treatments-xml/data/03/DD/A3/03DDA36B7D50FFECFF08FEFB01B1FB8C.xml
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<document id="04EEAFADEADE572477D08CB3EA67A8CA" ID-ISSN="1638-9387" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630324947137" checkinUser="marcus" docAuthor="Kitahara, Marcelo V. &amp; Cairns, Stephen D." docDate="2009" docId="03DDA36B7D50FFECFF08FEFB01B1FB8C" docLanguage="en" docName="Zoosystema.2009.31.2.233-248.pdf" docOrigin="Zoosystema 31 (2)" docStyle="DocumentStyle:E638A3C99FA728B529662B46F84D75FD.3:Zoosystema.2009-2010.journal_article" docStyleId="E638A3C99FA728B529662B46F84D75FD" docStyleName="Zoosystema.2009-2010.journal_article" docStyleVersion="3" docTitle="Deltocyathus crassiseptum Cairns 1999" docType="treatment" docVersion="3" lastPageNumber="244" masterDocId="FFE4DB137D5BFFE7FFAEFFAD0269FFA7" masterDocTitle="A revision of the genus Deltocyathus Milne Edwards &amp; Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species" masterLastPageNumber="248" masterPageNumber="233" pageNumber="244" updateTime="1699228079138" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="99B2D6D69728FD0BB2D8B0C621166836" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="4E2E8AA8A3CD6EB744A256EF729F3BD7">A revision of the genus Deltocyathus Milne Edwards &amp; Haime, 1848 (Scleractinia, Caryophylliidae) from New Caledonia, with the description of a new species</mods:title>
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<mods:name id="754F6CED73DA2C6841CD1D8FD02C7B64" type="personal">
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<mods:roleTerm id="6B9DE7897031BFF3E1DC4705318B6BB8">Author</mods:roleTerm>
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<mods:namePart id="83299A6DA0D72039EFD0B90D0E704578">Kitahara, Marcelo V.</mods:namePart>
<mods:affiliation id="65F0574913A8F0BB8A69B0EDBAE14162">ARC Centre of Excellence for Coral Reef Studies and Coral Genomics Group, Molecular Science Bld, Annex, James Cook University, Douglas Campus, Townsville, Qld 4811 (Australia) marcelo. visentinikitahara @ jcu. edu. au</mods:affiliation>
</mods:name>
<mods:name id="7403508D8F31252E49BD1A618D7A159E" type="personal">
<mods:role id="33F1A24E780AAFBB5BC12627E0EFB229">
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<mods:namePart id="42934FD46B5B8762AB6E5AE2581345A8">Cairns, Stephen D.</mods:namePart>
<mods:affiliation id="AF89000BC9FBACA5AD6AE917619B7DFE">National Museum of Natural History, Smithsonian Institution, Department of Zoology (Invertebrate Zoology), P. O. Box 37012, Washington, DC, 20013 - 7012 (USA) cairnss @ si. edu</mods:affiliation>
<mods:nameIdentifier id="752C55D46B5325E11CC7435C7F1C3E7A" type="email">cairnss@si.edu</mods:nameIdentifier>
</mods:name>
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<mods:title id="060080C05E0B898C19C3E430F986984F">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="3E9708A3FBDD73CE89E2AD09DD679F8E">
<mods:date id="389AAE69BAFBE07C31527BF51C920BBD">2009</mods:date>
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<mods:number id="7E114B8B7A4FC77A0B3476D3B8A4D38F">31</mods:number>
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<mods:classification id="04C5FE41FC5E171D861AB2F3BF7A403E">journal article</mods:classification>
<mods:identifier id="DBB06A1B13AD1D12F431671F8363FEB4" type="ISSN">1638-9387</mods:identifier>
