415 lines
37 KiB
XML
415 lines
37 KiB
XML
<document ID-DOI="10.37520/fi.2021.022" ID-ISSN="2533-4069" ID-Zenodo-Dep="7167583" approvalRequired="51" approvalRequired_for_taxonomicNames="7" approvalRequired_for_textStreams="44" checkinTime="1665260545767" checkinUser="felipe" docAuthor="Wheeler, Elisabeth A. & Manchester, Steven R." docDate="2021" docId="039787B7FF86FF88FC87DF7FB1F2F8C7" docLanguage="en" docName="FossilImprint.77.2.299-329.pdf" docOrigin="Fossil Imprint 77 (2)" docSource="http://dx.doi.org/10.37520/fi.2021.022" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.2:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="2" docTitle="Trochodendron beckii R. A. SCOTT et E. A. WHEELER 1982" docType="treatment" docVersion="2" lastPageNumber="323" masterDocId="FFAEFFCFFF91FF90FFFED964B118FFA2" masterDocTitle="A Diverse Assemblage Of Late Eocene Woods From Oregon, Western Usa" masterLastPageNumber="329" masterPageNumber="299" pageNumber="322" updateTime="1665273515484" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A Diverse Assemblage Of Late Eocene Woods From Oregon, Western Usa</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wheeler, Elisabeth A.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Manchester, Steven R.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Fossil Imprint</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-02-18</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>77</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>299</mods:start>
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<mods:end>329</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.37520/fi.2021.022</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.37520/fi.2021.022</mods:identifier>
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<mods:identifier type="ISSN">2533-4069</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7167583</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:039787B7FF86FF88FC87DF7FB1F2F8C7" httpUri="http://treatment.plazi.org/id/039787B7FF86FF88FC87DF7FB1F2F8C7" lastPageId="24" lastPageNumber="323" pageId="23" pageNumber="322">
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<subSubSection pageId="23" pageNumber="322" type="nomenclature">
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<paragraph blockId="23.[889,1435,1563,1619]" pageId="23" pageNumber="322">
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<heading centered="true" fontSize="8" level="2" pageId="23" pageNumber="322" reason="2">
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<taxonomicName authority="(HERGERT et H. K. PHINNEY) R. A. SCOTT et E. A. WHEELER, 1982" authorityName="R. A. SCOTT et E. A. WHEELER" authorityYear="1982" baseAuthorityName="HERGERT et H. K. PHINNEY" class="Magnoliopsida" family="Trochodendraceae" genus="Trochodendron" kingdom="Plantae" order="Trochodendrales" pageId="23" pageNumber="322" phylum="Tracheophyta" rank="species" species="beckii">
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<emphasis bold="true" pageId="23" pageNumber="322">
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<emphasis bold="true" box="[889,1132,1563,1587]" italics="true" pageId="23" pageNumber="322">Trochodendron beckii</emphasis>
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(HERGERT et H.K.PHINNEY) R.A.SCOTT et E.A.WHEELER, 1982
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</emphasis>
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</taxonomicName>
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</heading>
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||
</paragraph>
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<paragraph blockId="23.[1080,1244,1627,1651]" box="[1080,1244,1627,1651]" pageId="23" pageNumber="322">
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<figureCitation box="[1080,1244,1627,1651]" captionStart="Text-fig" captionStartId="22.[125,203,1885,1906]" captionTargetBox="[125,1440,144,1869]" captionTargetId="figure-3@22.[125,1449,144,1869]" captionTargetPageId="22" captionText="Text-fig. 11. Acer Post Hammer species 1, UF 279-34456. a, b: Diffuse-porous wood with distinct growth rings, marked by marginal parenchyma, vessels solitary and in short radial multiples, TS. c: Crowded alternate intervessel pits, TLS. d: Simple perforation plates, helical thickenings in vessel elements, TLS. e: Rays 2–3-seriate, gum deposit in vessel element, TLS. Acer Post Hammer species 2. UF279-34466. f: Diffuse-porous wood with distinct growth rings, marked by marginal parenchyma, vessels solitary and in short radial multiples, TS. g: Alternate intervessel pits, helical thickenings in vessel elements, TLS. h: Rays 1–4(–5)-seriate, TLS. i: Crystalliferous strand, multiseriate ray, TLS. Trochodendron beckii, UF 279-24558. j, k: Distinct growth rings, abrupt transition from earlywood to latewood, vesselless, wide raysnoded atgrowth ring boundaries, TS. l:Rays of two distinct sizes, uniseriate and multiseriates>10-seriate, TLS. m. Scalariform intertracheary pits. RLS. Scale bars: 500 µm in j; 200 µm in a, f, k, l; 100 µm in b, h; 50 µm in c, d, e, g, i, m." figureDoi="http://doi.org/10.5281/zenodo.7167605" httpUri="https://zenodo.org/record/7167605/files/figure.png" pageId="23" pageNumber="322">Text-fig. 11j–m</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="24" lastPageNumber="323" pageId="23" pageNumber="322" type="description">
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<paragraph blockId="23.[843,1482,1674,1763]" pageId="23" pageNumber="322">
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<materialsCitation collectionCode="USNM" pageId="23" pageNumber="322" specimenCode="UF279-24558" specimenCount="1">
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M a t e r i a l.
