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<document id="DF469981ECDF3F15708AB33B9394D49D" ID-DOI="10.11646/zootaxa.3920.1.1" ID-GBIF-Dataset="7289c86d-0ed1-4e93-b7b3-bc596042d854" ID-ISSN="1175-5326" ID-Zenodo-Dep="287886" ID-ZooBank="025F9918-F9F6-4049-B7CD-5AC632094F5E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460994190417" checkinUser="plazi" docAuthor="Kataev, Boris M. &amp; Liang, Hongbin" docDate="2015" docId="039787812E40FFCF94FD4B44ED58E955" docLanguage="en" docName="zt03920p039.pdf" docOrigin="Zootaxa 3920 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpalus (Pseudoophonus) aenigma Tschitscherine 1897" docType="treatment" docVersion="7" lastPageNumber="15" masterDocId="FFAEFFF92E4DFFC1946A4A2BE927EF1A" masterDocTitle="Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)" masterLastPageNumber="39" masterPageNumber="1" pageNumber="14" updateTime="1698562432667" updateUser="plazi">
<mods:mods id="7724EEB95C33985089378007880CFCC8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="323FCC54495A449E7CD1F7006C52549E">
<mods:title id="B4FADC2A1E117E5ACCF1C43E91DE95F4">Taxonomic review of Chinese species of ground beetles of the subgenus Pseudoophonus (genus Harpalus) (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="68B9D448272EBC090F4E7CC10DC99635">Kataev, Boris M.</mods:namePart>
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<mods:namePart id="6C518C9A7D4D41E7D75AE6362E7D7128">Liang, Hongbin</mods:namePart>
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<mods:title id="349908B4A1DBF78BF147AF64C8296ECA">Zootaxa</mods:title>
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<mods:part id="73B4F1B0BF295B1F31733C00C18B694E">
<mods:date id="CDC8662B277C88DC51D20E0B47861189">2015</mods:date>
<mods:detail id="851857C26CB8C6535C924380DA32114B" type="volume">
<mods:number id="DB641F2F4A99FFD85119733CF616E197">3920</mods:number>
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<mods:identifier id="337B9151CFBE3458E7E58C1D28352C20" type="DOI">10.11646/zootaxa.3920.1.1</mods:identifier>
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<treatment id="039787812E40FFCF94FD4B44ED58E955" ID-DOI="http://doi.org/10.5281/zenodo.6114256" ID-GBIF-Taxon="127671663" ID-Zenodo-Dep="6114256" LSID="urn:lsid:plazi:treatment:039787812E40FFCF94FD4B44ED58E955" httpUri="http://treatment.plazi.org/id/039787812E40FFCF94FD4B44ED58E955" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<subSubSection id="C324651C2E40FFCC94FD4B44E800EEB1" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8B8136972E40FFCC94FD4B44EA4CEE90" blockId="13.[151,875,367,427]" box="[151,875,367,394]" pageId="13" pageNumber="14">
<heading id="D0C981FB2E40FFCC94FD4B44EA4CEE90" bold="true" box="[151,875,367,394]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<taxonomicName id="4C3E4D142E40FFCC94FD4B44EA4CEE90" ID-CoL="3JST6" authority="Tschitscherine, 1897" authorityName="Tschitscherine" authorityYear="1897" box="[151,875,367,394]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="aenigma" subGenus="Pseudoophonus">
<emphasis id="B94AEA852E40FFCC94FD4B44EA4CEE90" bold="true" box="[151,875,367,394]" pageId="13" pageNumber="14">
<emphasis id="B94AEA852E40FFCC94FD4B44E82EEE93" bold="true" box="[151,265,367,393]" italics="true" pageId="13" pageNumber="14">Harpalus</emphasis>
(
<emphasis id="B94AEA852E40FFCC95704B44E8FDEE93" bold="true" box="[282,474,367,393]" italics="true" pageId="13" pageNumber="14">Pseudoophonus</emphasis>
)
<emphasis id="B94AEA852E40FFCC95814B44EB74EE93" bold="true" box="[491,595,367,393]" italics="true" pageId="13" pageNumber="14">aenigma</emphasis>
(
<bibRefCitation id="EFAF4B662E40FFCC96084B5BEA45EE90" author="Tschitscherine" box="[610,866,368,394]" pageId="13" pageNumber="14" refString="Tschitscherine, T. (1897) Carabiques nouveaux ou peu connus. L'Abeille, 29, 45 - 75." type="journal article" year="1897">Tschitschérine, 1897</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8136972E40FFCC94FD4BB9E800EEB1" blockId="13.[151,875,367,427]" box="[151,295,402,427]" pageId="13" pageNumber="14">
(
<figureCitation id="13052A122E40FFCC94F54BB9E838EEB1" box="[159,287,402,427]" captionStart="FIGURES 28 – 31" captionStartId="13.[151,263,1817,1840]" captionTargetBox="[337,1254,711,1790]" captionTargetId="figure@13.[325,1262,702,1796]" captionTargetPageId="13" captionText="FIGURES 28 – 31. Harpalus aenigma, median lobe of aedeagus (28 – 29, Koya; 30 – 31, Unzen). 28, 30, dorsal view; 29, 31, view from the left side. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287891/files/figure.png" pageId="13" pageNumber="14">Figs 28–31</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E40FFCC94FD4BF0EC1CED90" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="8B8136972E40FFCC94FD4BF0EAC6EEEA" blockId="13.[151,1436,474,650]" box="[151,993,474,496]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="AB2E11B92E40FFCC94FD4BF0EAC6EEEA" box="[151,993,474,496]" pageId="13" pageNumber="14">