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<treatment id="03DDA36B7D50FFECFF08FEFB01B1FB8C" ID-DOI="http://doi.org/10.5281/zenodo.5470122" ID-Zenodo-Dep="5470122" LSID="urn:lsid:plazi:treatment:03DDA36B7D50FFECFF08FEFB01B1FB8C" httpUri="http://treatment.plazi.org/id/03DDA36B7D50FFECFF08FEFB01B1FB8C" lastPageNumber="244" pageId="11" pageNumber="244">
<subSubSection id="C36E41F67D50FFECFF08FEFB03C6FE36" pageId="11" pageNumber="244" type="nomenclature">
<paragraph id="8BCB127D7D50FFECFF08FEFB03C6FE36" blockId="11.[166,594,342,401]" pageId="11" pageNumber="244">
<heading id="D083A5117D50FFECFF08FEFB03C6FE36" centered="true" fontSize="11" level="2" pageId="11" pageNumber="244" reason="2">
<taxonomicName id="4C7469FE7D50FFECFF08FEFB003BFED5" ID-CoL="34KF6" authority="Cairns, 1999" authorityName="Cairns" authorityYear="1999" box="[166,594,342,370]" class="Anthozoa" family="Deltocyathidae" genus="Deltocyathus" kingdom="Animalia" order="Scleractinia" pageId="11" pageNumber="244" phylum="Cnidaria" rank="species" species="crassiseptum">
<emphasis id="B900CE6F7D50FFECFF08FEFB03D7FED7" bold="true" box="[166,446,342,369]" italics="true" pageId="11" pageNumber="244">Deltocyathus crassiseptum</emphasis>
Cairns, 1999
</taxonomicName>
(
<figureCitation id="134F0EF87D50FFECFEFEFEDB03CCFE36" box="[336,421,374,401]" captionStart="FIG" captionStartId="7.[113,124,1499,1516]" captionText="FIG. 1. — A, Deltocyathus magnificus Moseley,1876 (MNHN-Scl.2008-0001);A1, calicular;A2, basal;and A3, lateral views;B, D. rotulus (Alcock,1898) (MNHN-Scl.2008-0004);B1, calicular;B2, basal;and B3, lateral views;C, D.suluensis Alcock,1902 (MNHN-Scl.2008-0006); C1, calicular; C2, basal; and C3, lateral views; D, D. vaughani Yabe &amp; Eguchi, 1932 (MNHN-Scl.2008-0009); D1, calicular; D2, basal; and D3, lateral views; E, D. ornatus Gardiner, 1899 (USNM 1114145); E1, calicular; E2, basal; and E3, lateral views; F, D. heteroclitus Wells, 1984 (USNM 1114160); F1, calicular; F2, basal; and F3, lateral views; G, D. corrugatus Cairns, 1999 (MNHN-Scl.2008-0023); G1, calicular; G2, basal; and G3, lateral views; H, D. crassiseptum Cairns, 1999 (MNHN-Scl.2008-0026); H1, calicular; H2, basal; and H3, lateral views; I, D. cameratus Cairns, 1999 (USNM 1114123); I1, calicular; I2, basal; and I3, lateral views; J, D. inusitatus n. sp.; J1-J3, paratype USNM 1114137;J1, calicular;J2, basal;and J3, lateral views;J4-J7, holotype MNHN-Scl.2008-0034;J4, calicular stereo pair; J5, basal; J6, oblique; and J7, lateral views. All lateral views are not presented in life position. Scale bars: 1 cm." pageId="11" pageNumber="244">Fig. 1H</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36E41F67D50FFECFFDFFE02001EFE60" box="[113,631,431,455]" pageId="11" pageNumber="244" type="reference_group">
<paragraph id="8BCB127D7D50FFECFFDFFE02001EFE60" blockId="11.[113,631,431,455]" box="[113,631,431,455]" pageId="11" pageNumber="244">
<taxonomicName id="4C7469FE7D50FFECFFDFFE020067FE60" ID-CoL="34KF6" authority="Cairns, 1999: 94" authorityName="Cairns" authorityPageNumber="94" authorityYear="1999" box="[113,526,431,455]" class="Anthozoa" family="Deltocyathidae" genus="Deltocyathus" kingdom="Animalia" order="Scleractinia" pageId="11" pageNumber="244" phylum="Cnidaria" rank="species" species="crassiseptum">
<emphasis id="B900CE6F7D50FFECFFDFFE020336FE60" box="[113,351,431,455]" italics="true" pageId="11" pageNumber="244">Deltocyathus crassiseptum</emphasis>
Cairns, 1999: 94
</taxonomicName>
, fig. 12c-f.
