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<document id="FB0C5D177258291DA72119691B9F5475" ID-CLB-Dataset="45350" ID-DOI="10.5281/zenodo.210988" ID-GBIF-Dataset="e0c02d22-8f7e-4893-82bf-3d2ed3b48c4d" ID-ISSN="1175-5326" ID-Zenodo-Dep="210988" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460446097030" checkinUser="plazi" docAuthor="Pham, Nhi Thi, Broad, Gavin R., Matsumoto, Rikio &amp; Böhme, Wolfgang" docDate="2012" docId="8F5F87F4FF80187A5CACFB96FF17F62A" docLanguage="en" docName="zt03207p053.pdf" docOrigin="Zootaxa 3207" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Acrodactyla shawi Pham, Broad, Matsumoto &amp; Bohme, sp. nov." docType="treatment" docVersion="10" lastPageNumber="52" masterDocId="7366FF8CFF8B18765C3BFFA9FFD7F53E" masterDocTitle="First record of the genus Acrodactyla Haliday (Hymenoptera: Ichneumonidae: Pimplinae) from Vietnam, with descriptions of six new species" masterLastPageNumber="53" masterPageNumber="40" pageNumber="51" updateTime="1698333383792" updateUser="ExternalLinkService">
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<mods:title id="4071B7DB984CCFEC0D08083B73A1069B">First record of the genus Acrodactyla Haliday (Hymenoptera: Ichneumonidae: Pimplinae) from Vietnam, with descriptions of six new species</mods:title>
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<mods:namePart id="546B4D8B713ED6B5118DB299D480E420">Pham, Nhi Thi</mods:namePart>
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<mods:namePart id="48ADD69AF7FD7204B584354CEF91E86E">Broad, Gavin R.</mods:namePart>
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<mods:namePart id="7D024AF293C9C5FD5916BB58811B105D">Matsumoto, Rikio</mods:namePart>
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<mods:namePart id="9BB469C3B5FC184B16D2D32A29099C5D">Böhme, Wolfgang</mods:namePart>
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<mods:date id="D4B40007F0E4F97F7B4C74AF15775B52">2012</mods:date>
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<treatment id="8F5F87F4FF80187A5CACFB96FF17F62A" ID-DOI="http://doi.org/10.5281/zenodo.6171025" ID-GBIF-Taxon="119509446" ID-Zenodo-Dep="6171025" LSID="urn:lsid:plazi:treatment:8F5F87F4FF80187A5CACFB96FF17F62A" httpUri="http://treatment.plazi.org/id/8F5F87F4FF80187A5CACFB96FF17F62A" lastPageId="12" lastPageNumber="52" pageId="11" pageNumber="51">
<subSubSection id="4FEC6569FF80187D5CACFB96FE15F144" pageId="11" pageNumber="51" type="nomenclature">
<paragraph id="074936E2FF80187D5CACFB96FC7DF167" blockId="11.[151,938,1087,1146]" box="[151,938,1087,1113]" pageId="11" pageNumber="51">
<heading id="5C01818EFF80187D5CACFB96FC7DF167" bold="true" box="[151,938,1087,1113]" fontSize="11" level="1" pageId="11" pageNumber="51" reason="1">
<emphasis id="3582EAF0FF80187D5CACFB96FC7DF167" bold="true" box="[151,938,1087,1113]" pageId="11" pageNumber="51">
<taxonomicName id="C0F64D61FF80187D5CACFB96FC97F167" authority="Pham, Broad, Matsumoto &amp; Bohme" authorityName="Pham, Broad, Matsumoto &amp; Bohme" box="[151,832,1087,1113]" class="Insecta" family="Ichneumonidae" genus="Acrodactyla" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="51" phylum="Arthropoda" rank="species" species="shawi" status="sp. nov.">
<emphasis id="3582EAF0FF80187D5CACFB96FEA2F167" bold="true" box="[151,373,1087,1113]" italics="true" pageId="11" pageNumber="51">Acrodactyla shawi</emphasis>
Pham, Broad, Matsumoto &amp; Böhme
</taxonomicName>
,
<taxonomicNameLabel id="2EB1578BFF80187D5F76FB96FC7DF167" box="[845,938,1087,1113]" pageId="11" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="074936E2FF80187D5CACFBCBFE15F144" blockId="11.[151,938,1087,1146]" box="[151,450,1122,1146]" pageId="11" pageNumber="51">
(
<figureCitation id="9FCD2A67FF80187D5CA4FBCBFF35F144" box="[159,226,1122,1146]" captionStart="FIGURE 1" captionStartId="2.[151,250,1738,1761]" captionTargetBox="[151,1436,195,1700]" captionTargetId="figure@2.[151,1436,195,1715]" captionTargetPageId="2" captionText="FIGURE 1. Lateral views of Acrodactyla species (scales: 1 mm): a. A. elongata sp. nov.; b. A. lachryma sp. nov.; c. A. tami sp. nov.; d. A. maiphuquyi sp. nov.; e. A. phuthoensis sp. nov.; f. A. shawi sp. nov." httpUri="https://zenodo.org/record/210989/files/figure.png" pageId="11" pageNumber="51">Figs 1</figureCitation>
f, 2e, 3e, 4c, 5e, 6e)
</paragraph>
</subSubSection>
<subSubSection id="4FEC6569FF80187D5CACFB0EFF0FF019" pageId="11" pageNumber="51" type="diagnosis">
<paragraph id="074936E2FF80187D5CACFB0EFF0FF019" blockId="11.[151,1437,1191,2013]" pageId="11" pageNumber="51">
<emphasis id="3582EAF0FF80187D5CACFB0EFEC6F181" bold="true" box="[151,273,1191,1216]" pageId="11" pageNumber="51">Diagnosis.</emphasis>
Body black, except tergites 2 onwards reddish brown; mesoscutum polished, almost without hairs; metapleuron coarsely rugose-punctate; propodeum with additional transverse carinae, superomedia area closed, petiolar divided into three subareas by longitudinal carinae; fore and mid femora normal, without median ventral tooth.
</paragraph>
</subSubSection>
<subSubSection id="4FEC6569FF80187A5CF2FA9BFB74F729" lastPageId="12" lastPageNumber="52" pageId="11" pageNumber="51" type="description">
<paragraph id="074936E2FF80187D5CF2FA9BFB4BF324" blockId="11.[151,1437,1191,2013]" pageId="11" pageNumber="51">
<emphasis id="3582EAF0FF80187D5CF2FA9BFE8DF075" bold="true" box="[201,346,1330,1355]" pageId="11" pageNumber="51">Description.</emphasis>
(
<typeStatus id="D84D8840FF80187D5D53FA9BFE05F074" box="[360,466,1330,1354]" pageId="11" pageNumber="51" type="holotype">Holotype</typeStatus>
male). Body length
<quantity id="C00E9B07FF80187D5E8CFA9BFCD8F074" box="[695,783,1330,1354]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="11" pageNumber="51" unit="mm" value="6.2">6.2 mm</quantity>
, fore wing
<quantity id="C00E9B07FF80187D5FB6FA9BFC31F074" box="[909,998,1330,1354]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="11" pageNumber="51" unit="mm" value="4.7">4.7 mm</quantity>
.
<emphasis id="3582EAF0FF80187D5FCAFA9AFBF9F075" box="[1009,1070,1331,1355]" italics="true" pageId="11" pageNumber="51">Head</emphasis>
. Antenna with 24 flagellomeres, first flagellomere 1.4x length of second; diameter of lateral ocellus 0.7x ocellar-ocular distance; frons polished, with sparse hairs; inner margins of eyes weakly concave above antennal sockets, parallel ventrally; face 0.8x as high as wide, pubescent; clypeus convex, about 0.5x as high as wide, separated from face by weak clypeofacial suture, apical margin round; malar space equal to basal width of mandible; mandible upper tooth slightly longer than lower tooth; palpi formula 5:4; occipital carina complete, meeting hypostomal carina about 1.4x basal mandible width from base of mandible, forming flange ventrally after meeting hypostomal carina.
</paragraph>
<paragraph id="074936E2FF80187D5CF2F98FFD99F2E3" blockId="11.[151,1437,1191,2013]" pageId="11" pageNumber="51">
<taxonomicName id="C0F64D61FF80187D5CF2F98FFE96F303" box="[201,321,1574,1597]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="3582EAF0FF80187D5CF2F98FFE96F303" box="[201,321,1574,1597]" italics="true" pageId="11" pageNumber="51">Mesosoma</emphasis>
</taxonomicName>
. Pronotum polished, with some hairs on posterior corner; mesoscutum polished, impunctate; mesoscutum 1.2x as long as wide at anterior level of tegulae; scutellum strongly convex, pubescent, lateral carina present at base; mesopleuron densely setose except concave area near mesopleural suture bare and polished, epicnemial carina present on lower half of mesopleuron; metapleuron coarsely rugose-punctate, pubescent, submetapleural carina complete, forming small lobe anteriorly; propodeum moderately convex, dorsally striate, lateral longitudinal, lateromedian longitudinal and posterior transverse carinae present, many transverse striations attached to lateromedian and lateral longitudinal carinae, petiolar area foveolate by carinae, pleural area rugose, pubescent. Legs unmodified, fore and mid femora without teeth ventrally; fore femur
<date id="73481022FF80187D585BF8B1FBABF20E" box="[1120,1148,1816,1840]" pageId="11" pageNumber="51">4x</date>
as long as wide; hind leg with femur 4.7x as long as wide, length 0.7x tibia, tibia
<date id="73481022FF80187D5F3DF893FCE7F26C" box="[774,816,1850,1874]" pageId="11" pageNumber="51" value="2010-10">10x</date>
as long as apical width; basitarsus 0.3x tibia, 0.35x tarsus, 1.6x second tarsomere, fifth tarsomere longer than third. Fore wing with vein
<emphasis id="3582EAF0FF80187D5801F8F7FB54F24B" box="[1082,1155,1886,1909]" italics="true" pageId="11" pageNumber="51">Rs&amp;M</emphasis>
basad of
<emphasis id="3582EAF0FF80187D58C9F8F7FAF3F24B" box="[1266,1316,1886,1909]" italics="true" pageId="11" pageNumber="51">cu-a</emphasis>
, vein
<emphasis id="3582EAF0FF80187D5951F8F7FA4CF24B" box="[1386,1435,1886,1909]" italics="true" pageId="11" pageNumber="51">cu-a</emphasis>
strongly inclivous, 2
<emphasis id="3582EAF0FF80187D5D47F828FE67F2A6" box="[380,432,1921,1944]" italics="true" pageId="11" pageNumber="51">rs-m</emphasis>
about 0.8x distance between 2
<emphasis id="3582EAF0FF80187D5F33F828FCECF2A6" box="[776,827,1921,1944]" italics="true" pageId="11" pageNumber="51">rs-m</emphasis>
and 2
<emphasis id="3582EAF0FF80187D5F44F828FC60F2A6" box="[895,951,1921,1944]" italics="true" pageId="11" pageNumber="51">m-cu</emphasis>
, vein
<emphasis id="3582EAF0FF80187D5FC7F829FBCAF2A9" box="[1020,1053,1920,1943]" italics="true" pageId="11" pageNumber="51">Cu</emphasis>
1
<emphasis id="3582EAF0FF80187D5810F828FBEEF2A6" box="[1067,1081,1921,1944]" italics="true" pageId="11" pageNumber="51">a</emphasis>
separated from 1
<emphasis id="3582EAF0FF80187D58C0F828FAE4F2A6" box="[1275,1331,1921,1944]" italics="true" pageId="11" pageNumber="51">m-cu</emphasis>
by 1.25x length of vein
<emphasis id="3582EAF0FF80187D5D03F80AFE8FF284" box="[312,344,1955,1978]" italics="true" pageId="11" pageNumber="51">Cu</emphasis>
1
<emphasis id="3582EAF0FF80187D5D5DF80AFEA3F285" box="[358,372,1955,1979]" italics="true" pageId="11" pageNumber="51">b</emphasis>
; hind wing with vein
<emphasis id="3582EAF0FF80187D5E52F80AFD63F284" box="[617,692,1955,1978]" italics="true" pageId="11" pageNumber="51">M+Cu</emphasis>
strongly curved, first abscissa of vein
<emphasis id="3582EAF0FF80187D585AF80AFB56F284" box="[1121,1153,1955,1978]" italics="true" pageId="11" pageNumber="51">Cu</emphasis>
1 1.3x as long as vein
<emphasis id="3582EAF0FF80187D5943F80DFF72F2E3" italics="true" pageId="11" pageNumber="51">cu- a</emphasis>
, second abscissa of vein
<emphasis id="3582EAF0FF80187D5DFBF86FFE36F2E3" box="[448,481,1990,2013]" italics="true" pageId="11" pageNumber="51">Cu</emphasis>
1 present.
</paragraph>
<paragraph id="074936E2FF87187A5CF2FF30FA4CF422" blockId="12.[151,1436,151,788]" pageId="12" pageNumber="52">
<emphasis id="3582EAF0FF87187A5CF2FF30FEE9F58E" box="[201,318,153,176]" italics="true" pageId="12" pageNumber="52">Metasoma</emphasis>
. Tergites polished, with scattered short hairs; first tergite 1.7x as long as apical width, dorsolateral carina complete, median longitudinal carina extending over oblique groove; second tergite 0.8x length of first tergite, 0.85x as long as apical width; third tergite shorter than second, basal and apical oblique grooves of tergites 23 moderately deep, rhombic areas convex; first sternite rugose, extending to level of spiracle, round convex apically.
</paragraph>
<paragraph id="074936E2FF87187A5CF2FE81FBB4F45A" blockId="12.[151,1436,151,788]" pageId="12" pageNumber="52">
<emphasis id="3582EAF0FF87187A5CF2FE81FEC0F47E" box="[201,279,296,320]" italics="true" pageId="12" pageNumber="52">Colour</emphasis>
. Black, except mandible, palpi, tegula, subalar prominence, legs reddish brown; metasomal tergite 2 onwards reddish except smooth transverse apical bands black; wings yellowish brown.
</paragraph>
<paragraph id="074936E2FF87187A5CF2FEC6FE48F4B9" blockId="12.[151,1436,151,788]" box="[201,415,367,391]" pageId="12" pageNumber="52">Female. Unknown.</paragraph>
<paragraph id="074936E2FF87187A5CF2FE3AFB74F729" blockId="12.[151,1436,151,788]" pageId="12" pageNumber="52">
<emphasis id="3582EAF0FF87187A5CF2FE3AFEB4F495" bold="true" box="[201,355,403,428]" pageId="12" pageNumber="52">Comparison.</emphasis>
This species belongs to the
<taxonomicName id="C0F64D61FF87187A5E96FE3DFC9EF492" box="[685,841,404,428]" class="Insecta" family="Ichneumonidae" genus="Acrodactyla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="52" phylum="Arthropoda" rank="species" species="quadrisculpta">
<emphasis id="3582EAF0FF87187A5E96FE3DFC9EF492" box="[685,841,404,428]" italics="true" pageId="12" pageNumber="52">quadrisculpta</emphasis>
</taxonomicName>
group (
<bibRefCitation id="63674B13FF87187A5F9FFE3DFB15F492" author="Townes" box="[932,1218,404,428]" pageId="12" pageNumber="52" refString="Townes, H. &amp; Townes, M. (1960) Ichneumon-flies of America North of Mexico: 2. Subfamilies Ephialtinae, Xoridinae, Acaenitinae. U. S. National Museum Bulletin, 216 (2), 1 - 676." type="journal article" year="1960">Townes &amp; Townes 1960</bibRefCitation>
). It is similar to
<taxonomicName id="C0F64D61FF87187A59BFFE3DFEBEF4F1" authority="Aubert" authorityName="Aubert" class="Insecta" family="Ichneumonidae" genus="Acrodactyla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="52" phylum="Arthropoda" rank="species" species="carinator">
<emphasis id="3582EAF0FF87187A59BFFE3DFED6F4EE" italics="true" pageId="12" pageNumber="52">A. carinator</emphasis>
(Aubert)
</taxonomicName>
in its petiolar area divided into three sub-areas by carinae. However, the propodeum of the new species has an additional, anterior transverse carina, the metasoma is partly red (all black or dark brown in
<taxonomicName id="C0F64D61FF87187A597DFE75FF03F726" class="Insecta" family="Ichneumonidae" genus="Acrodactyla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="52" phylum="Arthropoda" rank="species" species="carinator">
<emphasis id="3582EAF0FF87187A597DFE75FF03F726" italics="true" pageId="12" pageNumber="52">A. carinator</emphasis>
</taxonomicName>
) and the fore femur is slenderer than in
<taxonomicName id="C0F64D61FF87187A5EADFDA9FCCCF729" box="[662,795,511,536]" class="Insecta" family="Ichneumonidae" genus="Acrodactyla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="52" phylum="Arthropoda" rank="species" species="carinator">A. carinator</taxonomicName>
(strongly swollen in
<taxonomicName id="C0F64D61FF87187A5832FDA9FB45F726" box="[1033,1170,512,536]" class="Insecta" family="Ichneumonidae" genus="Acrodactyla" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="52" phylum="Arthropoda" rank="species" species="carinator">
<emphasis id="3582EAF0FF87187A5832FDA9FB45F726" box="[1033,1170,512,536]" italics="true" pageId="12" pageNumber="52">A. carinator</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="4FEC6569FF87187A5CF2FD8DFD0EF761" pageId="12" pageNumber="52" type="etymology">
<paragraph id="074936E2FF87187A5CF2FD8DFD0EF761" blockId="12.[151,1436,151,788]" pageId="12" pageNumber="52">
<emphasis id="3582EAF0FF87187A5CF2FD8DFE87F705" bold="true" box="[201,336,548,572]" pageId="12" pageNumber="52">Etymology.</emphasis>
This species is named after Mark Shaw, who has reared so many British polysphinctines and substantially improved our knowledge of their biology.
</paragraph>
</subSubSection>
<subSubSection id="4FEC6569FF87187A5CF2FDC2FF1CF7F2" pageId="12" pageNumber="52" type="distribution">
<paragraph id="074936E2FF87187A5CF2FDC2FB3CF7BA" blockId="12.[151,1436,151,788]" box="[201,1259,619,644]" pageId="12" pageNumber="52">
<emphasis id="3582EAF0FF87187A5CF2FDC2FEB5F7BD" bold="true" box="[201,354,619,644]" pageId="12" pageNumber="52">Distribution.</emphasis>
Currently known only from Sa Pa, Lao Cai Province, North of
<collectingCountry id="7FE17672FF87187A5811FDC5FB5EF7BA" box="[1066,1161,620,644]" name="Vietnam" pageId="12" pageNumber="52">Vietnam</collectingCountry>
(
<figureCitation id="9FCD2A67FF87187A58A3FDC5FB0AF7BA" box="[1176,1245,620,644]" captionStart="FIGURE 8" captionStartId="10.[151,250,1972,1995]" captionTargetBox="[188,1385,373,1933]" captionTargetId="figure@10.[151,1436,313,1991]" captionTargetPageId="10" captionText="FIGURE 8. Map showing the type localities of Acrodactyla in Vietnam: (1) A. elongata sp. nov.; (2) A. lachryma sp. nov.; (3) A. maiphuquyi sp. nov.; (4) A. phuthoensis sp. nov.; (5) A. shawi sp. nov.; (6) A. tami sp. nov." httpUri="https://zenodo.org/record/210996/files/figure.png" pageId="12" pageNumber="52">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="074936E2FF87187A5CF2FD26FF1CF7F2" blockId="12.[151,1436,151,788]" pageId="12" pageNumber="52">
<emphasis id="3582EAF0FF87187A5CF2FD26FE46F796" bold="true" box="[201,401,655,680]" pageId="12" pageNumber="52">Ecological notes.</emphasis>
The single specimen was collected in montane evergreen forest at an elevation of
<quantity id="C00E9B07FF87187A5904FD26FA4BF799" box="[1343,1436,655,679]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="52" unit="m" value="1700.0">1,700 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="4FEC6569FF87187A5CF2FD7EFF17F62A" pageId="12" pageNumber="52" type="materials_examined">
<paragraph id="074936E2FF87187A5CF2FD7EFF17F62A" blockId="12.[151,1436,151,788]" pageId="12" pageNumber="52">
<emphasis id="3582EAF0FF87187A5CF2FD7EFE65F7CE" bold="true" box="[201,434,727,752]" pageId="12" pageNumber="52">Material examined.</emphasis>
<typeStatus id="D84D8840FF87187A5D81FD7EFDF0F7D1" box="[442,551,727,751]" pageId="12" pageNumber="52" type="holotype">Holotype</typeStatus>
: 3 (
<collectionCode id="61E7AE27FF87187A5E6EFD7EFD67F7D1" box="[597,688,727,751]" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Sam Noble Oklahoma Museum of Natural History" pageId="12" pageNumber="52">OMNH</collectionCode>
), Lao Cai, Sa Pa,
<quantity id="C00E9B07FF87187A5F42FD7EFC19F7D1" box="[889,974,727,751]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="12" pageNumber="52" unit="m" value="1700.0">1700 m</quantity>
a.s.l,
<date id="73481022FF87187A582AFD7EFB56F7D1" box="[1041,1153,727,751]" pageId="12" pageNumber="52" value="1997-05-30">30.v.1997</date>
, hand net, R. Matsumoto leg.
</paragraph>
</subSubSection>
</treatment>
</document>