treatments-xml/data/03/CE/9F/03CE9F1CFF9F0C41FF65F8CBEECEFCBE.xml

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<mods:title id="A86E3612FADE55DEB718CA5C5A3F9976">The Colubrariidae, Eosiphonidae, Melongenidae, Pisaniidae, Prodotiidae and Tudiclidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea</mods:title>
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<mods:namePart id="7BAEEE261B5F698C52A6B821453175E9">Harzhauser, Mathias</mods:namePart>
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<mods:affiliation id="A9F76016607C4BB3EE30210E14F75118">Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655</mods:affiliation>
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<mods:namePart id="01FA5D1C0076D2A5C46E212B1AE7CE63">Landau, Bernard M.</mods:namePart>
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<mods:affiliation id="D127C266E5C23C8218EC9C7A18B44CD0">Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655 &amp; Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal bernardmlandau @ gmail. com; https: // orcid. org / 0000 - 0002 - 7768 - 8494 &amp; Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria mathias. harzhauser @ nhm-wien. ac. at; https: // orcid. org / 0000 - 0002 - 4471 - 6655</mods:affiliation>
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<paragraph id="8BD82E0AFF9F0C42FF65F8CBEE58F8C7" blockId="5.[151,557,1830,1857]" box="[151,557,1830,1857]" pageId="5" pageNumber="6">
<heading id="D0909966FF9F0C42FF65F8CBEE58F8C7" bold="true" box="[151,557,1830,1857]" fontSize="11" level="2" pageId="5" pageNumber="6" reason="3">
<emphasis id="B913F218FF9F0C42FF65F8CBEE58F8C7" bold="true" box="[151,557,1830,1857]" pageId="5" pageNumber="6">
Family 
<taxonomicName id="4C675589FF9F0C42FF05F8CBEE58F8C7" ID-CoL="7NJ3W" authority="Dall, 1904" authorityName="Dall" authorityYear="1904" box="[247,557,1830,1857]" class="Gastropoda" family="Colubrariidae" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="6" phylum="Mollusca" rank="family">
Colubrariidae 
<bibRefCitation id="EFF653FBFF9F0C42FE5DF8CAEE58F8C7" author="Dall, W. H." box="[431,557,1830,1857]" pageId="5" pageNumber="6" pagination="114 - 144" refId="ref80084" refString="Dall, W. H. (1904) An historical and systematic review of the frog-shells and tritons. Smithsonian Miscellaneous Collections, 47, 114 - 144. [https: // www. biodiversitylibrary. org / item / 35633 # page / 166 / mode / 1 up]" type="journal article" year="1904">Dall, 1904</bibRefCitation>
</taxonomicName>
</emphasis>
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</paragraph>
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<subSubSection id="C37D7D81FF9F0C41FF65F883EEB1FE9A" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8BD82E0AFF9F0C41FF65F883EEB1FE9A" blockId="5.[151,1436,1902,2037]" lastBlockId="6.[151,1437,151,393]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="B913F218FF9F0C42FF65F883ED1FF80F" bold="true" box="[151,362,1902,1928]" pageId="5" pageNumber="6">Revised diagnosis</emphasis>
. “ 
<emphasis id="B913F218FF9F0C42FE77F883EDA2F856" italics="true" pageId="5" pageNumber="6">
Shell medium-large to large, reaching over 
<quantity id="4C9F83EFFF9F0C42FC80F882EFB9F80F" box="[882,972,1903,1929]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="6" unit="mm" value="110.0">110 mm</quantity>
, from narrow fusiform to broad fusiform, with 
<taxonomicName id="4C675589FF9F0C42FF22F87FED5CF82B" box="[208,297,1938,1964]" class="Gastropoda" family="Colubrariidae" genus="Aperture" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="medium">medium</taxonomicName>
to high spire and short but distinct siphonal canal. Protoconch usually paucispiral of 1–2.5 smooth whorls, up to four whorls (
</emphasis>
<taxonomicName id="4C675589FF9F0C42FE25F85BEE53F857" box="[471,550,1974,2000]" class="Gastropoda" family="Colubrariidae" genus="Metula" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="6" phylum="Mollusca" rank="genus">Metula</taxonomicName>
<emphasis id="B913F218FF9F0C41FDD4F85AEDFEFE9A" italics="true" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
). Spire whorls evenly convex, without pronounced shoulder. Axial sculpture present or absent; when present, of distinct narrow and closely spaced axial ribs, crossing the spiral cords and forming reticulated sculpture pattern, often with regularly or irregularly spaced varices. Spiral sculpture of weak threads or fine, regularly set, beaded cords or strong spiral cords forming knobs at intersection with axials. 