<taxonomicName id="4C3E4D142E40FFCC94FD4BF0EA27EEEA" ID-CoL="8KTZP" authority="Tschitscherine, 1897: 47" authorityName="Tschitscherine" authorityPageNumber="47" authorityYear="1897" box="[151,768,474,496]" class="Insecta" family="Carabidae" genus="Ophonus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="aenigma" subGenus="Migadophonus">
<emphasis id="B94AEA852E40FFCC94FD4BF0E9D4EEEA" box="[151,243,475,496]" italics="true" pageId="13" pageNumber="14">Ophonus</emphasis>
(
<emphasis id="B94AEA852E40FFCC956B4BF0E8BEEEEA" box="[257,409,475,496]" italics="true" pageId="13" pageNumber="14">Migadophonus</emphasis>
)
<emphasis id="B94AEA852E40FFCC95CD4BF0E8D9EEEA" box="[423,510,475,496]" italics="true" pageId="13" pageNumber="14">aenigma</emphasis>
<treatmentCitation id="0A9F10862E40FFCC966F4BF1EA27EEEA" author="Tschitscherine" box="[517,768,474,496]" page="47" pageId="13" pageNumber="14" year="1897">
<bibRefCitation id="EFAF4B662E40FFCC966F4BF1EBFEEEEA" author="Tschitscherine" box="[517,729,474,496]" pageId="13" pageNumber="14" refString="Tschitscherine, T. (1897) Carabiques nouveaux ou peu connus. L'Abeille, 29, 45 - 75." type="journal article" year="1897">Tschitschérine, 1897</bibRefCitation>
: 47
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="548588352E40FFCC97604BF0EA1BEEEA" box="[778,828,475,496]" pageId="13" pageNumber="14">Type</typeStatus>
locality:
<collectingCountry id="F32976072E40FFCC97F14BF0EAFBEEEA" box="[923,988,475,496]" name="South Korea" pageId="13" pageNumber="14">Korea</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8136972E40FFCC94FD4BD2E84EED34" blockId="13.[151,1436,474,650]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="AB2E11B92E40FFCC94FD4BD2E84EED34" pageId="13" pageNumber="14">
<taxonomicName id="4C3E4D142E40FFCC94FD4BD2EA2EED15" ID-CoL="3JTG3" authority="Schauberger, 1933: 67" authorityName="Schauberger" authorityPageNumber="67" authorityYear="1933" box="[151,777,505,527]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="kinfushanicus" subGenus="Pardileus">
<emphasis id="B94AEA852E40FFCC94FD4BD2E9D0ED14" box="[151,247,505,526]" italics="true" pageId="13" pageNumber="14">Harpalus</emphasis>
(
<emphasis id="B94AEA852E40FFCC95614BD2E84BED14" box="[267,364,505,526]" italics="true" pageId="13" pageNumber="14">Pardileus</emphasis>
)
<emphasis id="B94AEA852E40FFCC95EB4BD2EB2BED14" box="[385,524,505,526]" italics="true" pageId="13" pageNumber="14">kinfushanicus</emphasis>
<treatmentCitation id="0A9F10862E40FFCC96724BD2EA2EED15" author="Schauberger" box="[536,777,505,527]" page="67" pageId="13" pageNumber="14" year="1933">
<bibRefCitation id="EFAF4B662E40FFCC96724BD2EBF9ED15" author="Schauberger" box="[536,734,505,527]" pageId="13" pageNumber="14" refString="Schauberger, E. (1933) Zur Kenntnis der palaarktischen Harpalinen (XII. Beitrag). Wiener Entomologische Zeitung, 50 (1 - 2), 64 - 78." type="journal article" year="1933">Schauberger, 1933</bibRefCitation>
: 67
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="548588352E40FFCC97724BD1EA6DED15" box="[792,842,506,527]" pageId="13" pageNumber="14">Type</typeStatus>
locality: &quot;Mts. Kinfushan [=Jinfo Shan] Sung-Kanho&quot;, Sichuan,
<collectingCountry id="F32976072E40FFCC954F4833E842ED34" box="[293,357,536,558]" name="China" pageId="13" pageNumber="14">China</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8136972E40FFCC94FD481CEC3AED56" blockId="13.[151,1436,474,650]" box="[151,1309,566,588]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="AB2E11B92E40FFCC94FD481CEC3AED56" box="[151,1309,566,588]" pageId="13" pageNumber="14">
<taxonomicName id="4C3E4D142E40FFCC94FD481CEBF1ED56" ID-CoL="3JTFT" authority="Jedlicka, 1958: 224" authorityName="Jedlicka" authorityPageNumber="224" authorityYear="1958" box="[151,726,566,588]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="japonicus" subGenus="Pseudophonus">
<emphasis id="B94AEA852E40FFCC94FD481CE9D0ED56" box="[151,247,567,588]" italics="true" pageId="13" pageNumber="14">Harpalus</emphasis>
(
<emphasis id="B94AEA852E40FFCC956F481CE8BEED56" box="[261,409,567,588]" italics="true" pageId="13" pageNumber="14">Pseudophonus</emphasis>
)
<emphasis id="B94AEA852E40FFCC95CD481CEB2EED56" box="[423,521,567,588]" italics="true" pageId="13" pageNumber="14">japonicus</emphasis>
<treatmentCitation id="0A9F10862E40FFCC967A481DEBF1ED56" author="Jedlicka" box="[528,726,566,588]" page="224" pageId="13" pageNumber="14" year="1958">
<bibRefCitation id="EFAF4B662E40FFCC967A481DEB81ED56" author="Jedlicka" box="[528,678,566,588]" pageId="13" pageNumber="14" refString="Jedlicka, A. (1958) Beitrag zur Kenntnis der palaarktischen Carabiden (Coleoptera). Acta Entomologica Musei Nationalis Pragae, 32 (498), 223 - 246." type="journal article" year="1958">Jedlička, 1958</bibRefCitation>
: 224
</treatmentCitation>
</taxonomicName>
(non A. Morawitz, 1862).