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<specimenCode box="[1053,1203,1674,1698]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="23" pageNumber="322" type="Museum">UF279-24558</specimenCode>
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(estimated minimum diameter of stem
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<quantity box="[1065,1147,1707,1731]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="322" unit="cm" value="15.0">15 cm</quantity>
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),
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<collectionCode box="[1178,1255,1707,1731]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="23" pageNumber="322" type="Museum">USNM</collectionCode>
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</materialsCitation>
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326709 (
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<typeStatus box="[1380,1478,1707,1731]" pageId="23" pageNumber="322">holotype</typeStatus>
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, estimated minimum diameter
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<quantity box="[1162,1232,1739,1763]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="23" pageNumber="322" unit="cm" value="18.0">18 cm</quantity>
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).
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</paragraph>
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<paragraph blockId="23.[843,1482,1786,2100]" pageId="23" pageNumber="322">D e s c r i p t i o n. Growth rings distinct, marked by noded rays and abrupt transition in radial diameters of latewood tracheids to the subsequent ring’s earlywood tracheids. Vessels absent. Transition from earlywood to latewood abrupt. Tangential diameter of earlywood tracheids 29 (7), 15–51 µm; tangential diameter of latewood tracheids 26 (5) µm, 17–42; scalariform intertracheary pitting observed on radial walls; axial parenchyma rare, occasional isolated cells; rays of two distinct sizes: uniseriates and multiseriates up to 15 cells wide; uniseriates numerous, composed</paragraph>
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<caption pageId="24" pageNumber="323" startId="24.[106,162,152,173]">
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<paragraph blockId="24.[106,1459,152,261]" pageId="24" pageNumber="323">
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<emphasis bold="true" pageId="24" pageNumber="323">
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Table 5. Comparison of
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<taxonomicName box="[354,483,152,173]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="24" pageNumber="323" phylum="Tracheophyta" rank="species" species="undetermined">
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<emphasis bold="true" box="[354,401,152,173]" italics="true" pageId="24" pageNumber="323">Acer</emphasis>
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species.
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</taxonomicName>
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Arranged by age, oldest to youngest. VTD = vessel tangential diameter in µm, mean (standard deviation) or minimum-average-maximum; V/mm
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<superScript attach="left" box="[623,630,180,193]" fontSize="5" pageId="24" pageNumber="323">2</superScript>
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= vessels per mm
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<superScript attach="left" box="[813,820,180,193]" fontSize="5" pageId="24" pageNumber="323">2</superScript>
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</emphasis>
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, range or average as a single number; RW = ray width in cell number; MsRH = multiseriate ray height in µm, mean (standard deviation) or minimum-average-maximum; Cry = crystals present in long chambered strands; Eo = Eocene, Mio = Miocene, e = early, m = middle, l = late; OR = Oregon, WA = Washington.
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</paragraph>
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</caption>
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<paragraph pageId="24" pageNumber="323">
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<table box="[115,1414,308,849]" gridcols="8" gridrows="15" pageId="24" pageNumber="323">
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<tr box="[115,1414,308,327]" gridrow="0" pageId="24" pageNumber="323">
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<th box="[115,426,308,327]" gridcol="0" gridrow="0" pageId="24" pageNumber="323">
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<emphasis bold="true" box="[197,358,308,327]" pageId="24" pageNumber="323">Taxon (reference)</emphasis>
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</th>
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<th box="[467,587,308,327]" gridcol="1" gridrow="0" pageId="24" pageNumber="323">
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<emphasis bold="true" box="[480,574,308,327]" pageId="24" pageNumber="323">VTD (µm)</emphasis>
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</th>
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<th box="[628,712,308,327]" gridcol="2" gridrow="0" pageId="24" pageNumber="323">
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<emphasis bold="true" box="[637,704,308,327]" pageId="24" pageNumber="323">V/mm2</emphasis>
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</th>
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<th box="[761,829,308,327]" gridcol="3" gridrow="0" pageId="24" pageNumber="323">
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<emphasis bold="true" box="[777,813,308,327]" pageId="24" pageNumber="323">RW</emphasis>
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</th>
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<th box="[869,1046,308,327]" gridcol="4" gridrow="0" pageId="24" pageNumber="323">
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<emphasis bold="true" box="[903,1012,308,327]" pageId="24" pageNumber="323">MsRH (µm)</emphasis>
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</th>
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<th box="[1092,1132,308,327]" gridcol="5" gridrow="0" pageId="24" pageNumber="323">
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<emphasis bold="true" box="[1095,1131,308,327]" pageId="24" pageNumber="323">Cry</emphasis>
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</th>
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<th box="[1193,1251,308,327]" gridcol="6" gridrow="0" pageId="24" pageNumber="323">
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<emphasis bold="true" box="[1204,1240,308,327]" pageId="24" pageNumber="323">Age</emphasis>
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</th>
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<th box="[1327,1414,308,327]" gridcol="7" gridrow="0" pageId="24" pageNumber="323">
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<emphasis bold="true" box="[1331,1408,308,327]" pageId="24" pageNumber="323">Country</emphasis>
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</th>
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</tr>
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<tr box="[115,1414,343,362]" gridrow="1" pageId="24" pageNumber="323">
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<th box="[115,426,343,362]" gridcol="0" gridrow="1" pageId="24" pageNumber="323">
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<taxonomicName box="[115,302,343,362]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="24" pageNumber="323" phylum="Tracheophyta" rank="species" species="integrifolioxylon">
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<emphasis box="[115,302,343,362]" italics="true" pageId="24" pageNumber="323">Acer integrifolioxylon</emphasis>
