then tapered to rounded tip, dorsoventral breadth at midlength 0.3
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<normalizedToken id="ABED253600BD0CAEF0E5BD6C5E023F31" originalValue="×">x</normalizedToken>
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distance from tip to base of closed basal bulb (Fig.
|
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<figureCitation id="9A9A757ED4821F75BED1904998D2EBE4" captionStart="Figures 86–98" captionStartId="F16" captionText="Figures 86 - 98. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 86, Cyphocoleus burwelli, internal sac everted, Ningua Res.; 87, Cyphocoleus angustatus, dextroventral view, internal sac everted, Mt. Panie; 88, Cyphocoleus monteithi, Parque Prov. Riviere Bleue, Haute Pourina; 89, Cyphocoleus miricollis, Mt. Mou; 90, Cyphocoleus fasciatus, dextral view, sac partially everted, Mt. Panie; 91 - 98, Cyphocoleus cychroides; 91, internal sac everted, Mandjelia; 92 - 93, Mandjelia; 94, internal sac everted, Hienghene; 95, Touho; 96, Tionaka Tal; 97, Ateou; 98, Pic d'Amoa. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation." figureDoi="10.3897/dez.63.10241.figures86-98" httpUri="https://binary.pensoft.net/fig/111553" pageId="22" pageNumber="233">86</figureCitation>
|
||
); lobe shaft brunneous, the internal sac obscured in uneverted position by the moderately melanized lobe wall; lobe apex with tapered tip, length distad ostial opening subequal to breadth; internal sac broadest at midlength, with extremely large sclerotized apical tooth and a broad ring of melanized microtrichia over middle half of sac (Fig.
|
||
<figureCitation id="5F4A1892ACADACD0BBC9D0E9E3DC51DD" captionStart="Figures 86–98" captionStartId="F16" captionText="Figures 86 - 98. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 86, Cyphocoleus burwelli, internal sac everted, Ningua Res.; 87, Cyphocoleus angustatus, dextroventral view, internal sac everted, Mt. Panie; 88, Cyphocoleus monteithi, Parque Prov. Riviere Bleue, Haute Pourina; 89, Cyphocoleus miricollis, Mt. Mou; 90, Cyphocoleus fasciatus, dextral view, sac partially everted, Mt. Panie; 91 - 98, Cyphocoleus cychroides; 91, internal sac everted, Mandjelia; 92 - 93, Mandjelia; 94, internal sac everted, Hienghene; 95, Touho; 96, Tionaka Tal; 97, Ateou; 98, Pic d'Amoa. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation." figureDoi="10.3897/dez.63.10241.figures86-98" httpUri="https://binary.pensoft.net/fig/111553" pageId="22" pageNumber="233">86</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A6A701C9CB917A705C642C1A1D14C1A8" pageId="22" pageNumber="233">
|
||
<emphasis id="C34EE8F8C3DDFB8C4A5508B3BFA5AB5A" italics="true" pageId="22" pageNumber="233">Female reproductive tract</emphasis>
|
||
(n = 1). Bursa copulatrix vase-shaped, basally stalked, distance from base of gonocoxites to spermathecal basal sclerite 1.4
|
||
<normalizedToken id="EC3373E09AA14C5630DC5CD5C1AEE5EA" originalValue="×">x</normalizedToken>
|
||
maximum breadth (dissection compressed under cover slip) (as in Fig.
|
||
<figureCitation id="CA5E944BA3C5538602CA54E4C92D0854" captionStart="Figures 60–65" captionStartId="F12" captionText="Figures 60 - 65. Female reproductive tract, ventral view, for Cyphocoleus spp.; 60, Cyphocoleus lissus, Mt. Humboldt; 61, Cyphocoleus prolixus, Mt. Humboldt; 62, Cyphocoleus heterogenus, Yahoue; 63, Cyphocoleus parovicollis, Mandjelia; 64, Cyphocoleus monteithi, Mt. Mou; 65, Cyphocoleus miricollis, Foret Nord. For abbreviations see Table 1." figureDoi="10.3897/dez.63.10241.figures60-65" httpUri="https://binary.pensoft.net/fig/111549" pageId="22" pageNumber="233">63</figureCitation>
|
||
); bursal walls thick, heavily pigmented by Chlorazol Black stain, with broad band of thick, densely packed spicules lining lumen over broadest part of bursa; basal gonocoxite with apical fringe of six to seven setae (Fig.