<taxonomicName id="4C675589FF9C0C41FAC8FF56EC85FF7E" class="Gastropoda" family="Colubrariidae" genus="Aperture" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="medium">Aperture medium</taxonomicName>
to high, lanceolate due to sharp posterior corner. Outer lip usually lirate inside […]. Columella smooth, often heavily callused
</emphasis>
.” (
<bibRefCitation id="EFF653FBFF9C0C41FE5FFEEEEECDFE9A" author="Kantor, Yu. I. &amp; Fedosov, A. E. &amp; Koysan, A. R. &amp; Puillandre, N. &amp; Sorokin, P. A. &amp; Kano, Y. &amp; Clark, R. &amp; Bouchet, P." box="[429,696,259,285]" pageId="6" pageNumber="7" pagination="789 - 857" refId="ref85869" refString="Kantor, Yu. I., Fedosov, A. E., Koysan, A. R., Puillandre, N., Sorokin, P. A., Kano, Y., Clark, R. &amp; Bouchet, P. (2022) Molecular phylogeny and revised classification of the Buccinoidea (Neogastropoda). Zoological Journal of the Linnean Society, 194, 789 - 857. [published in a journal issue in 2022, but made available online in 2021; because the paper is registred in Zoobank, the year of taxa description is fixed as 2021] https: // doi. org / 10.1093 / zoolinnean / zlab 031" type="journal article" year="2022">
Kantor 
<emphasis id="B913F218FF9C0C41FDF6FEEEEE49FE9A" box="[516,572,259,285]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
2022: 806
</bibRefCitation>
).
</paragraph>
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<emphasis id="B913F218FF9C0C41FF35FECBED39FEC7" bold="true" box="[199,332,294,320]" pageId="6" pageNumber="7">Discussion.</emphasis>
The relationships of this family were discussed by 
<bibRefCitation id="EFF653FBFF9C0C41FC43FECAE8E8FEC6" author="Kantor, Yu. I. &amp; Fedosov, A. E. &amp; Koysan, A. R. &amp; Puillandre, N. &amp; Sorokin, P. A. &amp; Kano, Y. &amp; Clark, R. &amp; Bouchet, P." box="[945,1181,295,321]" pageId="6" pageNumber="7" pagination="789 - 857" refId="ref85869" refString="Kantor, Yu. I., Fedosov, A. E., Koysan, A. R., Puillandre, N., Sorokin, P. A., Kano, Y., Clark, R. &amp; Bouchet, P. (2022) Molecular phylogeny and revised classification of the Buccinoidea (Neogastropoda). Zoological Journal of the Linnean Society, 194, 789 - 857. [published in a journal issue in 2022, but made available online in 2021; because the paper is registred in Zoobank, the year of taxa description is fixed as 2021] https: // doi. org / 10.1093 / zoolinnean / zlab 031" type="journal article" year="2022">
Kantor 
<emphasis id="B913F218FF9C0C41FBF8FECAE832FEC6" box="[1034,1095,295,321]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
(2022)
</bibRefCitation>
in their buccinoidean molecular phylogeny. The genera 
<taxonomicName id="4C675589FF9C0C41FDE1FEA6EF12FEE2" authority="Schumacher, 1817" authorityName="Schumacher" authorityYear="1817" box="[531,871,331,357]" class="Gastropoda" family="Colubrariidae" genus="Colubraria" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus">
<emphasis id="B913F218FF9C0C41FDE1FEA6EEE4FEE2" box="[531,657,331,357]" italics="true" pageId="6" pageNumber="7">Colubraria</emphasis>
<bibRefCitation id="EFF653FBFF9C0C41FD6AFEA6EF12FEE2" author="Schumacher, H. C. F." box="[664,871,331,357]" pageId="6" pageNumber="7" refId="ref91904" refString="Schumacher, H. C. F. (1817) Essais d'un nouveau systeme des habitations des vers testaces. Schultz, Copenhagen, 287 pp. [https: // www. biodiversitylibrary. org / item / 81329 # page / 7 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 35863" type="book" year="1817">Schumacher, 1817</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C675589FF9C0C41FC6FFEA6E816FEE2" authority="H. Adams" authorityName="H. Adams" box="[925,1123,331,357]" class="Gastropoda" family="Colubrariidae" genus="Metula" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus">
<emphasis id="B913F218FF9C0C41FC6FFEA6EF99FEE2" box="[925,1004,331,357]" italics="true" pageId="6" pageNumber="7">Metula</emphasis>
H. Adams
</taxonomicName>
&amp; A. 
