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<document id="663404C929208CAEDC37B06095591E90" ID-DOI="10.5281/zenodo.13203160" ID-ISSN="1407-8619" ID-Zenodo-Dep="13203160" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657382176" checkinUser="felipe" docAuthor="Fanti, Fabrizio &amp; Müller, Patrick" docDate="2022" docId="03DAD52BFFBBFFCAFF0DB95A5D72A26B" docLanguage="en" docName="BalticJColeopterol.22.2.331-380.pdf" docOrigin="Baltic Journal of Coleopterology 22 (2)" docStyle="DocumentStyle:9720869F1FB170B2A509257D1FA394CD.2:BalticJColeop.2022.journal_article" docStyleId="9720869F1FB170B2A509257D1FA394CD" docStyleName="BalticJColeop.2022.journal_article" docStyleVersion="2" docTitle="Sanaungulus nalae Fanti &amp; Müller, 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="363" masterDocId="FFE3AD53FFA5FFEAFF9ABD325F46A170" masterDocTitle="Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information" masterLastPageNumber="380" masterPageNumber="331" pageNumber="361" updateTime="1722697113453" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="DF1AFD432EF708B746E3D0EF4BD300B9">Fossil Cantharidae from the Cretaceous Burmese (Kachin) amber of the Patrick Müller collection, and taxonomic information</mods:title>
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<mods:namePart id="76FC5426D1B6D583FC97270382089129">Fanti, Fabrizio</mods:namePart>
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<mods:namePart id="1930120D0555C6217CE7678C837E67EE">Müller, Patrick</mods:namePart>
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<mods:title id="77740340EAEBEB798AB42EBF7D1717BD">Baltic Journal of Coleopterology</mods:title>
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<mods:date id="C17EE20BA065B3961EF8037F68D5A6C9">2022</mods:date>
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<treatment id="03DAD52BFFBBFFCAFF0DB95A5D72A26B" ID-DOI="http://doi.org/10.5281/zenodo.13199831" ID-Zenodo-Dep="13199831" LSID="urn:lsid:plazi:treatment:03DAD52BFFBBFFCAFF0DB95A5D72A26B" httpUri="http://treatment.plazi.org/id/03DAD52BFFBBFFCAFF0DB95A5D72A26B" lastPageId="32" lastPageNumber="363" pageId="30" pageNumber="361">
<subSubSection id="C36937B6FFBBFFF4FF0DB95A5E8FA5F0" box="[151,457,1128,1152]" pageId="30" pageNumber="361" type="nomenclature">
<paragraph id="8BCC643DFFBBFFF4FF0DB95A5E8FA5F0" blockId="30.[151,457,1128,1183]" box="[151,457,1128,1152]" pageId="30" pageNumber="361">
<heading id="D084D351FFBBFFF4FF0DB95A5E8FA5F0" bold="true" box="[151,457,1128,1152]" fontSize="10" level="2" pageId="30" pageNumber="361" reason="4">
<emphasis id="B907B82FFFBBFFF4FF0DB95A5E8FA5F0" bold="true" box="[151,457,1128,1152]" pageId="30" pageNumber="361">
<taxonomicName id="4C731FBEFFBBFFF4FF0DB95A5E2BA5F0" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[151,365,1128,1152]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="361" phylum="Arthropoda" rank="species" species="nalae" status="sp. nov.">
<emphasis id="B907B82FFFBBFFF4FF0DB95A5E2BA5F0" bold="true" box="[151,365,1128,1152]" italics="true" pageId="30" pageNumber="361">Sanaungulus nalae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFBBFFF4FEEEB95A5E8FA5F0" box="[372,457,1128,1152]" pageId="30" pageNumber="361" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFBBFFF4FF0DB9BA5FB2A5EF" box="[151,244,1160,1183]" pageId="30" pageNumber="361" type="description">
<paragraph id="8BCC643DFFBBFFF4FF0DB9BA5FB2A5EF" blockId="30.[151,457,1128,1183]" box="[151,244,1160,1183]" pageId="30" pageNumber="361">
(
<figureCitation id="134878B8FFBBFFF4FF05B9BA5FAAA5EF" box="[159,236,1160,1183]" captionStart="Fig" captionStartId="31.[113,152,1636,1659]" captionTargetBox="[223,1026,151,1606]" captionTargetId="figure-37@31.[223,1026,637,1120]" captionTargetPageId="31" captionText="Fig. 17. Sanaungulus nalae sp. nov. in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view; C: Holotype, detail of head and pronotum." figureDoi="http://doi.org/10.5281/zenodo.13203213" httpUri="https://zenodo.org/record/13203213/files/figure.png" pageId="30" pageNumber="361">Fig. 17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFBBFFF4FF0DB98D5EF0A773" pageId="30" pageNumber="361" type="materials_examined">
<paragraph id="8BCC643DFFBBFFF4FF0DB98D5E53A426" blockId="30.[151,644,1215,1366]" pageId="30" pageNumber="361">
<materialsCitation id="3B1B6E60FFBBFFF4FF0DB98D5E53A426" collectionCode="BUB" county="Male" location="Catalog" municipality="Burmese" pageId="30" pageNumber="361" specimenCode="BUB4033" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
<emphasis id="B907B82FFFBBFFF4FF0DB98D5E40A5A7" bold="true" box="[151,262,1215,1239]" pageId="30" pageNumber="361">
<typeStatus id="54C8DA9FFFBBFFF4FF0DB98D5E47A5A7" box="[151,257,1215,1239]" pageId="30" pageNumber="361" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62AD1CB1FFBBFFF4FE96B98D5E03A5A6" box="[268,325,1215,1238]" pageId="30" pageNumber="361">Male</collectingCounty>
, adult specimen in a
<collectingMunicipality id="6BA8FE47FFBBFFF4FDBEB98D5DC5A5A6" box="[548,643,1215,1238]" pageId="30" pageNumber="361">Burmese</collectingMunicipality>
(
<collectingRegion id="49B7AADFFFBBFFF4FF05B9ED5FA8A586" box="[159,238,1247,1270]" country="Myanmar" name="Kachin" pageId="30" pageNumber="361">Kachin</collectingRegion>
) amber piece:
<location id="8EAC32E6FFBBFFF4FE19B9ED5E93A586" LSID="urn:lsid:plazi:treatment:03DAD52BFFBBFFCAFF0DB95A5D72A26B:8EAC32E6FFBBFFF4FE19B9ED5E93A586" box="[387,469,1247,1270]" county="Male" municipality="Burmese" name="Catalog" pageId="30" pageNumber="361" stateProvince="Kachin">Catalog</location>
number
<quantity id="4C8BC9D8FFBBFFF4FDB4B9ED5FA7A466" metricMagnitude="2" metricUnit="m" metricValue="1.526794" pageId="30" pageNumber="361" unit="in" value="6011.0">SNHM­6011 in</quantity>
the
<location id="8EAC32E6FFBBFFF4FE9DB9CD5E69A446" LSID="urn:lsid:plazi:treatment:03DAD52BFFBBFFCAFF0DB95A5D72A26B:8EAC32E6FFBBFFF4FE9DB9CD5E69A446" county="Male" municipality="Burmese" name="Staatliches Naturhistorisches Museum Braunschweig" pageId="30" pageNumber="361" stateProvince="Kachin">Staatliches Naturhistorisches Museum Braunschweig</location>
(ex
<location id="8EAC32E6FFBBFFF4FEC6B82D5EF1A446" LSID="urn:lsid:plazi:treatment:03DAD52BFFBBFFCAFF0DB95A5D72A26B:8EAC32E6FFBBFFF4FEC6B82D5EF1A446" box="[348,439,1311,1334]" county="Male" municipality="Burmese" name="Muller's" pageId="30" pageNumber="361" stateProvince="Kachin">Müllers</location>
collection number:
<specimenCode id="DBD5CC46FFBBFFF4FF0DB80D5E4FA426" box="[151,265,1343,1366]" collectionCode="BUB" pageId="30" pageNumber="361">BUB4033</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BCC643DFFBBFFF4FF0DB8475EACA4DD" blockId="30.[151,643,1397,1453]" pageId="30" pageNumber="361">
<emphasis id="B907B82FFFBBFFF4FF0DB8475E76A4FD" bold="true" box="[151,304,1397,1421]" pageId="30" pageNumber="361">Type locality.</emphasis>
<collectingCountry id="F36424ADFFBBFFF4FEA2B8445EE2A4FD" box="[312,420,1398,1421]" name="Myanmar" pageId="30" pageNumber="361">Myanmar</collectingCountry>
:
<collectingRegion id="49B7AADFFFBBFFF4FE2AB8445D7CA4FD" box="[432,570,1398,1421]" country="Myanmar" name="Kachin" pageId="30" pageNumber="361">Kachin state</collectingRegion>
, Myitkyina District, Hukawng Valley.
</paragraph>
<paragraph id="8BCC643DFFBBFFF4FF0DB8FE5EF0A773" blockId="30.[151,643,1484,1539]" pageId="30" pageNumber="361">
<emphasis id="B907B82FFFBBFFF4FF0DB8FE5E6CA494" bold="true" box="[151,298,1484,1508]" pageId="30" pageNumber="361">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid­Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFBBFFF4FF0DBB115B4BA182" pageId="30" pageNumber="361" type="diagnosis">
<paragraph id="8BCC643DFFBBFFF4FF0DBB115B4BA182" blockId="30.[151,644,1571,1690]" lastBlockId="30.[680,1172,155,242]" pageId="30" pageNumber="361">
<emphasis id="B907B82FFFBBFFF4FF0DBB115ED6A74B" bold="true" box="[151,400,1571,1595]" pageId="30" pageNumber="361">Differential diagnosis.</emphasis>
<taxonomicName id="4C731FBEFFBBFFF4FE0FBB115D19A74A" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[405,607,1571,1594]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="361" phylum="Arthropoda" rank="species" species="nalae" status="sp. nov.">
<emphasis id="B907B82FFFBBFFF4FE0FBB115D19A74A" box="[405,607,1571,1594]" italics="true" pageId="30" pageNumber="361">Sanaungulus nalae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFBBFFF4FDFEBB115F83A72A" pageId="30" pageNumber="361" rank="species">sp. nov.</taxonomicNameLabel>
differs from other
<taxonomicName id="4C731FBEFFBBFFF4FE08BB715D5AA72A" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" box="[402,540,1603,1626]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="361" phylum="Arthropoda" rank="genus">
<emphasis id="B907B82FFFBBFFF4FE08BB715D5AA72A" box="[402,540,1603,1626]" italics="true" pageId="30" pageNumber="361">Sanaungulus</emphasis>
</taxonomicName>
with filiform antennae for a combination of characters: Head partially covered by pronotum, smooth elytra, and a pronotal shape with the sides wider near the posterior margin and narrower from the anterior margin to the middle.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FFBBFFCAFD32BC235F88A0A4" lastPageId="32" lastPageNumber="363" pageId="30" pageNumber="361" type="description">
<paragraph id="8BCC643DFFBBFFF4FD32BC235B73A0F8" blockId="30.[680,1172,273,392]" pageId="30" pageNumber="361">
<emphasis id="B907B82FFFBBFFF4FD32BC235C68A059" bold="true" box="[680,814,273,297]" pageId="30" pageNumber="361">Description.</emphasis>
Adult, winged, slender. Male, defined on the shape of the last sternite which is triangular, and narrower than the last tergite. Body length
<quantity id="4C8BC9D8FFBBFFF4FCABBC435CC3A0F8" box="[817,901,369,392]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="30" pageNumber="361" unit="mm" value="5.2">5.2 mm</quantity>
. Entirely brown.
</paragraph>
<paragraph id="8BCC643DFFBBFFCAFD32BC9A5F88A0A4" blockId="30.[680,1173,424,1692]" lastBlockId="32.[151,643,155,468]" lastPageId="32" lastPageNumber="363" pageId="30" pageNumber="361">Head transverse, wide, slightly convex in the middle, constricted behind the eyes, smooth with short pubescence. Eyes prominent, very large, roundish and slightly subelliptic, inserted in the lateral part of the head, inter­ocular dorsal distance about 2.1 times greater than the eye diameter. Mandibles falciform. Maxillary palps 4­ segmented with the last palpomere very elongated, securiform. Labial palps 3­ segmented. Antennae 11­ segmented, filiform, short, slightly surpassing the half of elytra and almost reaching the half of abdomen, antennal insertion in the eyes proximity; scape elongated, robust; antennomere II very short, subovate; antennomere III thinner and about 1.8 times longer than second; antennomeres IVV subequal, slightly longer than antennomere III; antennomere VI as long as the previous ones, slightly thinner; antennomeres VIIX filiform, subequal; antennomere XI filiform, slightly sturdier than previous ones, rounded at apex; all the antennomeres with short pubescence. Pronotum transverse, wider than head, narrow anteriorly, anterior margin rounded and strongly bordered, posterior margin almost straight, sides enlarged near the posterior margin and narrower from the anterior margin to the middle, posterior corners rounded, pronotum disc smooth and pubescent. Scutellum triangular­shaped. Elytra short which reveals various abdominal segments, at humeri wider than pronotum, humeral zone slightly gibbous, posteriorly parallel­sided, rounded apexes, surface smooth and sligthly pubescent equipped with short setae. Metathoracic wings infuscate, very longer than elytra, not reaching the apex of abdomen. Sternum elongated with posterior margin rounded, sternites narrow and transverse, last tergite wide with posterior margin rounded, last sternite triangular and narrower than last tergite. Legs thin, relatively short, pubescent; coxae massive, elongated, curved; trochanters elongated, curved, roundish at apex; femora more robust than tibiae, nearly straight; tibiae cylindrical, thin, without spurs near the apex, longer than femora. Tarsal formula 5­ 5­ 5; first tarsomere long, about 2.8 times longer than second; third tarsomere short, triangular; fourth strongly bilobed; fifth tarsomere elongated, thin, flat and curved; claws simple, without denticle at the base.</paragraph>
</subSubSection>
<caption id="DF0C34B5FFBAFFF5FFEBBB565D80A7EB" ID-DOI="http://doi.org/10.5281/zenodo.13203213" ID-Zenodo-Dep="13203213" httpUri="https://zenodo.org/record/13203213/files/figure.png" pageId="31" pageNumber="362" startId="31.[113,152,1636,1659]" targetBox="[223,1026,151,1606]" targetPageId="31" targetType="figure">
<paragraph id="8BCC643DFFBAFFF5FFEBBB565D80A7EB" blockId="31.[113,1134,1636,1691]" pageId="31" pageNumber="362">
Fig. 17.
<taxonomicName id="4C731FBEFFBAFFF5FF51BB565ED1A70B" authorityName="Fanti &amp; Müller" authorityYear="2022" box="[203,407,1636,1659]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="362" phylum="Arthropoda" rank="species" species="nalae" status="sp. nov.">
<emphasis id="B907B82FFFBAFFF5FF51BB565ED1A70B" box="[203,407,1636,1659]" italics="true" pageId="31" pageNumber="362">Sanaungulus nalae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2340554FFBAFFF5FE04BB565EB7A70B" box="[414,497,1636,1659]" pageId="31" pageNumber="362" rank="species">sp. nov.</taxonomicNameLabel>
in Burmese amber. A: Holotype, dorsal view; B: Holotype, ventral view; C: Holotype, detail of head and pronotum.
</paragraph>
</caption>
<subSubSection id="C36937B6FF85FFCAFF0DBCC65FA9A3F3" pageId="32" pageNumber="363" type="etymology">
<paragraph id="8BCC643DFF85FFCAFF0DBCC65D78A35C" blockId="32.[151,643,500,556]" pageId="32" pageNumber="363">
<emphasis id="B907B82FFF85FFCAFF0DBCC65E5EA37C" bold="true" box="[151,280,500,524]" pageId="32" pageNumber="363">Etymology.</emphasis>
Named in honor of Nala Müller, daughter of the second author (Patrick).
</paragraph>
<paragraph id="8BCC643DFF85FFCAFF0DBF795FA9A3F3" blockId="32.[151,643,587,643]" pageId="32" pageNumber="363">
<emphasis id="B907B82FFF85FFCAFF0DBF795E7CA313" bold="true" box="[151,314,587,611]" pageId="32" pageNumber="363">Syninclusions.</emphasis>
<taxonomicName id="4C731FBEFF85FFCAFED9BF7E5ED5A313" box="[323,403,588,611]" class="Insecta" kingdom="Animalia" order="Diptera" pageId="32" pageNumber="363" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
(partial), air bubbles, detritus.
</paragraph>
</subSubSection>
<subSubSection id="C36937B6FF85FFCAFF0DBF905D72A26B" pageId="32" pageNumber="363" type="discussion">
<paragraph id="8BCC643DFF85FFCAFF0DBF905D72A26B" blockId="32.[151,643,674,795]" pageId="32" pageNumber="363">
<emphasis id="B907B82FFF85FFCAFF0DBF905E40A3CA" bold="true" box="[151,262,674,698]" pageId="32" pageNumber="363">Remarks.</emphasis>
Piece of clear, honey colored amber that measures 10x10x
<quantity id="4C8BC9D8FF85FFCAFEE6BFF15E94A3AA" box="[380,466,707,730]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="32" pageNumber="363" unit="mm" value="4.5">4.5 mm</quantity>
. The inclusion is complete. The specimen is ventrally covered by a white and large cotton­like emulsion.
</paragraph>
</subSubSection>
</treatment>
</document>