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</taxonomicName>
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(3)
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</th>
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<td box="[467,587,343,362]" gridcol="1" gridrow="1" pageId="24" pageNumber="323">48 (5)–50 (7)</td>
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<td box="[628,712,343,362]" gridcol="2" gridrow="1" pageId="24" pageNumber="323">88–97</td>
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<td box="[761,829,343,362]" gridcol="3" gridrow="1" pageId="24" pageNumber="323">1–3(–4)</td>
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<td box="[869,1046,343,362]" gridcol="4" gridrow="1" pageId="24" pageNumber="323">297 (137)–546 (159)</td>
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<td box="[1092,1132,343,362]" gridcol="5" gridrow="1" pageId="24" pageNumber="323">Yes</td>
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<td box="[1193,1251,343,362]" gridcol="6" gridrow="1" pageId="24" pageNumber="323">m Eo</td>
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<td box="[1327,1414,343,362]" gridcol="7" gridrow="1" pageId="24" pageNumber="323">OR, USA</td>
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</tr>
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<tr box="[115,1414,377,397]" gridrow="2" pageId="24" pageNumber="323">
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<th box="[115,426,377,397]" gridcol="0" gridrow="2" pageId="24" pageNumber="323">
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<taxonomicName box="[115,323,377,397]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="24" pageNumber="323" phylum="Tracheophyta" rank="species" species="undefined-1">
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<emphasis box="[115,155,378,397]" italics="true" pageId="24" pageNumber="323">Acer</emphasis>
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Post Hammer sp. 1
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</taxonomicName>
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(this paper)
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</th>
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<td box="[467,587,377,397]" gridcol="1" gridrow="2" pageId="24" pageNumber="323">24-40-63</td>
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<td box="[628,712,377,397]" gridcol="2" gridrow="2" pageId="24" pageNumber="323">50–56</td>
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<td box="[761,829,377,397]" gridcol="3" gridrow="2" pageId="24" pageNumber="323">1–3(–4)</td>
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<td box="[869,1046,377,397]" gridcol="4" gridrow="2" pageId="24" pageNumber="323">238 (69)</td>
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<td box="[1092,1132,377,397]" gridcol="5" gridrow="2" pageId="24" pageNumber="323">No</td>
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<td box="[1193,1251,377,397]" gridcol="6" gridrow="2" pageId="24" pageNumber="323">l Eo</td>
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<td box="[1327,1414,377,397]" gridcol="7" gridrow="2" pageId="24" pageNumber="323">OR, USA</td>
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</tr>
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<tr box="[115,1414,412,431]" gridrow="3" pageId="24" pageNumber="323">
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<th box="[115,426,412,431]" gridcol="0" gridrow="3" pageId="24" pageNumber="323">
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<taxonomicName box="[115,323,412,431]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="24" pageNumber="323" phylum="Tracheophyta" rank="species" species="undefined-2">
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<emphasis box="[115,155,412,431]" italics="true" pageId="24" pageNumber="323">Acer</emphasis>
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Post Hammer sp. 2
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</taxonomicName>
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(this paper)
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</th>
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<td box="[467,587,412,431]" gridcol="1" gridrow="3" pageId="24" pageNumber="323">39-55-73</td>
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<td box="[628,712,412,431]" gridcol="2" gridrow="3" pageId="24" pageNumber="323">64–78</td>
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<td box="[761,829,412,431]" gridcol="3" gridrow="3" pageId="24" pageNumber="323">1–4(–5)</td>
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<td box="[869,1046,412,431]" gridcol="4" gridrow="3" pageId="24" pageNumber="323">512 (206)</td>
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<td box="[1092,1132,412,431]" gridcol="5" gridrow="3" pageId="24" pageNumber="323">Yes</td>
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<td box="[1193,1251,412,431]" gridcol="6" gridrow="3" pageId="24" pageNumber="323">l Eo</td>
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<td box="[1327,1414,412,431]" gridcol="7" gridrow="3" pageId="24" pageNumber="323">OR, USA</td>
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</tr>
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<tr box="[115,1414,447,501]" gridrow="4" pageId="24" pageNumber="323">
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<th box="[115,426,447,501]" gridcol="0" gridrow="4" pageId="24" pageNumber="323">
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<taxonomicName box="[115,271,465,484]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="24" pageNumber="323" phylum="Tracheophyta" rank="species" species="momijiyamense">
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<emphasis box="[115,271,465,484]" italics="true" pageId="24" pageNumber="323">A. momijiyamense</emphasis>
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</taxonomicName>
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(2, 5)
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</th>
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<td box="[467,587,447,501]" gridcol="1" gridrow="4" pageId="24" pageNumber="323">20–75 (range) 24-50-79</td>
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<td box="[628,712,447,501]" gridcol="2" gridrow="4" pageId="24" pageNumber="323">20–70 numerous</td>
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<td box="[761,829,447,501]" gridcol="3" gridrow="4" pageId="24" pageNumber="323">1–4(–5) 1–4</td>
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<td box="[869,1046,447,501]" gridcol="4" gridrow="4" pageId="24" pageNumber="323">50–600(–750)* 47-500-932</td>
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<td box="[1092,1132,447,501]" gridcol="5" gridrow="4" pageId="24" pageNumber="323">Rare No</td>
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<td box="[1193,1251,447,501]" gridcol="6" gridrow="4" pageId="24" pageNumber="323">Ol e Mio</td>
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<td box="[1327,1414,447,501]" gridcol="7" gridrow="4" pageId="24" pageNumber="323">Japan Korea</td>
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</tr>
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<tr box="[115,1414,517,536]" gridrow="5" pageId="24" pageNumber="323">
|
||
<th box="[115,426,517,536]" gridcol="0" gridrow="5" pageId="24" pageNumber="323">
|
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<taxonomicName box="[115,249,517,536]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" isUncertain="true" kingdom="Plantae" order="Sapindales" pageId="24" pageNumber="323" phylum="Tracheophyta" rank="species" species="amoenum">
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<emphasis box="[115,134,517,536]" italics="true" pageId="24" pageNumber="323">A.</emphasis>
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cf.