<typeStatus id="548588352E40FFCC978D481CED3EED56" box="[999,1049,567,588]" pageId="13" pageNumber="14">Type</typeStatus>
locality: &quot;Kiuriu&quot;,
<collectingCountry id="F32976072E40FFCC90B6481CEC3EED56" box="[1244,1305,567,588]" name="Japan" pageId="13" pageNumber="14">Japan</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8136972E40FFCC94FD487EEC7AED71" blockId="13.[151,1436,474,650]" box="[151,1373,597,619]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="AB2E11B92E40FFCC94FD487EEC7AED71" box="[151,1373,597,619]" pageId="13" pageNumber="14">
<taxonomicName id="4C3E4D142E40FFCC94FD487EEBD6ED71" ID-CoL="3JTCG" authority="Jedlicka, 1958: 225" authorityName="Jedlicka" authorityPageNumber="225" authorityYear="1958" box="[151,753,597,619]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ginfushanus" subGenus="Pseudophonus">
<emphasis id="B94AEA852E40FFCC94FD487EE9D0ED70" box="[151,247,597,618]" italics="true" pageId="13" pageNumber="14">Harpalus</emphasis>
(
<emphasis id="B94AEA852E40FFCC956F487EE8BEED70" box="[261,409,597,618]" italics="true" pageId="13" pageNumber="14">Pseudophonus</emphasis>
)
<emphasis id="B94AEA852E40FFCC95CD487EEB06ED70" box="[423,545,597,618]" italics="true" pageId="13" pageNumber="14">ginfushanus</emphasis>
<treatmentCitation id="0A9F10862E40FFCC9642487EEBD6ED71" author="Jedlicka" box="[552,753,597,619]" page="225" pageId="13" pageNumber="14" year="1958">
<bibRefCitation id="EFAF4B662E40FFCC9642487EEB98ED71" author="Jedlicka" box="[552,703,597,619]" pageId="13" pageNumber="14" refString="Jedlicka, A. (1958) Beitrag zur Kenntnis der palaarktischen Carabiden (Coleoptera). Acta Entomologica Musei Nationalis Pragae, 32 (498), 223 - 246." type="journal article" year="1958">Jedlička, 1958</bibRefCitation>
: 225
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="548588352E40FFCC9691487DEA0AED71" box="[763,813,598,619]" pageId="13" pageNumber="14">Type</typeStatus>
locality: &quot;Ginfu-Shan&quot; [= Jinfo Shan], Sichuan,
<collectingCountry id="F32976072E40FFCC9173487EEC7EED71" box="[1305,1369,597,619]" name="China" pageId="13" pageNumber="14">China</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B8136972E40FFCC94FD485FEC1CED90" blockId="13.[151,1436,474,650]" box="[151,1339,628,650]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="AB2E11B92E40FFCC94FD485FEC1CED90" box="[151,1339,628,650]" pageId="13" pageNumber="14">
<taxonomicName id="4C3E4D142E40FFCC94FD485FEBD2ED90" ID-CoL="6LDYK" authority="Mlynar, 1974: 116" authorityName="Mlynar" authorityPageNumber="116" authorityYear="1974" box="[151,757,628,650]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="kuangcensis" subGenus="Pseudoophonus">
<emphasis id="B94AEA852E40FFCC94FD485FE9D0ED93" box="[151,247,628,649]" italics="true" pageId="13" pageNumber="14">Harpalus</emphasis>
(
<emphasis id="B94AEA852E40FFCC956F485FE882ED93" box="[261,421,628,649]" italics="true" pageId="13" pageNumber="14">Pseudoophonus</emphasis>
)
<emphasis id="B94AEA852E40FFCC95DE485FEB17ED93" box="[436,560,628,649]" italics="true" pageId="13" pageNumber="14">kuangcensis</emphasis>
<treatmentCitation id="0A9F10862E40FFCC965C485FEBD2ED90" author="Mlynar" box="[566,757,628,650]" page="116" pageId="13" pageNumber="14" year="1974">
<bibRefCitation id="EFAF4B662E40FFCC965C485FEBE4ED90" author="Mlynar" box="[566,707,628,650]" pageId="13" pageNumber="14" refString="Mlynar, Z. (1974) Beitrag zur Kenntnis der Ostasiatischen Harpalinae s. str. (Coleoptera, Carabidae). Acta Zoologica Cracoviensia, 19 (6), 105 - 124." type="journal article" year="1974">Mlynář, 1974</bibRefCitation>
: 116
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="548588352E40FFCC9695485EEA16ED90" box="[767,817,629,650]" pageId="13" pageNumber="14">Type</typeStatus>
locality: &quot;Kwangtsen (Kuang-ce)&quot;, Fujiang,
<collectingCountry id="F32976072E40FFCC9092485FEC1FED90" box="[1272,1336,628,650]" name="China" pageId="13" pageNumber="14">China</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF41661F2E40FFCC94FD4D32EB08E854" httpUri="https://zenodo.org/record/287891/files/figure.png" pageId="13" pageNumber="14" targetBox="[337,1254,711,1790]" targetPageId="13">
<paragraph id="8B8136972E40FFCC94FD4D32EB08E854" blockId="13.[151,1435,1817,1870]" pageId="13" pageNumber="14">
<emphasis id="B94AEA852E40FFCC94FD4D32E872E835" bold="true" box="[151,341,1817,1840]" pageId="13" pageNumber="14">FIGURES 28–31.</emphasis>
<taxonomicName id="4C3E4D142E40FFCC95344D31EB3AE82A" box="[350,541,1818,1840]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="aenigma">
<emphasis id="B94AEA852E40FFCC95344D31EB3AE82A" box="[350,541,1818,1840]" italics="true" pageId="13" pageNumber="14">Harpalus aenigma</emphasis>
</taxonomicName>
, median lobe of aedeagus (28–29, Koya; 30–31, Unzen). 28, 30, dorsal view; 29, 31, view from the left side. Scale = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C324651C2E40FFCF94FD4D5EE870EC46" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8B8136972E40FFCC94FD4D5EE890E8A8" blockId="13.[151,1436,1909,2006]" pageId="13" pageNumber="14">
<emphasis id="B94AEA852E40FFCC94FD4D5EE861E894" bold="true" box="[151,326,1909,1934]" pageId="13" pageNumber="14">
<typeStatus id="548588352E40FFCC94FD4D5EE9F5E894" box="[151,210,1909,1934]" pageId="13" pageNumber="14">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="548588352E40FFCC95364D5EE8E3E894" box="[348,452,1909,1934]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C3E4D142E40FFCC95904D5DEBE8E894" box="[506,719,1910,1934]" class="Insecta" family="Carabidae" genus="Ophonus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="aenigma">
<emphasis id="B94AEA852E40FFCC95904D5DEBE8E894" box="[506,719,1910,1934]" italics="true" pageId="13" pageNumber="14">Ophonus aenigma</emphasis>
</taxonomicName>
: ♀, labeled &quot;
<collectingCountry id="F32976072E40FFCC97104D5DEAE6E894" box="[890,961,1910,1934]" name="South Korea" pageId="13" pageNumber="14">Korea</collectingCountry>
(Staud.–Rttr.)&quot;, and &quot;
<taxonomicName id="4C3E4D142E40FFCC90A44D5CEC08E894" box="[1230,1327,1911,1934]" class="Insecta" family="Carabidae" genus="Ophonus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="aenigma">
<emphasis id="B94AEA852E40FFCC90A44D5CEC08E894" box="[1230,1327,1911,1934]" italics="true" pageId="13" pageNumber="14">aenigma</emphasis>
</taxonomicName>
m. Typ. Tschitscherin det.&quot; (
<collectionCode id="ED2FAE522E40FFCC951D4DB1E88CE8A8" box="[375,427,1946,1970]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="13" pageNumber="14">ZIN</collectionCode>
).
</paragraph>
<paragraph id="8B8136972E40FFCF94AD4D96EBB3EFCF" blockId="13.[151,1436,1909,2006]" lastBlockId="14.[151,1437,151,1616]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<typeStatus id="548588352E40FFCC94AD4D96E808E8CC" box="[199,303,1981,2006]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C3E4D142E40FFCC95324D94EB31E8CC" box="[344,534,1982,2006]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="kinfushanicus">
<emphasis id="B94AEA852E40FFCC95324D94EB31E8CC" box="[344,534,1982,2006]" italics="true" pageId="13" pageNumber="14">H. kinfushanicus</emphasis>
</taxonomicName>
: ♀, labeled &quot;Szechwan mer., Mts. Kinfushan,
<quantity id="4CC69B722E40FFCC90414D95EDA3E8CC" box="[1067,1156,1982,2006]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="14" unit="m" value="2000.0">2000 m</quantity>
, pr. fluss. Sung-Kanho&quot;, &quot;
<typeStatus id="548588352E43FFCF94CA4AB3E9F8EFAA" box="[160,223,152,176]" pageId="14" pageNumber="15">Type</typeStatus>
&quot;, &quot;
<taxonomicName id="4C3E4D142E43FFCF956E4AB2EB9EEFAA" authority="Schaub." authorityName="Schaub." box="[260,697,151,176]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="kinfushanicus" subGenus="Pardileus">
<emphasis id="B94AEA852E43FFCF956E4AB2E83FEFAA" box="[260,280,153,176]" italics="true" pageId="14" pageNumber="15">H</emphasis>
. (
<emphasis id="B94AEA852E43FFCF955E4AB3E887EFAA" box="[308,416,152,176]" italics="true" pageId="14" pageNumber="15">Pardileus</emphasis>
)
<emphasis id="B94AEA852E43FFCF95DC4AB3EB77EFAA" box="[438,592,152,176]" italics="true" pageId="14" pageNumber="15">kinfushanicus</emphasis>
Schaub.
</taxonomicName>
, loc. class.&quot;, &quot;coll. Dr. E. Schauberger&quot;, &quot;
<taxonomicName id="4C3E4D142E43FFCF90C64AB3EC31EFAA" box="[1196,1302,152,176]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA852E43FFCF90C64AB3EC31EFAA" box="[1196,1302,152,176]" italics="true" pageId="14" pageNumber="15">Harpalus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3E4D142E43FFCF91434AB3ECB2EFAA" box="[1321,1429,152,176]" class="Insecta" family="Carabidae" genus="Pardileus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA852E43FFCF91434AB3ECB2EFAA" box="[1321,1429,152,176]" italics="true" pageId="14" pageNumber="15">Pardileus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3E4D142E43FFCF94FD4A97E8B1EFCF" authority="Schaub." authorityName="Schaub." box="[151,406,188,213]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="kinfushanicus">
<emphasis id="B94AEA852E43FFCF94FD4A97E816EFCE" box="[151,305,188,212]" italics="true" pageId="14" pageNumber="15">kinfushanicus</emphasis>
Schaub.
</taxonomicName>
&quot;, and &quot;
<typeStatus id="548588352E43FFCF95894A97EB07EFCE" box="[483,544,188,212]" pageId="14" pageNumber="15">Type</typeStatus>
&quot; (
<collectionCode id="ED2FAE522E43FFCF965D4A97EBAFEFCF" box="[567,648,188,213]" pageId="14" pageNumber="15">OÖLL</collectionCode>
).
</paragraph>
<paragraph id="8B8136972E43FFCF94AD4AF4E81FEE5A" blockId="14.[151,1437,151,1616]" pageId="14" pageNumber="15">
<typeStatus id="548588352E43FFCF94AD4AF4E808EFE2" box="[199,303,223,248]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C3E4D142E43FFCF953D4ACAE8CFEFED" box="[343,488,224,248]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B94AEA852E43FFCF953D4ACAE8CFEFED" box="[343,488,224,248]" italics="true" pageId="14" pageNumber="15">H. japonicus</emphasis>
</taxonomicName>
: ♀, labeled &quot;Unzen, Kiuriu, Jap. T. Baum&quot;, &quot;
<typeStatus id="548588352E43FFCF97844ACBED1EEFE2" box="[1006,1081,224,248]" pageId="14" pageNumber="15">Typus</typeStatus>
&quot;, &quot;
<taxonomicName id="4C3E4D142E43FFCF90314ACBEC54EFED" box="[1115,1395,224,248]" class="Insecta" family="Carabidae" genus="Pseudophonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="japonicus" status="sp. nov.">
<emphasis id="B94AEA852E43FFCF90314ACBEC54EFED" box="[1115,1395,224,248]" italics="true" pageId="14" pageNumber="15">Pseudophonus japonicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27957FE2E43FFCF91164ACBE98AEE06" pageId="14" pageNumber="15" rank="species">sp. n.</taxonomicNameLabel>
, det. Ing. Jedlička&quot;, &quot;Mus. Nat. Pragae. Inv. 20719&quot;, &quot;
<taxonomicName id="4C3E4D142E43FFCF97664B2EEC42EE07" authority="Schaubg., Z. Mlynar" authorityName="Schaubg., Z. Mlynar" box="[780,1381,260,285]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pseudophonoides" subGenus="Pardileus">
<emphasis id="B94AEA852E43FFCF97664B2EEA07EE06" box="[780,800,261,284]" italics="true" pageId="14" pageNumber="15">H</emphasis>
. (
<emphasis id="B94AEA852E43FFCF975D4B2FEA84EE06" box="[823,931,260,284]" italics="true" pageId="14" pageNumber="15">Pardileus</emphasis>
)
<emphasis id="B94AEA852E43FFCF97DE4B2FED51EE06" box="[948,1142,260,284]" italics="true" pageId="14" pageNumber="15">pseudophonoides</emphasis>
Schaubg., Z. Mlynář
</taxonomicName>
det., 1973&quot; (
<collectionCode id="ED2FAE522E43FFCF94834B03E80BEE5A" box="[233,300,296,320]" httpUri="http://grbio.org/cool/fx6k-enrh" name="Natal Museum" pageId="14" pageNumber="15">NMP</collectionCode>
).
</paragraph>
<paragraph id="8B8136972E43FFCF94AD4B67ED22EE92" blockId="14.[151,1437,151,1616]" pageId="14" pageNumber="15">
<typeStatus id="548588352E43FFCF94AD4B67E808EE7F" box="[199,303,332,357]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C3E4D142E43FFCF95084B66EB34EE7E" box="[354,531,332,356]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ginfushanus">
<emphasis id="B94AEA852E43FFCF95084B66EB34EE7E" box="[354,531,332,356]" italics="true" pageId="14" pageNumber="15">H. ginfushanus</emphasis>
</taxonomicName>
: ♂, labeled &quot;Ginfu-Shan, Szechuan, Em. Reitter&quot;, &quot;
<typeStatus id="548588352E43FFCF90ED4B67EDF5EE7E" box="[1159,1234,332,356]" pageId="14" pageNumber="15">Typus</typeStatus>
&quot;, &quot;
<taxonomicName id="4C3E4D142E43FFCF90934B67E839EE92" class="Insecta" family="Carabidae" genus="Pseudophonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ginfushanus" status="sp. nov.">
<emphasis id="B94AEA852E43FFCF90934B67E839EE92" italics="true" pageId="14" pageNumber="15">Pseudophonus ginfushanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27957FE2E43FFCF954F4B5BE845EE9D" box="[293,354,368,391]" pageId="14" pageNumber="15" rank="species">sp. n.</taxonomicNameLabel>
, det. Ing. Jedlička&quot;, &quot;Mus. Nat. Pragae, Inv. 20720&quot; (
<collectionCode id="ED2FAE522E43FFCF97DC4B5BEADEEE92" box="[950,1017,368,392]" httpUri="http://grbio.org/cool/fx6k-enrh" name="Natal Museum" pageId="14" pageNumber="15">NMP</collectionCode>
).
</paragraph>
<paragraph id="8B8136972E43FFCF94AD4BBEE9C9EEEE" blockId="14.[151,1437,151,1616]" pageId="14" pageNumber="15">
<typeStatus id="548588352E43FFCF94AD4BBEE80EEEB7" box="[199,297,405,429]" pageId="14" pageNumber="15" type="paratype">Paratype</typeStatus>
of
<taxonomicName id="4C3E4D142E43FFCF953B4BBEE8DAEEB6" box="[337,509,404,428]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="kuangcensis">
<emphasis id="B94AEA852E43FFCF953B4BBEE8DAEEB6" box="[337,509,404,428]" italics="true" pageId="14" pageNumber="15">H. kuangcensis</emphasis>
</taxonomicName>
: ♂, labeled &quot;Kwangtseh-Fukien, J. Klapperich O.,
<date id="FF8010572E43FFCF902E4BBFED9EEEB6" box="[1092,1209,404,428]" pageId="14" pageNumber="15" value="1937-03-10">3.10.1937</date>
&quot;, &quot;
<taxonomicName id="4C3E4D142E43FFCF90B34BBFE9D3EECA" authority="Schaub." authorityName="Schaub." class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pseudophonoides">
<emphasis id="B94AEA852E43FFCF90B34BBFECBCEEB6" box="[1241,1435,404,428]" italics="true" pageId="14" pageNumber="15">pseudophonoides</emphasis>
Schaub.
</taxonomicName>
, det. Ing. Jedlička&quot;, &quot;
<typeStatus id="548588352E43FFCF95964B93EB57EECA" box="[508,624,440,464]" pageId="14" pageNumber="15" type="paratype">Paratypus</typeStatus>
,
<taxonomicName id="4C3E4D142E43FFCF96E84B93ED73EECA" box="[642,1108,440,464]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="kuangcensis" subGenus="Pseudoophonus">
<emphasis id="B94AEA852E43FFCF96E84B93EBCCEECA" box="[642,747,440,464]" italics="true" pageId="14" pageNumber="15">Harpalus</emphasis>
(
<emphasis id="B94AEA852E43FFCF97694B93EA93EECA" box="[771,948,440,464]" italics="true" pageId="14" pageNumber="15">Pseudoophonus</emphasis>
)
<emphasis id="B94AEA852E43FFCF97A14B93ED73EECA" box="[971,1108,440,464]" italics="true" pageId="14" pageNumber="15">kuangcensis</emphasis>
</taxonomicName>
m., Z. Mlynář det., 1973&quot; (
<collectionCode id="ED2FAE522E43FFCF94F54BF7E9C5EEEE" box="[159,226,476,500]" httpUri="http://grbio.org/cool/fx6k-enrh" name="Natal Museum" pageId="14" pageNumber="15">NMP</collectionCode>
).
</paragraph>
<paragraph id="8B8136972E43FFCF94AD4BD4E870EC46" blockId="14.[151,1437,151,1616]" pageId="14" pageNumber="15">
<emphasis id="B94AEA852E43FFCF94AD4BD4EB09ED02" bold="true" box="[199,558,511,536]" pageId="14" pageNumber="15">Additional material examined</emphasis>
.
<specimenCount id="9D38FD1E2E43FFCF96574BD4EBFFED02" box="[573,728,511,536]" pageId="14" pageNumber="15" type="generic">55 specimens</specimenCount>
, including the following specimens from
<collectingCountry id="F32976072E43FFCF90DB4BD4EDD3ED02" box="[1201,1268,511,536]" name="China" pageId="14" pageNumber="15">China</collectingCountry>
[in addition to those listed by
<bibRefCitation id="EFAF4B662E43FFCF9555480EE8C1ED26" author="Kataev" box="[319,486,548,572]" pageId="14" pageNumber="15" refString="Kataev, B. M. (1997) Ground beetles of the genus Harpalus (Insecta, Coleoptera, Carabidae) from East Asia. Steenstrupia, 23, 123 - 160." type="journal article" year="1997">Kataev (1997)</bibRefCitation>
].
<emphasis id="B94AEA852E43FFCF959D480FEB67ED27" bold="true" box="[503,576,548,573]" pageId="14" pageNumber="15">
<collectingCountry id="F32976072E43FFCF959D480FEB67ED27" box="[503,576,548,573]" name="China" pageId="14" pageNumber="15">China</collectingCountry>
</emphasis>
.
<collectionCode id="ED2FAE522E43FFCF9624480EEB84ED26" box="[590,675,549,572]" pageId="14" pageNumber="15">HUBEI</collectionCode>
:
<specimenCount id="9D38FD1E2E43FFCF96D9480FEBF8ED26" box="[691,735,548,572]" pageId="14" pageNumber="15" type="generic">1 ♂</specimenCount>
, Shennongjia,
<quantity id="4CC69B722E43FFCF97EF480FEAFBED26" box="[901,988,548,573]" metricMagnitude="3" metricUnit="m" metricValue="1.64" pageId="14" pageNumber="15" unit="m" value="1640.0">1640 m</quantity>
,
<date id="FF8010572E43FFCF978D480FED54ED26" box="[999,1139,548,572]" pageId="14" pageNumber="15" value="1980-07-24">24.VII.1980</date>
, Yu Peiyu leg. (
<collectionCode id="ED2FAE522E43FFCF9149480FEC70ED26" box="[1315,1367,548,572]" pageId="14" pageNumber="15">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E43FFCF9101480FECB0ED26" box="[1387,1431,548,572]" pageId="14" pageNumber="15" type="generic">1 ♂</specimenCount>
, same data as preceding, but
<date id="FF8010572E43FFCF95B04863EB44ED7A" box="[474,611,584,608]" pageId="14" pageNumber="15" value="1980-07-20">20.VII.1980</date>
(
<collectionCode id="ED2FAE522E43FFCF961E4863EB8FED7A" box="[628,680,584,608]" pageId="14" pageNumber="15">IOZ</collectionCode>
). SICHUAN:
<specimenCount id="9D38FD1E2E43FFCF97234863EA50ED45" box="[841,887,584,608]" pageId="14" pageNumber="15" type="generic">1 ♂</specimenCount>
,
<quantity id="4CC69B722E43FFCF97EE4863EAEAED7A" box="[900,973,583,608]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="15" unit="km" value="70.0">70 km</quantity>
W Chengdu, Qingcheng Hou Shan Mts,
<quantity id="4CC69B722E43FFCF94FD4847E9D7ED9E" box="[151,240,620,645]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="15" unit="m" value="1500.0">1500 m</quantity>
,
<date id="FF8010572E43FFCF94974847E88DED9F" box="[253,426,620,645]" pageId="14" pageNumber="15" value="2005-06-21" valueMax="2005-06-25" valueMin="2005-06-21">21–25.VI.2005</date>
, S. &amp; V. Murzin leg. (
<collectionCode id="ED2FAE522E43FFCF96C14847EBF8ED9E" box="[683,735,620,644]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="14" pageNumber="15">ZIN</collectionCode>
); 1 ♀, &quot;Mou Pin, Thib. or., David&quot; (
<collectionCode id="ED2FAE522E43FFCF90EB4847ED92ED9E" box="[1153,1205,620,644]" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="14" pageNumber="15">ZIN</collectionCode>
);
<specimenCount id="9D38FD1E2E43FFCF90A14847EDDEED9E" box="[1227,1273,620,644]" pageId="14" pageNumber="15" type="generic">1 ♂</specimenCount>
, &quot;Ginfu-Shan, Szechuan, Em. Reitter&quot; (
<collectionCode id="ED2FAE522E43FFCF95D648BBE8D7EDB2" box="[444,496,656,680]" pageId="14" pageNumber="15">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E43FFCF966348BBEB1DEDBD" box="[521,570,656,680]" pageId="14" pageNumber="15" type="generic">1 ♂</specimenCount>
, Chongqing, Dabashan Mt. R., NE Heyu Vill.,
<geoCoordinate id="EE0A50502E43FFCF901C48A4EDDDEDB2" box="[1142,1274,655,680]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="15" value="31.921389">31°55'17&quot;N</geoCoordinate>
<geoCoordinate id="EE0A50502E43FFCF916C48BBECB0EDB2" box="[1286,1431,656,680]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="15" value="109.07694">109°04'37&quot;E</geoCoordinate>
,
<quantity id="4CC69B722E43FFCF94FD489FE812EDD6" box="[151,309,692,717]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.51" metricValueMin="1.7899999999999998" pageId="14" pageNumber="15" unit="m" value="2150.0" valueMax="2510.0" valueMin="1790.0">1790–2510 m</quantity>
,
<date id="FF8010572E43FFCF9529489FE89EEDD7" box="[323,441,692,717]" pageId="14" pageNumber="15" value="2010-05-16">16.V.2010</date>
, I. Belousov, I. Kabak &amp; A. Korolev leg. (cIB&amp;IK). SHAANXI/SICHUAN (border):
<specimenCount id="9D38FD1E2E43FFCF91E4489FE989EDF5" pageId="14" pageNumber="15" type="generic">1 ♂</specimenCount>
, Daba Shan, pass
<quantity id="4CC69B722E43FFCF95EB48F3E8ECEDEA" box="[385,459,727,752]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="15" unit="km" value="20.0">20 km</quantity>
SSE Zhenping
<quantity id="4CC69B722E43FFCF96EB48F3EA07EDF5" box="[641,800,728,752]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="14" pageNumber="15" unit="m" value="1750.0" valueMax="1800.0" valueMin="1700.0">1700–1800 m</quantity>
,
<geoCoordinate id="EE0A50502E43FFCF974448F3EAADEDEA" box="[814,906,728,752]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="31.733334">31°44'N</geoCoordinate>
<geoCoordinate id="EE0A50502E43FFCF97FE48FCEADAEDEA" box="[916,1021,727,752]" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="109.583336">109°35'E</geoCoordinate>
, (small creek vall./young dry mixed forest/field edge),
<date id="FF8010572E43FFCF950E48D7EB2CEC0E" box="[356,523,764,788]" pageId="14" pageNumber="15" value="2001-07-09" valueMax="2001-07-12" valueMin="2001-07-09">9–12.VII.2001</date>
, D.W. Wrase leg. (cWR). JIANGXI: 1 ♀, &quot;Ku-ling [= Guling],
<date id="FF8010572E43FFCF90A548D7EC64EC0F" box="[1231,1347,764,789]" pageId="14" pageNumber="15" value="1935-05-13">13.V.1935</date>
, O. Piel leg.&quot;(
<collectionCode id="ED2FAE522E43FFCF94B84934E82FEC22" box="[210,264,799,824]" pageId="14" pageNumber="15">IOZ</collectionCode>
);
<specimenCount id="9D38FD1E2E43FFCF9575490BE868EC2D" box="[287,335,800,824]" pageId="14" pageNumber="15" type="generic">1 ♂</specimenCount>
, same as preceding, but
<date id="FF8010572E43FFCF96194934EA20EC22" box="[627,775,799,824]" pageId="14" pageNumber="15" value="1935-08-20">20.VIII.1935</date>
&quot; (
<collectionCode id="ED2FAE522E43FFCF9749490BEA70EC22" box="[803,855,800,824]" pageId="14" pageNumber="15">IOZ</collectionCode>
). GUANGXI:
<specimenCount id="9D38FD1E2E43FFCF9069490BED15EC2D" box="[1027,1074,800,824]" pageId="14" pageNumber="15" type="generic">1 ♂</specimenCount>
, unreadable,
<date id="FF8010572E43FFCF90A44934EC7DEC22" box="[1230,1370,799,824]" pageId="14" pageNumber="15" value="1985-07-29">29.VII.1985</date>
, Liao Subai leg. (
<collectionCode id="ED2FAE522E43FFCF957C496FE86CEC46" box="[278,331,836,860]" pageId="14" pageNumber="15">IOZ</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E43FFCF94AD494CEA47EB2E" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="8B8136972E43FFCF94AD494CEA47EB2E" blockId="14.[151,1437,151,1616]" pageId="14" pageNumber="15">
<emphasis id="B94AEA852E43FFCF94AD494CE81DEC9A" bold="true" box="[199,314,871,896]" pageId="14" pageNumber="15">Diagnosis</emphasis>
. This species is similar in external characters to
<taxonomicName id="4C3E4D142E43FFCF97144942EAD5EC9A" box="[894,1010,872,896]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="B94AEA852E43FFCF97144942EAD5EC9A" box="[894,1010,872,896]" italics="true" pageId="14" pageNumber="15">H. rufipes</emphasis>
</taxonomicName>
, but differs from the latter in shiny, clearly iridescent elytra, with microsculpture on disc in baso-medial portion consisting of transverse meshes. Subapical sinuation of elytra somewhat deep (as in Fig. 7). Labial basal palpomere has somewhat straight longitudinal carina underneath. Internal sac of median lobe of aedeagus is with a medial spiny patch on the right side (
<figureCitation id="13052A122E43FFCF94BF49DCE87EEB0A" box="[213,345,1015,1040]" captionStart="FIGURES 28 – 31" captionStartId="13.[151,263,1817,1840]" captionTargetBox="[337,1254,711,1790]" captionTargetId="figure@13.[325,1262,702,1796]" captionTargetPageId="13" captionText="FIGURES 28 – 31. Harpalus aenigma, median lobe of aedeagus (28 – 29, Koya; 30 – 31, Unzen). 28, 30, dorsal view; 29, 31, view from the left side. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287891/files/figure.png" pageId="14" pageNumber="15">Figs 30–31</figureCitation>
), often also with an apical spiny patch on the left side, and rarely with additional medial spiny patch on the left side (
<figureCitation id="13052A122E43FFCF95E54E37EB34EB2E" box="[399,531,1052,1077]" captionStart="FIGURES 28 – 31" captionStartId="13.[151,263,1817,1840]" captionTargetBox="[337,1254,711,1790]" captionTargetId="figure@13.[325,1262,702,1796]" captionTargetPageId="13" captionText="FIGURES 28 – 31. Harpalus aenigma, median lobe of aedeagus (28 – 29, Koya; 30 – 31, Unzen). 28, 30, dorsal view; 29, 31, view from the left side. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287891/files/figure.png" pageId="14" pageNumber="15">Figs 28–29</figureCitation>
). Body length
<quantity id="4CC69B722E43FFCF96DC4E37EA7BEB2E" box="[694,860,1052,1077]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" metricValueMax="1.3599999999999999" metricValueMin="1.02" pageId="14" pageNumber="15" unit="mm" value="11.899999999999999" valueMax="13.6" valueMin="10.2">10.2–13.6 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E43FFCF94AD4E14EA4BEBDF" pageId="14" pageNumber="15" type="distribution">
<paragraph id="8B8136972E43FFCF94AD4E14EC54EB67" blockId="14.[151,1437,151,1616]" pageId="14" pageNumber="15">
<emphasis id="B94AEA852E43FFCF94AD4E14E87EEB42" bold="true" box="[199,345,1087,1112]" pageId="14" pageNumber="15">Distribution</emphasis>
.
<collectingCountry id="F32976072E43FFCF95034E14E896EB42" box="[361,433,1087,1112]" name="China" pageId="14" pageNumber="15">China</collectingCountry>
: the eastern and south-eastern parts. The species was previously known from Zhejiang, Sichuan and Fujian (
<bibRefCitation id="EFAF4B662E43FFCF95144E4FEB76EB66" author="Kataev" box="[382,593,1124,1148]" pageId="14" pageNumber="15" refString="Kataev, B. M., Wrase, D. W. &amp; Ito, N. (2003) Harpalina. In: Lobl, L. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Apollo Books, Stenstrup, pp. 367 - 397." type="book" year="2003">
Kataev
<emphasis id="B94AEA852E43FFCF95BF4E4EEB20EB66" box="[469,519,1124,1148]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. 2003
</bibRefCitation>
). It is recorded here from Hubei, Jiangxi and Guangxi for the first time.
</paragraph>
<paragraph id="8B8136972E43FFCF94AD4EACEA4BEBDF" blockId="14.[151,1437,151,1616]" pageId="14" pageNumber="15">
The species also occurs in North and
<collectingCountry id="F32976072E43FFCF96074EACEBDDEBBA" box="[621,762,1159,1184]" name="South Korea" pageId="14" pageNumber="15">South Korea</collectingCountry>
including Jejudo, and
<collectingCountry id="F32976072E43FFCF97904EA2ED1DEBBA" box="[1018,1082,1161,1184]" name="Japan" pageId="14" pageNumber="15">Japan</collectingCountry>
(Honshu, Kyushu, and Ryukyu Islands) (
<bibRefCitation id="EFAF4B662E43FFCF94944E87E8B6EBDE" author="Kataev" box="[254,401,1196,1220]" pageId="14" pageNumber="15" refString="Kataev, B. M. (1997) Ground beetles of the genus Harpalus (Insecta, Coleoptera, Carabidae) from East Asia. Steenstrupia, 23, 123 - 160." type="journal article" year="1997">Kataev 1997</bibRefCitation>
;
<bibRefCitation id="EFAF4B662E43FFCF95F74E87EB5FEBDF" author="Paik" box="[413,632,1196,1221]" pageId="14" pageNumber="15" refString="Paik, J. - Ch. &amp; Moon, Ch. - S. (2005) Some ground bettles (Coleoptera, Carabidae) from Korea (15). Korean Journal of Soil Zoology, 10 (1 - 2), 42 - 54." type="journal article" year="2005">Paik &amp; Moon 2005</bibRefCitation>
; Moon &amp;
<bibRefCitation id="EFAF4B662E43FFCF96804E87EA47EBDF" author="Paik" box="[746,864,1196,1221]" pageId="14" pageNumber="15" refString="Moon, Ch. - S. &amp; Paik, J. - Ch. (2006) Notes on Harpalini (Coleoptera, Carabidae) from Korea (1 - 3). Korean Journal of Soil Zoology, 11 (1 - 2), 30 - 84." type="journal article" year="2006">Paik 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E43FFCF94AD4EE4EBAAEA2A" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8B8136972E43FFCF94AD4EE4EBAAEA2A" blockId="14.[151,1437,151,1616]" pageId="14" pageNumber="15">
<emphasis id="B94AEA852E43FFCF94AD4EE4E813EBF2" bold="true" box="[199,308,1231,1256]" pageId="14" pageNumber="15">Remarks</emphasis>
. The geographic and individual variation of
<taxonomicName id="4C3E4D142E43FFCF972C4EFAEAE9EBFD" box="[838,974,1232,1256]" class="Insecta" family="Carabidae" genus="Ophonus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="aenigma">
<emphasis id="B94AEA852E43FFCF972C4EFAEAE9EBFD" box="[838,974,1232,1256]" italics="true" pageId="14" pageNumber="15">H. aenigma</emphasis>
</taxonomicName>
was considered by
<bibRefCitation id="EFAF4B662E43FFCF90D14EFAEC47EBF2" author="Kataev" box="[1211,1376,1232,1256]" pageId="14" pageNumber="15" refString="Kataev, B. M. (1997) Ground beetles of the genus Harpalus (Insecta, Coleoptera, Carabidae) from East Asia. Steenstrupia, 23, 123 - 160." type="journal article" year="1997">Kataev (1997)</bibRefCitation>
who synonymized with this species
<taxonomicName id="4C3E4D142E43FFCF959E4EDEEB96EA16" box="[500,689,1268,1292]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="kinfushanicus">
<emphasis id="B94AEA852E43FFCF959E4EDEEB96EA16" box="[500,689,1268,1292]" italics="true" pageId="14" pageNumber="15">H. kinfushanicus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3E4D142E43FFCF96D54EDEEA68EA16" box="[703,847,1269,1292]" class="Insecta" family="Carabidae" genus="Pseudophonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B94AEA852E43FFCF96D54EDEEA68EA16" box="[703,847,1269,1292]" italics="true" pageId="14" pageNumber="15">H. japonicus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3E4D142E43FFCF97374EDEED21EA16" box="[861,1030,1268,1292]" class="Insecta" family="Carabidae" genus="Pseudophonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ginfushanus">
<emphasis id="B94AEA852E43FFCF97374EDEED21EA16" box="[861,1030,1268,1292]" italics="true" pageId="14" pageNumber="15">H. ginfushanus</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C3E4D142E43FFCF902E4EDEEDD7EA16" box="[1092,1264,1268,1292]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="kuangcensis">
<emphasis id="B94AEA852E43FFCF902E4EDEEDD7EA16" box="[1092,1264,1268,1292]" italics="true" pageId="14" pageNumber="15">H. kuangcensis</emphasis>
</taxonomicName>
. The species is rather rare in
<collectingCountry id="F32976072E43FFCF95444F3CE853EA2A" box="[302,372,1303,1328]" name="China" pageId="14" pageNumber="15">China</collectingCountry>
, more common in
<collectingCountry id="F32976072E43FFCF962C4F32EBAEEA2A" box="[582,649,1305,1328]" name="Japan" pageId="14" pageNumber="15">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C324651C2E43FFCF94AD4F17ED58E955" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="8B8136972E43FFCF94AD4F17ED58E955" blockId="14.[151,1437,151,1616]" pageId="14" pageNumber="15">
<taxonomicName id="4C3E4D142E43FFCF94AD4F17E8BBEA4E" ID-CoL="3JST6" box="[199,412,1340,1364]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="aenigma">
<emphasis id="B94AEA852E43FFCF94AD4F17E8BBEA4E" box="[199,412,1340,1364]" italics="true" pageId="14" pageNumber="15">Harpalus aenigma</emphasis>
</taxonomicName>
is most similar morphologically to
<taxonomicName id="4C3E4D142E43FFCF97284F16EC2EEA4E" ID-CoL="6LDTL" authority="Schauberger, 1930" authorityName="Schauberger" authorityYear="1930" box="[834,1289,1340,1365]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pseudophonoides">
<emphasis id="B94AEA852E43FFCF97284F16ED0DEA4E" box="[834,1066,1340,1364]" italics="true" pageId="14" pageNumber="15">H. pseudophonoides</emphasis>
<bibRefCitation id="EFAF4B662E43FFCF905F4F17EC2EEA4E" author="Schauberger" box="[1077,1289,1340,1365]" pageId="14" pageNumber="15" refString="Schauberger, E. (1930) Zur Kenntnis der palaarktischen Harpalinen (VII. Beitrag). Coleopterologisches Centralblatt, 4, 169 - 218." type="journal article" year="1930">Schauberger, 1930</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F32976072E43FFCF913F4F16ECBFEA4E" box="[1365,1432,1341,1364]" name="Japan" pageId="14" pageNumber="15">Japan</collectingCountry>
, which is distinguished from the former species in the following characters: head relatively smaller; pronotum, on average, less strongly narrowed posteriad and less markedly rounded at sides, with basal angles lacking distinct denticles; subapical sinuation of elytra shallower (as in Fig. 8); elytral basal edge more strongly sinuate, meeting lateral margin of elytra usually at almost right angle; median lobe of aedeagus (
<figureCitation id="13052A122E43FFCF907B4FE7EDB5EAFF" box="[1041,1170,1484,1509]" captionStart="FIGURES 24 – 27" captionStartId="12.[151,263,1422,1445]" captionTargetBox="[324,1262,195,1396]" captionTargetId="figure@12.[314,1273,180,1404]" captionTargetPageId="12" captionText="FIGURES 24 – 27. Harpalus, median lobe of aedeagus. 24, 25, H. rufipes (Xinjiang); 26 – 27, H. pseudoophonoides (Osaka). 24, 26, dorsal view; 25, 27, view from the left side. Scale = 1.0 mm." httpUri="https://zenodo.org/record/287890/files/figure.png" pageId="14" pageNumber="15">Figs 26–27</figureCitation>
) stouter, with relatively larger basal bulb; sides of terminal lamella converging more markedly to apex; and internal sac with only left longitudinal spiny patch (
<bibRefCitation id="EFAF4B662E43FFCF95D44C3FEB72E936" author="Kataev" box="[446,597,1556,1580]" pageId="14" pageNumber="15" refString="Kataev, B. M. (1997) Ground beetles of the genus Harpalus (Insecta, Coleoptera, Carabidae) from East Asia. Steenstrupia, 23, 123 - 160." type="journal article" year="1997">Kataev 1997</bibRefCitation>
). All records of
<taxonomicName id="4C3E4D142E43FFCF97724C3EED26E936" ID-CoL="6LDTL" box="[792,1025,1556,1580]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="pseudophonoides">
<emphasis id="B94AEA852E43FFCF97724C3EED26E936" box="[792,1025,1556,1580]" italics="true" pageId="14" pageNumber="15">H. pseudophonoides</emphasis>
</taxonomicName>
from mainland
<collectingCountry id="F32976072E43FFCF90D44C3FEC23E937" box="[1214,1284,1556,1581]" name="China" pageId="14" pageNumber="15">China</collectingCountry>
, for example from Hubei, Fujian, and Sichuan (
<bibRefCitation id="EFAF4B662E43FFCF967F4C13EBAFE94A" author="Hua" box="[533,648,1592,1616]" pageId="14" pageNumber="15" refString="Hua, L. (2002) List of Chinese Insects. Vol. II. Zhongshan (Sun Yat-sen) University Press, Guangzhou, 612 pp." type="book" year="2002">Hua 2002</bibRefCitation>
), should probably be referred to
<emphasis id="B94AEA852E43FFCF979F4C12ED58E955" box="[1013,1151,1592,1616]" italics="true" pageId="14" pageNumber="15">
<taxonomicName id="4C3E4D142E43FFCF979F4C12ED5CE955" ID-CoL="3JST6" box="[1013,1147,1592,1616]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="aenigma">H. aenigma</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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