|
||
<figureCitation id="9477AA7DFC9F8F714EB6B2B032BA4EBB" captionStart="Figures 66–75" captionStartId="F13" captionText="Figures 66 - 75. Female right gonocoxae, ventral view, for Cyphocoleus spp. illustrating apical fringe setae of basal gonocoxite 1, and two or three lateral ensiform setae, one dorsal ensiform seta, and two apical nematiform setae on apical gonocoxite 2; 66, Cyphocoleus lissus, Mt. Humboldt; 67, Cyphocoleus prolixus, Mt. Humboldt; 68, Cyphocoleus heterogenus, Yahoue; 69, Cyphocoleus parovicollis, Mandjelia; 70, Cyphocoleus ovicollis, Ningua Res.; 71, Cyphocoleus burwelli, Mt. Humboldt; 72, Cyphocoleus monteithi, Mt. Mou; 73, Cyphocoleus miricollis, Foret Nord; 74, Cyphocoleus mirabilis, Mt. Koghi; 75, Cyphocoleus lescheni, apical gonocoxite damaged, Aoupinie. Scale bar, 0.10 mm." figureDoi="10.3897/dez.63.10241.figures66-75" httpUri="https://binary.pensoft.net/fig/111550" pageId="22" pageNumber="233">71</figureCitation>
|
||
); apical gonocoxite triangular with rounded apex (due to wear?) and three lateral ensiform setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8B7F67BCA460EBDD94228B985AE42A70" pageId="22" pageNumber="233" type="types">
|
||
<paragraph id="31DDE91B39F2C8C9C66DF1E1B813A11B" pageId="22" pageNumber="233">Types.</paragraph>
|
||
<paragraph id="7422B178D3AE8ECEE385D9282C6CBBF5" pageId="22" pageNumber="233">
|
||
Holotype female (MNHB deposited in MNHN): Drs. F. Sarasin&J. Roux / Neukaledonien / Mt. Humboldt (on obverse of blue label) / Gipsel, 1600 m. / 18.9.1911 (on reverse of blue label) //
|
||
<taxonomicName id="A203E1C3DA3F6F5F7FBDB0818FBBBD0D" authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus" order="Diptera" pageId="22" pageNumber="233" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4247A24C257851E5DC74FB165EF0AF4F" italics="true" pageId="22" pageNumber="233">Cyphocoleus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="834D20ED68ACAFDB69EE1A2DF2F70ACF" authorityName="Fauvel" authorityYear="1882" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="ovicollis" order="Diptera" pageId="22" pageNumber="233" phylum="Arthropoda" rank="species" species="ovicollis">
|
||
<emphasis id="08DAB4BD7AD6279FDD3E7873E4BC47E6" italics="true" pageId="22" pageNumber="233">ovicollis</emphasis>
|
||
</taxonomicName>
|
||
Fauv. / Det. K.M. Heller 1914 //
|
||
<taxonomicName id="7932B127BFE01B4AC91E73EF12E28E0A" authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus" order="Diptera" pageId="22" pageNumber="233" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6FACE7A55E95473A53A5EBCE17D9694F" italics="true" pageId="22" pageNumber="233">Cyphocoleus</emphasis>
|
||
</taxonomicName>
|
||
revision / measured specimen 1 / J.K.
|
||
<bibRefCitation id="2B28348E58CCE5822063E510C61C3B32" author="Liebherr, JK" journalOrPublisher="ZooKeys" pageId="49" pageNumber="260" pagination="1 - 407" publicationUrl="10.3897/zookeys.544.6074" refId="B33" refString="Liebherr, JK, 2015. The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1 - 407, DOI: 10.3897/zookeys.544.6074" title="The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala, Maui: Keystone of a hyperdiverse Hawaiian radiation." url="10.3897/zookeys.544.6074" volume="544" year="2015">Liebherr 2015</bibRefCitation>
|
||
// HOLOTYPE /
|
||
<taxonomicName id="E0BFB56504F66078C9923D78F7DEEF02" authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus" order="Diptera" pageId="22" pageNumber="233" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="44C70A2A3C9EE9EBD89B25D2F470C693" italics="true" pageId="22" pageNumber="233">Cyphocoleus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="B391DE31AABDE201EABC097B5A81CD2C" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="burwelli" order="Diptera" pageId="22" pageNumber="233" phylum="Arthropoda" rank="species" species="burwelli">
|
||
<emphasis id="0E00B00148EFAD4B879A506A6E50E6B8" italics="true" pageId="22" pageNumber="233">burwelli</emphasis>
|
||
</taxonomicName>
|
||
/ J.K. Liebherr 2016 (black-bordered red label).