|
||
<emphasis box="[166,249,517,536]" italics="true" pageId="24" pageNumber="323">amoenum</emphasis>
|
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</taxonomicName>
|
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(1)
|
||
</th>
|
||
<td box="[467,587,517,536]" gridcol="1" gridrow="5" pageId="24" pageNumber="323">up to 75 µm</td>
|
||
<td box="[628,712,517,536]" gridcol="2" gridrow="5" pageId="24" pageNumber="323">?</td>
|
||
<td box="[761,829,517,536]" gridcol="3" gridrow="5" pageId="24" pageNumber="323">1–6(–9)</td>
|
||
<td box="[869,1046,517,536]" gridcol="4" gridrow="5" pageId="24" pageNumber="323">up to 1200</td>
|
||
<td box="[1092,1132,517,536]" gridcol="5" gridrow="5" pageId="24" pageNumber="323">Yes</td>
|
||
<td box="[1193,1251,517,536]" gridcol="6" gridrow="5" pageId="24" pageNumber="323">e Mio</td>
|
||
<td box="[1327,1414,517,536]" gridcol="7" gridrow="5" pageId="24" pageNumber="323">Japan</td>
|
||
</tr>
|
||
<tr box="[115,1414,551,570]" gridrow="6" pageId="24" pageNumber="323">
|
||
<th box="[115,426,551,570]" gridcol="0" gridrow="6" pageId="24" pageNumber="323">
|
||
<taxonomicName box="[115,196,551,570]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="24" pageNumber="323" phylum="Tracheophyta" rank="species" species="undetermined">
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<emphasis box="[115,155,551,570]" italics="true" pageId="24" pageNumber="323">Acer</emphasis>
|
||
spp.
|
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</taxonomicName>
|
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(7)
|
||
</th>
|
||
<td box="[467,587,551,570]" gridcol="1" gridrow="6" pageId="24" pageNumber="323">37</td>
|
||
<td box="[628,712,551,570]" gridcol="2" gridrow="6" pageId="24" pageNumber="323">132</td>
|
||
<td box="[761,829,551,570]" gridcol="3" gridrow="6" pageId="24" pageNumber="323">1–5</td>
|
||
<td box="[869,1046,551,570]" gridcol="4" gridrow="6" pageId="24" pageNumber="323">214</td>
|
||
<td box="[1092,1132,551,570]" gridcol="5" gridrow="6" pageId="24" pageNumber="323">No</td>
|
||
<td box="[1193,1251,551,570]" gridcol="6" gridrow="6" pageId="24" pageNumber="323">e Mio</td>
|
||
<td box="[1327,1414,551,570]" gridcol="7" gridrow="6" pageId="24" pageNumber="323">Turkey</td>
|
||
</tr>
|
||
<tr box="[115,1414,586,605]" gridrow="7" pageId="24" pageNumber="323">
|
||
<th box="[115,426,586,605]" gridcol="0" gridrow="7" pageId="24" pageNumber="323">
|
||
<taxonomicName box="[115,190,586,605]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="24" pageNumber="323" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis box="[115,155,586,605]" italics="true" pageId="24" pageNumber="323">Acer</emphasis>
|
||
spp
|
||
</taxonomicName>
|
||
(6)
|
||
</th>
|
||
<td box="[467,587,586,605]" gridcol="1" gridrow="7" pageId="24" pageNumber="323">55</td>
|
||
<td box="[628,712,586,605]" gridcol="2" gridrow="7" pageId="24" pageNumber="323">206</td>
|
||
<td box="[761,829,586,605]" gridcol="3" gridrow="7" pageId="24" pageNumber="323">1–5</td>
|
||
<td box="[869,1046,586,605]" gridcol="4" gridrow="7" pageId="24" pageNumber="323">220</td>
|
||
<td box="[1092,1132,586,605]" gridcol="5" gridrow="7" pageId="24" pageNumber="323">No</td>
|
||
<td box="[1193,1251,586,605]" gridcol="6" gridrow="7" pageId="24" pageNumber="323">e Mio</td>
|
||
<td box="[1327,1414,586,605]" gridcol="7" gridrow="7" pageId="24" pageNumber="323">Turkey</td>
|
||
</tr>
|
||
<tr box="[115,1414,621,640]" gridrow="8" pageId="24" pageNumber="323">
|
||
<th box="[115,426,621,640]" gridcol="0" gridrow="8" pageId="24" pageNumber="323">
|
||
<taxonomicName box="[115,269,621,640]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="24" pageNumber="323" phylum="Tracheophyta" rank="species" species="minokamoensis">
|
||
<emphasis box="[115,269,621,640]" italics="true" pageId="24" pageNumber="323">A. minokamoensis</emphasis>
|
||
</taxonomicName>
|
||
(5)
|
||
</th>
|
||
<td box="[467,587,621,640]" gridcol="1" gridrow="8" pageId="24" pageNumber="323">24-43-55</td>
|
||
<td box="[628,712,621,640]" gridcol="2" gridrow="8" pageId="24" pageNumber="323">numerous</td>
|
||
<td box="[761,829,621,640]" gridcol="3" gridrow="8" pageId="24" pageNumber="323">1–3</td>
|
||
