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<mods:title id="8BA6F9784576D03BF254D97AFD2407F2">Cobitis nalbanti, a new species of spined loach from South Korea, and redescription of Cobitis lutheri (Teleostei: Cobitidae)</mods:title>
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<paragraph id="6A6A369E3D7F9B12B6982F13FE1E074C" blockId="4.[151,511,1336,1397]" box="[151,511,1336,1363]" pageId="4" pageNumber="581">
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<taxonomicName id="ADD54D1D3D7F9B12B6982F13FE1E074C" ID-CoL="WJZD" authority="Rendahl, 1935" authorityName="Rendahl" authorityYear="1935" box="[151,511,1336,1363]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="581" phylum="Chordata" rank="species" species="lutheri">
<emphasis id="58A1EA8C3D7F9B12B6982F13FE1E074C" bold="true" box="[151,511,1336,1363]" pageId="4" pageNumber="581">
<emphasis id="58A1EA8C3D7F9B12B6982F13FEA5074D" bold="true" box="[151,324,1336,1362]" italics="true" pageId="4" pageNumber="581">Cobitis lutheri</emphasis>
<bibRefCitation id="0E444B6F3D7F9B12B7432F12FE1E074C" author="Rendahl" box="[332,511,1337,1363]" pageId="4" pageNumber="581" refString="Rendahl, H. (1935) Ein paar neue Unterarten von Cobitis taenia. Memoranda Societatis pro Fauna et Flora Fennica, 10 (1933 - 1934), 329 - 336." type="journal article" year="1935">Rendahl, 1935</bibRefCitation>
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(
<figureCitation id="F2EE2A1B3D7F9B12B6902F77FEED076B" box="[159,268,1372,1397]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="5.[151,250,890,912]" captionStartId-1="7.[151,250,1550,1572]" captionStartId-2="8.[151,250,1424,1446]" captionTargetBox-0="[226,1360,553,869]" captionTargetBox-1="[163,1428,280,1513]" captionTargetBox-2="[178,1377,833,1373]" captionTargetId-0="figure@5.[226,1360,553,869]" captionTargetId-1="figure@7.[151,1436,265,1528]" captionTargetId-2="figure@8.[151,1436,769,1403]" captionTargetPageId-0="5" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURE 2. The lectotype of Cobitis lutheri — NRM 10813, male, 64.5 mm SL (copyright under Creative Commons License CC BY NC)." captionText-1="FIGURE 3. Male (a) SL 45 mm and female (b) SL 50.5 mm of Cobitis lutheri from the Khanka Lake system, Russia (ZMMU P- 20036) and conspecific male (c) SL 53.5 mm and female (d) SL 74 mm from the Tygda River, Russia (ZMMU P- 21545)." captionText-2="FIGURE 4. Characteristic stripes on the head of Cobitis lutheri (a): 1 — stripe extending across the eye; 2 — suborbital stripe; 3 and 4 — stripes on opercle; lamina circularis (b) in male from the Lyanchihe River, Russia (ZMMU P- 3830)." httpUri-0="https://zenodo.org/record/215096/files/figure.png" httpUri-1="https://zenodo.org/record/215097/files/figure.png" httpUri-2="https://zenodo.org/record/215098/files/figure.png" pageId="4" pageNumber="581">Figs. 2–4</figureCitation>
; 
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)
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<emphasis id="58A1EA8C3D7F9B12B6982F8FFEC907A5" box="[151,296,1444,1466]" italics="true" pageId="4" pageNumber="581">Cobitis taenia</emphasis>
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(not of Linnaeus, 1758): 
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: 47
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; 
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: 153
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(partim); 
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: 438
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(partim); 
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: 169
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: 328
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(partim); 
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: 243
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(partim); 
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: 577
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(partim); 
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: 82
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(partim); 
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: 398
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(partim); 
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: 81
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: 63
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(partim); 
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: 891
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(partim, Fig. 642); 
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: 382
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(partim); 
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: 89
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(partim); 
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Novikov 
<emphasis id="58A1EA8C3D7F9B12B4482C0BFD9B042B" box="[583,634,1566,1589]" italics="true" pageId="4" pageNumber="581">et al.</emphasis>
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(partim); 
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<bibRefCitation id="0E444B6F3D7F9B12B55C2C34FBF5042B" author="Zavertanova" box="[851,1044,1566,1588]" pageId="4" pageNumber="581" refString="Zavertanova, Yu. V. (2007) Species composition of ichthyoplankton in the Artemovka River. In: Ichthyological investigations in inner reservoirs. Materialy mezhdunarodnoi nauchnoi konferentsii, Saransk, 2007. Saransk. pp. 44 - 46. [in Russian]" type="journal article" year="2007">Zavertanova, 2007</bibRefCitation>
: 45
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="6A6A369E3D7F9B12B6982C16FCFF044C" blockId="4.[151,1436,1444,1834]" box="[151,798,1597,1619]" pageId="4" pageNumber="581">
<treatmentCitationGroup id="4AC511B03D7F9B12B6982C16FCFF044C" box="[151,798,1597,1619]" pageId="4" pageNumber="581">
<taxonomicName id="ADD54D1D3D7F9B12B6982C16FDDC044C" ID-CoL="WJZD" authority="Rendahl, 1935: 330" authorityName="Rendahl" authorityPageNumber="330" authorityYear="1935" box="[151,573,1597,1619]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="581" phylum="Chordata" rank="subSpecies" species="taenia" subSpecies="lutheri">
<emphasis id="58A1EA8C3D7F9B12B6982C16FE91044C" box="[151,368,1597,1619]" italics="true" pageId="4" pageNumber="581">Cobitis taenia lutheri</emphasis>
<treatmentCitation id="EB74108F3D7F9B12B7792C16FDDC044C" author="Rendahl" box="[374,573,1597,1619]" page="330" pageId="4" pageNumber="581" year="1935">
<bibRefCitation id="0E444B6F3D7F9B12B7792C16FDEC044C" author="Rendahl" box="[374,525,1597,1619]" pageId="4" pageNumber="581" refString="Rendahl, H. (1935) Ein paar neue Unterarten von Cobitis taenia. Memoranda Societatis pro Fauna et Flora Fennica, 10 (1933 - 1934), 329 - 336." type="journal article" year="1935">Rendahl, 1935</bibRefCitation>
: 330
</treatmentCitation>
</taxonomicName>
(Khanka Lake basin).
</treatmentCitationGroup>
</paragraph>
<paragraph id="6A6A369E3D7F9B12B6982C77FC000535" blockId="4.[151,1436,1444,1834]" pageId="4" pageNumber="581">
<treatmentCitationGroup id="4AC511B03D7F9B12B6982C77FC000535" pageId="4" pageNumber="581">
<taxonomicName id="ADD54D1D3D7F9B12B6982C77FECD046D" ID-CoL="WJZD" box="[151,300,1628,1650]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="581" phylum="Chordata" rank="species" species="lutheri">
<emphasis id="58A1EA8C3D7F9B12B6982C77FECD046D" box="[151,300,1628,1650]" italics="true" pageId="4" pageNumber="581">Cobitis lutheri</emphasis>
</taxonomicName>
: Vasil’eva, 1988: 1029; Vasil’eva, 1994: 839; 
<treatmentCitation id="EB74108F3D7F9B12B51E2C77FB85046D" author="Bogutskaya" box="[785,1124,1628,1650]" page="48" pageId="4" pageNumber="581" year="1996">
<bibRefCitation id="0E444B6F3D7F9B12B51E2C77FBDD046D" author="Bogutskaya" box="[785,1084,1628,1650]" pageId="4" pageNumber="581" refString="Bogutskaya, N. G. &amp; Naseka, A. M. (1996) Cyclostomes and Khanka Lake fish (Amur River system): Annotated checklist of species with comments on their taxonomy and zoogeography of the region. St. Petersburg. Izdatel'stvo Gosudarstvennogo nauchno-issledovatel'skogo instituta ozernogo i rechnogo rybnogo khozyaistva, 89 pp. [in Rissian]" type="book" year="1996">Bogutskaya &amp; Naseka, 1996</bibRefCitation>
: 48
</treatmentCitation>
; Vasil’eva, 1998: 97 (partim); Vasil’eva, 1999: 155; Vasil’ev 
<emphasis id="58A1EA8C3D7F9B12B4052C50FDA1048F" box="[522,576,1658,1680]" italics="true" pageId="4" pageNumber="581">et al.</emphasis>
, 1999: 355; 
<treatmentCitation id="EB74108F3D7F9B12B4C72C51FC71048F" author="Tagirova" box="[712,912,1658,1680]" page="24" pageId="4" pageNumber="581" year="2000">
<bibRefCitation id="0E444B6F3D7F9B12B4C72C51FC88048F" author="Tagirova" box="[712,873,1658,1680]" pageId="4" pageNumber="581" refString="Tagirova, V. T. (2000) Fertebrate animals of Khabarovsk district. Khabarovsk. Khabarovskiy gos. ped. Universitet, 30 pp. [in Russian]" type="book" year="2000">Tagirova, 2000</bibRefCitation>
: 24
</treatmentCitation>
; 
<treatmentCitation id="EB74108F3D7F9B12B5912C51FB84048F" author="Shedko" box="[926,1125,1658,1680]" page="233" pageId="4" pageNumber="581" year="2001">
<bibRefCitation id="0E444B6F3D7F9B12B5912C51FBD1048F" author="Shedko" box="[926,1072,1658,1680]" pageId="4" pageNumber="581" refString="Shedko, S. V. (2001) List of Cyclostomata and fishes of fresh waters of the coast of Primorye. In: Chteniya pamyati Fladimira Yakovlevicha Levanidova, 1. Vladivostok. Dalnauka. pp. 229 - 249 [in Russian]." type="journal article" year="2001">Shedko, 2001</bibRefCitation>
: 233
</treatmentCitation>
; 
<treatmentCitation id="EB74108F3D7F9B12B27D2C51FA79048F" author="Bogutskaya" box="[1138,1432,1658,1680]" page="44" pageId="4" pageNumber="581" year="2001">
<bibRefCitation id="0E444B6F3D7F9B12B27D2C51FA8E048F" author="Bogutskaya" box="[1138,1391,1658,1680]" pageId="4" pageNumber="581" refString="Bogutskaya, N. G., Naseka, A. M. &amp; Komlev, A. M. (2001) Freshwater fishes of Russia: preliminary results of the fauna revision. Proceedings of Zoological Institute of Russian Academy of Sciences, 289, 39 - 50." type="journal article" year="2001">
Bogutskaya 
<emphasis id="58A1EA8C3D7F9B12B2FB2C50FACB048F" box="[1268,1322,1658,1680]" italics="true" pageId="4" pageNumber="581">et al.</emphasis>
, 2001
</bibRefCitation>
: 44
</treatmentCitation>
; 
<treatmentCitation id="EB74108F3D7F9B12B6C82CB2FE1304B0" author="Zemnukhov" box="[199,498,1689,1711]" page="420" pageId="4" pageNumber="581" year="2001">
<bibRefCitation id="0E444B6F3D7F9B12B6C82CB2FE5F04B0" author="Zemnukhov" box="[199,446,1689,1711]" pageId="4" pageNumber="581" refString="Zemnukhov, V. V., Sobolevskii, E. I., Panchenko, V. V. &amp; Antonenko, D. V. (2001) List of fish species from Pil'tun Bay (northeastern Sakhalin). Foprosy Ikhtiologii, 41 (3), 420 - 421 [in Russian]" type="journal article" year="2001">
Zemnukhov 
<emphasis id="58A1EA8C3D7F9B12B7472CB1FE9D04B0" box="[328,380,1689,1711]" italics="true" pageId="4" pageNumber="581">et al.</emphasis>
, 2001
</bibRefCitation>
: 420
</treatmentCitation>
; 
<treatmentCitation id="EB74108F3D7F9B12B7F22CB2FD0F04B0" author="Reshetnikov" box="[509,750,1689,1711]" page="361" pageId="4" pageNumber="581" year="2002">
<bibRefCitation id="0E444B6F3D7F9B12B7F22CB2FD5F04B0" author="Reshetnikov" box="[509,702,1689,1711]" pageId="4" pageNumber="581" refString="Reshetnikov, Yu. S. (Ed.) (2002) Atlas of freshwater fishes of Russia. In two volumes. V. 2. Moscow. Nauka. 253 pp. [in Russian]" type="book" year="2002">Reshetnikov, 2002</bibRefCitation>
: 361
</treatmentCitation>
(the characters and the species range are presented with mistakes); 
<treatmentCitation id="EB74108F3D7F9B12B6C82C93FDEC04D1" author="Safronov" box="[199,525,1720,1742]" page="46" pageId="4" pageNumber="581" year="2003">
<bibRefCitation id="0E444B6F3D7F9B12B6C82C93FE0604D1" author="Safronov" box="[199,487,1720,1742]" pageId="4" pageNumber="581" refString="Safronov, S. N. &amp; Nikiforov, S. N. (2003) A list of fishlike vertebrates and fishes of the fresh and brackish waters of Sakhalin. Foprosy Ikhtiologii, 43 (1), 42 - 53. [in Russian]" type="journal article" year="2003">Safronov &amp; Nikiforov, 2003</bibRefCitation>
: 46
</treatmentCitation>
; Vasil’eva, 2004: 161; Novomodny 
<emphasis id="58A1EA8C3D7F9B12B5772C92FC4A04D1" box="[888,939,1720,1742]" italics="true" pageId="4" pageNumber="581">et al.</emphasis>
, 2004: 12, 49; 
<treatmentCitation id="EB74108F3D7F9B12B24D2C93FA7904D1" author="Bogutskaya" box="[1090,1432,1720,1742]" page="105" pageId="4" pageNumber="581" year="2004">
<bibRefCitation id="0E444B6F3D7F9B12B24D2C93FA8404D1" author="Bogutskaya" box="[1090,1381,1720,1742]" pageId="4" pageNumber="581" refString="Bogutskaya, N. G. &amp; Naseka, A. M. (2004) Catalogue of Agnathans and fishes of fresh and brackish waters of Russia with comments on nomenclature and taxonomy. Moskva. Tovarishchestvo nauchnykh izdaniy KMK, 389 pp. [in Rusian]" type="book" year="2004">Bogutskaya &amp; Naseka, 2004</bibRefCitation>
: 105
</treatmentCitation>
; Novomodny, 2004: 313; 
<treatmentCitation id="EB74108F3D7F9B12B7C92CFDFD5B04F3" author="Novomodny" box="[454,698,1750,1772]" page="103" pageId="4" pageNumber="581" year="2005">
<bibRefCitation id="0E444B6F3D7F9B12B7C92CFDFD6604F3" author="Novomodny" box="[454,647,1750,1772]" pageId="4" pageNumber="581" refString="Novomodny, G. V. (2005) Preliminary results of modern researches of diversity of fishes in the Amur River basin: species composition at a turn of the XX-XXI centuries. In: Biodiversity of fishes of fresh waters of the Amur River and adjacent territories. Materials of the First international conference, Khabarovsk, October 29 - November 1, 2002. Khabarovsk. Magellan. pp. 98 - 114." type="journal article" year="2005">Novomodny, 2005</bibRefCitation>
: 103
</treatmentCitation>
; 
<treatmentCitation id="EB74108F3D7F9B12B4CA2CFCFC4004F3" author="Tarazanov" box="[709,929,1750,1772]" page="196" pageId="4" pageNumber="581" year="2005">
<bibRefCitation id="0E444B6F3D7F9B12B4CA2CFCFC9104F3" author="Tarazanov" box="[709,880,1750,1772]" pageId="4" pageNumber="581" refString="Tarazanov, V. I. (2005) Ichthyoplankton of the Sungacha River. In: Biodiversity of fishes of freshwaters of the Amur River and adjacent territories. Materials of the first international conference, Khabarovsk, October 29 - November 1, 2002. Khabarovsk. Magellan. pp. 33 - 40." type="journal article" year="2005">Tarazanov, 2005</bibRefCitation>
: 196
</treatmentCitation>
(partim?); 
<treatmentCitation id="EB74108F3D7F9B12B21B2CFDFAC204F3" author="Safronov" box="[1044,1315,1750,1773]" page="172" pageId="4" pageNumber="581" year="2005">
<bibRefCitation id="0E444B6F3D7F9B12B21B2CFDFB0E04F3" author="Safronov" box="[1044,1263,1750,1773]" pageId="4" pageNumber="581" refString="Safronov, S. N., Nikiforov, S. N., Zvezdov, T. V., Safronov, A. S. &amp; Nikitin, V. D. (2005). New and rare species of fishes of internal reservoirs of Sakhalin and problem of preservation of biological diversity. In: Biodiversity of fishes of fresh waters of Amur River and adjacent territories. Materials of the First international conference, Khabarovsk, October 29 - November 1, 2002. Khabarovsk. Publishing house Magellan. pp. 144 - 152." type="journal article" year="2005">
Safronov 
<emphasis id="58A1EA8C3D7F9B12B2772CF3FB4D04F3" box="[1144,1196,1750,1773]" italics="true" pageId="4" pageNumber="581">et al.</emphasis>
, 2005
</bibRefCitation>
: 172
</treatmentCitation>
; Vasil’ev &amp; Vasil’eva, 2008a: 1, 2; Vasil’ev &amp; Vasil’eva, 2008b: 51, 52; 
<treatmentCitation id="EB74108F3D7F9B12B5492CDEFB990514" author="Bogutskaya" box="[838,1144,1781,1803]" page="337" pageId="4" pageNumber="581" year="2008">
<bibRefCitation id="0E444B6F3D7F9B12B5492CDEFBA30514" author="Bogutskaya" box="[838,1090,1781,1803]" pageId="4" pageNumber="581" refString="Bogutskaya, N. G., Naseka, A. M., Shedko, S. V., Vasil'eva, E. D. &amp; Chereshnev, I. A. (2008) The fishes of the Amur River: updated check-list and zoogeography. Ichthyological Exploration of Freshwaters, 19 (4), 301 - 366." type="journal article" year="2008">
Bogutskaya 
<emphasis id="58A1EA8C3D7F9B12B5C82CDDFC1C0514" box="[967,1021,1781,1803]" italics="true" pageId="4" pageNumber="581">et al.</emphasis>
, 2008
</bibRefCitation>
: 337
</treatmentCitation>
; 
<treatmentCitation id="EB74108F3D7F9B12B2892CDEFA790514" author="Kolpakov" box="[1158,1432,1781,1803]" page="52" pageId="4" pageNumber="581" year="2008">
<bibRefCitation id="0E444B6F3D7F9B12B2892CDEFA8E0514" author="Kolpakov" box="[1158,1391,1781,1803]" pageId="4" pageNumber="581" refString="Kolpakov, N. V., Barabanshchikov, E. I. &amp; Chepurnoi, A. Yu. (2008) Species composition, distribution and biological condition of alien fish species in estuary of the Razdolnaya River (Peter the Great Bay, Sea of Japan). Rossiiskiy zhurnal biologicheskikh invasiy, 2, 47 - 57. [in Russian]" type="journal article" year="2008">
Kolpakov 
<emphasis id="58A1EA8C3D7F9B12B2FC2CDDFAC80514" box="[1267,1321,1781,1803]" italics="true" pageId="4" pageNumber="581">et al.</emphasis>
, 2008
</bibRefCitation>
: 52
</treatmentCitation>
; 
<treatmentCitation id="EB74108F3D7F9B12B6C82D3FFDD60535" author="Kolpakov" box="[199,567,1812,1834]" page="499" pageId="4" pageNumber="581" year="2010">
<bibRefCitation id="0E444B6F3D7F9B12B6C82D3FFDE40535" author="Kolpakov" box="[199,517,1812,1834]" pageId="4" pageNumber="581" refString="Kolpakov, N. V. &amp; Milovankin, P. G. (2010) Distribution and seasonal changes of fish abundance in eastuary of Razdolnaya River (Peter the Great Bay, Sea of Japan). Foprosy Ikhtiologii, 50 (4), 495 - 509. [in Russian]" type="journal article" year="2010">Kolpakov &amp; Milovankin, 2010</bibRefCitation>
: 499
</treatmentCitation>
; 
<treatmentCitation id="EB74108F3D7F9B12B44E2D3FFD1A0535" author="Kottelat" box="[577,763,1812,1834]" page="26" pageId="4" pageNumber="581" year="2012">
<bibRefCitation id="0E444B6F3D7F9B12B44E2D3FFD370535" author="Kottelat" box="[577,726,1812,1834]" pageId="4" pageNumber="581" refString="Kottelat, M. (2012) Conspectus Cobitinum: an inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei). Raffles Bulletin of Zoology, Supplement 36, 1 - 199." type="journal article" year="2012">Kottelat, 2012</bibRefCitation>
: 26
</treatmentCitation>
; 
<treatmentCitation id="EB74108F3D7F9B12B5092D3FFC3C0535" author="Parin" box="[774,989,1812,1834]" page="98" pageId="4" pageNumber="581" year="2014">
<bibRefCitation id="0E444B6F3D7F9B12B5092D3FFC560535" author="Parin" box="[774,951,1812,1834]" pageId="4" pageNumber="581" refString="Parin, N. V., Evseenko, S. A. &amp; Vasil'eva, E. D. (2014) Fishes of Russian seas: Annotated catalogue. Moscow. KMK Scientific Press, 733 pp." type="book" year="2014">
Parin 
<emphasis id="58A1EA8C3D7F9B12B54D2D3EFC940535" box="[834,885,1812,1834]" italics="true" pageId="4" pageNumber="581">et al.</emphasis>
, 2014
</bibRefCitation>
: 98
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="22CF65153D7F9B13B6982D7DFCD703EA" lastPageId="5" lastPageNumber="582" pageId="4" pageNumber="581" type="materials_examined">
<paragraph id="6A6A369E3D7F9B13B6982D7DFCD703EA" blockId="4.[151,1436,1878,2011]" lastBlockId="5.[151,1436,151,501]" lastPageId="5" lastPageNumber="582" pageId="4" pageNumber="581">
<materialsCitation id="DABD3CC33D7F9B13B6982D7DFD8902CA" ID-GBIF-Occurrence="1417501499" collectionCode="MZH, NRM" county="Consequently" lastPageId="5" lastPageNumber="582" location="Santakheza River" municipality="Rendahl" pageId="4" pageNumber="581" specimenCount="7" typeStatus="holotype">
<emphasis id="58A1EA8C3D7F9B12B6982D7DFEA50570" bold="true" box="[151,324,1878,1903]" pageId="4" pageNumber="581">
<typeStatus id="B56E883C3D7F9B12B6982D7DFF330570" box="[151,210,1878,1903]" pageId="4" pageNumber="581">Type</typeStatus>
material
</emphasis>
. The original description of 
<taxonomicName id="ADD54D1D3D7F9B12B4AC2D7CFCF40571" box="[675,789,1878,1902]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="581" phylum="Chordata" rank="species" species="lutheri">
<emphasis id="58A1EA8C3D7F9B12B4AC2D7CFCF40571" box="[675,789,1878,1902]" italics="true" pageId="4" pageNumber="581">C. lutheri</emphasis>
</taxonomicName>
was based on five specimens. One specimen initially deposited in the Finnish Museum of Natural History, Zoological Collection (
<collectionCode id="0CC4AE5B3D7F9B12B5FD2D50FBD4058C" box="[1010,1077,1915,1939]" country="Finland" httpUri="http://grbio.org/cool/2vmj-fp0v" name="Finnish Museum of Natural History" pageId="4" pageNumber="581">MZH</collectionCode>
), collected from the 
<location id="6F0A60453D7F9B12B3122D51FF3405A8" LSID="urn:lsid:plazi:treatment:E27C87883D7F9B1FB6982F13FD8502CB:6F0A60453D7F9B12B3122D51FF3405A8" county="Consequently" municipality="Rendahl" name="Santakheza River" pageId="4" pageNumber="581">Santakheza River</location>
(= 
<location id="6F0A60453D7F9B12B6FC2DB5FE8505A8" LSID="urn:lsid:plazi:treatment:E27C87883D7F9B1FB6982F13FD8502CB:6F0A60453D7F9B12B6FC2DB5FE8505A8" box="[243,356,1950,1975]" county="Consequently" municipality="Rendahl" name="Spasovka" pageId="4" pageNumber="581">Spasovka</location>
), 46.0 mm SL, was indicated as “ 
<typeStatus id="B56E883C3D7F9B12B4D82DB4FCF905A8" box="[727,792,1951,1975]" pageId="4" pageNumber="581">typus</typeStatus>
” for males, whereas another specimen, in the 
<collectionCode id="0CC4AE5B3D7F9B12B3172DB4FABA05A9" box="[1304,1371,1951,1974]" country="Sweden" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="4" pageNumber="581">NRM</collectionCode>
, from the 
<location id="6F0A60453D7F9B12B6CF2DE9FEB905C4" LSID="urn:lsid:plazi:treatment:E27C87883D7F9B1FB6982F13FD8502CB:6F0A60453D7F9B12B6CF2DE9FEB905C4" box="[192,344,1986,2011]" county="Consequently" municipality="Rendahl" name="Odarka River" pageId="4" pageNumber="581">Odarka River</location>
(tributary of the 
<location id="6F0A60453D7F9B12B4172DE9FD0005C4" LSID="urn:lsid:plazi:treatment:E27C87883D7F9B1FB6982F13FD8502CB:6F0A60453D7F9B12B4172DE9FD0005C4" box="[536,737,1986,2011]" county="Consequently" municipality="Rendahl" name="Santakheza River" pageId="4" pageNumber="581">Santakheza River</location>
), 90.0 mm SL, was specified as “ 
<typeStatus id="B56E883C3D7F9B12B2592DE8FB7705C4" box="[1110,1174,1987,2011]" pageId="4" pageNumber="581">typus</typeStatus>
” for females (
<bibRefCitation id="0E444B6F3D7F9B13B3392DE9FF3202AF" author="Rendahl" lastPageId="5" lastPageNumber="582" pageId="4" pageNumber="581" refString="Rendahl, H. (1935) Ein paar neue Unterarten von Cobitis taenia. Memoranda Societatis pro Fauna et Flora Fennica, 10 (1933 - 1934), 329 - 336." type="journal article" year="1935">
<collectingMunicipality id="8A0EACE43D7F9B12B3392DE9FA7905C4" box="[1334,1432,1986,2011]" pageId="4" pageNumber="581">Rendahl</collectingMunicipality>
, 1935
</bibRefCitation>
). 
<collectingCounty id="830B4E123D7E9B13B6E22ABCFE6902AF" box="[237,392,151,176]" pageId="5" pageNumber="582">Consequently</collectingCounty>
, no single specimen was designated as name-bearing 
<typeStatus id="B56E883C3D7E9B13B2122AB3FBAC02AF" box="[1053,1101,152,176]" pageId="5" pageNumber="582">type</typeStatus>
(
<typeStatus id="B56E883C3D7E9B13B26C2AB3FB7402AF" box="[1123,1173,152,176]" pageId="5" pageNumber="582">type</typeStatus>
or 
<typeStatus id="B56E883C3D7E9B13B2C62ABCFAD102AF" box="[1225,1328,151,176]" pageId="5" pageNumber="582" type="holotype">holotype</typeStatus>
), and all specimens in the description are 
<typeStatus id="B56E883C3D7E9B13B40D2A96FD8902CA" box="[514,616,189,213]" pageId="5" pageNumber="582" type="syntype">syntypes</typeStatus>
</materialsCitation>
. 
<materialsCitation id="DABD3CC33D7E9B13B47D2A97FCD303EA" ID-GBIF-Occurrence="1417501500" collectionCode="MZH, NRM" county="By" location="Rendahl" municipality="We" pageId="5" pageNumber="582" specimenCode="NRM 10810, 64, NRM 10811, 90, NRM 10812, 63.6" specimenCount="4" typeStatus="syntype">
The 
<collectionCode id="0CC4AE5B3D7E9B13B4AB2A96FD0302CB" box="[676,738,189,212]" country="Finland" httpUri="http://grbio.org/cool/2vmj-fp0v" name="Finnish Museum of Natural History" pageId="5" pageNumber="582">MZH</collectionCode>
specimen mentioned by 
<location id="6F0A60453D7E9B13B5F72A97FBB402CA" LSID="urn:lsid:plazi:treatment:E27C87883D7F9B1FB6982F13FD8502CB:6F0A60453D7E9B13B5F72A97FBB402CA" box="[1016,1109,188,213]" county="By" municipality="We" name="Rendahl" pageId="5" pageNumber="582">Rendahl</location>
has been lost (
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, pers. comm.). 
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specimens in the 
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collection are bleached (see 
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search). 
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, the 
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of 
<quantity id="AD2D9B7B3D7E9B13B71B2B2FFE990303" box="[276,376,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="6.45" pageId="5" pageNumber="582" unit="mm" value="64.5">64.5 mm</quantity>
SL, male, 
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10813 (
<figureCitation id="F2EE2A1B3D7E9B13B4802B2FFD320303" box="[655,723,260,285]" captionStart="FIGURE 2" captionStartId="5.[151,250,890,912]" captionTargetBox="[226,1360,553,869]" captionTargetId="figure@5.[226,1360,553,869]" captionTargetPageId="5" captionText="FIGURE 2. The lectotype of Cobitis lutheri — NRM 10813, male, 64.5 mm SL (copyright under Creative Commons License CC BY NC)." httpUri="https://zenodo.org/record/215096/files/figure.png" pageId="5" pageNumber="582">Fig. 2</figureCitation>
) retains the traces of species-specific color pattern on the head, previously documented on 
<figureCitation id="F2EE2A1B3D7E9B13B7D32B0CFDC4035F" box="[476,549,295,320]" captionStart="FIGURE 4" captionStartId="8.[151,250,1424,1446]" captionTargetBox="[178,1377,833,1373]" captionTargetId="figure@8.[151,1436,769,1403]" captionTargetPageId="8" captionText="FIGURE 4. Characteristic stripes on the head of Cobitis lutheri (a): 1 — stripe extending across the eye; 2 — suborbital stripe; 3 and 4 — stripes on opercle; lamina circularis (b) in male from the Lyanchihe River, Russia (ZMMU P- 3830)." httpUri="https://zenodo.org/record/215098/files/figure.png" pageId="5" pageNumber="582">
Fig. 
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</figureCitation>
in 
<bibRefCitation id="0E444B6F3D7E9B13B4582B0CFCF1035F" author="Rendahl" box="[599,784,295,320]" pageId="5" pageNumber="582" refString="Rendahl, H. (1935) Ein paar neue Unterarten von Cobitis taenia. Memoranda Societatis pro Fauna et Flora Fennica, 10 (1933 - 1934), 329 - 336." type="journal article" year="1935">
<location id="6F0A60453D7E9B13B4582B0CFD55035F" LSID="urn:lsid:plazi:treatment:E27C87883D7F9B1FB6982F13FD8502CB:6F0A60453D7E9B13B4582B0CFD55035F" box="[599,692,295,320]" county="By" municipality="We" name="Rendahl" pageId="5" pageNumber="582">Rendahl</location>
(1935)
</bibRefCitation>
. 
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, this specimen defines the other diagnostic characteristics of the species, as the shape of 
<emphasis id="58A1EA8C3D7E9B13B4AF2B67FC83037B" box="[672,866,332,356]" italics="true" pageId="5" pageNumber="582">lamina circularis</emphasis>
, the snout shape, the position of suborbital spine, and the mandibular barbel length (
<figureCitation id="F2EE2A1B3D7E9B13B4262B44FD900397" box="[553,625,367,392]" captionStart="FIGURE 2" captionStartId="5.[151,250,890,912]" captionTargetBox="[226,1360,553,869]" captionTargetId="figure@5.[226,1360,553,869]" captionTargetPageId="5" captionText="FIGURE 2. The lectotype of Cobitis lutheri — NRM 10813, male, 64.5 mm SL (copyright under Creative Commons License CC BY NC)." httpUri="https://zenodo.org/record/215096/files/figure.png" pageId="5" pageNumber="582">Fig. 2</figureCitation>
). 
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designate this specimen as 
<typeStatus id="B56E883C3D7E9B13B5FA2B44FBBA0397" box="[1013,1115,367,392]" pageId="5" pageNumber="582" type="lectotype">lectotype</typeStatus>
of 
<taxonomicName id="ADD54D1D3D7E9B13B2882B5BFF3303B2" authority="Rendahl, 1935" authorityName="Rendahl" authorityYear="1935" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="582" phylum="Chordata" rank="species" species="lutheri">
<emphasis id="58A1EA8C3D7E9B13B2882B5BFACF0397" box="[1159,1326,368,392]" italics="true" pageId="5" pageNumber="582">Cobitis lutheri</emphasis>
<bibRefCitation id="0E444B6F3D7E9B13B3372B44FF3303B2" author="Rendahl" pageId="5" pageNumber="582" refString="Rendahl, H. (1935) Ein paar neue Unterarten von Cobitis taenia. Memoranda Societatis pro Fauna et Flora Fennica, 10 (1933 - 1934), 329 - 336." type="journal article" year="1935">Rendahl, 1935</bibRefCitation>
</taxonomicName>
, to fix the name of the species to an available zoological entity to extend or modify the description in the future for any taxonomic purposes. 
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that action the other 
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(
<specimenCode id="3A739EE53D7E9B13B5B62B93FB9403CF" box="[953,1141,439,464]" pageId="5" pageNumber="582">
<collectionCode id="0CC4AE5B3D7E9B13B5B62B93FC1B03CF" box="[953,1018,440,464]" country="Sweden" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="5" pageNumber="582">NRM</collectionCode>
10810, 64
</specimenCode>
mm SL, 
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<collectionCode id="0CC4AE5B3D7E9B13B2EC2B93FAC203D0" box="[1251,1315,440,463]" country="Sweden" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="5" pageNumber="582">NRM</collectionCode>
10811, 90
</specimenCode>
mm SL, 
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<collectionCode id="0CC4AE5B3D7E9B13B6F72BF6FED903EB" box="[248,312,477,500]" country="Sweden" httpUri="http://grbio.org/cool/4m0z-xq14" name="Swedish Museum of Natural History - Zoological Collections" pageId="5" pageNumber="582">NRM</collectionCode>
10812, 
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</specimenCode>
mm SL) become 
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</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="3EAA66163D7E9B13B6982951FEFA01B1" httpUri="https://zenodo.org/record/215096/files/figure.png" pageId="5" pageNumber="582" targetBox="[226,1360,553,869]" targetPageId="5">
<paragraph id="6A6A369E3D7E9B13B6982951FEFA01B1" blockId="5.[151,1436,890,942]" pageId="5" pageNumber="582">
<emphasis id="58A1EA8C3D7E9B13B6982951FEF50190" bold="true" box="[151,276,890,912]" pageId="5" pageNumber="582">FIGURE 2.</emphasis>
The lectotype of 
<taxonomicName id="ADD54D1D3D7E9B13B7C42951FD81018F" box="[459,608,890,912]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="582" phylum="Chordata" rank="species" species="lutheri">
<emphasis id="58A1EA8C3D7E9B13B7C42951FD81018F" box="[459,608,890,912]" italics="true" pageId="5" pageNumber="582">Cobitis lutheri</emphasis>
</taxonomicName>
—NRM 10813, male, 64.5 mm SL (copyright under Creative Commons License CC BY NC).
</paragraph>
</caption>
<subSubSection id="22CF65153D7E9B13B6C829FDFC8306D8" pageId="5" pageNumber="582" type="diagnosis">
<paragraph id="6A6A369E3D7E9B13B6C829FDFC8306D8" blockId="5.[151,1437,982,2015]" pageId="5" pageNumber="582">
<emphasis id="58A1EA8C3D7E9B13B6C829FDFEA001F0" bold="true" box="[199,321,982,1007]" pageId="5" pageNumber="582">Diagnosis.</emphasis>
<taxonomicName id="ADD54D1D3D7E9B13B74529FDFE0E01F1" box="[330,495,982,1006]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="582" phylum="Chordata" rank="species" species="lutheri">
<emphasis id="58A1EA8C3D7E9B13B74529FDFE0E01F1" box="[330,495,982,1006]" italics="true" pageId="5" pageNumber="582">Cobitis lutheri</emphasis>
</taxonomicName>
is distinguished by a combination of the following character states: wide poleaxshaped 
<emphasis id="58A1EA8C3D7E9B13B6E229D1FE51060D" box="[237,432,1018,1042]" italics="true" pageId="5" pageNumber="582">lamina circularis</emphasis>
; round scales with slightly-displaced focal zone equal to 61–65% of the scale diameter; short (usually less than 42% of head length) obtuse snout in adults; suborbital spine mostly reaching the center of the eye, and mandibular barbel reaching the anterior edge of the eye in adult specimens; usually fewer than 12 dark brown blotches on the side at the middle of the body; two pronounced spots at the base of caudal fin; commonly several dark streaks on the head, across the eye, suborbital, and two crossed stripes on the opercle; 2n=50 (20 meta- and submetacentrics, and 30 subtelo- and acrocentrics); NF=70.
</paragraph>
</subSubSection>
<subSubSection id="22CF65153D7E9B1FB6C82EF9FD8502CB" lastPageId="9" lastPageNumber="586" pageId="5" pageNumber="582" type="description">
<paragraph id="6A6A369E3D7E9B13B6C82EF9FC9B072D" blockId="5.[151,1437,982,2015]" pageId="5" pageNumber="582">
<emphasis id="58A1EA8C3D7E9B13B6C82EF9FEB806F4" bold="true" box="[199,345,1234,1259]" pageId="5" pageNumber="582">Description.</emphasis>
Dorsal-fin rays 
<date id="1E6B105E3D7E9B13B41F2EF9FDDF06F4" box="[528,574,1234,1259]" pageId="5" pageNumber="582">iii 7</date>
½(8), anal-fin rays 
<date id="1E6B105E3D7E9B13B5192EF9FCA206F4" box="[790,835,1234,1259]" pageId="5" pageNumber="582">iii 5</date>
½, pectoral-fin rays 
<date id="1E6B105E3D7E9B13B2282EF9FB8306F4" box="[1063,1122,1234,1259]" pageId="5" pageNumber="582">i 7–8</date>
½, pelvic fin (V) 
<date id="1E6B105E3D7E9B13B32A2EF9FAA306F4" box="[1317,1346,1234,1259]" pageId="5" pageNumber="582">i 5</date>
½–6(7), caudal-fin rays i (13) 
<date id="1E6B105E3D7E9B13B7702EDCFE4D0710" box="[383,428,1270,1295]" pageId="5" pageNumber="582" value="2014-01">14 i</date>
, total vertebrae in five specimens from the Karasik River 41–44. Morphometric characters of adult specimens from the 
<typeStatus id="B56E883C3D7E9B13B7D92F30FDE7072C" box="[470,518,1307,1331]" pageId="5" pageNumber="582">type</typeStatus>
locality are presented in 
<tableCitation id="275703253D7E9B13B52E2F31FC96072D" box="[801,887,1306,1331]" captionStart="TABLE 2" captionStartId="6.[151,239,652,675]" captionTargetBox="[159,1309,702,1840]" captionTargetPageId="6" captionText="TABLE 2. Selected morphometric characters of Cobitis lutheri from the type locality." httpUri="http://table.plazi.org/id/3EAA66163D7D9B10B69828A7FBCD00BC" pageId="5" pageNumber="582" tableUuid="3EAA66163D7D9B10B69828A7FBCD00BC">Table 2</tableCitation>
.
</paragraph>
<paragraph id="6A6A369E3D7E9B13B6C82F15FB350550" blockId="5.[151,1437,982,2015]" pageId="5" pageNumber="582">
Body elongate, rather deep, laterally compressed. Largest adult females, from 
<collectingRegion id="A811F87C3D7E9B13B2212F14FB8E0748" box="[1070,1135,1343,1367]" country="Russia" name="Amur" pageId="5" pageNumber="582">Amur</collectingRegion>
River, reaching 
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SL (ZMMU P-7566, P-7572). Head elongated, compressed, its upper profile convex; eyes large, superiorly located; eye diameter usually (80–100% of adult individuals in sample) greater than 20% of the head length; interorbital space narrow, convex. Snout mostly short, obtuse, its length in males usually (80–100%) less than 42% of head length; specimens with elongated snout observed among large adult females. Suborbital spine bifid, thick, slightly curved, mostly reaching center of eye (see below). Mouth small, inferior, with fleshy lips; lower lip divided in two well developed parts, its mental lobes a little longer than rest of folded portion. Three short pairs of barbels: rostral, maxillary, and mandibular barbels; mandibular barbel often reaches anterior edge of eye or extends further, primarily in adult males (see below). Caudal peduncle obviously shorter than head, well compressed, with poorly developed fleshy keels on dorsal and ventral sides. Pectoral fin elongated in males, its first ray not more elongated than others, not filamentous (
<figureCitation id="F2EE2A1B3D7E9B13B7EB2C8DFDC804A0" box="[484,553,1702,1727]" captionStart="FIGURE 3" captionStartId="7.[151,250,1550,1572]" captionTargetBox="[163,1428,280,1513]" captionTargetId="figure@7.[151,1436,265,1528]" captionTargetPageId="7" captionText="FIGURE 3. Male (a) SL 45 mm and female (b) SL 50.5 mm of Cobitis lutheri from the Khanka Lake system, Russia (ZMMU P- 20036) and conspecific male (c) SL 53.5 mm and female (d) SL 74 mm from the Tygda River, Russia (ZMMU P- 21545)." httpUri="https://zenodo.org/record/215097/files/figure.png" pageId="5" pageNumber="582">Fig. 3</figureCitation>
). Males with a single 
<emphasis id="58A1EA8C3D7E9B13B5282C8DFC0804A1" box="[807,1001,1702,1726]" italics="true" pageId="5" pageNumber="582">lamina circularis</emphasis>
(
<emphasis id="58A1EA8C3D7E9B13B5F52C8DFB2704A1" box="[1018,1222,1702,1726]" italics="true" pageId="5" pageNumber="582">lamina Canestrini</emphasis>
) at base of second pectoral-fin ray, broad, branched; plate of 
<emphasis id="58A1EA8C3D7E9B13B47F2CE1FCD004FD" box="[624,817,1738,1762]" italics="true" pageId="5" pageNumber="582">lamina circularis</emphasis>
wide, poleax-shaped, its external edge crossed by very short dense lines (
<figureCitation id="F2EE2A1B3D7E9B13B7502CC5FE420518" box="[351,419,1774,1799]" captionStart="FIGURE 4" captionStartId="8.[151,250,1424,1446]" captionTargetBox="[178,1377,833,1373]" captionTargetId="figure@8.[151,1436,769,1403]" captionTargetPageId="8" captionText="FIGURE 4. Characteristic stripes on the head of Cobitis lutheri (a): 1 — stripe extending across the eye; 2 — suborbital stripe; 3 and 4 — stripes on opercle; lamina circularis (b) in male from the Lyanchihe River, Russia (ZMMU P- 3830)." httpUri="https://zenodo.org/record/215098/files/figure.png" pageId="5" pageNumber="582">Fig. 4</figureCitation>
b); end of plate reaching the 2–5th segments of attached ray. Pelvic fin originating posterior to origin of dorsal fin. Body covered with small scales; scales at base of dorsal fin more or less round, with a rather large, slightly-displaced focal zone; diameter of this zone being about 61–65% of scale diameter.
</paragraph>
<paragraph id="6A6A369E3D7E9B10B6C82D71FC65007F" blockId="5.[151,1437,982,2015]" lastBlockId="6.[151,1437,151,608]" lastPageId="6" lastPageNumber="583" pageId="5" pageNumber="582">
Coloration in alcohol. Background color beige in preserved individuals. Head with small dark brown speckles or stripes on dorsal and lateral sides; apart from typical stripe extending across eye from tip of snout to nape, most specimens in preservative with three well developed stripes: first one (suborbital) below and parallel to aforementioned stripe across eye; second and third crossed stripes on opercle (
<figureCitation id="F2EE2A1B3D7E9B13B2322DEDFB6605C0" box="[1085,1159,1990,2015]" captionStart="FIGURE 4" captionStartId="8.[151,250,1424,1446]" captionTargetBox="[178,1377,833,1373]" captionTargetId="figure@8.[151,1436,769,1403]" captionTargetPageId="8" captionText="FIGURE 4. Characteristic stripes on the head of Cobitis lutheri (a): 1 — stripe extending across the eye; 2 — suborbital stripe; 3 and 4 — stripes on opercle; lamina circularis (b) in male from the Lyanchihe River, Russia (ZMMU P- 3830)." httpUri="https://zenodo.org/record/215098/files/figure.png" pageId="5" pageNumber="582">Fig. 4</figureCitation>
a). Gambetta’s zones of pigmentation (
<bibRefCitation id="0E444B6F3D7D9B10B7332ABCFE1402AF" author="Gambetta" box="[316,501,151,176]" pageId="6" pageNumber="583" refString="Gambetta, L. (1934) Sulla variabilita del cobite fluviatile (Cobitis taenia) e sul rapporto numerico dei sessi. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Universita di Torino, 44, 297 - 324." type="journal article" year="1934">Gambetta, 1934</bibRefCitation>
) nearly complete; first zone, at middle of back, represented by large dark-brown spots of variable shape, surrounded by small brown speckles. Second zone consisting of large brown horizontallyelongated spots partially combined in one line or completely forming a relatively wide streak from head to base of caudal fin in males and some females. Third zone consisting of small dark-brown speckles; fourth zone on side at middle of body represented by large dark-brown blotches (larger than eye), square, round or elongated; their number varying from 8 to 16, usually fewer than 12 (often fewer than 10) (see below). Two spots at base of caudal fin: one elongated vivid black spot at upper part of caudal-fin base, another dark brown or black spot of irregular shape at lower part of caudal-fin base. Specimens with only one (upper) visible spot at base of caudal fin very rare. The faded specimens of 
<taxonomicName id="ADD54D1D3D7D9B10B7A92B93FDF003CF" box="[422,529,440,464]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="583" phylum="Chordata" rank="species" species="lutheri">
<emphasis id="58A1EA8C3D7D9B10B7A92B93FDF003CF" box="[422,529,440,464]" italics="true" pageId="6" pageNumber="583">C. lutheri</emphasis>
</taxonomicName>
from the samples collected more than 30 years ago usually possess only one spot at the upper base of the caudal fin in the surface layer sensu 
<bibRefCitation id="0E444B6F3D7D9B10B54D2BF7FBB903EB" author="Saitoh" box="[834,1112,476,501]" pageId="6" pageNumber="583" refString="Saitoh, K. &amp; Aizawa, H. (1987) Local differentiation within the striated spined loach (the striata type of Cobitis taenia complex). Japanese Journal of Ichthyology, 34 (3), 334 - 345." type="journal article" year="1987">Saitoh &amp; Aizawa (1987)</bibRefCitation>
, but both spots are visible in the deeper layer, in the connective tissue surrounding the base of fin rays. Dorsal fin with dark brown pigmentation along its rays; caudal fin with elongated brown speckles along its rays, forming usually four transverse lines; small brown scattered speckles on pectoral, pelvic, and anal fins (
<figureCitation id="F2EE2A1B3D7D9B10B53D286CFC97007F" box="[818,886,583,608]" captionStart="FIGURE 3" captionStartId="7.[151,250,1550,1572]" captionTargetBox="[163,1428,280,1513]" captionTargetId="figure@7.[151,1436,265,1528]" captionTargetPageId="7" captionText="FIGURE 3. Male (a) SL 45 mm and female (b) SL 50.5 mm of Cobitis lutheri from the Khanka Lake system, Russia (ZMMU P- 20036) and conspecific male (c) SL 53.5 mm and female (d) SL 74 mm from the Tygda River, Russia (ZMMU P- 21545)." httpUri="https://zenodo.org/record/215097/files/figure.png" pageId="6" pageNumber="583">Fig. 3</figureCitation>
).
</paragraph>
<caption id="3EAA66163D7D9B10B69828A7FBCD00BC" ID-Table-UUID="3EAA66163D7D9B10B69828A7FBCD00BC" box="[151,1068,651,675]" httpUri="http://table.plazi.org/id/3EAA66163D7D9B10B69828A7FBCD00BC" pageId="6" pageNumber="583" targetBox="[159,1309,702,1840]" targetIsTable="true" targetPageId="6">
<paragraph id="6A6A369E3D7D9B10B69828A7FBCD00BC" blockId="6.[151,1068,651,675]" box="[151,1068,651,675]" pageId="6" pageNumber="583">
<emphasis id="58A1EA8C3D7D9B10B69828A7FEE200BC" bold="true" box="[151,259,652,675]" pageId="6" pageNumber="583">TABLE 2</emphasis>
. Selected morphometric characters of 
<taxonomicName id="ADD54D1D3D7D9B10B49028A0FCD800BC" box="[671,825,651,675]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="583" phylum="Chordata" rank="species" species="lutheri">
<emphasis id="58A1EA8C3D7D9B10B49028A0FCD800BC" box="[671,825,651,675]" italics="true" pageId="6" pageNumber="583">Cobitis lutheri</emphasis>
</taxonomicName>
from the type locality.
</paragraph>
</caption>
<paragraph id="6A6A369E3D7D9B10B6902895FAFD052F" pageId="6" pageNumber="583">
<table id="18D5C43E3D7D64E9B6902895FAFC052F" box="[159,1309,702,1840]" gridcols="4" gridrows="28" pageId="6" pageNumber="583">
<tr id="D4E534DC3D7D64E9B6902895FAFC00CB" box="[159,1309,702,724]" gridrow="0" pageId="6" pageNumber="583" rowspan-3="1">
<th id="97345DA03D7D64E9B6902895FD6900CB" box="[159,648,702,724]" gridcol="0" gridrow="0" pageId="6" pageNumber="583">Characters</th>
<th id="97345DA03D7D64E9B4A82895FCB100CB" box="[679,848,702,724]" gridcol="1" gridrow="0" pageId="6" pageNumber="583">Females (n=6)</th>
<th id="97345DA03D7D64E9B5D02895FB8900CB" box="[991,1128,702,724]" gridcol="2" gridrow="0" pageId="6" pageNumber="583">Males (n=10)</th>
</tr>
<tr id="D4E534DC3D7D64E9B69028C3FAFC00E1" box="[159,1309,744,766]" gridrow="1" pageId="6" pageNumber="583" rowspan-0="1">
<td id="97345DA03D7D64E9B4A828C3FCB100E1" box="[679,848,744,766]" gridcol="1" gridrow="1" pageId="6" pageNumber="583">Mean (range)</td>
<td id="97345DA03D7D64E9B5D028C3FB8900E1" box="[991,1128,744,766]" gridcol="2" gridrow="1" pageId="6" pageNumber="583">Mean+SD</td>
<td id="97345DA03D7D64E9B2B828C3FAFC00E1" box="[1207,1309,744,766]" gridcol="3" gridrow="1" pageId="6" pageNumber="583">Range</td>
</tr>
<tr id="D4E534DC3D7D64E9B690293AFAFC0138" box="[159,1309,785,807]" gridrow="2" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B690293AFD690138" box="[159,648,785,807]" gridcol="0" gridrow="2" pageId="6" pageNumber="583">Total body length (mm)</th>
<td id="97345DA03D7D64E9B4A8293AFCB10138" box="[679,848,785,807]" gridcol="1" gridrow="2" pageId="6" pageNumber="583">63.3 (57.5–69.5)</td>
<td id="97345DA03D7D64E9B5D0293AFB890138" box="[991,1128,785,807]" gridcol="2" gridrow="2" pageId="6" pageNumber="583">56.9+3.46</td>
<td id="97345DA03D7D64E9B2B8293AFAFC0138" box="[1207,1309,785,807]" gridcol="3" gridrow="2" pageId="6" pageNumber="583">50.5–62.3</td>
</tr>
<tr id="D4E534DC3D7D64E9B6902911FAFC014F" box="[159,1309,826,848]" gridrow="3" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B6902911FD69014F" box="[159,648,826,848]" gridcol="0" gridrow="3" pageId="6" pageNumber="583">Standard body length (mm)</th>
<td id="97345DA03D7D64E9B4A82911FCB1014F" box="[679,848,826,848]" gridcol="1" gridrow="3" pageId="6" pageNumber="583">55.7 (49.3–61.0)</td>
<td id="97345DA03D7D64E9B5D02911FB89014F" box="[991,1128,826,848]" gridcol="2" gridrow="3" pageId="6" pageNumber="583">48.5+3.25</td>
<td id="97345DA03D7D64E9B2B82911FAFC014F" box="[1207,1309,826,848]" gridcol="3" gridrow="3" pageId="6" pageNumber="583">43.0–53.6</td>
</tr>
<tr id="D4E534DC3D7D64E9B690294FFAFC0165" box="[159,1309,868,890]" gridrow="4" pageId="6" pageNumber="583" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="97345DA03D7D64E9B690294FFD690165" box="[159,648,868,890]" gridcol="0" gridrow="4" pageId="6" pageNumber="583">In % of standard length</th>
</tr>
<tr id="D4E534DC3D7D64E9B69029A6FAFC01BC" box="[159,1309,909,931]" gridrow="5" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B69029A6FD6901BC" box="[159,648,909,931]" gridcol="0" gridrow="5" pageId="6" pageNumber="583">Predorsal length</th>
<td id="97345DA03D7D64E9B4A829A6FCB101BC" box="[679,848,909,931]" gridcol="1" gridrow="5" pageId="6" pageNumber="583">49.5 (45.9–51.2)</td>
<td id="97345DA03D7D64E9B5D029A6FB8901BC" box="[991,1128,909,931]" gridcol="2" gridrow="5" pageId="6" pageNumber="583">50.1+1.48</td>
<td id="97345DA03D7D64E9B2B829A6FAFC01BC" box="[1207,1309,909,931]" gridcol="3" gridrow="5" pageId="6" pageNumber="583">46.7–52.5</td>
</tr>
<tr id="D4E534DC3D7D64E9B690299DFAFC01D3" box="[159,1309,950,972]" gridrow="6" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B690299DFD6901D3" box="[159,648,950,972]" gridcol="0" gridrow="6" pageId="6" pageNumber="583">Postdorsal lenght</th>
<td id="97345DA03D7D64E9B4A8299DFCB101D3" box="[679,848,950,972]" gridcol="1" gridrow="6" pageId="6" pageNumber="583">38.3 (35.4–40.2)</td>
<td id="97345DA03D7D64E9B5D0299DFB8901D3" box="[991,1128,950,972]" gridcol="2" gridrow="6" pageId="6" pageNumber="583">39.2+1.53</td>
<td id="97345DA03D7D64E9B2B8299DFAFC01D3" box="[1207,1309,950,972]" gridcol="3" gridrow="6" pageId="6" pageNumber="583">37.0–42.3</td>
</tr>
<tr id="D4E534DC3D7D64E9B69029CBFAFC01E9" box="[159,1309,992,1014]" gridrow="7" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B69029CBFD6901E9" box="[159,648,992,1014]" gridcol="0" gridrow="7" pageId="6" pageNumber="583">Preventral length</th>
<td id="97345DA03D7D64E9B4A829CBFCB101E9" box="[679,848,992,1014]" gridcol="1" gridrow="7" pageId="6" pageNumber="583">50.1 (46.4–51.5)</td>
<td id="97345DA03D7D64E9B5D029CBFB8901E9" box="[991,1128,992,1014]" gridcol="2" gridrow="7" pageId="6" pageNumber="583">51.5+2.07</td>
<td id="97345DA03D7D64E9B2B829CBFAFC01E9" box="[1207,1309,992,1014]" gridcol="3" gridrow="7" pageId="6" pageNumber="583">48.9–55.6</td>
</tr>
<tr id="D4E534DC3D7D64E9B6902E22FAFC0600" box="[159,1309,1033,1055]" gridrow="8" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B6902E22FD690600" box="[159,648,1033,1055]" gridcol="0" gridrow="8" pageId="6" pageNumber="583">Preanal length</th>
<td id="97345DA03D7D64E9B4A82E22FCB10600" box="[679,848,1033,1055]" gridcol="1" gridrow="8" pageId="6" pageNumber="583">76.3 (72.3–80.0)</td>
<td id="97345DA03D7D64E9B5D02E22FB890600" box="[991,1128,1033,1055]" gridcol="2" gridrow="8" pageId="6" pageNumber="583">77.3+1.96</td>
<td id="97345DA03D7D64E9B2B82E22FAFC0600" box="[1207,1309,1033,1055]" gridcol="3" gridrow="8" pageId="6" pageNumber="583">73.0–79.5</td>
</tr>
<tr id="D4E534DC3D7D64E9B6902E19FAFC0657" box="[159,1309,1074,1096]" gridrow="9" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B6902E19FD690657" box="[159,648,1074,1096]" gridcol="0" gridrow="9" pageId="6" pageNumber="583">Caudal peduncle length</th>
<td id="97345DA03D7D64E9B4A82E19FCB10657" box="[679,848,1074,1096]" gridcol="1" gridrow="9" pageId="6" pageNumber="583">13.3 (12.2–14.3)</td>
<td id="97345DA03D7D64E9B5D02E19FB890657" box="[991,1128,1074,1096]" gridcol="2" gridrow="9" pageId="6" pageNumber="583">14.5+1.41</td>
<td id="97345DA03D7D64E9B2B82E19FAFC0657" box="[1207,1309,1074,1096]" gridcol="3" gridrow="9" pageId="6" pageNumber="583">12.1–17.3</td>
</tr>
<tr id="D4E534DC3D7D64E9B6902E77FAFC066D" box="[159,1309,1116,1138]" gridrow="10" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B6902E77FD69066D" box="[159,648,1116,1138]" gridcol="0" gridrow="10" pageId="6" pageNumber="583">Maximum body depth</th>
<td id="97345DA03D7D64E9B4A82E77FCB1066D" box="[679,848,1116,1138]" gridcol="1" gridrow="10" pageId="6" pageNumber="583">13.5 (12.0–15.4)</td>
<td id="97345DA03D7D64E9B5D02E77FB89066D" box="[991,1128,1116,1138]" gridcol="2" gridrow="10" pageId="6" pageNumber="583">14.6+2.51</td>
<td id="97345DA03D7D64E9B2B82E77FAFC066D" box="[1207,1309,1116,1138]" gridcol="3" gridrow="10" pageId="6" pageNumber="583">10.6–17.7</td>
</tr>
<tr id="D4E534DC3D7D64E9B6902EAEFAFC0684" box="[159,1309,1157,1179]" gridrow="11" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B6902EAEFD690684" box="[159,648,1157,1179]" gridcol="0" gridrow="11" pageId="6" pageNumber="583">Caudal peduncle depth</th>
<td id="97345DA03D7D64E9B4A82EAEFCB10684" box="[679,848,1157,1179]" gridcol="1" gridrow="11" pageId="6" pageNumber="583">8.6 (7.6–9.5)</td>
<td id="97345DA03D7D64E9B5D02EAEFB890684" box="[991,1128,1157,1179]" gridcol="2" gridrow="11" pageId="6" pageNumber="583">9.0+1.11</td>
<td id="97345DA03D7D64E9B2B82EAEFAFC0684" box="[1207,1309,1157,1179]" gridcol="3" gridrow="11" pageId="6" pageNumber="583">7.0–10.4</td>
</tr>
<tr id="D4E534DC3D7D64E9B6902E85FAFC06DB" box="[159,1309,1198,1220]" gridrow="12" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B6902E85FD6906DB" box="[159,648,1198,1220]" gridcol="0" gridrow="12" pageId="6" pageNumber="583">Maximum body width</th>
<td id="97345DA03D7D64E9B4A82E85FCB106DB" box="[679,848,1198,1220]" gridcol="1" gridrow="12" pageId="6" pageNumber="583">9.8 (8.3–10.8)</td>
<td id="97345DA03D7D64E9B5D02E85FB8906DB" box="[991,1128,1198,1220]" gridcol="2" gridrow="12" pageId="6" pageNumber="583">8.6+3.16</td>
<td id="97345DA03D7D64E9B2B82E85FAFC06DB" box="[1207,1309,1198,1220]" gridcol="3" gridrow="12" pageId="6" pageNumber="583">3.6–15.7</td>
</tr>
<tr id="D4E534DC3D7D64E9B6902EF3FAFC06F1" box="[159,1309,1240,1262]" gridrow="13" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B6902EF3FD6906F1" box="[159,648,1240,1262]" gridcol="0" gridrow="13" pageId="6" pageNumber="583">Length from pectoral fin base to ventral fin base</th>
<td id="97345DA03D7D64E9B4A82EF3FCB106F1" box="[679,848,1240,1262]" gridcol="1" gridrow="13" pageId="6" pageNumber="583">30.2 (28.5–31.9)</td>
<td id="97345DA03D7D64E9B5D02EF3FB8906F1" box="[991,1128,1240,1262]" gridcol="2" gridrow="13" pageId="6" pageNumber="583">30.4+1.54</td>
<td id="97345DA03D7D64E9B2B82EF3FAFC06F1" box="[1207,1309,1240,1262]" gridcol="3" gridrow="13" pageId="6" pageNumber="583">28.3–33.3</td>
</tr>
<tr id="D4E534DC3D7D64E9B6902F2AFAFC0708" box="[159,1309,1281,1303]" gridrow="14" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B6902F2AFD690708" box="[159,648,1281,1303]" gridcol="0" gridrow="14" pageId="6" pageNumber="583">Pectoral fin length</th>
<td id="97345DA03D7D64E9B4A82F2AFCB10708" box="[679,848,1281,1303]" gridcol="1" gridrow="14" pageId="6" pageNumber="583">14.3 (13.0–15.0)</td>
<td id="97345DA03D7D64E9B5D02F2AFB890708" box="[991,1128,1281,1303]" gridcol="2" gridrow="14" pageId="6" pageNumber="583">18.1+1.92</td>
<td id="97345DA03D7D64E9B2B82F2AFAFC0708" box="[1207,1309,1281,1303]" gridcol="3" gridrow="14" pageId="6" pageNumber="583">14.4–20.9</td>
</tr>
<tr id="D4E534DC3D7D64E9B6902F01FAFC075F" box="[159,1309,1322,1344]" gridrow="15" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B6902F01FD69075F" box="[159,648,1322,1344]" gridcol="0" gridrow="15" pageId="6" pageNumber="583">Ventral fin length</th>
<td id="97345DA03D7D64E9B4A82F01FCB1075F" box="[679,848,1322,1344]" gridcol="1" gridrow="15" pageId="6" pageNumber="583">12.7 (11.8–13.6)</td>
<td id="97345DA03D7D64E9B5D02F01FB89075F" box="[991,1128,1322,1344]" gridcol="2" gridrow="15" pageId="6" pageNumber="583">15.3+1.44</td>
<td id="97345DA03D7D64E9B2B82F01FAFC075F" box="[1207,1309,1322,1344]" gridcol="3" gridrow="15" pageId="6" pageNumber="583">13.3–17.4</td>
</tr>
<tr id="D4E534DC3D7D64E9B6902F7FFAFC0775" box="[159,1309,1364,1386]" gridrow="16" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B6902F7FFD690775" box="[159,648,1364,1386]" gridcol="0" gridrow="16" pageId="6" pageNumber="583">Length of dorsal fin base</th>
<td id="97345DA03D7D64E9B4A82F7FFCB10775" box="[679,848,1364,1386]" gridcol="1" gridrow="16" pageId="6" pageNumber="583">11.6 (10.0–12.5)</td>
<td id="97345DA03D7D64E9B5D02F7FFB890775" box="[991,1128,1364,1386]" gridcol="2" gridrow="16" pageId="6" pageNumber="583">11.5+0.97</td>
<td id="97345DA03D7D64E9B2B82F7FFAFC0775" box="[1207,1309,1364,1386]" gridcol="3" gridrow="16" pageId="6" pageNumber="583">10.0–12.7</td>
</tr>
<tr id="D4E534DC3D7D64E9B6902F56FAFC078C" box="[159,1309,1405,1427]" gridrow="17" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B6902F56FD69078C" box="[159,648,1405,1427]" gridcol="0" gridrow="17" pageId="6" pageNumber="583">Dorsal fin depth</th>
<td id="97345DA03D7D64E9B4A82F56FCB1078C" box="[679,848,1405,1427]" gridcol="1" gridrow="17" pageId="6" pageNumber="583">16.0 (14.5–17.0)</td>
<td id="97345DA03D7D64E9B5D02F56FB89078C" box="[991,1128,1405,1427]" gridcol="2" gridrow="17" pageId="6" pageNumber="583">16.3+1.75</td>
<td id="97345DA03D7D64E9B2B82F56FAFC078C" box="[1207,1309,1405,1427]" gridcol="3" gridrow="17" pageId="6" pageNumber="583">13.8–20.0</td>
</tr>
<tr id="D4E534DC3D7D64E9B6902F8DFAFC07A3" box="[159,1309,1446,1468]" gridrow="18" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B6902F8DFD6907A3" box="[159,648,1446,1468]" gridcol="0" gridrow="18" pageId="6" pageNumber="583">Length of anal fin base</th>
<td id="97345DA03D7D64E9B4A82F8DFCB107A3" box="[679,848,1446,1468]" gridcol="1" gridrow="18" pageId="6" pageNumber="583">8.6 (7.3–9.9)</td>
<td id="97345DA03D7D64E9B5D02F8DFB8907A3" box="[991,1128,1446,1468]" gridcol="2" gridrow="18" pageId="6" pageNumber="583">8.8+0.41</td>
<td id="97345DA03D7D64E9B2B82F8DFAFC07A3" box="[1207,1309,1446,1468]" gridcol="3" gridrow="18" pageId="6" pageNumber="583">8.3–9.6</td>
</tr>
<tr id="D4E534DC3D7D64E9B6902FFBFAFC07F9" box="[159,1309,1488,1510]" gridrow="19" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B6902FFBFD6907F9" box="[159,648,1488,1510]" gridcol="0" gridrow="19" pageId="6" pageNumber="583">Head length</th>
<td id="97345DA03D7D64E9B4A82FFBFCB107F9" box="[679,848,1488,1510]" gridcol="1" gridrow="19" pageId="6" pageNumber="583">20.6 (18.7–21.8)</td>
<td id="97345DA03D7D64E9B5D02FFBFB8907F9" box="[991,1128,1488,1510]" gridcol="2" gridrow="19" pageId="6" pageNumber="583">21.5+1.22</td>
<td id="97345DA03D7D64E9B2B82FFBFAFC07F9" box="[1207,1309,1488,1510]" gridcol="3" gridrow="19" pageId="6" pageNumber="583">19.8–24.0</td>
</tr>
<tr id="D4E534DC3D7D64E9B6902FD2FAFC0410" box="[159,1309,1529,1551]" gridrow="20" pageId="6" pageNumber="583" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="97345DA03D7D64E9B6902FD2FD690410" box="[159,648,1529,1551]" gridcol="0" gridrow="20" pageId="6" pageNumber="583">In % of head length</th>
</tr>
<tr id="D4E534DC3D7D64E9B6902C09FAFC0427" box="[159,1309,1570,1592]" gridrow="21" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B6902C09FD690427" box="[159,648,1570,1592]" gridcol="0" gridrow="21" pageId="6" pageNumber="583">Snout length</th>
<td id="97345DA03D7D64E9B4A82C09FCB10427" box="[679,848,1570,1592]" gridcol="1" gridrow="21" pageId="6" pageNumber="583">41.5 (39.2–42.7)</td>
<td id="97345DA03D7D64E9B5D02C09FB890427" box="[991,1128,1570,1592]" gridcol="2" gridrow="21" pageId="6" pageNumber="583">38.3+3.09</td>
<td id="97345DA03D7D64E9B2B82C09FAFC0427" box="[1207,1309,1570,1592]" gridcol="3" gridrow="21" pageId="6" pageNumber="583">30.0–41.2</td>
</tr>
<tr id="D4E534DC3D7D64E9B6902C67FAFC047D" box="[159,1309,1612,1634]" gridrow="22" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B6902C67FD69047D" box="[159,648,1612,1634]" gridcol="0" gridrow="22" pageId="6" pageNumber="583">Eye diameter</th>
<td id="97345DA03D7D64E9B4A82C67FCB1047D" box="[679,848,1612,1634]" gridcol="1" gridrow="22" pageId="6" pageNumber="583">22.8 (20.5–25.2)</td>
<td id="97345DA03D7D64E9B5D02C67FB89047D" box="[991,1128,1612,1634]" gridcol="2" gridrow="22" pageId="6" pageNumber="583">23.2+2.61</td>
<td id="97345DA03D7D64E9B2B82C67FAFC047D" box="[1207,1309,1612,1634]" gridcol="3" gridrow="22" pageId="6" pageNumber="583">19.8–27.0</td>
</tr>
<tr id="D4E534DC3D7D64E9B6902C5EFAFC0494" box="[159,1309,1653,1675]" gridrow="23" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B6902C5EFD690494" box="[159,648,1653,1675]" gridcol="0" gridrow="23" pageId="6" pageNumber="583">Interorbital distance</th>
<td id="97345DA03D7D64E9B4A82C5EFCB10494" box="[679,848,1653,1675]" gridcol="1" gridrow="23" pageId="6" pageNumber="583">11.8 (9.2–13.5)</td>
<td id="97345DA03D7D64E9B5D02C5EFB890494" box="[991,1128,1653,1675]" gridcol="2" gridrow="23" pageId="6" pageNumber="583">13.9+2.35</td>
<td id="97345DA03D7D64E9B2B82C5EFAFC0494" box="[1207,1309,1653,1675]" gridcol="3" gridrow="23" pageId="6" pageNumber="583">9.9–18.0</td>
</tr>
<tr id="D4E534DC3D7D64E9B6902CB5FAFC04AB" box="[159,1309,1694,1716]" gridrow="24" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B6902CB5FD6904AB" box="[159,648,1694,1716]" gridcol="0" gridrow="24" pageId="6" pageNumber="583">Head depth</th>
<td id="97345DA03D7D64E9B4A82CB5FCB104AB" box="[679,848,1694,1716]" gridcol="1" gridrow="24" pageId="6" pageNumber="583">60.0 (55.8–62.7)</td>
<td id="97345DA03D7D64E9B5D02CB5FB8904AB" box="[991,1128,1694,1716]" gridcol="2" gridrow="24" pageId="6" pageNumber="583">57.4+4.39</td>
<td id="97345DA03D7D64E9B2B82CB5FAFC04AB" box="[1207,1309,1694,1716]" gridcol="3" gridrow="24" pageId="6" pageNumber="583">51.5–63.1</td>
</tr>
<tr id="D4E534DC3D7D64E9B6902CE3FAFC04C1" box="[159,1309,1736,1758]" gridrow="25" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B6902CE3FD6904C1" box="[159,648,1736,1758]" gridcol="0" gridrow="25" pageId="6" pageNumber="583">Rostral barbel length</th>
<td id="97345DA03D7D64E9B4A82CE3FCB104C1" box="[679,848,1736,1758]" gridcol="1" gridrow="25" pageId="6" pageNumber="583">9.0 (4.6–10.8)</td>
<td id="97345DA03D7D64E9B5D02CE3FB8904C1" box="[991,1128,1736,1758]" gridcol="2" gridrow="25" pageId="6" pageNumber="583">9.6*</td>
<td id="97345DA03D7D64E9B2B82CE3FAFC04C1" box="[1207,1309,1736,1758]" gridcol="3" gridrow="25" pageId="6" pageNumber="583">6.8–11.0</td>
</tr>
<tr id="D4E534DC3D7D64E9B6902CDAFAFC0518" box="[159,1309,1777,1799]" gridrow="26" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B6902CDAFD690518" box="[159,648,1777,1799]" gridcol="0" gridrow="26" pageId="6" pageNumber="583">Mandibular barbel length</th>
<td id="97345DA03D7D64E9B4A82CDAFCB10518" box="[679,848,1777,1799]" gridcol="1" gridrow="26" pageId="6" pageNumber="583">13.2 (9.1–15.5)</td>
<td id="97345DA03D7D64E9B5D02CDAFB890518" box="[991,1128,1777,1799]" gridcol="2" gridrow="26" pageId="6" pageNumber="583">15.8+4.64</td>
<td id="97345DA03D7D64E9B2B82CDAFAFC0518" box="[1207,1309,1777,1799]" gridcol="3" gridrow="26" pageId="6" pageNumber="583">8.7–22.0</td>
</tr>
<tr id="D4E534DC3D7D64E9B6902D31FAFC052F" box="[159,1309,1818,1840]" gridrow="27" pageId="6" pageNumber="583">
<th id="97345DA03D7D64E9B6902D31FD69052F" box="[159,648,1818,1840]" gridcol="0" gridrow="27" pageId="6" pageNumber="583">Caudal peduncle depth in % of its length</th>
<td id="97345DA03D7D64E9B4A82D31FCB1052F" box="[679,848,1818,1840]" gridcol="1" gridrow="27" pageId="6" pageNumber="583">65.0 (56.3–74.6)</td>
<td id="97345DA03D7D64E9B5D02D31FB89052F" box="[991,1128,1818,1840]" gridcol="2" gridrow="27" pageId="6" pageNumber="583">62.7+11.79</td>
<td id="97345DA03D7D64E9B2B82D31FAFC052F" box="[1207,1309,1818,1840]" gridcol="3" gridrow="27" pageId="6" pageNumber="583">40.2–75.0</td>
</tr>
</table>
</paragraph>
<paragraph id="6A6A369E3D7D9B10B6982D7AFDC90577" blockId="6.[151,552,1873,1896]" box="[151,552,1873,1896]" pageId="6" pageNumber="583">* based on eight examined specimens</paragraph>
<paragraph id="6A6A369E3D7D9B11B6C82DBEFC1F02CA" blockId="6.[151,1436,1941,2038]" lastBlockId="7.[151,1436,151,213]" lastPageId="7" lastPageNumber="584" pageId="6" pageNumber="583">
Sexual dimorphism. Females with larger, slenderer body than males. Males with 
<emphasis id="58A1EA8C3D7D9B10B26C2DBEFAC405B2" box="[1123,1317,1941,1965]" italics="true" pageId="6" pageNumber="583">lamina circularis</emphasis>
at base of second pectoral-fin ray, elongated pectoral and pelvic fins, usually shorter snout (
<tableCitation id="275703253D7D9B10B2112D92FB9405CD" box="[1054,1141,1977,2002]" captionStart="TABLE 2" captionStartId="6.[151,239,652,675]" captionTargetBox="[159,1309,702,1840]" captionTargetPageId="6" captionText="TABLE 2. Selected morphometric characters of Cobitis lutheri from the type locality." httpUri="http://table.plazi.org/id/3EAA66163D7D9B10B69828A7FBCD00BC" pageId="6" pageNumber="583" tableUuid="3EAA66163D7D9B10B69828A7FBCD00BC">Table 2</tableCitation>
); during spawning season, dark spots and blotches in second and fourth Gambetta’ zones in adult males usually merging with each other, forming two broad dark stripes (
<bibRefCitation id="0E444B6F3D7C9B11B4012ABCFD5902AF" author="Shedko" box="[526,696,151,176]" pageId="7" pageNumber="584" refString="Shedko, S. V. (2001) List of Cyclostomata and fishes of fresh waters of the coast of Primorye. In: Chteniya pamyati Fladimira Yakovlevicha Levanidova, 1. Vladivostok. Dalnauka. pp. 229 - 249 [in Russian]." type="journal article" year="2001">Shedko (2001)</bibRefCitation>
noted that these stripes split into series of independent spots or blotches during late autumn, as were observed in the specimens in aquarium).
</paragraph>
<caption id="3EAA66163D7C9B11B6982C25FA9A045D" httpUri="https://zenodo.org/record/215097/files/figure.png" pageId="7" pageNumber="584" targetBox="[163,1428,280,1513]" targetPageId="7">
<paragraph id="6A6A369E3D7C9B11B6982C25FA9A045D" blockId="7.[151,1435,1550,1602]" pageId="7" pageNumber="584">
<emphasis id="58A1EA8C3D7C9B11B6982C25FEF3043C" bold="true" box="[151,274,1550,1572]" pageId="7" pageNumber="584">FIGURE 3.</emphasis>
Male (a) SL 45 mm and female (b) SL 50.5 mm of 
<taxonomicName id="ADD54D1D3D7C9B11B52C2C25FC57043B" box="[803,950,1550,1572]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="584" phylum="Chordata" rank="species" species="lutheri">
<emphasis id="58A1EA8C3D7C9B11B52C2C25FC57043B" box="[803,950,1550,1572]" italics="true" pageId="7" pageNumber="584">Cobitis lutheri</emphasis>
</taxonomicName>
from the Khanka Lake system, Russia (ZMMU P-20036) and conspecific male (c) SL 53.5 mm and female (d) SL 74 mm from the Tygda River, Russia (ZMMU P-21545).
</paragraph>
</caption>
<paragraph id="6A6A369E3D7C9B1EB6C82C40FA75009B" blockId="7.[151,1437,1642,2027]" lastBlockId="8.[151,1437,151,717]" lastPageId="8" lastPageNumber="585" pageId="7" pageNumber="584">
Intra- and inter-population variability of diagnostic features. Among 
<specimenCount id="7CD3FD173D7C9B11B5F72C41FB45049C" box="[1016,1188,1642,1667]" pageId="7" pageNumber="584" type="generic">135 specimens</specimenCount>
examined from ZIN collection, only 6.7% (three females from the 
<collectingRegion id="A811F87C3D7C9B11B4912CA5FD3E04B9" box="[670,735,1678,1702]" country="Russia" name="Amur" pageId="7" pageNumber="584">Amur</collectingRegion>
River and 
<specimenCount id="7CD3FD173D7C9B11B5532CA6FC2904B9" box="[860,968,1677,1702]" pageId="7" pageNumber="584" type="female">6 females</specimenCount>
from the 
<collectingRegion id="A811F87C3D7C9B11B2382CA6FB6E04B9" box="[1079,1167,1677,1702]" country="Russia" name="Tyumen'" pageId="7" pageNumber="584">Tumen’</collectingRegion>
-Ula River) had none or only a vague dark suborbital stripe on the head. Similarly, 3.0% (three females from the 
<collectingRegion id="A811F87C3D7C9B11B2992C99FB3604D5" box="[1174,1239,1714,1738]" country="Russia" name="Amur" pageId="7" pageNumber="584">Amur</collectingRegion>
and 
<collectingRegion id="A811F87C3D7C9B11B31C2C99FA8A04D4" box="[1299,1387,1714,1739]" country="Russia" name="Tyumen'" pageId="7" pageNumber="584">Tumen’</collectingRegion>
-Ula rivers and one male from the Tunguska River) lacked the spot at the lower part of the caudal-fin base. All of 40 examined specimens from Khanka Lake basin had a conspicuous suborbital stripe and two spots at the caudal-fin base. Among 
<specimenCount id="7CD3FD173D7C9B11B73A2D35FE730529" box="[309,402,1821,1846]" pageId="7" pageNumber="584" type="adult">17 adult</specimenCount>
specimens from the 
<typeStatus id="B56E883C3D7C9B11B48D2D34FD530528" box="[642,690,1823,1847]" pageId="7" pageNumber="584">type</typeStatus>
locality in ZMMU collection, only one specimen (male, SL 
<quantity id="AD2D9B7B3D7C9B11B3642D36FF250545" metricMagnitude="-2" metricUnit="m" metricValue="4.95" pageId="7" pageNumber="584" unit="mm" value="49.5">49.5 mm</quantity>
) had no visible suborbital stripe; all specimens had two spots at the caudal-fin base; the number of blotches in the fourth Gambetta’s zone varied from 8 to 14, most specimens (68.8%) had fewer than 12 blotches; suborbital spine usually reached the center of the eye (58.8%) or extended further (35.3%); the mandibular barbel reached the anterior edge of the eye or extended further in 35.3% of the adults; one male (P-17837) had 
<emphasis id="58A1EA8C3D7C9B11B2AD2D85FA8205D9" box="[1186,1379,1966,1990]" italics="true" pageId="7" pageNumber="584">lamina circularis</emphasis>
with an atypical narrow bottle-shaped plate on the right pectoral fin. Only one specimen among 20 completely faded adults from the Lyanchihe River in the ZMMU collection had mandibular barbel not reaching the anterior edge of the eye; in all specimens the suborbital spine reached the center of the eye or extended further. Only two specimens among 
<specimenCount id="7CD3FD173D739B1EB6E22ACBFEB602E7" box="[237,343,223,248]" pageId="8" pageNumber="585" type="adult">34 adults</specimenCount>
from the Tsukanovka River (ZMMU P-23900) had a poorly visible suborbital stripe; in most specimens (67.7%) the mandibular barbel did not reach the anterior edge of the eye; the suborbital spine did not reach the center of the eye in 50% of specimens; one male had 
<emphasis id="58A1EA8C3D739B1EB56B2B03FBC7035F" box="[868,1062,296,320]" italics="true" pageId="8" pageNumber="585">lamina circularis</emphasis>
with an atypical narrow plate on the right pectoral fin; all specimens had two spots at the caudal fin base; among 
<specimenCount id="7CD3FD173D739B1EB2262B67FB67037A" box="[1065,1158,332,357]" pageId="8" pageNumber="585" type="adult">57 adult</specimenCount>
and juvenile specimens, 47.4% had fewer than 10 blotches in the fourth Gambetta’s zone, the total number of blotches varying from 8 to 16. In 
<specimenCount id="7CD3FD173D739B1EB6B42BBFFEB803B2" box="[187,345,404,429]" pageId="8" pageNumber="585" type="generic">12 specimens</specimenCount>
from the Razdolnaya River (ZMMU P-20041), 8–12 blotches were observed in the fourth Gambetta’s zone, and specimens with fewer than 10 blotches comprised 48.4%. Large adults (
<specimenCount id="7CD3FD173D739B1EB2DF2B9CFAD603CF" box="[1232,1335,439,464]" pageId="8" pageNumber="585" type="male">16 males</specimenCount>
with SL 52.5–58.0 mm, 
<specimenCount id="7CD3FD173D739B1EB7472BF7FE2203EA" box="[328,451,476,501]" pageId="8" pageNumber="585" type="female">20 females</specimenCount>
with SL 64.3–85.0 mm) from the Tygda River (ZMMU P-21545) had 7–12 blotches in the fourth Gambetta’s zone, specimens with 7–10 blotches were predominant (72.2%); only one female had no spot at the lower part of the caudal fin base; one male had suborbital spine not reaching the center of the eye; mandibular barbel not reaching the anterior edge of the eye was observed in six females; the snout length in females varied from 37.9 to 43.5% of the head length (mean 40.9%), in males, from 36.0 to 43.8% (mean 39.4%).
</paragraph>
<paragraph id="6A6A369E3D739B1EB6C828A4FB3900D2" blockId="8.[151,1437,151,717]" pageId="8" pageNumber="585">
The karyotype is characterized by the total chromosome number (2n) of 50 chromosomes with 12 meta-, 8 submeta- and 30 subtelo- and acrocentrics and NF=70 (
<tableCitation id="275703253D739B1EB50C289FFCB700D3" box="[771,854,692,717]" captionStart="TABLE 1" captionStartId="1.[151,236,980,1003]" captionTargetBox="[151,1425,1064,1743]" captionTargetPageId="1" captionText="TABLE 1. The karyotypes of Cobitis lutheri from the Amur River and Primoye, and C. nalbanti sp. nov from South Korea" httpUri="http://table.plazi.org/id/3EAA66163D7A9B17B69829FFFF390613" pageId="8" pageNumber="585" tableUuid="3EAA66163D7A9B17B69829FFFF390613">Table 1</tableCitation>
) (Vasil’ev &amp; Vasil’eva, 2008a, b).
</paragraph>
<caption id="3EAA66163D739B1EB6982FBBFB0307DA" httpUri="https://zenodo.org/record/215098/files/figure.png" pageId="8" pageNumber="585" targetBox="[178,1377,833,1373]" targetPageId="8">
<paragraph id="6A6A369E3D739B1EB6982FBBFB0307DA" blockId="8.[151,1435,1424,1477]" pageId="8" pageNumber="585">
<emphasis id="58A1EA8C3D739B1EB6982FBBFEEC07BA" bold="true" box="[151,269,1424,1446]" pageId="8" pageNumber="585">FIGURE 4</emphasis>
. Characteristic stripes on the head of 
<taxonomicName id="ADD54D1D3D739B1EB4842FBBFCFE07B9" box="[651,799,1424,1446]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="585" phylum="Chordata" rank="species" species="lutheri">
<emphasis id="58A1EA8C3D739B1EB4842FBBFCFE07B9" box="[651,799,1424,1446]" italics="true" pageId="8" pageNumber="585">Cobitis lutheri</emphasis>
</taxonomicName>
(a): 1—stripe extending across the eye; 2—suborbital stripe; 3 and 4—stripes on opercle; 
<emphasis id="58A1EA8C3D739B1EB7A82F84FDB707DA" box="[423,598,1455,1477]" italics="true" pageId="8" pageNumber="585">lamina circularis</emphasis>
(b) in male from the Lyanchihe River, Russia (ZMMU P-3830).
</paragraph>
</caption>
<paragraph id="6A6A369E3D739B1EB6C82FDAFA660535" blockId="8.[151,1436,1521,2018]" pageId="8" pageNumber="585">
<materialsCitation id="DABD3CC33D739B1EB6C82FDAFA660535" ID-GBIF-Occurrence="1417501501" country="Japan" county="Gastello village" location="Ussuri River" municipality="Lakh River" pageId="8" pageNumber="585" specimenCount="1" stateProvince="Amur">
<emphasis id="58A1EA8C3D739B1EB6C82FDAFDEB0415" bold="true" box="[199,522,1521,1546]" pageId="8" pageNumber="585">Geographical distribution.</emphasis>
<taxonomicName id="ADD54D1D3D739B1EB4192FDAFD5E0416" box="[534,703,1521,1545]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="585" phylum="Chordata" rank="species" species="lutheri">
<emphasis id="58A1EA8C3D739B1EB4192FDAFD5E0416" box="[534,703,1521,1545]" italics="true" pageId="8" pageNumber="585">Cobitis lutheri</emphasis>
</taxonomicName>
occurs in the lower and middle 
<collectingRegion id="A811F87C3D739B1EB2412FD9FB6E0415" box="[1102,1167,1522,1546]" country="Russia" name="Amur" pageId="8" pageNumber="585">Amur</collectingRegion>
River drainage (up to Blagoveshchensk), including the 
<location id="6F0A60453D739B1EB4192C3EFD490431" LSID="urn:lsid:plazi:treatment:E27C87883D7F9B1FB6982F13FD8502CB:6F0A60453D739B1EB4192C3EFD490431" box="[534,680,1557,1582]" country="Japan" county="Gastello village" municipality="Lakh River" name="Ussuri River" pageId="8" pageNumber="585" stateProvince="Amur">Ussuri River</location>
and 
<location id="6F0A60453D739B1EB4EA2C3EFC9E0431" LSID="urn:lsid:plazi:treatment:E27C87883D7F9B1FB6982F13FD8502CB:6F0A60453D739B1EB4EA2C3EFC9E0431" box="[741,895,1557,1582]" country="Japan" county="Gastello village" municipality="Lakh River" name="Khanka Lake" pageId="8" pageNumber="585" stateProvince="Amur">Khanka Lake</location>
systems, rivers flowing into 
<collectingRegion id="A811F87C3D739B1EB2DB2C3DFAF70431" box="[1236,1302,1558,1582]" country="Russia" name="Amur" pageId="8" pageNumber="585">Amur</collectingRegion>
and Ussuri bays in Primorye, and adjacent parts of the 
<location id="6F0A60453D739B1EB48A2C11FD04044D" LSID="urn:lsid:plazi:treatment:E27C87883D7F9B1FB6982F13FD8502CB:6F0A60453D739B1EB48A2C11FD04044D" box="[645,741,1594,1618]" country="Japan" county="Gastello village" municipality="Lakh River" name="East Sea" pageId="8" pageNumber="585" stateProvince="Amur">East Sea</location>
/Sea of 
<collectingCountry id="12C2760E3D739B1EB5312C11FC9F044E" box="[830,894,1594,1617]" name="Japan" pageId="8" pageNumber="585">Japan</collectingCountry>
(e.g., 
<location id="6F0A60453D739B1EB5C92C11FB6E044D" LSID="urn:lsid:plazi:treatment:E27C87883D7F9B1FB6982F13FD8502CB:6F0A60453D739B1EB5C92C11FB6E044D" box="[966,1167,1593,1618]" country="Japan" county="Gastello village" municipality="Lakh River" name="Tumannaya River" pageId="8" pageNumber="585" stateProvince="Amur">Tumannaya River</location>
at 
<collectingCountry id="12C2760E3D739B1EB2BB2C12FB1F044D" box="[1204,1278,1593,1618]" name="Russia" pageId="8" pageNumber="585">Russia</collectingCountry>
/ 
<location id="6F0A60453D739B1EB3012C12FA7D044E" LSID="urn:lsid:plazi:treatment:E27C87883D7F9B1FB6982F13FD8502CB:6F0A60453D739B1EB3012C12FA7D044E" box="[1294,1436,1593,1618]" country="Japan" county="Gastello village" municipality="Lakh River" name="North Korea" pageId="8" pageNumber="585" stateProvince="Amur">North Korea</location>
border), and in rivers in northern 
<collectingRegion id="A811F87C3D739B1EB4042C76FD8C0469" box="[523,621,1629,1654]" country="Russia" name="Sakhalin" pageId="8" pageNumber="585">Sakhalin</collectingRegion>
Island (
<figureCitation id="F2EE2A1B3D739B1EB4C82C76FCF70469" box="[711,790,1629,1654]" captionStart="FIGURE 1" captionStartId="2.[151,250,1829,1851]" captionTargetBox="[188,1398,881,1808]" captionTargetId="figure@2.[188,1398,881,1808]" captionTargetPageId="2" captionText="FIGURE 1 a. The range of Cobitis lutheri covered by the samples from the museum collections marked by mottled sites (a) and sampling localities of Cobitis nalbanti sp. nov. indicated by circles with numbers (b). Locality names in (b) are the following: 1, Cheongmi River, Han River drainage (type locality), 2, Geumdang River, Han River drainage, and 3, Baekgok River, Geum River drainage." httpUri="https://zenodo.org/record/215095/files/figure.png" pageId="8" pageNumber="585">Fig. 1a</figureCitation>
). 
<bibRefCitation id="0E444B6F3D739B1EB5232C76FC300469" author="Shedko" box="[812,977,1629,1654]" pageId="8" pageNumber="585" refString="Shedko, S. V. (2001) List of Cyclostomata and fishes of fresh waters of the coast of Primorye. In: Chteniya pamyati Fladimira Yakovlevicha Levanidova, 1. Vladivostok. Dalnauka. pp. 229 - 249 [in Russian]." type="journal article" year="2001">Shedko (2001)</bibRefCitation>
found this species in rivers of the central part of Primorye coast, namely Kievka, Chernaya, Margaritovka, and Rudnaya. 
<bibRefCitation id="0E444B6F3D739B1EB2452CAAFA7D0485" author="Safronov" box="[1098,1436,1665,1690]" pageId="8" pageNumber="585" refString="Safronov, A. S. &amp; Safronov, S. N. (2002) Luther's spined loach Cobitis lutheri Rendahl (Cobitidae, Pisces) from the Sakhalin Island. In: Biologiya vnutrennikh vod: problemy ecologii i bioraznoobraziya. Tezisy mezhdunarodnoi konferentsii molodykh uchenykh. Borok. pp. 97 - 97. [in Russian]" type="book" year="2002">Safronov &amp; Safronov (2002)</bibRefCitation>
recorded 
<taxonomicName id="ADD54D1D3D739B1EB70E2C8DFE8C04A2" box="[257,365,1701,1725]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="585" phylum="Chordata" rank="species" species="lutheri">
<emphasis id="58A1EA8C3D739B1EB70E2C8DFE8C04A2" box="[257,365,1701,1725]" italics="true" pageId="8" pageNumber="585">C. lutheri</emphasis>
</taxonomicName>
from rivers of northeastern 
<collectingRegion id="A811F87C3D739B1EB4A32C8EFCEF04A1" box="[684,782,1701,1726]" country="Russia" name="Sakhalin" pageId="8" pageNumber="585">Sakhalin</collectingRegion>
Island (from 
<location id="6F0A60453D739B1EB5A52C8EFB9204A1" LSID="urn:lsid:plazi:treatment:E27C87883D7F9B1FB6982F13FD8502CB:6F0A60453D739B1EB5A52C8EFB9204A1" box="[938,1139,1701,1726]" country="Japan" county="Gastello village" municipality="Lakh River" name="Uspenskoye Lake" pageId="8" pageNumber="585" stateProvince="Amur">Uspenskoye Lake</location>
to 
<collectingMunicipality id="8A0EACE43D739B1EB2962C8EFAF904A1" box="[1177,1304,1701,1726]" pageId="8" pageNumber="585">Lakh River</collectingMunicipality>
drainages), from the eastern 
<collectingRegion id="A811F87C3D739B1EB7532CE2FE5F04FD" box="[348,446,1737,1762]" country="Russia" name="Sakhalin" pageId="8" pageNumber="585">Sakhalin</collectingRegion>
coast (including Tym’ and Poronai rivers), and from the open pits at 
<collectingCounty id="830B4E123D739B1EB2E62CE2FA7D04FD" box="[1257,1436,1737,1762]" pageId="8" pageNumber="585">Gastello village</collectingCounty>
(southeastern 
<collectingRegion id="A811F87C3D739B1EB73A2CC6FE7D0519" box="[309,412,1773,1798]" country="Russia" name="Sakhalin" pageId="8" pageNumber="585">Sakhalin</collectingRegion>
). However, the morphological description of the species by 
<bibRefCitation id="0E444B6F3D739B1EB25D2CC6FA7D0519" author="Safronov" box="[1106,1436,1773,1798]" pageId="8" pageNumber="585" refString="Safronov, A. S. &amp; Safronov, S. N. (2002) Luther's spined loach Cobitis lutheri Rendahl (Cobitidae, Pisces) from the Sakhalin Island. In: Biologiya vnutrennikh vod: problemy ecologii i bioraznoobraziya. Tezisy mezhdunarodnoi konferentsii molodykh uchenykh. Borok. pp. 97 - 97. [in Russian]" type="book" year="2002">Safronov &amp; Safronov (2002)</bibRefCitation>
includes features absent in 
<taxonomicName id="ADD54D1D3D739B1EB7C92D39FDD30536" box="[454,562,1809,1833]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="585" phylum="Chordata" rank="species" species="lutheri">
<emphasis id="58A1EA8C3D739B1EB7C92D39FDD30536" box="[454,562,1809,1833]" italics="true" pageId="8" pageNumber="585">C. lutheri</emphasis>
</taxonomicName>
(e.g., “narrow plate” of 
<emphasis id="58A1EA8C3D739B1EB54C2D3AFBE50536" box="[835,1028,1809,1833]" italics="true" pageId="8" pageNumber="585">lamina circularis</emphasis>
, and 18 branched caudal fin rays).
</materialsCitation>
</paragraph>
<paragraph id="6A6A369E3D739B1EB6C82D1EFD540569" blockId="8.[151,1436,1521,2018]" pageId="8" pageNumber="585">
<emphasis id="58A1EA8C3D739B1EB6C82D1EFE710551" bold="true" box="[199,400,1845,1870]" pageId="8" pageNumber="585">Common names.</emphasis>
Luther’s spined loach (English); shchipovka Ljutera [щиповка Лютера] (Russia); Ah-murjum-jool-jong-geh [ỌșäḏḵǭNj] (Korea).
</paragraph>
<paragraph id="6A6A369E3D739B1FB6C82DAAFD8502CB" blockId="8.[151,1436,1521,2018]" lastBlockId="9.[151,1436,151,213]" lastPageId="9" lastPageNumber="586" pageId="8" pageNumber="585">
<emphasis id="58A1EA8C3D739B1EB6C82DAAFE390585" bold="true" box="[199,472,1921,1946]" pageId="8" pageNumber="585">Comparative remarks.</emphasis>
M a l e s o f 
<taxonomicName id="ADD54D1D3D739B1EB4732DA9FD080586" box="[636,745,1921,1945]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="585" phylum="Chordata" rank="species" species="lutheri">
<emphasis id="58A1EA8C3D739B1EB4732DA9FD080586" box="[636,745,1921,1945]" italics="true" pageId="8" pageNumber="585">C. lutheri</emphasis>
</taxonomicName>
are easy to distinguish from males of sympatric in the 
<collectingRegion id="A811F87C3D739B1EB3542DA9FA7D0585" box="[1371,1436,1922,1946]" country="Russia" name="Amur" pageId="8" pageNumber="585">Amur</collectingRegion>
River drainage, 
<taxonomicName id="ADD54D1D3D739B1EB7432D8DFE1D05A2" box="[332,508,1957,1981]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="585" phylum="Chordata" rank="species" species="melanoleuca">
<emphasis id="58A1EA8C3D739B1EB7432D8DFE1D05A2" box="[332,508,1957,1981]" italics="true" pageId="8" pageNumber="585">C. melanoleuca</emphasis>
</taxonomicName>
and 
<taxonomicName id="ADD54D1D3D739B1EB43B2D8DFCFB05A1" authority="Kim &amp; Son" authorityName="Kim &amp; Son" box="[564,794,1957,1982]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="585" phylum="Chordata" rank="species" species="choii">
<emphasis id="58A1EA8C3D739B1EB43B2D8DFD6C05A2" box="[564,653,1957,1981]" italics="true" pageId="8" pageNumber="585">C. choii</emphasis>
Kim &amp; Son
</taxonomicName>
, by the shape of 
<emphasis id="58A1EA8C3D739B1EB5D52D8EFB7A05A2" box="[986,1179,1957,1981]" italics="true" pageId="8" pageNumber="585">lamina circularis</emphasis>
: 
<taxonomicName id="ADD54D1D3D739B1EB2A52D8DFABA05A2" box="[1194,1371,1957,1981]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="585" phylum="Chordata" rank="species" species="melanoleuca">
<emphasis id="58A1EA8C3D739B1EB2A52D8DFABA05A2" box="[1194,1371,1957,1981]" italics="true" pageId="8" pageNumber="585">C. melanoleuca</emphasis>
</taxonomicName>
has a narrow bottle-shaped plate, whereas 
<taxonomicName id="ADD54D1D3D739B1EB4382DE1FD7005FE" box="[567,657,1993,2017]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="585" phylum="Chordata" rank="species" species="choii">
<emphasis id="58A1EA8C3D739B1EB4382DE1FD7005FE" box="[567,657,1993,2017]" italics="true" pageId="8" pageNumber="585">C. choii</emphasis>
</taxonomicName>
has a very narrow knife-shaped plate with serrated outer edge. Both males and females of 
<taxonomicName id="ADD54D1D3D729B1FB79E2AB3FDA302AF" box="[401,578,152,176]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="586" phylum="Chordata" rank="species" species="melanoleuca">
<emphasis id="58A1EA8C3D729B1FB79E2AB3FDA302AF" box="[401,578,152,176]" italics="true" pageId="9" pageNumber="586">C. melanoleuca</emphasis>
</taxonomicName>
and 
<taxonomicName id="ADD54D1D3D729B1FB4732AB3FD3702AF" box="[636,726,152,176]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="586" phylum="Chordata" rank="species" species="choii">
<emphasis id="58A1EA8C3D729B1FB4732AB3FD3702AF" box="[636,726,152,176]" italics="true" pageId="9" pageNumber="586">C. choii</emphasis>
</taxonomicName>
lack the suborbital stripe and additional stripes on the opercle that are diagnostic features for 
<taxonomicName id="ADD54D1D3D729B1FB7FD2A97FDBF02CB" box="[498,606,188,212]" class="Actinopterygii" family="Cobitidae" genus="Cobitis" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="586" phylum="Chordata" rank="species" species="lutheri">
<emphasis id="58A1EA8C3D729B1FB7FD2A97FDBF02CB" box="[498,606,188,212]" italics="true" pageId="9" pageNumber="586">C. lutheri</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>