</paragraph>
</subSubSection>
<subSubSection id="C36E41F67D50FFECFFDFFE72000BFC09" pageId="11" pageNumber="244" type="materials_examined">
<paragraph id="8BCB127D7D50FFECFFDFFE72033BFDE1" blockId="11.[113,646,479,582]" pageId="11" pageNumber="244">
TYPE MATERIAL. — The
<typeStatus id="54CFACDF7D50FFECFEF1FE7203DCFE50" box="[351,437,479,503]" pageId="11" pageNumber="244" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D72D9F47D50FFECFE49FE72000AFE50" box="[487,611,479,503]" pageId="11" pageNumber="244" type="generic" typeStatus="paratypes">78 paratypes</specimenCount>
are deposited at the
<collectionCode id="ED658AB87D50FFECFEBEFE54030AFDB6" box="[272,355,505,529]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="244">MNHN</collectionCode>
(uncatalogued), and 39 at the
<collectionCode id="ED658AB87D50FFECFFDFFDB902D2FD8B" box="[113,187,532,556]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="244" type="Museum">USNM</collectionCode>
(98681 [2], 98682 [9], 98683 [13], 98684 [8], 98685 [2], 98686 [5]).
</paragraph>
<paragraph id="8BCB127D7D50FFECFFDFFDF302D0FD37" blockId="11.[113,646,606,656]" pageId="11" pageNumber="244">
<typeStatus id="54CFACDF7D50FFECFFDFFDF302CAFDD3" box="[113,163,606,629]" pageId="11" pageNumber="244">TYPE</typeStatus>
LOCALITY. —
<geoCoordinate id="EE4074BA7D50FFECFE96FDF303E1FDD1" box="[312,392,606,630]" degrees="19" direction="south" minutes="21" orientation="latitude" pageId="11" pageNumber="244" precision="925" value="-19.35">19°21S</geoCoordinate>
,
<geoCoordinate id="EE4074BA7D50FFECFE3BFDF3039BFDD1" box="[405,498,606,630]" degrees="169" direction="east" minutes="25" orientation="longitude" pageId="11" pageNumber="244" precision="925" value="169.41667">169°25E</geoCoordinate>
(Tanna),
<quantity id="4C8CBF987D50FFECFDF6FDF302DCFD37" metricMagnitude="2" metricUnit="m" metricValue="4.415" metricValueMax="4.5" metricValueMin="4.33" pageId="11" pageNumber="244" unit="m" value="441.5" valueMax="450.0" valueMin="433.0">
433-
<specimenCount id="9D72D9F47D50FFECFFDFFDD502DCFD37" box="[113,181,632,656]" pageId="11" pageNumber="244" type="generic">450 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph id="8BCB127D7D50FFECFFDFFD05000BFC09" blockId="11.[111,648,680,942]" pageId="11" pageNumber="244">
NEW RECORDS. — BATHUS 3, stn CP 818,
<specimenCount id="9D72D9F47D50FFECFDBAFD0500EFFD67" box="[532,646,680,704]" pageId="11" pageNumber="244" type="generic">4 specimens</specimenCount>
(MNHN-Scl.2008-0024). — BATHUS 4, stn CP 889,
<specimenCount id="9D72D9F47D50FFECFFDEFD700288FD52" box="[112,225,733,757]" pageId="11" pageNumber="244" type="generic">1 specimen</specimenCount>
(MNHN-Scl.2008-0026). — HALIPRO 1, stn CP 877,
<specimenCount id="9D72D9F47D50FFECFE84FD5A03CEFCA8" box="[298,423,759,783]" pageId="11" pageNumber="244" type="generic">1 specimen</specimenCount>
(MNHN-Scl.2008- 0025). — NORFOLK 2, stn DW 2041,
<specimenCount id="9D72D9F47D50FFECFDA1FCBF00EFFC8D" box="[527,646,786,810]" pageId="11" pageNumber="244" type="generic">2 specimens</specimenCount>
(MNHN-Scl.2008-0027); stn DW 2024,
<specimenCount id="9D72D9F47D50FFECFDBEFC8100EFFCE3" box="[528,646,812,836]" pageId="11" pageNumber="244" type="generic">6 specimens</specimenCount>
(MNHN-Scl.2008-0028);
<materialsCitation id="3B1C18207D50FFECFEC8FCEA00EDFCF8" box="[358,644,839,863]" collectionCode="USNM" pageId="11" pageNumber="244" specimenCode="USNM 1114122" specimenCount="6">