<td box="[869,1046,621,640]" gridcol="4" gridrow="8" pageId="24" pageNumber="323">79-290-505</td>
|
||
<td box="[1092,1132,621,640]" gridcol="5" gridrow="8" pageId="24" pageNumber="323">No</td>
|
||
<td box="[1193,1251,621,640]" gridcol="6" gridrow="8" pageId="24" pageNumber="323">e Mio</td>
|
||
<td box="[1327,1414,621,640]" gridcol="7" gridrow="8" pageId="24" pageNumber="323">Korea</td>
|
||
</tr>
|
||
<tr box="[115,1414,656,675]" gridrow="9" pageId="24" pageNumber="323">
|
||
<th box="[115,426,656,675]" gridcol="0" gridrow="9" pageId="24" pageNumber="323">
|
||
<taxonomicName box="[115,245,656,675]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="24" pageNumber="323" phylum="Tracheophyta" rank="species" species="pohangensis">
|
||
<emphasis box="[115,245,656,675]" italics="true" pageId="24" pageNumber="323">A. pohangensis</emphasis>
|
||
</taxonomicName>
|
||
(5)
|
||
</th>
|
||
<td box="[467,587,656,675]" gridcol="1" gridrow="9" pageId="24" pageNumber="323">24-39-55</td>
|
||
<td box="[628,712,656,675]" gridcol="2" gridrow="9" pageId="24" pageNumber="323">numerous</td>
|
||
<td box="[761,829,656,675]" gridcol="3" gridrow="9" pageId="24" pageNumber="323">1–4</td>
|
||
<td box="[869,1046,656,675]" gridcol="4" gridrow="9" pageId="24" pageNumber="323">32-600-1185</td>
|
||
<td box="[1092,1132,656,675]" gridcol="5" gridrow="9" pageId="24" pageNumber="323">No</td>
|
||
<td box="[1193,1251,656,675]" gridcol="6" gridrow="9" pageId="24" pageNumber="323">e Mio</td>
|
||
<td box="[1327,1414,656,675]" gridcol="7" gridrow="9" pageId="24" pageNumber="323">Korea</td>
|
||
</tr>
|
||
<tr box="[115,1414,690,710]" gridrow="10" pageId="24" pageNumber="323">
|
||
<th box="[115,426,690,710]" gridcol="0" gridrow="10" pageId="24" pageNumber="323">
|
||
<taxonomicName box="[115,241,691,710]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="24" pageNumber="323" phylum="Tracheophyta" rank="species" species="watarianum">
|
||
<emphasis box="[115,241,691,710]" italics="true" pageId="24" pageNumber="323">A. watarianum</emphasis>
|
||
</taxonomicName>
|
||
(2)
|
||
</th>
|
||
<td box="[467,587,690,710]" gridcol="1" gridrow="10" pageId="24" pageNumber="323">30–90 (range)</td>
|
||
<td box="[628,712,690,710]" gridcol="2" gridrow="10" pageId="24" pageNumber="323">15-35</td>
|
||
<td box="[761,829,690,710]" gridcol="3" gridrow="10" pageId="24" pageNumber="323">1–5(–6)</td>
|
||
<td box="[869,1046,690,710]" gridcol="4" gridrow="10" pageId="24" pageNumber="323">70–460 (range)</td>
|
||
<td box="[1092,1132,690,710]" gridcol="5" gridrow="10" pageId="24" pageNumber="323">Yes</td>
|
||
<td box="[1193,1251,690,710]" gridcol="6" gridrow="10" pageId="24" pageNumber="323">e Mio</td>
|
||
<td box="[1327,1414,690,710]" gridcol="7" gridrow="10" pageId="24" pageNumber="323">Japan</td>
|
||
</tr>
|
||
<tr box="[115,1414,725,744]" gridrow="11" pageId="24" pageNumber="323">
|
||
<th box="[115,426,725,744]" gridcol="0" gridrow="11" pageId="24" pageNumber="323">
|
||
<taxonomicName box="[115,230,725,744]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="24" pageNumber="323" phylum="Tracheophyta" rank="species" species="beckianum">
|
||
<emphasis box="[115,230,725,744]" italics="true" pageId="24" pageNumber="323">A. beckianum</emphasis>
|
||
</taxonomicName>
|
||
(4)
|
||
</th>
|
||
<td box="[467,587,725,744]" gridcol="1" gridrow="11" pageId="24" pageNumber="323">31-60-82</td>
|
||
<td box="[628,712,725,744]" gridcol="2" gridrow="11" pageId="24" pageNumber="323">23–35</td>
|
||
<td box="[761,829,725,744]" gridcol="3" gridrow="11" pageId="24" pageNumber="323">1–3</td>
|
||
<td box="[869,1046,725,744]" gridcol="4" gridrow="11" pageId="24" pageNumber="323">237 (65)</td>
|
||
<td box="[1092,1132,725,744]" gridcol="5" gridrow="11" pageId="24" pageNumber="323">Yes</td>
|
||
<td box="[1193,1251,725,744]" gridcol="6" gridrow="11" pageId="24" pageNumber="323">m Mio</td>
|
||
<td box="[1327,1414,725,744]" gridcol="7" gridrow="11" pageId="24" pageNumber="323">WA, USA</td>
|
||
</tr>
|
||
<tr box="[115,1414,760,779]" gridrow="12" pageId="24" pageNumber="323">
|
||
<th box="[115,426,760,779]" gridcol="0" gridrow="12" pageId="24" pageNumber="323">
|
||
<taxonomicName box="[115,221,760,779]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="24" pageNumber="323" phylum="Tracheophyta" rank="species" species="berkhoffii">
|
||
<emphasis box="[115,221,760,779]" italics="true" pageId="24" pageNumber="323">A. berkhoffii</emphasis>
|
||
</taxonomicName>
|
||
(4)
|
||
</th>
|
||
<td box="[467,587,760,779]" gridcol="1" gridrow="12" pageId="24" pageNumber="323">35-65-90</td>
|
||
<td box="[628,712,760,779]" gridcol="2" gridrow="12" pageId="24" pageNumber="323">17–24</td>
|
||
<td box="[761,829,760,779]" gridcol="3" gridrow="12" pageId="24" pageNumber="323">1–3</td>
|
||
<td box="[869,1046,760,779]" gridcol="4" gridrow="12" pageId="24" pageNumber="323">365 (135)</td>
|
||
<td box="[1092,1132,760,779]" gridcol="5" gridrow="12" pageId="24" pageNumber="323">No</td>
|
||
<td box="[1193,1251,760,779]" gridcol="6" gridrow="12" pageId="24" pageNumber="323">m Mio</td>
|
||
<td box="[1327,1414,760,779]" gridcol="7" gridrow="12" pageId="24" pageNumber="323">WA, USA</td>
|
||
</tr>
|
||
<tr box="[115,1414,795,814]" gridrow="13" pageId="24" pageNumber="323">
|
||
<th box="[115,426,795,814]" gridcol="0" gridrow="13" pageId="24" pageNumber="323">
|
||
<emphasis box="[115,217,795,814]" italics="true" pageId="24" pageNumber="323">
|
||
<taxonomicName box="[115,204,795,814]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="24" pageNumber="323" phylum="Tracheophyta" rank="species" species="olearyii">A. olearyii</taxonomicName>
|
||
(
|
||
</emphasis>
|
||
4)
|
||
</th>
|
||
<td box="[467,587,795,814]" gridcol="1" gridrow="13" pageId="24" pageNumber="323">30-59-80</td>
|
||
<td box="[628,712,795,814]" gridcol="2" gridrow="13" pageId="24" pageNumber="323">52–71</td>
|
||
<td box="[761,829,795,814]" gridcol="3" gridrow="13" pageId="24" pageNumber="323">1–7</td>
|
||
<td box="[869,1046,795,814]" gridcol="4" gridrow="13" pageId="24" pageNumber="323">437 (239)</td>
|
||
<td box="[1092,1132,795,814]" gridcol="5" gridrow="13" pageId="24" pageNumber="323">No</td>
|
||
<td box="[1193,1251,795,814]" gridcol="6" gridrow="13" pageId="24" pageNumber="323">m Mio</td>
|
||
<td box="[1327,1414,795,814]" gridcol="7" gridrow="13" pageId="24" pageNumber="323">WA, USA</td>
|
||
</tr>
|
||
<tr box="[115,1414,829,849]" gridrow="14" pageId="24" pageNumber="323">
|
||
<th box="[115,426,829,849]" gridcol="0" gridrow="14" pageId="24" pageNumber="323">
|
||
<taxonomicName box="[115,232,830,849]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="24" pageNumber="323" phylum="Tracheophyta" rank="species" species="puratanum">
|
||
<emphasis box="[115,232,830,849]" italics="true" pageId="24" pageNumber="323">A. puratanum</emphasis>
|
||
</taxonomicName>
|
||
(4)
|
||
</th>
|
||
<td box="[467,587,829,849]" gridcol="1" gridrow="14" pageId="24" pageNumber="323">30-53-72</td>
|
||
<td box="[628,712,829,849]" gridcol="2" gridrow="14" pageId="24" pageNumber="323">18–26</td>
|
||
<td box="[761,829,829,849]" gridcol="3" gridrow="14" pageId="24" pageNumber="323">1–4(–5)</td>
|
||
<td box="[869,1046,829,849]" gridcol="4" gridrow="14" pageId="24" pageNumber="323">230 (60)</td>
|
||
<td box="[1092,1132,829,849]" gridcol="5" gridrow="14" pageId="24" pageNumber="323">Yes</td>
|
||
<td box="[1193,1251,829,849]" gridcol="6" gridrow="14" pageId="24" pageNumber="323">m Mio</td>
|
||
<td box="[1327,1414,829,849]" gridcol="7" gridrow="14" pageId="24" pageNumber="323">WA, USA</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph blockId="24.[106,1464,881,952]" pageId="24" pageNumber="323">
|
||
1.
|
||
<bibRefCitation author="Watari, S." box="[127,248,881,900]" pageId="24" pageNumber="323" pagination="97 - 134" refId="ref28139" refString="Watari, S. (1952): Dicotyledonous woods from the Miocene along the Japan-Sea of Honsyu. - Journal of the Faculty of Sciences, University of Tokyo, Sect. III, Botany, 6: 97 - 134." type="journal article" year="1952">Watari (1952)</bibRefCitation>
|
||
, 2.
|
||
<bibRefCitation author="Takahashi, A. & Suzuki, M." box="[272,530,881,900]" pageId="24" pageNumber="323" pagination="474 - 481" refId="ref27815" refString="Takahashi, A., Suzuki, M. (1988): Two new fossil woods of Acer and a new combination of Prunus from the Tertiary of Japan. - Botanical Magazine, Tokyo, 101: 474 - 481. https: // doi. org / 10.1007 / BF 02488089" type="journal article" year="1988">Takahashi and Suzuki (1988)</bibRefCitation>
|
||
, 3.
|
||
<bibRefCitation author="Wheeler, E. A. & Manchester, S. R." box="[553,839,881,900]" pageId="24" pageNumber="323" pagination="1 - 188" refId="ref28521" refString="Wheeler, E. A., Manchester, S. R. (2002): Woods of the Middle Eocene Nut Beds Flora, Clarno Formation, Oregon, USA. - IAWA Journal, Supplement 3: 1 - 188." type="journal article" year="2002">
|
||
Wheeler and
|
||
<collectingRegion box="[674,773,881,900]" country="United Kingdom" name="Manchester" pageId="24" pageNumber="323">Manchester</collectingRegion>
|
||
(2002)
|
||
</bibRefCitation>
|
||
, 4.Wheeler and Dillhoff (2008), 5.
|
||
<bibRefCitation author="Jeong, K. & Kim, K. & Suzuki, M. & Kim, J. W." box="[1150,1315,881,900]" pageId="24" pageNumber="323" pagination="124 - 138" refId="ref24792" refString="Jeong, K., Kim, K., Suzuki, M., Kim, J. W. (2009): Fossil woods from the lower coal-bearing formation of the Janggi Group (Early Miocene) in the Pohang Basin, Korea. - Review of Palaeobotany and Palynology, 153: 124 - 138. https: // doi. org / 10.1016 / j. revpalbo. 2008.07.006" type="journal article" year="2009">Jeong et al. (2009)</bibRefCitation>
|
||
, 6.
|
||
<bibRefCitation author="Bayam, N. N. A. & Akkemik, U. & Poole, I. & Akarsu, F." pageId="24" pageNumber="323" refId="ref22840" refString="Bayam, N. N. A., Akkemik, U., Poole, I., Akarsu, F. (2018): Further contributions to the early Miocene forest vegetation of the Galatian Volcanic Province, Turkey. - Palaeontologica Electronica, 21 (3): 40 (42 pp.). https: // doi. org / 10.26879 / 816" type="book" year="2018">Bayam et al. (2018)</bibRefCitation>
|
||
, 7.
|
||
<bibRefCitation author="Akkemik, U. & Arslan, M. & Poole, I. & Tosun, S. & Kose, N. & Kilic, N. K. & Aydin, A." box="[216,404,907,926]" pageId="24" pageNumber="323" pagination="31 - 51" refId="ref22419" refString="Akkemik, U., Arslan, M., Poole, I., Tosun, S., Kose, N., Kilic, N. K., Aydin, A. (2016): Silicified woods from two previously undescribed early Miocene forest sites near Seben, northwest Turkey. - Review of Palaeobotany and Palynology, 235: 31 - 51." type="journal article" year="2016">Akkemik et al. (2016)</bibRefCitation>
|
||
</paragraph>
|
||
<paragraph blockId="24.[106,1464,881,952]" box="[106,604,933,952]" pageId="24" pageNumber="323">* 50–600 reported as range, with 750 µm given as outlier.</paragraph>
|
||
<paragraph blockId="24.[106,745,1015,1200]" pageId="24" pageNumber="323">
|
||
exclusively of upright cells; central cells of multiseriate rays procumbent, with up to 10 marginal rows of upright cells, heights of multiseriate portions of the rays
|
||
<quantity box="[564,709,1079,1103]" metricMagnitude="-3" metricUnit="m" metricValue="1.13" metricValueMax="1.56" metricValueMin="0.7" pageId="24" pageNumber="323" unit="mm" value="1.13" valueMax="1.56" valueMin="0.7">0.7–1.56 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="24.[106,745,1015,1200]" pageId="24" pageNumber="323">Note: We did not measure total ray height including the uniseriate margins because it was difficult to be sure of where the uniseriate margins ended.</paragraph>
|
||
<paragraph blockId="24.[106,745,1224,1893]" pageId="24" pageNumber="323">
|
||
R e m a r k s. Vesselless woods are rare among angiosperms, and this specimen with its wide heterocellular rays and scalariform bordered pits has features of
|
||
<taxonomicName authorityName="EICHLER" authorityYear="1865" box="[549,745,1288,1312]" class="Magnoliopsida" family="Trochodendraceae" kingdom="Plantae" order="Trochodendrales" pageId="24" pageNumber="323" phylum="Tracheophyta" rank="family">Trochodendraceae</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Bailey, I. W. & Nast, C. G." box="[113,362,1320,1344]" pageId="24" pageNumber="323" pagination="143 - 154" refId="ref22675" refString="Bailey, I. W., Nast, C. G. (1945): Morphology and relationships of Trochodendron and Tetracentron, I. Stem, root, and leaf. - Journal of the Arnold Arboretum, 26 (2): 143 - 154. https: // doi. org / 10.5962 / p. 185529" type="journal article" year="1945">Bailey and Nast 1945</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Hergert, H. & Phinney, H. K." box="[377,676,1320,1345]" pageId="24" pageNumber="323" pagination="118 - 122" refId="ref24458" refString="Hergert, H., Phinney, H. K. (1954): Trochodendroxylon beckii gen. et sp. nov. from the Tertiary of Oregon. - Bulletin of the Torrey Botanical Club, 81: 118 - 122. https: // doi. org / 10.2307 / 2481846" type="journal article" year="1954">Hergert and Phinney 1954</bibRefCitation>
|
||
). This specimen from the Post Hammer locality (UF 279) further documents the occurrence of
|
||
<taxonomicName box="[437,599,1385,1409]" class="Magnoliopsida" family="Trochodendraceae" genus="Trochodendron" kingdom="Plantae" order="Trochodendrales" pageId="24" pageNumber="323" phylum="Tracheophyta" rank="genus">
|
||
<emphasis box="[437,599,1385,1409]" italics="true" pageId="24" pageNumber="323">Trochodendron</emphasis>
|
||
</taxonomicName>
|
||
wood in the Eocene of
|
||
<collectingRegion box="[225,305,1417,1441]" country="United States of America" name="Oregon" pageId="24" pageNumber="323">Oregon</collectingRegion>
|
||
(
|
||
<bibRefCitation author="Scott, R. A. & Wheeler, E. A." box="[321,591,1417,1441]" pageId="24" pageNumber="323" pagination="135 - 154" refId="ref27151" refString="Scott, R. A., Wheeler, E. A. (1982): Fossil woods from the Eocene Clarno Formation of Oregon. - IAWA Bulletin, n. s., 3: 135 - 154. https: // doi. org / 10.1163 / 22941932 - 90000829" type="journal article" year="1982">Scott and Wheeler 1982</bibRefCitation>
|
||
). This species was previously recognized from the Miocene of
|
||
<collectingRegion box="[665,745,1449,1473]" country="United States of America" name="Oregon" pageId="24" pageNumber="323">Oregon</collectingRegion>
|
||
(
|
||
<bibRefCitation author="Hergert, H. & Phinney, H. K." box="[113,426,1481,1506]" pageId="24" pageNumber="323" pagination="118 - 122" refId="ref24458" refString="Hergert, H., Phinney, H. K. (1954): Trochodendroxylon beckii gen. et sp. nov. from the Tertiary of Oregon. - Bulletin of the Torrey Botanical Club, 81: 118 - 122. https: // doi. org / 10.2307 / 2481846" type="journal article" year="1954">Hergert and Phinney 1954</bibRefCitation>
|
||
) from the same formation where fossil trochodendraceous infructescences and leaves have been recognized (
|
||
<bibRefCitation author="Manchester, S. R. & Pigg, K. B. & DeVore, M. L." box="[360,620,1546,1570]" pageId="24" pageNumber="323" pagination="45 - 54" refId="ref25965" refString="Manchester, S. R., Pigg, K. B., DeVore, M. L. (2018): Trochodendraceous fruits and foliage in the Miocene of western North America. - Fossil Imprint, 74: 45 - 54. https: // doi. org / 10.2478 / if- 2018 - 0004" type="journal article" year="2018">
|
||
<collectingRegion box="[360,486,1546,1570]" country="United Kingdom" name="Manchester" pageId="24" pageNumber="323">Manchester</collectingRegion>
|
||
et al. 2018
|
||
</bibRefCitation>
|
||
). There are differences in the quantitative features (tracheid diameters, ray width) between the woods from these separate occurrences, but we don’t consider them to be great enough to treat them as different species. Although the two extant genera of this family,
|
||
<taxonomicName box="[344,506,1707,1731]" class="Magnoliopsida" family="Trochodendraceae" genus="Trochodendron" kingdom="Plantae" order="Trochodendrales" pageId="24" pageNumber="323" phylum="Tracheophyta" rank="genus">
|
||
<emphasis box="[344,506,1707,1731]" italics="true" pageId="24" pageNumber="323">Trochodendron</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[561,697,1708,1731]" class="Magnoliopsida" family="Trochodendraceae" genus="Tetracentron" kingdom="Plantae" order="Trochodendrales" pageId="24" pageNumber="323" phylum="Tracheophyta" rank="genus">
|
||
<emphasis box="[561,697,1708,1731]" italics="true" pageId="24" pageNumber="323">Tetracentron</emphasis>
|
||
</taxonomicName>
|
||
, are confined to eastern Asia, the family is well represented in the Cenozoic of western North America and known from leaves and fruits of both modern and extinct genera ranging from Paleocene to Miocene (
|
||
<bibRefCitation author="Manchester, S. R. & Pigg, K. B. & DeVore, M. L." box="[358,609,1836,1860]" pageId="24" pageNumber="323" pagination="45 - 54" refId="ref25965" refString="Manchester, S. R., Pigg, K. B., DeVore, M. L. (2018): Trochodendraceous fruits and foliage in the Miocene of western North America. - Fossil Imprint, 74: 45 - 54. https: // doi. org / 10.2478 / if- 2018 - 0004" type="journal article" year="2018">
|
||
<collectingRegion box="[358,484,1836,1860]" country="United Kingdom" name="Manchester" pageId="24" pageNumber="323">Manchester</collectingRegion>
|
||
et al. 2018
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Manchester, S. R. & Kvacek, Z. & Judd, W. S." pageId="24" pageNumber="323" pagination="467 - 484" refId="ref25809" refString="Manchester, S. R., Kvacek, Z., Judd, W. S. (2021): Morphology, anatomy, phylogenetics and distribution of fossil and extant Trochodendraceae in the Northern Hemisphere. - Botanical Journal of the Linnean Society, 195 (3): 467 - 484. https: // doi. org / 10.1093 / botlinnean / boaa 046" type="journal article" year="2021">
|
||
<collectingRegion box="[620,744,1836,1860]" country="United Kingdom" name="Manchester" pageId="24" pageNumber="323">Manchester</collectingRegion>
|
||
et al. 2021
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |