treatments-xml/data/DA/13/A7/DA13A721FFB12F06CAD2B39EFBFF78C8.xml

<document id="13C6E6FC81E4C2D2439F2E2D991CF6E5" ID-CLB-Dataset="24806" ID-DOI="10.11646/zootaxa.4700.4.3" ID-GBIF-Dataset="406e61f4-d6e6-4e1e-ae43-67f46cfa2a9a" ID-ISSN="1175-5326" ID-Zenodo-Dep="3557570" ID-ZooBank="90BCD9FD-ED89-44B2-A81E-F72A5BE42720" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1575035340439" checkinUser="plazi" docAuthor="Santana, Larissa, Benetti, Cesar João, Clarkson, Bruno &amp; Pes, Ana Maria" docDate="2019" docId="DA13A721FFB12F06CAD2B39EFBFF78C8" docLanguage="en" docName="zootaxa.4700.4.2.pdf" docOrigin="Zootaxa 4700 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Berosus parvus Santana &amp; Benetti &amp; Clarkson &amp; Pes 2019, sp. n." docType="treatment" docVersion="8" lastPageNumber="457" masterDocId="262ADF59FFBA2F0ACA45B243FFC07F17" masterDocTitle="On the genus Berosus Leach (Coleoptera: Hydrophilidae) in the State of Roraima Brazil: description of three new species and new records" masterLastPageNumber="463" masterPageNumber="445" pageNumber="456" updateTime="1698769524145" updateUser="ExternalLinkService">
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<mods:title id="B6408946885D14A7581FEAD45CEAC81C">On the genus Berosus Leach (Coleoptera: Hydrophilidae) in the State of Roraima Brazil: description of three new species and new records</mods:title>
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<mods:namePart id="7DEE8A5FEDA8E315E2FBF7D26EE20662">Santana, Larissa</mods:namePart>
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<mods:namePart id="BF6E47A392351BB3F0166C3727BA7562">Benetti, Cesar João</mods:namePart>
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<mods:namePart id="930150C2D28A1AF3D78FC5CB0FCEABED">Clarkson, Bruno</mods:namePart>
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<mods:namePart id="22A4460B1FB1C9E2FB5A7EE8AC7A1F18">Pes, Ana Maria</mods:namePart>
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<treatment id="DA13A721FFB12F06CAD2B39EFBFF78C8" ID-DOI="http://doi.org/10.5281/zenodo.5942431" ID-GBIF-Taxon="160599077" ID-Zenodo-Dep="5942431" LSID="urn:lsid:plazi:treatment:DA13A721FFB12F06CAD2B39EFBFF78C8" httpUri="http://treatment.plazi.org/id/DA13A721FFB12F06CAD2B39EFBFF78C8" lastPageId="12" lastPageNumber="457" pageId="11" pageNumber="456">
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<paragraph id="52051637FFB12F01CAD2B39EFE5B7EE0" blockId="11.[151,411,477,540]" box="[151,411,477,503]" pageId="11" pageNumber="456">
<heading id="094DA15BFFB12F01CAD2B39EFE5B7EE0" bold="true" box="[151,411,477,503]" fontSize="11" level="1" pageId="11" pageNumber="456" reason="1">
<emphasis id="60CECA25FFB12F01CAD2B39EFE5B7EE0" bold="true" box="[151,411,477,503]" pageId="11" pageNumber="456">
<taxonomicName id="95BA6DB4FFB12F01CAD2B39EFE917EE0" ID-CoL="8XVQZ" authority="Santana &amp; Benetti &amp; Clarkson &amp; Pes, 2019" authorityName="Santana &amp; Benetti &amp; Clarkson &amp; Pes" authorityYear="2019" box="[151,337,477,503]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="456" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="60CECA25FFB12F01CAD2B39EFE917EE0" bold="true" box="[151,337,477,503]" italics="true" pageId="11" pageNumber="456">Berosus parvus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7BFD775EFFB12F01CB1CB39EFE5B7EE0" box="[345,411,477,503]" pageId="11" pageNumber="456" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1AA045BCFFB12F01CAD2B042FEDF7D0B" box="[151,287,513,540]" pageId="11" pageNumber="456" type="description">
<paragraph id="52051637FFB12F01CAD2B042FEDF7D0B" blockId="11.[151,411,477,540]" box="[151,287,513,540]" pageId="11" pageNumber="456">
(
<figureCitation id="CA810AB2FFB12F01CADAB042FF2C7D0B" box="[159,236,513,540]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Figs. 6</figureCitation>
a–h)
</paragraph>
</subSubSection>
<subSubSection id="1AA045BCFFB12F01CAD2B00AFDF67CDC" pageId="11" pageNumber="456" type="materials_examined">
<paragraph id="52051637FFB12F01CAD2B00AFDB17D90" blockId="11.[151,1437,584,2016]" pageId="11" pageNumber="456">
<materialsCitation id="E2D21C6AFFB12F01CAD2B00AFDAD7D90" ID-GBIF-Occurrence="2459985274" country="Brazil" latitude="2.787611" location="Boa Vista" longLatPrecision="1" longitude="-60.777027" pageId="11" pageNumber="456" specimenCount="1" stateProvince="Roraima" typeStatus="holotype">
<emphasis id="60CECA25FFB12F01CAD2B00AFEFC7D74" bold="true" box="[151,316,584,611]" pageId="11" pageNumber="456">
<typeStatus id="8D01A895FFB12F01CAD2B00AFF127D74" box="[151,210,585,611]" pageId="11" pageNumber="456">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="2AAD56A7FFB12F01CB0CB00AFE737D74" box="[329,435,585,611]" name="Brazil" pageId="11" pageNumber="456">BRAZIL</collectingCountry>
: 
<collectingRegion id="907ED8D5FFB12F01CB86B00AFDAC7D74" box="[451,620,585,611]" country="Brazil" name="Roraima" pageId="11" pageNumber="456">Roraima State</collectingRegion>
, 
<location id="576540ECFFB12F01C838B00AFD307D74" LSID="urn:lsid:plazi:treatment:DA13A721FFB12F06CAD2B39EFBFF78C8:576540ECFFB12F01C838B00AFD307D74" box="[637,752,585,611]" country="Brazil" latitude="2.787611" longLatPrecision="1" longitude="-60.777027" name="Boa Vista" pageId="11" pageNumber="456" stateProvince="Roraima">Boa Vista</location>
municipality, pond near the “Estrada do Contorno” road (
<geoCoordinate id="378E70F0FFB12F01CAE5B02FFE887D90" box="[160,328,620,647]" degrees="02" direction="north" minutes="47" orientation="latitude" pageId="11" pageNumber="456" precision="1" seconds="15.4" value="2.787611">02°47’15.4’’ N</geoCoordinate>
/ 
<geoCoordinate id="378E70F0FFB12F01CB1BB02EFDC77D9F" box="[350,519,621,648]" degrees="60" direction="west" minutes="46" orientation="longitude" pageId="11" pageNumber="456" precision="1" seconds="37.3" value="-60.777027">60°46’37.3’’W</geoCoordinate>
) (
<figureCitation id="CA810AB2FFB12F01C85AB02EFDA37D90" box="[543,611,621,647]" captionStart="FIGURES 1–3" captionStartId="1.[151,264,1933,1958]" captionTargetBox="[151,1436,877,1902]" captionTargetId="figure@1.[151,1436,877,1913]" captionTargetPageId="1" captionText="FIGURES 1–3. Habitat and type locality of the Berosus new species: 1) Brazil: Roraima State: Boa Vista County, pond near the RR-205 road, 2) detail of the habitat. 3) Type locality of Berosus andreazzei sp. n. (triangle), Berosus illuviosus sp. n. (square) and Berosus parvus sp. n. (circle)." figureDoi="http://doi.org/10.5281/zenodo.3557572" httpUri="https://zenodo.org/record/3557572/files/figure.png" pageId="11" pageNumber="456">Fig. 3</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="52051637FFB12F01CA82B0D2FDF67CDC" blockId="11.[151,1437,584,2016]" pageId="11" pageNumber="456">
<emphasis id="60CECA25FFB12F01CA82B0D2FEB37DBC" bold="true" box="[199,371,656,683]" pageId="11" pageNumber="456">Type material.</emphasis>
<materialsCitation id="E2D21C6AFFB12F01CB3DB0D2FCC37DE4" ID-GBIF-Occurrence="2459985279" collectingDate="2015-06-01" collectionCode="INPA" collectorName="K. Dias &amp; Dias, C" country="Brazil" county="Boa Vista County" latitude="2.787611" location="Boa Vista County" longLatPrecision="1" longitude="-60.777027" pageId="11" pageNumber="456" specimenCount="2" stateProvince="Roraima" typeStatus="holotype">
<emphasis id="60CECA25FFB12F01CB3DB0D2FE257DBC" bold="true" box="[376,485,657,683]" pageId="11" pageNumber="456">
<typeStatus id="8D01A895FFB12F01CB3DB0D2FE257DBC" box="[376,485,657,683]" pageId="11" pageNumber="456" type="holotype">Holotype</typeStatus>
</emphasis>
male. 
<emphasis id="60CECA25FFB12F01C869B0D3FC9A7DBC" bold="true" box="[556,858,656,683]" pageId="11" pageNumber="456">
<collectingCountry id="2AAD56A7FFB12F01C869B0D3FD5C7DBC" box="[556,668,656,683]" name="Brazil" pageId="11" pageNumber="456">BRAZIL</collectingCountry>
: 
<collectingRegion id="907ED8D5FFB12F01C8E0B0D3FC957DBC" box="[677,853,656,683]" country="Brazil" name="Roraima" pageId="11" pageNumber="456">Roraima State</collectingRegion>
:
</emphasis>
<collectingCounty id="BB646EBBFFB12F01C91AB0D2FBE17DBC" box="[863,1057,657,683]" pageId="11" pageNumber="456">Boa Vista County</collectingCounty>
(
<geoCoordinate id="378E70F0FFB12F01CE6AB0D3FB127DBC" box="[1071,1234,656,683]" degrees="02" direction="north" minutes="47" orientation="latitude" pageId="11" pageNumber="456" precision="1" seconds="15.4" value="2.787611">02°47’15.4’’N</geoCoordinate>
/ 
<geoCoordinate id="378E70F0FFB12F01CEA6B0D2FA507DBB" box="[1251,1424,657,684]" degrees="60" direction="west" minutes="46" orientation="longitude" pageId="11" pageNumber="456" precision="1" seconds="37.3" value="-60.777027">60°46’37.3’’W</geoCoordinate>
), 
<date id="260430F7FFB12F01CAD2B0F7FED27DD8" box="[151,274,692,719]" pageId="11" pageNumber="456" value="2015-06-01">
<collectingDate id="3640C91FFFB12F01CAD2B0F7FED27DD8" box="[151,274,692,719]" pageId="11" pageNumber="456" value="2015-06-01">01.vi.2015</collectingDate>
</date>
, leg. 
<collectorName id="FF4F73E1FFB12F01CB08B0F6FE657DD8" box="[333,421,693,719]" pageId="11" pageNumber="456">K. Dias</collectorName>
, C. Benetti. Condition of 
<typeStatus id="8D01A895FFB12F01C881B0F6FCEA7DD8" box="[708,810,693,719]" pageId="11" pageNumber="456" type="holotype">holotype</typeStatus>
: stored in 80% ethanol with the dissected male genitalia stored in microvials with glycerin, deposited at 
<collectionCode id="34AB8EF2FFB12F01C885B09AFCC37DE4" box="[704,771,729,755]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="11" pageNumber="456">INPA</collectionCode>
</materialsCitation>
. 
<materialsCitation id="E2D21C6AFFB12F01C955B09AFDF27CDC" ID-GBIF-Occurrence="2459985273" collectingDate="2015-06-02" collectingDateMax="2015-06-07" collectingDateMin="2015-06-02" collectionCode="INPA, MZSP, MNRJ, SEMC, DZUP" collectorName="K. Dias &amp; Dias, C" country="Brazil" county="Alto Alegre County" latitude="2.8538055" location="Alto Alegre County" longLatPrecision="1" longitude="-60.84244" municipality="Same" pageId="11" pageNumber="456" specimenCount="6" specimenCount-male="6" stateProvince="Roraima" typeStatus="paratype">
<emphasis id="60CECA25FFB12F01C955B09AFC487DE4" bold="true" box="[784,904,729,755]" pageId="11" pageNumber="456">
<typeStatus id="8D01A895FFB12F01C955B09AFC487DE4" box="[784,904,729,755]" pageId="11" pageNumber="456" type="paratype">Paratypes</typeStatus>
</emphasis>
(6). 
<emphasis id="60CECA25FFB12F01C986B09BFB3A7DE4" bold="true" box="[963,1274,728,755]" pageId="11" pageNumber="456">
<collectingCountry id="2AAD56A7FFB12F01C986B09BFBF37DE4" box="[963,1075,728,755]" name="Brazil" pageId="11" pageNumber="456">BRAZIL</collectingCountry>
: 
<collectingRegion id="907ED8D5FFB12F01CE04B09BFB367DE4" box="[1089,1270,728,755]" country="Brazil" name="Roraima" pageId="11" pageNumber="456">Roraima State</collectingRegion>
:
</emphasis>
<collectingMunicipality id="B2618C4DFFB12F01CF41B09AFA827DE4" box="[1284,1346,729,755]" pageId="11" pageNumber="456">Same</collectingMunicipality>
data as 
<typeStatus id="8D01A895FFB12F01CAD2B0BEFF3C7C00" box="[151,252,765,791]" pageId="11" pageNumber="456" type="holotype">holotype</typeStatus>
, except (
<geoCoordinate id="378E70F0FFB12F01CB1AB0BFFDC37C00" box="[351,515,764,791]" degrees="02" direction="north" minutes="54" orientation="latitude" pageId="11" pageNumber="456" precision="1" seconds="36.5" value="2.910139">02°54’36.5’’N</geoCoordinate>
/ 
<geoCoordinate id="378E70F0FFB12F01C85DB0BFFD047C00" box="[536,708,764,791]" degrees="60" direction="west" minutes="57" orientation="longitude" pageId="11" pageNumber="456" precision="1" seconds="30.9" value="-60.958584">60°57’30.9’’W</geoCoordinate>
), 
<date id="260430F7FFB12F01C89DB0BFFC907C00" box="[728,848,764,791]" pageId="11" pageNumber="456" value="2015-06-03">
<collectingDate id="3640C91FFFB12F01C89DB0BFFC907C00" box="[728,848,764,791]" pageId="11" pageNumber="456" value="2015-06-03">03.vi.2015</collectingDate>
</date>
[
<specimenCount id="44BCDDBEFFB12F01C91BB0BEFC767C00" box="[862,950,765,791]" pageId="11" pageNumber="456" type="male">2 males</specimenCount>
stored in 80% ethanol, deposited at 
<collectionCode id="34AB8EF2FFB12F01CF09B0BFFA517C00" box="[1356,1425,764,791]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="11" pageNumber="456">INPA</collectionCode>
]; same data as holotype, except (
<geoCoordinate id="378E70F0FFB12F01CBB3B163FD5A7C2C" box="[502,666,800,827]" degrees="02" direction="north" minutes="52" orientation="latitude" pageId="11" pageNumber="456" precision="1" seconds="29.7" value="2.8749168">02°52’29.7’’N</geoCoordinate>
/ 
<geoCoordinate id="378E70F0FFB12F01C8F5B163FC9D7C2C" box="[688,861,800,827]" degrees="60" direction="west" minutes="51" orientation="longitude" pageId="11" pageNumber="456" precision="1" seconds="48.9" value="-60.863583">60°51’48.9’’W</geoCoordinate>
), 
<date id="260430F7FFB12F01C934B163FC297C2C" box="[881,1001,800,827]" pageId="11" pageNumber="456" value="2015-06-05">
<collectingDate id="3640C91FFFB12F01C934B163FC297C2C" box="[881,1001,800,827]" pageId="11" pageNumber="456" value="2015-06-05">05.vi.2015</collectingDate>
</date>
[
<specimenCount id="44BCDDBEFFB12F01C9BFB162FB857C2C" box="[1018,1093,801,827]" pageId="11" pageNumber="456" type="male">1 male</specimenCount>
stored in 80% ethanol, deposited at 
<collectionCode id="34AB8EF2FFB12F01CAA0B107FEF47C48" box="[229,308,836,863]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="11" pageNumber="456" type="Museum">MZSP</collectionCode>
]; same data as 
<typeStatus id="8D01A895FFB12F01CB9BB106FD837C48" box="[478,579,837,863]" pageId="11" pageNumber="456" type="holotype">holotype</typeStatus>
, except (
<geoCoordinate id="378E70F0FFB12F01C8EDB107FC8B7C48" box="[680,843,836,863]" degrees="03" direction="north" minutes="10" orientation="latitude" pageId="11" pageNumber="456" precision="1" seconds="53.9" value="3.181639">03°10’53.9’’N</geoCoordinate>
/ 
<geoCoordinate id="378E70F0FFB12F01C927B107FBCD7C48" box="[866,1037,836,863]" degrees="60" direction="west" minutes="50" orientation="longitude" pageId="11" pageNumber="456" precision="1" seconds="00.8" value="-60.833553">60°50’00.8’’W</geoCoordinate>
), 
<date id="260430F7FFB12F01CE66B107FB5B7C48" box="[1059,1179,836,863]" pageId="11" pageNumber="456" value="2015-06-07">
<collectingDate id="3640C91FFFB12F01CE66B107FB5B7C48" box="[1059,1179,836,863]" pageId="11" pageNumber="456" value="2015-06-07">07.vi.2015</collectingDate>
</date>
[
<specimenCount id="44BCDDBEFFB12F01CEE9B106FB377C48" box="[1196,1271,837,863]" pageId="11" pageNumber="456" type="male">1 male</specimenCount>
stored in 80% ethanol, deposited at 
<collectionCode id="34AB8EF2FFB12F01CBCFB12BFE1B7C94" box="[394,475,872,899]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="11" pageNumber="456">MNRJ</collectionCode>
]; 
<collectingCounty id="BB646EBBFFB12F01CBA9B12AFD0C7C94" box="[492,716,873,899]" pageId="11" pageNumber="456">Alto Alegre County</collectingCounty>
(
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/ 
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), 
<date id="260430F7FFB12F01CE1FB12BFB127C94" box="[1114,1234,872,899]" pageId="11" pageNumber="456" value="2015-06-05">
<collectingDate id="3640C91FFFB12F01CE1FB12BFB127C94" box="[1114,1234,872,899]" pageId="11" pageNumber="456" value="2015-06-05">05.vi.2015</collectingDate>
</date>
[
<specimenCount id="44BCDDBEFFB12F01CEA0B12AFAF17C94" box="[1253,1329,873,899]" pageId="11" pageNumber="456" type="male">1 male</specimenCount>
stored in 80% ethanol, deposited at 
<collectionCode id="34AB8EF2FFB12F01CBF8B1CFFDCF7CB0" box="[445,527,908,935]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="11" pageNumber="456" type="Museum">SEMC</collectionCode>
]; 
<location id="576540ECFFB12F01C85AB1CEFD3A7CB0" LSID="urn:lsid:plazi:treatment:DA13A721FFB12F06CAD2B39EFBFF78C8:576540ECFFB12F01C85AB1CEFD3A7CB0" box="[543,762,909,935]" country="Brazil" county="Alto Alegre County" latitude="2.8538055" longLatPrecision="1" longitude="-60.84244" municipality="Same" name="Alto Alegre County" pageId="11" pageNumber="456" stateProvince="Roraima">Alto Alegre County</location>
(
<geoCoordinate id="378E70F0FFB12F01C94CB1CFFC6C7CB0" box="[777,940,908,935]" degrees="02" direction="north" minutes="51" orientation="latitude" pageId="11" pageNumber="456" precision="1" seconds="13.7" value="2.8538055">02°51’13.7’’N</geoCoordinate>
/ 
<geoCoordinate id="378E70F0FFB12F01C985B1CFFBAD7CB0" box="[960,1133,908,935]" degrees="60" direction="west" minutes="50" orientation="longitude" pageId="11" pageNumber="456" precision="1" seconds="32.8" value="-60.84244">60°50’32.8’’W</geoCoordinate>
), 
<date id="260430F7FFB12F01CE3AB1CFFB377CB0" box="[1151,1271,908,935]" pageId="11" pageNumber="456" value="2015-06-02">
<collectingDate id="3640C91FFFB12F01CE3AB1CFFB377CB0" box="[1151,1271,908,935]" pageId="11" pageNumber="456" value="2015-06-02">02.vi.2015</collectingDate>
</date>
[
<specimenCount id="44BCDDBEFFB12F01CF42B1CEFA917CB0" box="[1287,1361,909,935]" pageId="11" pageNumber="456" type="male">1 male</specimenCount>
stored in 80% ethanol, deposited at 
<collectionCode id="34AB8EF2FFB12F01CB99B1F3FDEC7CDC" box="[476,556,944,971]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="11" pageNumber="456" type="University or college">DZUP</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="1AA045BCFFB12F01CA82B197FC357A07" pageId="11" pageNumber="456" type="diagnosis">
<paragraph id="52051637FFB12F01CA82B197FC357A07" blockId="11.[151,1437,584,2016]" pageId="11" pageNumber="456">
<emphasis id="60CECA25FFB12F01CA82B197FE827CF8" bold="true" box="[199,322,980,1007]" pageId="11" pageNumber="456">Diagnosis.</emphasis>
<taxonomicName id="95BA6DB4FFB12F01CB02B195FE337CF8" authority="Santana &amp; Benetti &amp; Clarkson &amp; Pes, 2019" authorityName="Santana &amp; Benetti &amp; Clarkson &amp; Pes" authorityYear="2019" box="[327,499,982,1007]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="456" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="60CECA25FFB12F01CB02B195FE337CF8" box="[327,499,982,1007]" italics="true" pageId="11" pageNumber="456">Berosus parvus</emphasis>
</taxonomicName>
<emphasis id="60CECA25FFB12F01CBBCB196FDF77CF8" bold="true" box="[505,567,981,1007]" pageId="11" pageNumber="456">
<taxonomicNameLabel id="7BFD775EFFB12F01CBBCB196FDF77CF8" box="[505,567,981,1007]" pageId="11" pageNumber="456" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be distinguished from other Neotropical species of 
<taxonomicName id="95BA6DB4FFB12F01CEE5B195FB397CF8" authorityName="Leach" authorityYear="1817" box="[1184,1273,982,1007]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="456" phylum="Arthropoda" rank="genus">
<emphasis id="60CECA25FFB12F01CEE5B195FB397CF8" box="[1184,1273,982,1007]" italics="true" pageId="11" pageNumber="456">Berosus</emphasis>
</taxonomicName>
by the following combination of characteristics: small size (
<quantity id="9542BBD2FFB12F01C8DDB1BBFCFF7B04" box="[664,831,1016,1043]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="11" pageNumber="456" unit="mm" value="2.45" valueMax="2.5" valueMin="2.4">2.40–2.50 mm</quantity>
); clypeus yellow with small dark brown triangle on the middle of posterior margin, frons entirely dark brown or with large dark-brown spot on the median area; pronotum yellow with one discal dark-brown spot and elytra yellow with small dark-brown spots, without metallic luster (
<figureCitation id="CA810AB2FFB12F01CF2AB602FF747B97" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Fig. 6a</figureCitation>
); pronotum with fine, round punctures (
<figureCitation id="CA810AB2FFB12F01C8C5B626FD137B97" box="[640,723,1125,1152]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Fig. 6a</figureCitation>
); elytral, striae well-impressed with round punctures ca. 2× as large as those on the pronotum (
<figureCitation id="CA810AB2FFB12F01CBB2B6CAFD887BB3" box="[503,584,1161,1188]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Fig. 6a</figureCitation>
); mesoventral process laminar with very short curved anterior tooth, directed downwards and backwards, with smooth margins (
<figureCitation id="CA810AB2FFB12F01C88EB6EEFCDD7BDF" box="[715,797,1197,1224]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Fig. 6d</figureCitation>
); abdominal ventrites not crenulate along lateral margins, posterior margins crenulate; first ventrite medially carinate along its whole length; fifth ventrite with apical notch ca. one-quarter the total length, bearing a single short median tooth (
<figureCitation id="CA810AB2FFB12F01C9D3B6B6FC287A07" box="[918,1000,1269,1296]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Fig. 6e</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="1AA045BCFFB12F06CA82B75BFCBF7D0B" lastPageId="12" lastPageNumber="457" pageId="11" pageNumber="456" type="description">
<paragraph id="52051637FFB12F01CA82B75BFCF07A4F" blockId="11.[151,1437,584,2016]" pageId="11" pageNumber="456">
<emphasis id="60CECA25FFB12F01CA82B75BFE997A24" bold="true" box="[199,345,1304,1331]" pageId="11" pageNumber="456">Description.</emphasis>
<emphasis id="60CECA25FFB12F01CB24B75AFE3F7A24" box="[353,511,1305,1331]" italics="true" pageId="11" pageNumber="456">Size and form</emphasis>
. Total length: 
<quantity id="9542BBD2FFB12F01C8E3B75AFC8E7A24" box="[678,846,1305,1331]" metricMagnitude="-3" metricUnit="m" metricValue="2.455" metricValueMax="2.47" metricValueMin="2.44" pageId="11" pageNumber="456" unit="mm" value="2.455" valueMax="2.47" valueMin="2.44">2.44–2.47 mm</quantity>
. Body short, nearly 2.2× longer than wide in dorsal view (
<figureCitation id="CA810AB2FFB12F01CA98B77EFEEE7A4F" box="[221,302,1341,1368]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Fig. 6a</figureCitation>
), moderately convex in lateral view (
<figureCitation id="CA810AB2FFB12F01C88AB77EFCE37A4F" box="[719,803,1341,1368]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Fig. 6b</figureCitation>
).
</paragraph>
<paragraph id="52051637FFB12F01CA82B722FEE47927" blockId="11.[151,1437,584,2016]" pageId="11" pageNumber="456">
<emphasis id="60CECA25FFB12F01CA82B722FEC87A6C" box="[199,264,1377,1403]" italics="true" pageId="11" pageNumber="456">Color</emphasis>
. Labrum slightly yellow; clypeus yellow with small dark-brown triangle on the middle of posterior margin; frons entirely dark brown or with a large dark-brown spot on median area, without metallic luster (
<figureCitation id="CA810AB2FFB12F01CF75B7C6FA4E7AB7" box="[1328,1422,1413,1440]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Figs. 6a</figureCitation>
); maxillary palpi yellow with fourth palpomere dark brown at apex (
<figureCitation id="CA810AB2FFB12F01C9D7B7EAFC207AD3" box="[914,992,1449,1476]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Figs. 6</figureCitation>
a–c); pronotum yellow, with one discal dark-brown spot without metallic luster; scutellar shield slightly darkened (
<figureCitation id="CA810AB2FFB12F01C9ACB78EFBFB7AFF" box="[1001,1083,1485,1512]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Fig. 6a</figureCitation>
); elytra yellow with small dark brown spots (
<figureCitation id="CA810AB2FFB12F01CB6BB7B2FEBF791B" box="[302,383,1521,1548]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Fig. 6a</figureCitation>
); venter of thorax and abdomen dark brown (
<figureCitation id="CA810AB2FFB12F01C933B7B2FC07791B" box="[886,967,1521,1548]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Fig. 6c</figureCitation>
); femora, tibiae and tarsus entirely yellow (
<figureCitation id="CA810AB2FFB12F01CADAB456FF2D7927" box="[159,237,1557,1584]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Figs. 6</figureCitation>
a–c).
</paragraph>
<paragraph id="52051637FFB12F01CA82B47AFDDD798B" blockId="11.[151,1437,584,2016]" pageId="11" pageNumber="456">
<emphasis id="60CECA25FFB12F01CA82B47AFECB7944" box="[199,267,1593,1619]" italics="true" pageId="11" pageNumber="456">Head.</emphasis>
Clypeus and frons sparsely and finely punctate, punctures ca. 2× as large as ommatidia, round in shape (
<figureCitation id="CA810AB2FFB12F01CAE5B41EFF30796F" box="[160,240,1629,1656]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Fig. 6a</figureCitation>
). Frontal carina absent. Eyes slightly prominent (
<figureCitation id="CA810AB2FFB12F01C954B41EFCA2796F" box="[785,866,1629,1656]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Fig. 6a</figureCitation>
). Maxillary palpi short, nearly half as long as width of the head, and thick (
<figureCitation id="CA810AB2FFB12F01CBDDB4C2FE26798B" box="[408,486,1665,1692]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Figs. 6</figureCitation>
a–c).
</paragraph>
<paragraph id="52051637FFB12F06CA82B4E6FC957FEB" blockId="11.[151,1437,584,2016]" lastBlockId="12.[151,1437,153,2015]" lastPageId="12" lastPageNumber="457" pageId="11" pageNumber="456">
<emphasis id="60CECA25FFB12F01CA82B4E6FED879A8" box="[199,280,1701,1727]" italics="true" pageId="11" pageNumber="456">Thorax</emphasis>
. Posterior margin of pronotum about as wide as basal margin of elytra, with fine, round, and moderately dense punctures, the same size as those on head, slightly coarser along sides (
<figureCitation id="CA810AB2FFB12F01CE64B48AFBB579F3" box="[1057,1141,1737,1764]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Fig. 6a</figureCitation>
). Scutellar shield densely punctate with punctures similar in size to those on elytra. Elytral striae well-impressed with round punctures ca. 2× larger than those on pronotum; interstriae flat on elytral disc, ca. 2× as wide as striae, sparsely and finely punctate (punctures smaller than those on pronotum); humeral hump not prominent (
<figureCitation id="CA810AB2FFB12F01C9B4B576FB847847" box="[1009,1092,1845,1872]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Fig. 6a</figureCitation>
); elytral apices rounded (
<figureCitation id="CA810AB2FFB12F01CF21B576FF657863" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Figs. 6</figureCitation>
a–c); spine-like hairs absent (
<figureCitation id="CA810AB2FFB12F01CBAFB51AFDFB7863" box="[490,571,1881,1908]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Fig. 6a</figureCitation>
). Mesoventral process laminar with very short, curved anterior tooth, directed downwards and backwards with smooth margins; posterior angle of the mesoventral process weakly raised in lateral view, not prominent (
<figureCitation id="CA810AB2FFB12F01CBFBB5E2FDD378AB" box="[446,531,1953,1980]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Fig. 6d</figureCitation>
). Metaventral process narrow, carinate before median depression, posterolateral angles produced into small rounded laminae; posterior angle not raised (
<figureCitation id="CA810AB2FFB12F01C988B586FBE078F7" box="[973,1056,1989,2016]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="11" pageNumber="456">Fig. 6c</figureCitation>
). Basal pubescence on two-thirds of mesofemora and three-fourths of metafemora, limit oblique (
<figureCitation id="CA810AB2FFB62F06C92FB2DAFC7D7FA3" box="[874,957,153,180]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">Fig. 6c</figureCitation>
). Protarsus of male with adhesive soles in tarsomeres 1–2, first tarsomere ca. 1.5× longer than second, fourth tarsomere elongate, almost as long as tarsomeres 1–3 combined (
<figureCitation id="CA810AB2FFB62F06CB03B2A2FE7D7FEC" box="[326,445,225,251]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">Figs. 6a, c</figureCitation>
). Claws weakly arched (
<figureCitation id="CA810AB2FFB62F06C895B2A2FCDD7FEB" box="[720,797,225,252]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">Figs. 6</figureCitation>
a–c).
</paragraph>
<paragraph id="52051637FFB62F06CA82B346FCBF7D0B" blockId="12.[151,1437,153,2015]" pageId="12" pageNumber="457">
<emphasis id="60CECA25FFB62F06CA82B346FEF17E08" box="[199,305,261,287]" italics="true" pageId="12" pageNumber="457">Abdomen</emphasis>
. First ventrite medially carinate its whole length, without lateral depressions; abdominal ventrites 2–4 without central carina or tooth-like projection, not crenulate along lateral margins; posterior margins crenulate; fifth ventrite with apical notch ca. one-quarter the total length, bearing a single short median tooth (
<figureCitation id="CA810AB2FFB62F06CE82B30EFADA7E7F" box="[1223,1306,333,360]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">Fig. 6e</figureCitation>
). Aedeagus compressed laterally with basal piece ca. three-quarters of total length, 2.4× longer than its greatest width (
<figureCitation id="CA810AB2FFB62F06CF21B332FF657EA7" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">Figs. 6</figureCitation>
f–h); parameres symmetrical, as long as median lobe in dorsal view, subcylindrical (
<figureCitation id="CA810AB2FFB62F06CE00B3D6FB537EA7" box="[1093,1171,405,432]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">Fig. 6f</figureCitation>
); apical portion of parameres curved towards ventral face, obliquely directed regarding dorsal outline of aedeagus in lateral view, bearing a ventral row of short hairs along subapical portion (
<figureCitation id="CA810AB2FFB62F06C8A3B39EFCA17EEF" box="[742,865,477,504]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">
Figs. 
<quantity id="9542BBD2FFB62F06C963B39EFC847EE0" box="[806,836,477,503]" metricMagnitude="-3" metricUnit="kg" metricValue="6.0" pageId="12" pageNumber="457" unit="g" value="6.0">6g</quantity>
, h
</figureCitation>
); median lobe cylindrical, not swollen subapically, apex with small hole in the median region in dorsal view (
<figureCitation id="CA810AB2FFB62F06C966B042FCB27D0B" box="[803,882,513,540]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">Fig. 6f</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="1AA045BCFFB62F06CA82B066FF257D73" pageId="12" pageNumber="457" type="etymology">
<paragraph id="52051637FFB62F06CA82B066FF257D73" blockId="12.[151,1437,153,2015]" pageId="12" pageNumber="457">
<emphasis id="60CECA25FFB62F06CA82B066FE8E7D28" bold="true" box="[199,334,549,575]" pageId="12" pageNumber="457">Etymology.</emphasis>
The specific epithet, 
<taxonomicName id="95BA6DB4FFB62F06C87EB065FD477D28" authorityName="Santana &amp; Benetti &amp; Clarkson &amp; Pes" authorityYear="2019" box="[571,647,550,575]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="60CECA25FFB62F06C87EB065FD477D28" box="[571,647,550,575]" italics="true" pageId="12" pageNumber="457">parvus</emphasis>
</taxonomicName>
, refers to the very small size of specimens (From Latin “ 
<taxonomicName id="95BA6DB4FFB62F06CEB9B065FA887D28" authorityName="Santana &amp; Benetti &amp; Clarkson &amp; Pes" authorityYear="2019" box="[1276,1352,550,575]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="60CECA25FFB62F06CEB9B065FA887D28" box="[1276,1352,550,575]" italics="true" pageId="12" pageNumber="457">parvus</emphasis>
</taxonomicName>
”, little, small).
</paragraph>
</subSubSection>
<subSubSection id="1AA045BCFFB62F06CA82B02FFDEE7D9F" box="[199,558,620,648]" pageId="12" pageNumber="457" type="distribution">
<paragraph id="52051637FFB62F06CA82B02FFDEE7D9F" blockId="12.[151,1437,153,2015]" box="[199,558,620,648]" pageId="12" pageNumber="457">
<emphasis id="60CECA25FFB62F06CA82B02FFEA17D90" bold="true" box="[199,353,620,647]" pageId="12" pageNumber="457">Distribution.</emphasis>
<collectingCountry id="2AAD56A7FFB62F06CB22B02EFE6B7D90" box="[359,427,621,647]" name="Brazil" pageId="12" pageNumber="457">Brazil</collectingCountry>
(
<collectingRegion id="907ED8D5FFB62F06CBFEB02EFDE27D9F" box="[443,546,621,648]" country="Brazil" name="Roraima" pageId="12" pageNumber="457">Roraima</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="1AA045BCFFB62F06CA82B0D3FBFF78C8" pageId="12" pageNumber="457" type="biology_ecology">
<paragraph id="52051637FFB62F06CA82B0D3FB257DBC" blockId="12.[151,1437,153,2015]" box="[199,1253,656,683]" pageId="12" pageNumber="457">
<emphasis id="60CECA25FFB62F06CA82B0D3FEE77DBC" bold="true" box="[199,295,656,683]" pageId="12" pageNumber="457">Biology.</emphasis>
The specimens were collected in ponds with abundant macrophyte cover (
<figureCitation id="CA810AB2FFB62F06CE2EB0D2FB187DBC" box="[1131,1240,657,683]" captionStart="FIGURES 1–3" captionStartId="1.[151,264,1933,1958]" captionTargetBox="[151,1436,877,1902]" captionTargetId="figure@1.[151,1436,877,1913]" captionTargetPageId="1" captionText="FIGURES 1–3. Habitat and type locality of the Berosus new species: 1) Brazil: Roraima State: Boa Vista County, pond near the RR-205 road, 2) detail of the habitat. 3) Type locality of Berosus andreazzei sp. n. (triangle), Berosus illuviosus sp. n. (square) and Berosus parvus sp. n. (circle)." figureDoi="http://doi.org/10.5281/zenodo.3557572" httpUri="https://zenodo.org/record/3557572/files/figure.png" pageId="12" pageNumber="457">Figs. 1, 2</figureCitation>
).
</paragraph>
<paragraph id="52051637FFB62F06CA82B0F7FBFF78C8" blockId="12.[151,1437,153,2015]" pageId="12" pageNumber="457">
<emphasis id="60CECA25FFB62F06CA82B0F7FE147DD8" bold="true" box="[199,468,692,719]" pageId="12" pageNumber="457">Taxonomic comments.</emphasis>
<taxonomicName id="95BA6DB4FFB62F06CB98B0F5FD8B7DD8" authority="Santana &amp; Benetti &amp; Clarkson &amp; Pes, 2019" authorityName="Santana &amp; Benetti &amp; Clarkson &amp; Pes" authorityYear="2019" box="[477,587,694,719]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="60CECA25FFB62F06CB98B0F5FD8B7DD8" box="[477,587,694,719]" italics="true" pageId="12" pageNumber="457">B. parvus</emphasis>
</taxonomicName>
<emphasis id="60CECA25FFB62F06C811B0F6FD567DD8" bold="true" box="[596,662,693,719]" pageId="12" pageNumber="457">
<taxonomicNameLabel id="7BFD775EFFB62F06C811B0F6FD567DD8" box="[596,662,693,719]" pageId="12" pageNumber="457" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be placed in the 
<taxonomicName id="95BA6DB4FFB62F06C9D5B0F5FC2C7DD8" authorityName="Boheman" authorityYear="1859" box="[912,1004,694,719]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="sticticus">
<emphasis id="60CECA25FFB62F06C9D5B0F5FC2C7DD8" box="[912,1004,694,719]" italics="true" pageId="12" pageNumber="457">sticticus</emphasis>
</taxonomicName>
-complex (
<bibRefCitation id="362B6BC6FFB62F06CE22B0F6FB2B7DD8" author="Oliva, A." box="[1127,1259,693,719]" pageId="12" pageNumber="457" pagination="57 - 237" refId="ref15731" refString="Oliva, A. (1989) El genero Berosus (Coleoptera: Hydrophilidae) en America del Sur. Revista el Museo Argentino de Ciencias Naturales, 6 (4), 57 - 237." type="journal article" year="1989">Oliva 1989</bibRefCitation>
; 
<bibRefCitation id="362B6BC6FFB62F06CEBCB0F6FF137DE4" author="Oliva, A. &amp; Short, A. E. Z." pageId="12" pageNumber="457" pagination="1 - 69" refId="ref15768" refString="Oliva, A. &amp; Short, A. E. Z. (2012) Review of the Berosus Leach of Venezuela (Coleoptera: Hydrophilidae: Berosini) with description of fourteen new species. Zookeys, 206, 1 - 69. https: // doi. org / 10.3897 / zookeys. 206.2587" type="journal article" year="2012">Oliva &amp; Short 2012</bibRefCitation>
), based on the following characteristics: dorsal sculpture fine (varying from coarse to fine in the complex); elytra with humeral humps not prominent and without spine-like hairs; abdominal ventrites medially carinate behind metacoxae without lateral depressions in first ventrite; protarsus of males with soles on the two basal tarsomeres, which are weakly swollen (weakly to moderately swollen in the complex); and male genitalia on which the aedeagus is somewhat compressed laterally, with parameres parallel and acuminate, with median lobe weakly curved (strongly or weakly curved in the complex). The new species is similar to 
<taxonomicName id="95BA6DB4FFB62F06C9EEB1CDFB767CB0" authority="Queney, 2006" authorityName="Queney" authorityYear="2006" box="[939,1206,909,935]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="olivae">
<emphasis id="60CECA25FFB62F06C9EEB1CDFBD07CB0" box="[939,1040,909,935]" italics="true" pageId="12" pageNumber="457">B. olivae</emphasis>
<bibRefCitation id="362B6BC6FFB62F06CE5CB1CEFB767CB0" author="Queney, P." box="[1049,1206,909,935]" pageId="12" pageNumber="457" pagination="457 - 464" refId="ref15825" refString="Queney, P. (2006) Description de trois especes nouvelles de Berosus de Guyane francaise (Coleoptera, Hydrophilidae). Bulletin de la societe entomologique de France, 111 (4), 457 - 464." type="journal article" year="2006">Queney, 2006</bibRefCitation>
</taxonomicName>
by the body length, dorsal coloration and shape of the apical emargination of the fifth ventrite, but they can be differentiated by the shape of male genitalia. In 
<taxonomicName id="95BA6DB4FFB62F06CB82B195FDF27CF8" authority="Santana &amp; Benetti &amp; Clarkson &amp; Pes, 2019" authorityName="Santana &amp; Benetti &amp; Clarkson &amp; Pes" authorityYear="2019" box="[455,562,982,1007]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="60CECA25FFB62F06CB82B195FDF27CF8" box="[455,562,982,1007]" italics="true" pageId="12" pageNumber="457">B. parvus</emphasis>
</taxonomicName>
<emphasis id="60CECA25FFB62F06C87FB196FDB97CF8" bold="true" box="[570,633,981,1007]" pageId="12" pageNumber="457">
<taxonomicNameLabel id="7BFD775EFFB62F06C87FB196FDB97CF8" box="[570,633,981,1007]" pageId="12" pageNumber="457" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
the aedeagus is subcylindrical in dorsal view (
<figureCitation id="CA810AB2FFB62F06CEC0B196FB127CE7" box="[1157,1234,981,1008]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">Fig. 6f</figureCitation>
) and sigmoidal in lateral view (
<figureCitation id="CA810AB2FFB62F06CB6EB1BAFEBF7B03" box="[299,383,1017,1044]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">
Fig. 
<quantity id="9542BBD2FFB62F06CB25B1BAFEBF7B03" box="[352,383,1017,1044]" metricMagnitude="-3" metricUnit="kg" metricValue="6.0" pageId="12" pageNumber="457" unit="g" value="6.0">6g</quantity>
</figureCitation>
); the basal piece corresponds to ca. three-quarters of the total length (
<figureCitation id="CA810AB2FFB62F06CECAB1BAFACB7B03" box="[1167,1291,1017,1044]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">
Figs. 
<quantity id="9542BBD2FFB62F06CE8AB1BAFB2D7B04" box="[1231,1261,1017,1043]" metricMagnitude="-3" metricUnit="kg" metricValue="6.0" pageId="12" pageNumber="457" unit="g" value="6.0">6g</quantity>
, h
</figureCitation>
); the median lobe is cylindrical and has a hole at apex (
<figureCitation id="CA810AB2FFB62F06C820B65EFD737B2F" box="[613,691,1053,1080]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">Fig. 6f</figureCitation>
); in 
<taxonomicName id="95BA6DB4FFB62F06C8A6B65DFC857B20" authorityName="Queney" authorityYear="2006" box="[739,837,1053,1079]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="olivae">
<emphasis id="60CECA25FFB62F06C8A6B65DFC857B20" box="[739,837,1053,1079]" italics="true" pageId="12" pageNumber="457">B. olivae</emphasis>
</taxonomicName>
, the aedeagus gradually narrows along its distal third; the basal piece corresponds to a little more than half the total genitalia length. 
<taxonomicName id="95BA6DB4FFB62F06CE4CB601FBB57B4C" authority="Santana &amp; Benetti &amp; Clarkson &amp; Pes, 2019" authorityName="Santana &amp; Benetti &amp; Clarkson &amp; Pes" authorityYear="2019" box="[1033,1141,1090,1115]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="60CECA25FFB62F06CE4CB601FBB57B4C" box="[1033,1141,1090,1115]" italics="true" pageId="12" pageNumber="457">B. parvus</emphasis>
</taxonomicName>
<emphasis id="60CECA25FFB62F06CE39B602FB7C7B4C" bold="true" box="[1148,1212,1089,1115]" pageId="12" pageNumber="457">
<taxonomicNameLabel id="7BFD775EFFB62F06CE39B602FB7C7B4C" box="[1148,1212,1089,1115]" pageId="12" pageNumber="457" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is also similar to 
<taxonomicName id="95BA6DB4FFB62F06CFC1B601FE607B68" authority="Oliva, 1989" authorityName="Oliva" authorityYear="1989" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="subandinus">
<emphasis id="60CECA25FFB62F06CFC1B601FED77B68" italics="true" pageId="12" pageNumber="457">B. subandinus</emphasis>
<bibRefCitation id="362B6BC6FFB62F06CB58B626FE607B68" author="Oliva, A." box="[285,416,1125,1151]" pageId="12" pageNumber="457" pagination="57 - 237" refId="ref15731" refString="Oliva, A. (1989) El genero Berosus (Coleoptera: Hydrophilidae) en America del Sur. Revista el Museo Argentino de Ciencias Naturales, 6 (4), 57 - 237." type="journal article" year="1989">Oliva, 1989</bibRefCitation>
</taxonomicName>
by the shape of the male genitalia in lateral view and length of the basal piece, but 
<taxonomicName id="95BA6DB4FFB62F06CF77B625FA5C7B68" authority="Santana &amp; Benetti &amp; Clarkson &amp; Pes, 2019" authorityName="Santana &amp; Benetti &amp; Clarkson &amp; Pes" authorityYear="2019" box="[1330,1436,1126,1151]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="60CECA25FFB62F06CF77B625FA5C7B68" box="[1330,1436,1126,1151]" italics="true" pageId="12" pageNumber="457">B. parvus</emphasis>
</taxonomicName>
<emphasis id="60CECA25FFB62F06CAD2B6CAFF177BB4" bold="true" box="[151,215,1161,1187]" pageId="12" pageNumber="457">
<taxonomicNameLabel id="7BFD775EFFB62F06CAD2B6CAFF177BB4" box="[151,215,1161,1187]" pageId="12" pageNumber="457" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has wide parameres in the distal three-quarters of its length (
<figureCitation id="CA810AB2FFB62F06C9C9B6CAFC197BB3" box="[908,985,1161,1188]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">Figs. 6</figureCitation>
f–h), while in 
<taxonomicName id="95BA6DB4FFB62F06CE39B6C9FADB7BB4" authorityName="Oliva" authorityYear="1989" box="[1148,1307,1161,1187]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="subandinus">
<emphasis id="60CECA25FFB62F06CE39B6C9FADB7BB4" box="[1148,1307,1161,1187]" italics="true" pageId="12" pageNumber="457">B. subandinus</emphasis>
</taxonomicName>
the parameres are narrow and have a marked notch in the distal 4/5 of its length. In addition, 
<taxonomicName id="95BA6DB4FFB62F06CE0BB6EDFB7A7BD0" authority="Santana &amp; Benetti &amp; Clarkson &amp; Pes, 2019" authorityName="Santana &amp; Benetti &amp; Clarkson &amp; Pes" authorityYear="2019" box="[1102,1210,1198,1223]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="60CECA25FFB62F06CE0BB6EDFB7A7BD0" box="[1102,1210,1198,1223]" italics="true" pageId="12" pageNumber="457">B. parvus</emphasis>
</taxonomicName>
<emphasis id="60CECA25FFB62F06CE87B6EEFAC37BD0" bold="true" box="[1218,1283,1197,1223]" pageId="12" pageNumber="457">
<taxonomicNameLabel id="7BFD775EFFB62F06CE87B6EEFAC37BD0" box="[1218,1283,1197,1223]" pageId="12" pageNumber="457" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has a narrow metaventral process, which is carinate before the median depression with posterolateral angles produced into small rounded laminae (
<figureCitation id="CA810AB2FFB62F06CB20B6B6FE767A07" box="[357,438,1269,1296]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">Fig. 6c</figureCitation>
) and the apical emargination of the fifth abdominal ventrite bears a single medial tooth (
<figureCitation id="CA810AB2FFB62F06CAE5B75AFF327A23" box="[160,242,1305,1332]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">Fig. 6e</figureCitation>
), while 
<taxonomicName id="95BA6DB4FFB62F06CB0AB759FE2F7A24" authorityName="Oliva" authorityYear="1989" box="[335,495,1305,1331]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="subandinus">
<emphasis id="60CECA25FFB62F06CB0AB759FE2F7A24" box="[335,495,1305,1331]" italics="true" pageId="12" pageNumber="457">B. subandinus</emphasis>
</taxonomicName>
has a wide metaventral process with posterolateral angles not produced and apical emargination of the fifth abdominal ventrite with two teeth in the medial region. The new species is similar to 
<taxonomicName id="95BA6DB4FFB62F06CFC1B77DFED37A6C" authorityName="Queney" authorityYear="2006" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="guyanensis">
<emphasis id="60CECA25FFB62F06CFC1B77DFED37A6C" italics="true" pageId="12" pageNumber="457">B. guyanensis</emphasis>
</taxonomicName>
in dorsal coloration, punctuation of the head, pronotum, elytra and shape of mesoventral process. However, 
<taxonomicName id="95BA6DB4FFB62F06CA91B7C5FEFF7A88" authority="Santana &amp; Benetti &amp; Clarkson &amp; Pes, 2019" authorityName="Santana &amp; Benetti &amp; Clarkson &amp; Pes" authorityYear="2019" box="[212,319,1414,1439]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="60CECA25FFB62F06CA91B7C5FEFF7A88" box="[212,319,1414,1439]" italics="true" pageId="12" pageNumber="457">B. parvus</emphasis>
</taxonomicName>
<emphasis id="60CECA25FFB62F06CB03B7C6FE467A88" bold="true" box="[326,390,1413,1439]" pageId="12" pageNumber="457">
<taxonomicNameLabel id="7BFD775EFFB62F06CB03B7C6FE467A88" box="[326,390,1413,1439]" pageId="12" pageNumber="457" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has the first abdominal ventrite carinate along its entire length medially and apical emargination of the fifth abdominal ventrite with a single medial tooth (
<figureCitation id="CA810AB2FFB62F06C914B7EAFC627AD3" box="[849,930,1449,1476]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">Fig. 6e</figureCitation>
), while 
<taxonomicName id="95BA6DB4FFB62F06C9BEB7E9FB567AD4" authorityName="Queney" authorityYear="2006" box="[1019,1174,1450,1475]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="guyanensis">
<emphasis id="60CECA25FFB62F06C9BEB7E9FB567AD4" box="[1019,1174,1450,1475]" italics="true" pageId="12" pageNumber="457">B. guyanensis</emphasis>
</taxonomicName>
has the first abdominal ventrite carinate on anterior one-third and apical emargination of the fifth abdominal ventrite with two medial teeth. In addition, 
<taxonomicName id="95BA6DB4FFB62F06CB65B7B1FE4B791C" authority="Santana &amp; Benetti &amp; Clarkson &amp; Pes, 2019" authorityName="Santana &amp; Benetti &amp; Clarkson &amp; Pes" authorityYear="2019" box="[288,395,1522,1547]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="60CECA25FFB62F06CB65B7B1FE4B791C" box="[288,395,1522,1547]" italics="true" pageId="12" pageNumber="457">B. parvus</emphasis>
</taxonomicName>
<emphasis id="60CECA25FFB62F06CBD6B7B2FE13791C" bold="true" box="[403,467,1521,1547]" pageId="12" pageNumber="457">
<taxonomicNameLabel id="7BFD775EFFB62F06CBD6B7B2FE13791C" box="[403,467,1521,1547]" pageId="12" pageNumber="457" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has a subcylindrical aedeagus with the same thickness from base to apex (
<figureCitation id="CA810AB2FFB62F06CF5CB7B2FA4F791B" box="[1305,1423,1521,1548]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">Figs. 6f, h</figureCitation>
), basal piece reaching three-quarter of the total length (
<figureCitation id="CA810AB2FFB62F06C8ACB456FCF47927" box="[745,820,1557,1584]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">Figs. 6</figureCitation>
g–h) and parameres with the same length as the median lobe (
<figureCitation id="CA810AB2FFB62F06CA92B47AFE837943" box="[215,323,1593,1620]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">Fig. 6f, h</figureCitation>
), while in 
<taxonomicName id="95BA6DB4FFB62F06CBFEB479FD977944" authorityName="Queney" authorityYear="2006" box="[443,599,1594,1619]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="guyanensis">
<emphasis id="60CECA25FFB62F06CBFEB479FD977944" box="[443,599,1594,1619]" italics="true" pageId="12" pageNumber="457">B. guyanensis</emphasis>
</taxonomicName>
the male genitalia narrows abruptly at four-fifths of total length, the basal piece reaches four-fifths of total length and the parameres are larger than the median lobe. The species also similar to 
<taxonomicName id="95BA6DB4FFB62F06CAF1B4C1FE2B798C" authority="Boheman, 1859" authorityName="Boheman" authorityYear="1859" box="[180,491,1664,1691]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="sticticus">
<emphasis id="60CECA25FFB62F06CAF1B4C1FEF0798C" box="[180,304,1666,1691]" italics="true" pageId="12" pageNumber="457">B. sticticus</emphasis>
Boheman, 1859
</taxonomicName>
, but differs from this species by the shape of the mesoventral process, carina of first ventrite and apical notch of fifth abdominal ventrite. 
<taxonomicName id="95BA6DB4FFB62F06C8A8B4E5FC9979A8" authority="Santana &amp; Benetti &amp; Clarkson &amp; Pes, 2019" authorityName="Santana &amp; Benetti &amp; Clarkson &amp; Pes" authorityYear="2019" box="[749,857,1702,1727]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="60CECA25FFB62F06C8A8B4E5FC9979A8" box="[749,857,1702,1727]" italics="true" pageId="12" pageNumber="457">B. parvus</emphasis>
</taxonomicName>
<emphasis id="60CECA25FFB62F06C924B4E6FC6179A8" bold="true" box="[865,929,1701,1727]" pageId="12" pageNumber="457">
<taxonomicNameLabel id="7BFD775EFFB62F06C924B4E6FC6179A8" box="[865,929,1701,1727]" pageId="12" pageNumber="457" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has the mesoventral process with very short, curved anterior tooth, directed downwards and backwards, posterior angle weakly raised in lateral view, not prominent (
<figureCitation id="CA810AB2FFB62F06CA9DB4AEFEED781F" box="[216,301,1773,1800]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">Fig. 6d</figureCitation>
), while 
<taxonomicName id="95BA6DB4FFB62F06CBCFB4ADFDC67810" authorityName="Boheman" authorityYear="1859" box="[394,518,1774,1799]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="sticticus">
<emphasis id="60CECA25FFB62F06CBCFB4ADFDC67810" box="[394,518,1774,1799]" italics="true" pageId="12" pageNumber="457">B. sticticus</emphasis>
</taxonomicName>
has a mesoventral process with a large and wide tooth, directed downwards and prominent posterior angle in lateral view. In addition, 
<taxonomicName id="95BA6DB4FFB62F06C8BDB551FCA4783C" authority="Santana &amp; Benetti &amp; Clarkson &amp; Pes, 2019" authorityName="Santana &amp; Benetti &amp; Clarkson &amp; Pes" authorityYear="2019" box="[760,868,1810,1835]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="60CECA25FFB62F06C8BDB551FCA4783C" box="[760,868,1810,1835]" italics="true" pageId="12" pageNumber="457">B. parvus</emphasis>
</taxonomicName>
<emphasis id="60CECA25FFB62F06C92EB552FC6B783C" bold="true" box="[875,939,1809,1835]" pageId="12" pageNumber="457">
<taxonomicNameLabel id="7BFD775EFFB62F06C92EB552FC6B783C" box="[875,939,1809,1835]" pageId="12" pageNumber="457" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has the first ventrite medially carinate in its whole length and the fifth ventrite has apical notch with single short median tooth (
<figureCitation id="CA810AB2FFB62F06CE0DB576FB5B7847" box="[1096,1179,1845,1872]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">Fig. 6e</figureCitation>
), while 
<taxonomicName id="95BA6DB4FFB62F06CEB3B575FAB27858" authorityName="Boheman" authorityYear="1859" box="[1270,1394,1846,1871]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="sticticus">
<emphasis id="60CECA25FFB62F06CEB3B575FAB27858" box="[1270,1394,1846,1871]" italics="true" pageId="12" pageNumber="457">B. sticticus</emphasis>
</taxonomicName>
the first ventrite is carinated in the anterior third and the apical notch of the fifth ventrite has two adjacent teeth. These species also differ in the shape of the male genitalia, in 
<taxonomicName id="95BA6DB4FFB62F06C8BFB53DFCA47880" authority="Santana &amp; Benetti &amp; Clarkson &amp; Pes, 2019" authorityName="Santana &amp; Benetti &amp; Clarkson &amp; Pes" authorityYear="2019" box="[762,868,1918,1943]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="60CECA25FFB62F06C8BFB53DFCA47880" box="[762,868,1918,1943]" italics="true" pageId="12" pageNumber="457">B. parvus</emphasis>
</taxonomicName>
<emphasis id="60CECA25FFB62F06C92FB53EFC687880" bold="true" box="[874,936,1917,1943]" pageId="12" pageNumber="457">
<taxonomicNameLabel id="7BFD775EFFB62F06C92FB53EFC687880" box="[874,936,1917,1943]" pageId="12" pageNumber="457" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
the length of the basal piece is three-quarters of the total length (
<figureCitation id="CA810AB2FFB62F06CB2EB5E2FE2278AB" box="[363,482,1953,1980]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">
Figs. 
<quantity id="9542BBD2FFB62F06CBEFB5E2FE0778AC" box="[426,455,1953,1979]" metricMagnitude="-3" metricUnit="kg" metricValue="6.0" pageId="12" pageNumber="457" unit="g" value="6.0">6g</quantity>
, h
</figureCitation>
) and the median lobe is as long as the parameres (
<figureCitation id="CA810AB2FFB62F06CE55B5E2FB4478AB" box="[1040,1156,1953,1980]" captionStart="FIGURE 6" captionStartId="13.[151,250,1965,1990]" captionTargetBox="[205,1382,166,1945]" captionTargetId="figure@13.[205,1382,166,1945]" captionTargetPageId="13" captionText="FIGURE 6. Berosus parvus sp. n. (A–C) Habitus, A) dorsal view; B) lateral view; C) ventral view; D) Mesoventral process, ventral view, E) Abdomen, ventral view; (F–H) Aedeagus, F) dorsal view; G) lateral view; H) ventral view." figureDoi="http://doi.org/10.5281/zenodo.3557578" httpUri="https://zenodo.org/record/3557578/files/figure.png" pageId="12" pageNumber="457">Figs. 6f, h</figureCitation>
), while in 
<taxonomicName id="95BA6DB4FFB62F06CEBCB5E1FAB478AC" authorityName="Boheman" authorityYear="1859" box="[1273,1396,1954,1979]" class="Insecta" family="Hydrophilidae" genus="Berosus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="457" phylum="Arthropoda" rank="species" species="sticticus">
<emphasis id="60CECA25FFB62F06CEBCB5E1FAB478AC" box="[1273,1396,1954,1979]" italics="true" pageId="12" pageNumber="457">B. sticticus</emphasis>
</taxonomicName>
the basal piece is about half of total length and median lobe shorter than the parameres.
</paragraph>
</subSubSection>
</treatment>
</document>