<bibRefCitation id="EFF653FBFF9C0C41FB54FEA6E94EFEE2" author="Adams, H. &amp; Adams, A." box="[1190,1339,330,357]" pageId="6" pageNumber="7" refId="ref76109" refString="Adams, H. &amp; Adams, A. (1853) The genera of Recent Mollusca; arranged according to their organization. Vol. 1. van Voorst, London, 256 pp. [https: // www. biodiversitylibrary. org / page / 3782650]" type="book" year="1853">Adams, 1853</bibRefCitation>
occur in Paratethyan assemblages.
</paragraph>
<paragraph id="8BD82E0AFF9C0C41FF65FE37EE2AFE73" blockId="6.[151,607,474,501]" box="[151,607,474,501]" pageId="6" pageNumber="7">
<emphasis id="B913F218FF9C0C41FF65FE37EE2AFE73" bold="true" box="[151,607,474,501]" pageId="6" pageNumber="7">
Genus 
<taxonomicName id="4C675589FF9C0C41FF1CFE37EE2AFE73" authority="Schumacher, 1817" authorityName="Schumacher" authorityYear="1817" box="[238,607,474,501]" class="Gastropoda" family="Colubrariidae" genus="Colubraria" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus">
<emphasis id="B913F218FF9C0C41FF1CFE37ED01FE72" bold="true" box="[238,372,474,501]" italics="true" pageId="6" pageNumber="7">Colubraria</emphasis>
<bibRefCitation id="EFF653FBFF9C0C41FE89FE37EE2AFE73" author="Schumacher, H. C. F." box="[379,607,474,500]" pageId="6" pageNumber="7" refId="ref91904" refString="Schumacher, H. C. F. (1817) Essais d'un nouveau systeme des habitations des vers testaces. Schultz, Copenhagen, 287 pp. [https: // www. biodiversitylibrary. org / item / 81329 # page / 7 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 35863" type="book" year="1817">Schumacher, 1817</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37D7D81FF9C0C41FF65FDCEEF03FD4A" pageId="6" pageNumber="7" type="type_taxon">
<paragraph id="8BD82E0AFF9C0C41FF65FDCEED8DFDE6" blockId="6.[151,1437,546,825]" pageId="6" pageNumber="7">
<emphasis id="B913F218FF9C0C41FF65FDCEED46FDBB" bold="true" box="[151,307,546,573]" pageId="6" pageNumber="7">
<typeStatus id="54DC90A8FF9C0C41FF65FDCEECA7FDBA" box="[151,210,547,573]" pageId="6" pageNumber="7">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C675589FF9C0C41FEC8FDCEEF70FDBA" ID-CoL="5ZQ7P" authority="Schumacher, 1817" authorityName="Schumacher" authorityYear="1817" box="[314,773,547,573]" class="Gastropoda" family="Colubrariidae" genus="Colubraria" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="granulata">
<emphasis id="B913F218FF9C0C41FEC8FDCEEE58FDBA" box="[314,557,547,573]" italics="true" pageId="6" pageNumber="7">Colubraria granulata</emphasis>
<bibRefCitation id="EFF653FBFF9C0C41FDC7FDCEEF70FDBA" author="Schumacher, H. C. F." box="[565,773,547,573]" pageId="6" pageNumber="7" refId="ref91904" refString="Schumacher, H. C. F. (1817) Essais d'un nouveau systeme des habitations des vers testaces. Schultz, Copenhagen, 287 pp. [https: // www. biodiversitylibrary. org / item / 81329 # page / 7 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 35863" type="book" year="1817">Schumacher, 1817</bibRefCitation>
</taxonomicName>
[= 
<taxonomicName id="4C675589FF9C0C41FCDFFDCEE862FDBB" ID-CoL="5ZPW2" baseAuthorityName="Lightfoot" baseAuthorityYear="1786" box="[813,1047,547,573]" class="Gastropoda" family="Colubrariidae" genus="Colubraria" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="muricata">
<emphasis id="B913F218FF9C0C41FCDFFDCEE862FDBB" box="[813,1047,547,573]" italics="true" pageId="6" pageNumber="7">Colubraria muricata</emphasis>
</taxonomicName>
([Lightfoot, 1786]), by monotypy. Present-day, Indo-West Pacific.
</paragraph>
<paragraph id="8BD82E0AFF9C0C41FF35FD87EF03FD4A" blockId="6.[151,1437,546,825]" pageId="6" pageNumber="7">
<emphasis id="B913F218FF9C0C41FF35FD87EDDCFD03" bold="true" box="[199,425,618,644]" pageId="6" pageNumber="7">Original diagnosis.</emphasis>
“ 
<emphasis id="B913F218FF9C0C41FE49FD86ED27FD4A" italics="true" pageId="6" pageNumber="7">Aperture oblong oval; outer lip with margin, internally plicate-dentate; inner lip callous, anteriorly thickened and detached. Columella varicose and with transverse folds. Rostrum short, open at both ends and canaliculate</emphasis>
.” (
<bibRefCitation id="EFF653FBFF9C0C41FE86FD5EEE1BFD4A" author="Schumacher, H. C. F." box="[372,622,691,717]" pageId="6" pageNumber="7" refId="ref91904" refString="Schumacher, H. C. F. (1817) Essais d'un nouveau systeme des habitations des vers testaces. Schultz, Copenhagen, 287 pp. [https: // www. biodiversitylibrary. org / item / 81329 # page / 7 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 35863" type="book" year="1817">Schumacher 1817: 76</bibRefCitation>
, translated from Latin).
</paragraph>
</subSubSection>
<subSubSection id="C37D7D81FF9C0C41FF35FD3BEECEFCBE" pageId="6" pageNumber="7" type="discussion">
<paragraph id="8BD82E0AFF9C0C41FF35FD3BEECEFCBE" blockId="6.[151,1437,546,825]" pageId="6" pageNumber="7">
<emphasis id="B913F218FF9C0C41FF35FD3BED39FD77" bold="true" box="[199,332,726,752]" pageId="6" pageNumber="7">Discussion.</emphasis>
<taxonomicName id="4C675589FF9C0C41FEA1FD3AEE5FFD76" box="[339,554,727,753]" pageId="6" pageNumber="7">
<emphasis id="B913F218FF9C0C41FEA1FD3AEDA4FD76" box="[339,465,727,753]" italics="true" pageId="6" pageNumber="7">Colubraria</emphasis>
species
</taxonomicName>
inhabit shallow marine, rocky and coral environments in tropical, sub-tropical and temperate seas where they feed on the blood of fishes (
<bibRefCitation id="EFF653FBFF9C0C41FCE8FD16EF9BFC92" author="Modica, M. V. &amp; Lombardo, F. &amp; Franchini, P. &amp; Oliverio, M." box="[794,1006,762,789]" pageId="6" pageNumber="7" pagination="441" refId="ref88243" refString="Modica, M. V., Lombardo, F., Franchini, P. &amp; Oliverio, M. (2015) The venomous cocktail of the vampire snail Colubraria reticulata (Mollusca, Gastropoda). BMC Genomics, 16, 441. https: // doi. org / 10.1186 / s 12864 - 015 - 1648 - 4" type="journal article" year="2015">
Modica 
<emphasis id="B913F218FF9C0C41FC85FD16EFDBFC92" box="[887,942,763,789]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
2015
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; 
<bibRefCitation id="EFF653FBFF9C0C41FC0AFD16E97BFC92" author="Oliverio, M. &amp; Modica, M. V." box="[1016,1294,763,789]" pageId="6" pageNumber="7" pagination="779 - 800" refId="ref89032" refString="Oliverio, M. &amp; Modica, M. V. (2010) Relationships of the haematophagous marine snail Colubraria (Rachiglossa: Colubrariidae), within the neogastropod phylogenetic framework. Zoological Journal of the Linnean Society, 158, 779 - 800. https: // doi. org / 10.1111 / j. 1096 - 3642.2009.00568. x" type="journal article" year="2010">Oliverio &amp; Modica 2010</bibRefCitation>
). 
<taxonomicName id="4C675589FF9C0C41FAECFD16E9E9FC92" authorityName="Schumacher" authorityYear="1817" box="[1310,1436,763,789]" class="Gastropoda" family="Colubrariidae" genus="Colubraria" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="7" phylum="Mollusca" rank="genus">
<emphasis id="B913F218FF9C0C41FAECFD16E9E9FC92" box="[1310,1436,763,789]" italics="true" pageId="6" pageNumber="7">Colubraria</emphasis>
</taxonomicName>
is a rare element in the Paratethyan assemblages.
</paragraph>
</subSubSection>
</treatment>
</document>