<specimenCount id="9D72D9F47D50FFECFEC8FCEA03B8FCF8" box="[358,465,839,863]" pageId="11" pageNumber="244" type="generic">6 specimens</specimenCount>
(
<specimenCode id="DBD2BA067D50FFECFE75FCEA0014FCF8" box="[475,637,839,863]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="244" type="Museum">USNM 1114122</specimenCode>
)
</materialsCitation>
; stn DW 2025,
<specimenCount id="9D72D9F47D50FFECFF54FCCC030AFCDE" box="[250,355,865,889]" pageId="11" pageNumber="244" type="generic">1 specimen</specimenCount>
(MNHN-Scl.2008-0029); stn DW 2097,
<specimenCount id="9D72D9F47D50FFECFF4FFCD10331FC33" box="[225,344,892,916]" pageId="11" pageNumber="244" type="generic">2 specimens</specimenCount>
(MNHN-Scl.2008-0030); stn DW 2117,
<specimenCount id="9D72D9F47D50FFECFF4DFC3B0330FC09" box="[227,345,918,942]" pageId="11" pageNumber="244" type="generic">2 specimens</specimenCount>
(MNHN-Scl.2008-0031).
</paragraph>
</subSubSection>
<subSubSection id="C36E41F67D50FFECFFDFFC6B000FFB8A" pageId="11" pageNumber="244" type="distribution">
<paragraph id="8BCB127D7D50FFECFFDFFC6B000FFB8A" blockId="11.[111,647,966,1069]" pageId="11" pageNumber="244">
<materialsCitation id="3B1C18207D50FFECFFDFFC6B02DDFB8A" country="New Caledonia" elevation="477" latitude="-24.733334" location="New Caledonia" longLatPrecision="1249" longitude="168.1" pageId="11" pageNumber="244" specimenCount="1">
DISTRIBUTION. —
<collectingCountry id="F36352ED7D50FFECFEB2FC6B03C0FC79" box="[284,425,966,990]" name="New Caledonia" pageId="11" pageNumber="244">New Caledonia</collectingCountry>
waters from
<geoCoordinate id="EE4074BA7D50FFECFDB5FC6B00EAFC79" box="[539,643,966,990]" degrees="21" direction="south" minutes="00.83" orientation="latitude" pageId="11" pageNumber="244" precision="9" value="-21.013834">21°00.83S</geoCoordinate>
,
<geoCoordinate id="EE4074BA7D50FFECFFC1FC4D0281FC5F" box="[111,232,992,1016]" degrees="164" direction="east" minutes="27.34" orientation="longitude" pageId="11" pageNumber="244" precision="9" value="164.45567">164°27.34E</geoCoordinate>
to
<geoCoordinate id="EE4074BA7D50FFECFEABFC4D033BFC5F" box="[261,338,992,1016]" degrees="24" direction="south" minutes="44" orientation="latitude" pageId="11" pageNumber="244" precision="925" value="-24.733334">24°44S</geoCoordinate>
,
<geoCoordinate id="EE4074BA7D50FFECFEF4FC4D03DFFC5F" box="[346,438,992,1016]" degrees="168" direction="east" minutes="06" orientation="longitude" pageId="11" pageNumber="244" precision="925" value="168.1">168°06E</geoCoordinate>
,
<quantity id="4C8CBF987D50FFECFE11FC4D0045FC5F" box="[447,556,992,1016]" metricMagnitude="2" metricUnit="m" metricValue="4.765" metricValueMax="5.83" metricValueMin="3.7" pageId="11" pageNumber="244" unit="m" value="476.5" valueMax="583.0" valueMin="370.0">
<elevation id="0059F54E7D50FFECFE11FC4D0045FC5F" box="[447,556,992,1016]" metricMagnitude="2" metricUnit="m" metricValue="4.765" metricValueMax="5.83" metricValueMin="3.7" pageId="11" pageNumber="244" unit="m" value="476.5" valueMax="583.0" valueMin="370.0">370-583 m</elevation>
</quantity>
. Previous records:
<collectingCountry id="F36352ED7D50FFECFF6CFC56031CFBB4" box="[194,373,1019,1043]" name="Wallis and Futuna" pageId="11" pageNumber="244">Wallis and Futuna</collectingCountry>
region (Cairns 1999),
<quantity id="4C8CBF987D50FFECFDF8FC5602DDFB8A" metricMagnitude="2" metricUnit="m" metricValue="4.65" metricValueMax="5.1" metricValueMin="4.2" pageId="11" pageNumber="244" unit="m" value="465.0" valueMax="510.0" valueMin="420.0">
<elevation id="0059F54E7D50FFECFDF8FC5602DDFB8A" metricMagnitude="2" metricUnit="m" metricValue="4.65" metricValueMax="5.1" metricValueMin="4.2" pageId="11" pageNumber="244" unit="m" value="465.0" valueMax="510.0" valueMin="420.0">420- 510 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation id="3B1C18207D50FFECFF10FBB8000FFB8A" box="[190,614,1045,1069]" country="Vanuatu" elevation="475" location="Vanuatu" pageId="11" pageNumber="244" specimenCount="1">
<collectingCountry id="F36352ED7D50FFECFF10FBB80379FB8A" box="[190,272,1045,1069]" name="Vanuatu" pageId="11" pageNumber="244">Vanuatu</collectingCountry>
region (Cairns 1999),
<quantity id="4C8CBF987D50FFECFE5FFBB8000BFB8A" box="[497,610,1045,1069]" metricMagnitude="2" metricUnit="m" metricValue="4.745" metricValueMax="5.36" metricValueMin="4.13" pageId="11" pageNumber="244" unit="m" value="474.5" valueMax="536.0" valueMin="413.0">
<elevation id="0059F54E7D50FFECFE5FFBB8000BFB8A" box="[497,610,1045,1069]" metricMagnitude="2" metricUnit="m" metricValue="4.745" metricValueMax="5.36" metricValueMin="4.13" pageId="11" pageNumber="244" unit="m" value="474.5" valueMax="536.0" valueMin="413.0">413-536 m</elevation>
</quantity>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36E41F67D50FFECFFDFFBE20115FD97" pageId="11" pageNumber="244" type="description">
<paragraph id="8BCB127D7D50FFECFFDFFBE2036BFBC0" blockId="11.[111,649,1103,1700]" box="[113,258,1103,1129]" pageId="11" pageNumber="244">DESCRIPTION</paragraph>
<paragraph id="8BCB127D7D50FFECFFDFFBC20004FA41" blockId="11.[111,649,1103,1700]" pageId="11" pageNumber="244">Corallum circular to slightly elliptical shaped as a shallow bowl.Theca thick.Base flat to slightly convex, usually with a scar, and sometimes protuberant. Largest specimen examined (BATHUS 3, stn CP 877) 14.2 mm in calicular diameter and 4.6 mm in height. However, smaller coralla show a CD:HT ratio near 2. Costae unequal and more prominent at calicular edge where they are separated by intercostal grooves bisected by a row of low and rounded granules that sometimes resemble a small ridge in the valley formed by the adjacent costae. C1-2 broader than other costae. Corallum white to reddish-brown.</paragraph>
<paragraph id="8BCB127D7D50FFECFF26FA460621FE57" blockId="11.[111,649,1103,1700]" lastBlockId="11.[675,1212,216,560]" pageId="11" pageNumber="244">
Septa hexamerally arranged in four cycles (48 septa) with septal formula: S1&gt;S2&gt;S3&gt;S4. However, in larger coralla up to 3 pairs of S5 are present resulting in 56 septa. S1 thick and most exsert septa, extending half distance to columella and bear a dimorphic lamellar paliform lobe. Those paliform lobes before the two principal S1 are narrower than the other four P1, but all reach the columella. S2 equal to slightly smaller and less exsert than S1, also bearing a paliform lobe that reaches columella by a small paliform teeth. S3 about ¾ width of S2, bearing a tall recessed pali that fuses P
<quantity id="4C8CBF987D50FFECFB84FEDB063BFE36" box="[1066,1106,374,401]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="11" pageNumber="244" unit="in" value="2.0">2 in</quantity>
a chevron arrangement typical of the genus. S4 smallest septa and in some coralla composed by a row of spines. All septal and palar faces very granular.
</paragraph>
<paragraph id="8BCB127D7D50FFECFD0BFE580115FD97" blockId="11.[675,1212,216,560]" pageId="11" pageNumber="244">Fossa shallow to moderately deep, containing a papillose columella.</paragraph>
</subSubSection>
<subSubSection id="C36E41F67D50FFECFD0BFDF801B1FB8C" pageId="11" pageNumber="244" type="discussion">
<paragraph id="8BCB127D7D50FFECFD0BFDF80160FDCA" blockId="11.[675,1211,597,1067]" box="[677,777,597,623]" pageId="11" pageNumber="244">REMARKS</paragraph>
<paragraph id="8BCB127D7D50FFECFD0BFDD901B1FB8C" blockId="11.[675,1211,597,1067]" pageId="11" pageNumber="244">
First time reported since the original description, the specimens of
<taxonomicName id="4C7469FE7D50FFECFCF0FD390194FD0A" authorityName="Cairns" authorityYear="1999" box="[862,1021,660,685]" class="Anthozoa" family="Deltocyathidae" genus="Deltocyathus" kingdom="Animalia" order="Scleractinia" pageId="11" pageNumber="244" phylum="Cnidaria" rank="species" species="crassiseptum">
<emphasis id="B900CE6F7D50FFECFCF0FD390194FD0A" box="[862,1021,660,685]" italics="true" pageId="11" pageNumber="244">D. crassiseptum</emphasis>
</taxonomicName>
are distinguished from the
<collectingCountry id="F36352ED7D50FFECFCA3FD1E01DFFD69" box="[781,950,691,718]" name="New Caledonia" pageId="11" pageNumber="244">New Caledonia</collectingCountry>
congeners by having: a slighty thicker S
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relation to the other septa, corallum shaped as a bowl, absence of costal spines, well-arranged crowns formed by the paliform lobes, and paliform lobes and columellar elements that terminate below the septa upper edges. Different pigmentation patterns can be found in
<collectingCountry id="F36352ED7D50FFECFB28FCDC017AFC0B" name="New Caledonia" pageId="11" pageNumber="244">New Caledonia</collectingCountry>
specimens of
<taxonomicName id="4C7469FE7D50FFECFC0AFC3F0629FC0C" authorityName="Cairns" authorityYear="1999" box="[932,1088,914,939]" class="Anthozoa" family="Deltocyathidae" genus="Deltocyathus" kingdom="Animalia" order="Scleractinia" pageId="11" pageNumber="244" phylum="Cnidaria" rank="species" species="crassiseptum">
<emphasis id="B900CE6F7D50FFECFC0AFC3F0629FC0C" box="[932,1088,914,939]" italics="true" pageId="11" pageNumber="244">D. crassiseptum</emphasis>
</taxonomicName>
: some have the entire calicular edge and septa reddish-brown, other specimens present only the S3 and paliform lobes pigmented, some are completely pigmented and some are enterely white.
</paragraph>
</subSubSection>
</treatment>
</document>