|
||
</paragraph>
|
||
<paragraph id="0A6A300A2782DE6AC3C9E53A0EE9ED33" pageId="22" pageNumber="233">
|
||
Paratypes (3 specimens). NEW CALEDONIA: Ningua Reserve, near summit, 1300 m el.,
|
||
<geoCoordinate id="266E3A0FD1AA772B9DA6FE1D7BBE2449" degrees="21" direction="south" minutes="45" orientation="latitude" precision="925" value="-21.75">21°45'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="121B08EC3265883A52EE3EF926F47995" degrees="166" direction="east" minutes="09" orientation="longitude" precision="925" value="166.15">166°09'E</geoCoordinate>
|
||
, 13-xi-2001, Burwell & Monteith (QMB, 1); Mt. Humboldt, 1600 m el.,
|
||
<geoCoordinate id="844FE1A36B73BD5DD5AE945DE5574351" degrees="21" direction="south" minutes="53" orientation="latitude" precision="925" value="-21.883333">21°53'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1C6D65B7C40833954A10C77D110061E5" degrees="166" direction="east" minutes="26" orientation="longitude" precision="925" value="166.43333">166°26'E</geoCoordinate>
|
||
, 18-ix-1911, Sarasin & Roux (SMTD, 1); Mt. Kanala, Wald, 800-1000 m el.,
|
||
<geoCoordinate id="CB9196ACA365FDAB541029082C53222E" degrees="21" direction="south" minutes="34" orientation="latitude" precision="925" value="-21.566668">21°34'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="C42792398A8B783BA86FF0BB95C4F7E0" degrees="165" direction="east" minutes="56" orientation="longitude" precision="925" value="165.93333">165°56'E</geoCoordinate>
|
||
, 04-xi-1911, Sarasin/Roux (MNHB, 1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4C2EFE138C59737FA877962F007232B4" pageId="22" pageNumber="233" type="etymology">
|
||
<paragraph id="18973050F1A23738CCA02FBF95639D85" pageId="22" pageNumber="233">Etymology.</paragraph>
|
||
<paragraph id="8B530A46D90C1C7CF6A69308429C1E6C" pageId="22" pageNumber="233">
|
||
This species is named for Dr. Chris Burwell, Senior Curator of Insects, Queensland Museum, in recognition of the numerous specimens of
|
||
<taxonomicName id="7347283181F041B355434F6C4AC8E08D" authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus" order="Diptera" pageId="22" pageNumber="233" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A14C446434E3E520282E8CB95BACA7B1" italics="true" pageId="22" pageNumber="233">Cyphocoleus</emphasis>
|
||
</taxonomicName>
|
||
he has collected: many representing previously undescribed species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CF706FC1295E1E4A9012ECDD29DC73B6" pageId="22" pageNumber="233" type="distribution">
|
||
<paragraph id="F39FF56DE157680C9384651D3BDA9226" pageId="22" pageNumber="233">Distribution and habitat.</paragraph>
|
||
<paragraph id="1682C58D68785863D032A441E69D7B12" pageId="22" pageNumber="233">
|
||
The distribution of this species ranges from Mt. Humboldt on the south, to Mt. Canala on the north (Fig.
|
||
<figureCitation id="81FED931AF22D5F68163B0E8B58ADE7D" captionStart="Figures 76–81" captionStartId="F14" captionText="Figures 76 - 81. Distribution maps of Cyphocoleus spp.: 76, Cyphocoleus lissus; 77, Cyphocoleus heterogenus, Cyphocoleus prolixus; 78, Cyphocoleus parovicollis, Cyphocoleus ovicollis; 79, Cyphocoleus burwelli, Cyphocoleus angustatus, Cyphocoleus monteithi; 80, Cyphocoleus miricollis, Cyphocoleus fasciatus; 81, Cyphocoleus mirabilis, Cyphocoleus moorei, Cyphocoleus lescheni." figureDoi="10.3897/dez.63.10241.figures76-81" httpUri="https://binary.pensoft.net/fig/111551" pageId="22" pageNumber="233">79</figureCitation>
|
||
). Collection sites range in elevation from 1000-1600 m.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |