treatments-xml/data/03/D8/A5/03D8A574FF8A9B340908FB5E5514D317.xml
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<document ID-DOI="http://doi.org/10.5281/zenodo.6103176" ID-GBIF-Dataset="a8aba141-9159-4d8c-9b5d-928358570431" ID-GBIF-Taxon="120701237" ID-Zenodo-Dep="6103176" checkinTime="1464156649483" checkinUser="plazi" docAuthor="Theodore W. Pietsch &amp; Tracey T. Sutton" docDate="2015" docId="03D8A574FF8A9B340908FB5E5514D317" docLanguage="en" docName="copeia.103.2.429.pdf" docOrigin="Copeia 103 (2)" docStyle="DocumentStyle{}" docTitle="Lasiognathus dinema Pietsch &amp; Sutton, 2015, new specIes" docType="treatment" docVersion="5" lastPageId="3" lastPageNumber="432" masterDocId="FFE1DD0CFF8B9B370A56FFB2511BD44F" masterDocTitle="A New Species of the Ceratioid Anglerfish Genus Lasiognathus Regan (Lophiiformes: Oneirodidae) from the Northern Gulf of Mexico" masterLastPageNumber="432" masterPageNumber="429" pageId="1" pageNumber="429" updateTime="1645000614884" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A New Species of the Ceratioid Anglerfish Genus Lasiognathus Regan (Lophiiformes: Oneirodidae) from the Northern Gulf of Mexico</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Theodore W. Pietsch</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tracey T. Sutton</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Copeia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>103</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>429</mods:start>
<mods:end>432</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1643/Ci-14-181</mods:identifier>
<mods:identifier type="GBIF-Dataset">a8aba141-9159-4d8c-9b5d-928358570431</mods:identifier>
<mods:identifier type="Zenodo-Dep">269572</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6103176" ID-GBIF-Taxon="120701237" ID-Zenodo-Dep="6103176" LSID="urn:lsid:plazi:treatment:03D8A574FF8A9B340908FB5E5514D317" httpUri="http://treatment.plazi.org/id/03D8A574FF8A9B340908FB5E5514D317" lastPageId="3" pageId="1">
<subSubSection box="[862,1263,1260,1282]" pageId="1" type="nomenclature">
<paragraph blockId="1.[862,1533,520,1870]" box="[862,1263,1260,1282]" pageId="1">
<emphasis bold="true" box="[862,1263,1260,1282]" pageId="1">
<taxonomicName box="[862,1103,1260,1282]" class="Actinopterygii" family="Thaumatichthyidae" genus="Lasiognathus" kingdom="Animalia" order="Lophiiformes" pageId="1" phylum="Chordata" rank="species" species="dinema" status="new specIes">
<emphasis bold="true" box="[862,1103,1260,1282]" italics="true" pageId="1">
Lasiognathus
<taxonomicName box="[1019,1103,1260,1282]" pageId="1" rank="species" species="dinema" status="new specIes">dinema</taxonomicName>
</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[1119,1263,1261,1282]" pageId="1" rank="species">new specIes</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection box="[862,984,1291,1314]" pageId="1" type="description">
<paragraph blockId="1.[862,1533,520,1870]" box="[862,984,1291,1314]" pageId="1">
Figures
<number box="[945,984,1291,1312]" pageId="1" value="2.0" valueMax="3.0" valueMin="1.0">13</number>
</paragraph>
</subSubSection>
<subSubSection pageId="1" type="materials_examined">
<paragraph blockId="1.[862,1533,520,1870]" pageId="1">
<emphasis bold="true" box="[862,986,1342,1364]" italics="true" pageId="1">
<typeStatus box="[862,957,1342,1364]" pageId="1">Holotype</typeStatus>
.—
</emphasis>
<accessionNumber box="[987,1125,1342,1365]" httpUri="http://www.uniprot.org/uniprot/UW152603" pageId="1">
<collectionCodeEnum box="[987,1031,1343,1364]" pageId="1">UW</collectionCodeEnum>
<number box="[1040,1125,1342,1365]" pageId="1" value="152603.0">152603</number>
</accessionNumber>
, female,
<quantity box="[1230,1315,1342,1365]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" pageId="1" unit="mm" value="95.0">95 mm</quantity>
,
<potCollectionCode box="[1329,1400,1343,1364]" pageId="1">
<collectionCode box="[1329,1400,1343,1364]" country="USA" httpUri="http://grbio.org/cool/qzhp-igg5" name="National Oceanic and Atmospeheric Administration" pageId="1">NOAA</collectionCode>
</potCollectionCode>
<collectionCode box="[1410,1455,1343,1364]" pageId="1">FRV</collectionCode>
<taxonomicName box="[1465,1525,1343,1364]" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="1" phylum="Craniata" rank="superClass" superClass="Pisces">
<emphasis box="[1465,1525,1343,1364]" italics="true" pageId="1">Pisces</emphasis>
</taxonomicName>
, Station B
<number box="[977,1020,1372,1393]" pageId="1" value="249.0">249</number>
,
<potCollectionCode box="[1040,1082,1372,1393]" pageId="1">DD</potCollectionCode>
<number box="[1082,1095,1372,1393]" pageId="1" value="2.0">2</number>
, northern Gulf of
<collectingCountry box="[1340,1427,1371,1394]" name="Mexico" pageId="1">Mexico</collectingCountry>
,
<geoCoordinate box="[1447,1528,1371,1394]" direction="north" orientation="latitude" pageId="1" precision="5555" value="27.5">
<number box="[1447,1496,1371,1394]" pageId="1" value="27.5">27.5</number>
°N
</geoCoordinate>
,
<geoCoordinate box="[862,948,1401,1424]" direction="west" orientation="longitude" pageId="1" precision="5555" value="-88.5">
<number box="[862,910,1401,1424]" pageId="1" value="88.5">88.5</number>
°W
</geoCoordinate>
, Irish Herring Trawl, depth of collection
<quantity box="[1414,1527,1401,1423]" metricMagnitude="2" metricUnit="m" metricValue="6.3549999999999995" metricValueMax="12.709999999999999" metricValueMin="0.0" pageId="1" unit="m" value="635.5" valueMax="1271.0" valueMin="0.0">
0
<date box="[1442,1494,1401,1422]" pageId="1" value="1271">1271</date>
m
</quantity>
, bottom depth
<quantity box="[1027,1115,1431,1452]" metricMagnitude="3" metricUnit="m" metricValue="2.104" pageId="1" unit="m" value="2104.0">
<date box="[1027,1082,1431,1452]" pageId="1" value="2104">2104</date>
m
</quantity>
,
<date box="[1127,1276,1430,1453]" pageId="1" value="2011-06-26">
<number box="[1127,1154,1430,1453]" pageId="1" value="26.0">26</number>
June
<number box="[1221,1276,1431,1452]" pageId="1" value="2011.0">
<date box="[1221,1276,1431,1452]" pageId="1" value="2011" year="2011">2011</date>
</number>
</date>
(
<figureCitation box="[1291,1354,1430,1453]" captionStart="Fig. 2" captionStartId="2.[815,842,1897,1917]" captionTargetBox="[815,1484,1302,1860]" captionTargetId="figure@2.[815,1485,1302,1862]" captionTargetPageId="2" captionText="Fig. 2. Escae of Lasiognathus dinema Pietsch and Sutton, new species, in preservation: (A) Holotype, UW 152603, female, 95 mm SL, left lateral view; (B) Paratype, UW 152604, female, 30 mm SL, right lateral view (prolongation of distal escal appendage rotated to show anterior face)." httpUri="https://zenodo.org/record/269574/files/figure.png" pageId="1">
Fig.
<number box="[1340,1354,1431,1453]" pageId="1" value="2.0">2</number>
</figureCitation>
A).
</paragraph>
<paragraph blockId="1.[862,1533,520,1870]" pageId="1">
<emphasis bold="true" box="[862,996,1493,1514]" italics="true" pageId="1">
<typeStatus box="[862,967,1493,1514]" pageId="1">Paratypes</typeStatus>
.—
</emphasis>
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UW
<number box="[1053,1136,1492,1515]" pageId="1" value="152604.0">152604</number>
</accessionNumber>
, female,
<quantity box="[1246,1333,1493,1514]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="1" unit="mm" value="30.0">30 mm</quantity>
, R/V
<emphasis box="[1405,1525,1492,1514]" italics="true" pageId="1">Meg Skansi</emphasis>
, Station SW
<number box="[997,1009,1522,1545]" pageId="1" value="6.0">6</number>
D-net
<number box="[1087,1103,1522,1543]" pageId="1" value="2.0">2</number>
, northern Gulf of
<collectingCountry box="[1344,1429,1522,1545]" name="Mexico" pageId="1">Mexico</collectingCountry>
,
<geoCoordinate box="[1449,1528,1522,1544]" direction="north" orientation="latitude" pageId="1" precision="5555" value="27.0">27.0°N</geoCoordinate>
,
<geoCoordinate box="[862,945,1551,1573]" direction="west" orientation="longitude" pageId="1" precision="5555" value="-90.1">
<number box="[862,909,1551,1572]" pageId="1" value="90.1">90.1</number>
°W
</geoCoordinate>
,
<quantity box="[956,1011,1551,1573]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="1" unit="m" value="10.0">10 m</quantity>
<number box="[1012,1021,1549,1562]" pageId="1" value="2.0">2</number>
MOCNESS midwater trawl, depth of collection
<quantity box="[862,1002,1581,1602]" metricMagnitude="3" metricUnit="m" metricValue="1.0995" metricValueMax="1.2" metricValueMin="0.999" pageId="1" unit="m" value="1099.5" valueMax="1200.0" valueMin="999.0">9991200 m</quantity>
, bottom depth
<quantity box="[1183,1272,1580,1603]" metricMagnitude="3" metricUnit="m" metricValue="2.559" pageId="1" unit="m" value="2559.0">2559 m</quantity>
,
<date box="[1287,1437,1581,1602]" pageId="1" value="2011-06-23">
<date box="[1287,1374,1581,1602]" day="23" month="06" monthName="June" pageId="1" year="1923">
<number box="[1287,1314,1581,1602]" pageId="1" value="23.0">23</number>
<date box="[1324,1437,1581,1602]" month="06" monthName="June" pageId="1" year="2011">
June
<number box="[1384,1437,1581,1602]" pageId="1" value="2011.0">2011</number>
</date>
</date>
</date>
(
<figureCitation box="[1455,1525,1580,1603]" captionStart="Fig. 1" captionStartId="2.[97,124,1130,1150]" captionTargetBox="[97,1486,133,1093]" captionTargetId="figure@2.[97,1486,133,1095]" captionText="Fig. 1. Lasiognathus dinema Pietsch and Sutton, new species, paratype, UW 152604, female, 30 mm SL." httpUri="https://zenodo.org/record/269573/files/figure.png" pageId="1">Figs. 1</figureCitation>
,
<figureCitation box="[862,875,1610,1632]" captionStart="Fig. 2" captionStartId="2.[815,842,1897,1917]" captionTargetBox="[815,1484,1302,1860]" captionTargetId="figure@2.[815,1485,1302,1862]" captionTargetPageId="2" captionText="Fig. 2. Escae of Lasiognathus dinema Pietsch and Sutton, new species, in preservation: (A) Holotype, UW 152603, female, 95 mm SL, left lateral view; (B) Paratype, UW 152604, female, 30 mm SL, right lateral view (prolongation of distal escal appendage rotated to show anterior face)." httpUri="https://zenodo.org/record/269574/files/figure.png" pageId="1">2</figureCitation>
B);
<accessionNumber box="[917,1054,1609,1632]" httpUri="http://www.uniprot.org/uniprot/UW154626" pageId="1">
UW
<number box="[972,1054,1609,1632]" pageId="1" value="154626.0">154626</number>
</accessionNumber>
, female,
<quantity box="[1162,1248,1609,1632]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="1" unit="mm" value="36.0">36 mm</quantity>
, R/V
<emphasis box="[1318,1435,1609,1632]" italics="true" pageId="1">Meg Skansi</emphasis>
, Station SE
<number box="[892,902,1639,1660]" pageId="1" value="3.0">3</number>
D-net
<number box="[970,984,1639,1660]" pageId="1" value="1.0">1</number>
, northern Gulf of
<collectingCountry box="[1189,1272,1639,1662]" name="Mexico" pageId="1">Mexico</collectingCountry>
,
<geoCoordinate box="[1283,1360,1639,1662]" direction="north" orientation="latitude" pageId="1" precision="5555" value="26.6">
<number box="[1283,1330,1639,1662]" pageId="1" value="26.6">26.6</number>
°N
</geoCoordinate>
,
<geoCoordinate box="[1372,1454,1639,1662]" direction="west" orientation="longitude" pageId="1" precision="5555" value="-86.8">
<number box="[1372,1419,1639,1662]" pageId="1" value="86.8">86.8</number>
°W
</geoCoordinate>
,
<quantity box="[1466,1523,1639,1661]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="1" unit="m" value="10.0">10 m</quantity>
<number box="[1523,1532,1637,1650]" pageId="1" value="2.0">2</number>
MOCNESS midwater trawl, depth of collection
<quantity box="[1400,1528,1669,1690]" metricMagnitude="2" metricUnit="m" metricValue="8.995" metricValueMax="9.99" metricValueMin="8.0" pageId="1" unit="m" value="899.5" valueMax="999.0" valueMin="800.0">800999 m</quantity>
, bottom depth
<quantity box="[1023,1109,1697,1720]" metricMagnitude="3" metricUnit="m" metricValue="2.9669999999999996" pageId="1" unit="m" value="2967.0">2967 m</quantity>
,
<date box="[1121,1331,1697,1720]" pageId="1" value="2011-09-11">
<number box="[1121,1148,1698,1719]" pageId="1" value="11.0">11</number>
September
<number box="[1277,1331,1698,1719]" pageId="1" value="2011.0">2011</number>
</date>
(
<figureCitation box="[1346,1407,1697,1720]" captionStart="Fig. 3" captionStartId="3.[145,172,1207,1227]" captionTargetBox="[145,814,133,1171]" captionTargetId="figure@3.[145,815,133,1171]" captionTargetPageId="3" captionText="Fig. 3. Esca of Lasiognathus dinema Pietsch and Sutton, new species, paratype, UW 154626, female, 36 mm SL, left lateral view, in preservation. Image courtesy of L. Malarky (NSUOC)." httpUri="https://zenodo.org/record/269575/files/figure.png" pageId="1">
Fig.
<number box="[1393,1407,1698,1720]" pageId="1" value="3.0">3</number>
</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="2" pageId="1" type="diagnosis">
<paragraph blockId="1.[862,1533,520,1870]" pageId="1">
<emphasis bold="true" box="[862,995,1760,1781]" italics="true" pageId="1">Diagnosis.—</emphasis>
Escal bulb with an elongate, cylindrical, internally pigmented anterior appendage; distal escal appendage a short cylindrical stalk, with an expanded distal end and a pair of tapering, filamentous prolongations emerging anteriorly from
</paragraph>
<footnote pageId="1">
<paragraph blockId="1.[145,1532,1899,2023]" pageId="1">
<number box="[145,154,1899,1911]" pageId="1" value="1.0">1</number>
School of Aquatic and Fishery Sciences and Burke Museum of Natural History and Culture, University of Washington, Campus Box
<number box="[1453,1529,1902,1922]" pageId="1" value="355020.0">355020</number>
, Seattle, Washington
<number box="[377,439,1927,1947]" pageId="1" value="98105.0">98105</number>
-
<number box="[445,495,1927,1947]" pageId="1" value="5020.0">5020</number>
; E-mail: twp@uw.edu. Send reprint requests to this address.
</paragraph>
<paragraph blockId="1.[145,1532,1899,2023]" box="[145,1521,1949,1973]" pageId="1">
<number box="[145,154,1949,1961]" pageId="1" value="2.0">2</number>
Nova Southeastern University Oceanographic Center,
<number box="[699,748,1953,1973]" pageId="1" value="8000.0">8000</number>
North Ocean Drive,
<collectingCountry box="[958,1017,1953,1973]" name="Denmark" pageId="1">Dania</collectingCountry>
Beach, Florida
<number box="[1171,1234,1953,1973]" pageId="1" value="33004.0">33004</number>
; E-mail: tsutton
<number box="[1396,1404,1953,1973]" pageId="1" value="1.0">1</number>
@nova.edu.
</paragraph>
<paragraph blockId="1.[145,1532,1899,2023]" box="[145,976,1978,1998]" pageId="1">
Submitted:
<date box="[260,430,1978,1998]" pageId="1" value="2014-10-29">
<number box="[260,284,1978,1998]" pageId="1" value="29.0">29</number>
October
<number box="[378,430,1978,1998]" pageId="1" value="2014.0">2014</number>
</date>
. Accepted:
<date box="[543,710,1978,1998]" pageId="1" value="2015-01-29">
<number box="[543,567,1978,1998]" pageId="1" value="29.0">29</number>
January
<number box="[658,710,1978,1998]" pageId="1" value="2015.0">2015</number>
</date>
. Associate Editor: D. Buth.
</paragraph>
<footnote box="[145,1428,2002,2023]" pageId="1">
<paragraph blockId="1.[145,1532,1899,2023]" box="[145,1428,2002,2023]" pageId="1">
©
<number box="[170,219,2003,2023]" pageId="1" value="2015.0">2015</number>
by the American Society of Ichthyologists and Herpetologists DOI:
<number box="[928,1009,2003,2023]" pageId="1" value="10.1643">10.1643</number>
/CI-
<number box="[1050,1114,2003,2023]" pageId="1" value="97.5" valueMax="181.0" valueMin="14.0">14-181</number>
Published online:
<date box="[1302,1428,2003,2023]" pageId="1" value="2015-06-04">June 4, 2015</date>
</paragraph>
</footnote>
</footnote>
<caption box="[97,1105,1130,1150]" httpUri="https://zenodo.org/record/269573/files/figure.png" pageId="2" targetBox="[97,1486,133,1093]" targetPageId="2">
<paragraph blockId="2.[97,151,1130,1150]" box="[97,151,1130,1150]" pageId="2">
<heading bold="true" box="[97,151,1130,1150]" centered="true" fontSize="8" level="5" pageId="2" reason="2">
<emphasis bold="true" box="[97,151,1130,1150]" pageId="2">Fig. 1.</emphasis>
</heading>
</paragraph>
<paragraph blockId="2.[172,1105,1130,1150]" box="[172,1105,1130,1150]" pageId="2">
<emphasis bold="true" box="[172,1105,1130,1150]" pageId="2">
<taxonomicName authority="Pietsch and Sutton" authorityName="Pietsch and Sutton" box="[172,565,1130,1150]" class="Actinopterygii" family="Thaumatichthyidae" genus="Lasiognathus" kingdom="Animalia" order="Lophiiformes" pageId="2" phylum="Chordata" rank="species" species="dinema" status="new species">
<emphasis bold="true" box="[172,379,1130,1150]" italics="true" pageId="2">Lasiognathus dinema</emphasis>
Pietsch and Sutton
</taxonomicName>
,
<taxonomicNameLabel box="[576,693,1131,1150]" pageId="2" rank="species">new species</taxonomicNameLabel>
, paratype, UW 152604, female, 30 mm SL.
</emphasis>
</paragraph>
</caption>
<paragraph blockId="2.[97,766,1201,1312]" pageId="2">
bases of escal hooks; distal escal appendage bearing three, lightly pigmented, bony, hook-like denticles; posterior escal appendage membranous, antero-posteriorly compressed, with a scalloped or rounded distal margin (
<figureCitation box="[514,583,1289,1312]" captionStart="Fig. 2" captionStartId="2.[815,842,1897,1917]" captionTargetBox="[815,1484,1302,1860]" captionTargetId="figure@2.[815,1485,1302,1862]" captionTargetPageId="2" captionText="Fig. 2. Escae of Lasiognathus dinema Pietsch and Sutton, new species, in preservation: (A) Holotype, UW 152603, female, 95 mm SL, left lateral view; (B) Paratype, UW 152604, female, 30 mm SL, right lateral view (prolongation of distal escal appendage rotated to show anterior face)." httpUri="https://zenodo.org/record/269574/files/figure.png" pageId="2">Figs. 2</figureCitation>
,
<figureCitation box="[597,610,1290,1311]" captionStart="Fig. 3" captionStartId="3.[145,172,1207,1227]" captionTargetBox="[145,814,133,1171]" captionTargetId="figure@3.[145,815,133,1171]" captionTargetPageId="3" captionText="Fig. 3. Esca of Lasiognathus dinema Pietsch and Sutton, new species, paratype, UW 154626, female, 36 mm SL, left lateral view, in preservation. Image courtesy of L. Malarky (NSUOC)." httpUri="https://zenodo.org/record/269575/files/figure.png" pageId="2">3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" pageId="2" type="description">
<paragraph blockId="2.[97,767,1352,2024]" lastBlockId="2.[815,1484,1201,1254]" pageId="2">
<emphasis bold="true" box="[97,246,1353,1374]" italics="true" pageId="2">Description.—</emphasis>
Length of illicium
<quantity box="[455,574,1352,1375]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="4.7" metricValueMin="1.5" pageId="2" unit="mm" value="31.0" valueMax="47.0" valueMin="15.0">1547 mm</quantity>
(
<number box="[588,635,1352,1375]" pageId="2" value="49.5">49.5</number>
50.0% SL); illicium of three known specimens fully extended anteriorly, cutaneous sheath surrounding posterior extension of illicial pterygiophore fully internalized (
<bibRefCitation author="Pietsch, T. W." box="[465,732,1440,1463]" journalOrPublisher="University of California Press, Berkeley and Los Angeles .." pageId="2" refString="Pietsch, T. W. 2009. Oceanic Anglerfishes: Extraordinary Diversity in the Deep-sea. University of California Press, Berkeley and Los Angeles .." title="Oceanic Anglerfishes: Extraordinary Diversity in the Deep-sea" type="book" year="2009">
Pietsch,
<number box="[558,618,1441,1462]" pageId="2" value="2009.0">2009</number>
:
<number box="[621,649,1441,1462]" pageId="2" value="84.0">84</number>
, fig.
<number box="[702,732,1440,1463]" pageId="2" value="85.0">85</number>
</bibRefCitation>
). A cylindrical, internally pigmented, anterior escal appendage (a light-guiding structure, without a homolog in any other known species of
<taxonomicName box="[299,436,1529,1551]" class="Actinopterygii" family="Thaumatichthyidae" genus="Lasiognathus" kingdom="Animalia" order="Lophiiformes" pageId="2" phylum="Chordata" rank="genus">
<emphasis box="[299,436,1529,1551]" italics="true" pageId="2">Lasiognathus</emphasis>
</taxonomicName>
;
<bibRefCitation author="Pietsch, T. W." box="[452,606,1529,1552]" journalOrPublisher="University of California Press, Berkeley and Los Angeles .." pageId="2" refString="Pietsch, T. W. 2009. Oceanic Anglerfishes: Extraordinary Diversity in the Deep-sea. University of California Press, Berkeley and Los Angeles .." title="Oceanic Anglerfishes: Extraordinary Diversity in the Deep-sea" type="book" year="2009">
Pietsch,
<number box="[547,606,1530,1551]" pageId="2" value="2009.0">2009</number>
</bibRefCitation>
:
<number box="[609,652,1530,1551]" pageId="2" value="237.0">237</number>
, figs.
<number pageId="2" value="245248.0">245, 248</number>
), rounded distally in
<typeStatus box="[416,519,1558,1581]" pageId="2">holotype</typeStatus>
, more elongate and tapering to a point in
<typeStatus box="[360,469,1589,1610]" pageId="2">paratypes</typeStatus>
, length
<quantity box="[566,702,1589,1610]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.8" metricValueMin="1.3" pageId="2" unit="mm" value="2.05" valueMax="2.8" valueMin="1.3">1.32.8 mm</quantity>
(
<number box="[719,752,1589,1610]" pageId="2" value="4.3">4.3</number>
<number box="[97,131,1618,1639]" pageId="2" value="2.9">2.9</number>
% SL). A thin, translucent, broad-based, antero-posteriorly compressed, posterior escal appendage, length
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="2.2" metricValueMin="0.9" pageId="2" unit="mm" value="1.55" valueMax="2.2" valueMin="0.9">0.92.2 mm</quantity>
(3.0
<number box="[214,247,1677,1698]" pageId="2" value="2.3">2.3</number>
% SL), with a scalloped or rounded distal margin, emerging from below escal pore. Distal escal appendage cylindrical, with narrow base and distal swelling, length
<quantity box="[185,326,1766,1787]" metricMagnitude="-3" metricUnit="m" metricValue="3.3499999999999996" metricValueMax="4.8" metricValueMin="1.9" pageId="2" unit="mm" value="3.3499999999999996" valueMax="4.8" valueMin="1.9">1.94.8 mm</quantity>
(
<number box="[348,381,1765,1788]" pageId="2" value="6.3">6.3</number>
5.0% SL), bearing three lightly pigmented, strongly curved, bony hooks: one somewhat larger, length 2.0
<quantity box="[313,404,1825,1846]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="2" unit="mm" value="2.2">2.2 mm</quantity>
(
<number box="[425,458,1824,1847]" pageId="2" value="6.7">6.7</number>
<number box="[472,506,1824,1847]" pageId="2" value="2.2">2.2</number>
% SL), emerging from distal tip, directed posteriorly; and a pair of near equal size, length
<number box="[181,215,1883,1906]" pageId="2" value="1.4">1.4</number>
/
<number box="[224,258,1883,1906]" pageId="2" value="1.6">1.6</number>
<number box="[273,307,1883,1906]" pageId="2" value="1.4">1.4</number>
/
<quantity box="[316,406,1883,1906]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="2" unit="mm" value="1.9">1.9 mm</quantity>
(5.0/
<number box="[468,502,1883,1906]" pageId="2" value="5.3">5.3</number>
<number box="[517,549,1883,1906]" pageId="2" value="1.9">1.9</number>
/2.0% SL), situated somewhat below, directed anteriorly. A pair of elongate, filamentous, tapering prolongations of distal appendage emerging slightly above bases of anteriorly directed escal hooks, considerably smaller in
<typeStatus box="[440,549,2002,2023]" pageId="2">paratypes</typeStatus>
, length
<number box="[638,671,2001,2024]" pageId="2" value="1.6">1.6</number>
/
<quantity box="[680,766,2001,2024]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="2" unit="mm" value="2.2">2.2 mm</quantity>
(
<number box="[822,857,1201,1224]" pageId="2" value="6.7">6.7</number>
/
<number box="[866,901,1201,1224]" pageId="2" value="7.3">7.3</number>
% SL); more highly developed in
<typeStatus box="[1294,1397,1201,1224]" pageId="2">holotype</typeStatus>
, length
<number box="[815,849,1231,1254]" pageId="2" value="6.3">6.3</number>
/
<quantity box="[857,946,1231,1254]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="2" unit="mm" value="6.4">6.4 mm</quantity>
(
<number box="[961,996,1231,1254]" pageId="2" value="6.6">6.6</number>
/
<number box="[1005,1040,1231,1254]" pageId="2" value="6.7">6.7</number>
% SL).
</paragraph>
<caption httpUri="https://zenodo.org/record/269574/files/figure.png" pageId="2" targetBox="[815,1484,1302,1860]" targetPageId="2">
<paragraph blockId="2.[815,1485,1897,2023]" pageId="2">
Fig. 2. Escae of
<taxonomicName authority="Pietsch and Sutton" authorityName="Pietsch and Sutton" box="[970,1356,1897,1917]" class="Actinopterygii" family="Thaumatichthyidae" genus="Lasiognathus" kingdom="Animalia" order="Lophiiformes" pageId="2" phylum="Chordata" rank="species" species="dinema" status="new species">
<emphasis bold="true" box="[970,1174,1897,1917]" italics="true" pageId="2">Lasiognathus dinema</emphasis>
Pietsch and Sutton
</taxonomicName>
,
<taxonomicNameLabel box="[1365,1479,1898,1917]" pageId="2" rank="species">new species</taxonomicNameLabel>
, in preservation: (A) Holotype, UW 152603, female, 95 mm SL, left lateral view; (B) Paratype, UW 152604, female, 30 mm SL, right lateral view (prolongation of distal escal appendage rotated to show anterior face).
</paragraph>
</caption>
<caption httpUri="https://zenodo.org/record/269575/files/figure.png" pageId="3" targetBox="[145,814,133,1171]" targetPageId="3">
<paragraph blockId="3.[145,815,1207,1280]" pageId="3">
Fig. 3. Esca of
<taxonomicName authority="Pietsch and Sutton" authorityName="Pietsch and Sutton" box="[292,683,1207,1227]" class="Actinopterygii" family="Thaumatichthyidae" genus="Lasiognathus" kingdom="Animalia" order="Lophiiformes" pageId="3" phylum="Chordata" rank="species" species="dinema" status="new species">
<emphasis bold="true" box="[292,499,1207,1227]" italics="true" pageId="3">Lasiognathus dinema</emphasis>
Pietsch and Sutton
</taxonomicName>
,
<taxonomicNameLabel box="[694,810,1207,1227]" pageId="3" rank="species">new species</taxonomicNameLabel>
, paratype, UW 154626, female, 36 mm SL, left lateral view, in preservation. Image courtesy of L. Malarky (NSUOC).
</paragraph>
</caption>
<paragraph blockId="3.[145,815,1320,1903]" pageId="3">
The esca of all three
<typeStatus box="[429,478,1321,1342]" pageId="3">type</typeStatus>
specimens is well preserved (
<figureCitation box="[153,227,1349,1372]" captionStart="Fig. 2" captionStartId="2.[815,842,1897,1917]" captionTargetBox="[815,1484,1302,1860]" captionTargetId="figure@2.[815,1485,1302,1862]" captionTargetPageId="2" captionText="Fig. 2. Escae of Lasiognathus dinema Pietsch and Sutton, new species, in preservation: (A) Holotype, UW 152603, female, 95 mm SL, left lateral view; (B) Paratype, UW 152604, female, 30 mm SL, right lateral view (prolongation of distal escal appendage rotated to show anterior face)." httpUri="https://zenodo.org/record/269574/files/figure.png" pageId="3">Figs. 2</figureCitation>
,
<figureCitation box="[239,254,1350,1372]" captionStart="Fig. 3" captionStartId="3.[145,172,1207,1227]" captionTargetBox="[145,814,133,1171]" captionTargetId="figure@3.[145,815,133,1171]" captionTargetPageId="3" captionText="Fig. 3. Esca of Lasiognathus dinema Pietsch and Sutton, new species, paratype, UW 154626, female, 36 mm SL, left lateral view, in preservation. Image courtesy of L. Malarky (NSUOC)." httpUri="https://zenodo.org/record/269575/files/figure.png" pageId="3">3</figureCitation>
). The head and body of the
<typeStatus box="[574,677,1349,1372]" pageId="3">holotype</typeStatus>
, however, is in extremely poor condition, so badly damaged upon capture that only a few of the standard ceratioid measurements can be made (the following are given in percent of standard length, values for the smaller
<typeStatus box="[576,673,1468,1489]" pageId="3">paratype</typeStatus>
are followed by those of the
<typeStatus box="[316,416,1497,1520]" pageId="3">holotype</typeStatus>
when available): tip of upper jaw to posteriormost margin of preopercle
<number box="[550,599,1527,1548]" pageId="3" value="41.7">41.7</number>
?; tip of upper jaw to anteriormost margin of opercular opening
<number box="[733,784,1556,1579]" pageId="3" value="53.3">53.3</number>
?; distance between tips of sphenotic spines 25.0?; tip of sphenotic spine to posttemporal
<number box="[536,584,1615,1636]" pageId="3" value="13.7">13.7</number>
?; tip of sphenotic spine to tip of quadrate spine 60.0?; length of sphenotic spine 6.0
<number box="[264,299,1674,1697]" pageId="3" value="2.2">2.2</number>
; length of quadrate spine
<number box="[610,644,1674,1697]" pageId="3" value="5.8">5.8</number>
<number box="[659,693,1674,1697]" pageId="3" value="2.3">2.3</number>
; length of premaxilla
<number box="[270,318,1703,1726]" pageId="3" value="36.7">36.7</number>
<number box="[333,381,1703,1726]" pageId="3" value="35.8">35.8</number>
; length of lower jaw
<number box="[612,660,1704,1725]" pageId="3" value="34.7">34.7</number>
<number box="[674,723,1704,1725]" pageId="3" value="28.4">28.4</number>
; longest tooth in upper jaw
<number box="[360,408,1733,1756]" pageId="3" value="11.7">11.7</number>
<number box="[423,457,1733,1756]" pageId="3" value="6.3">6.3</number>
; longest tooth in lower jaw
<number box="[768,801,1733,1754]" pageId="3" value="4.7">4.7</number>
<number box="[145,184,1762,1785]" pageId="3" value="2.6">2.6</number>
. Total number of teeth on premaxillae
<number box="[693,764,1762,1785]" pageId="3" value="55.0" valueMax="86.0" valueMin="24.0">2486</number>
, on dentaries
<number box="[257,326,1792,1813]" pageId="3" value="60.5" valueMax="98.0" valueMin="23.0">2398</number>
; vomerine teeth absent; dorsal-fin rays
<number pageId="3" value="5.5" valueMax="6.0" valueMin="5.0">5 6</number>
; anal-fin rays
<number box="[333,346,1821,1844]" pageId="3" value="5.0">5</number>
<number box="[362,375,1821,1844]" pageId="3" value="5.0">5</number>
; pectoral-fin rays
<number box="[589,616,1822,1843]" pageId="3" value="18.0">18</number>
/
<number box="[625,652,1822,1843]" pageId="3" value="19.0">19</number>
<number box="[668,695,1822,1843]" pageId="3" value="19.0">19</number>
/
<number box="[705,732,1822,1843]" pageId="3" value="19.0">19</number>
; pelvic fins absent; caudal-fin rays
<number box="[445,459,1851,1872]" pageId="3" value="9.0">9</number>
<number box="[474,488,1851,1872]" pageId="3" value="9.0">9</number>
. Remaining description as for the genus (
<bibRefCitation author="Bertelsen, E. &amp; T. W. Pietsch" box="[270,582,1880,1903]" journalOrPublisher="Copeia" pageId="3" pagination="401 - 409" part="1996" refString="Bertelsen, E., and T. W. Pietsch. 1996. A revision of the deep-sea anglerfish genus Lasiognathus (Lophiiformes: Thaumatichthyidae), with the description of a new species. Copeia 1996: 401 - 409." title="A revision of the deep-sea anglerfish genus Lasiognathus (Lophiiformes: Thaumatichthyidae), with the description of a new species" type="journal article" year="1996">
Bertelsen and Pietsch,
<number box="[525,582,1880,1903]" pageId="3" value="1996.0">1996</number>
</bibRefCitation>
;
<bibRefCitation author="Pietsch, T. W." box="[595,748,1880,1903]" journalOrPublisher="University of California Press, Berkeley and Los Angeles .." pageId="3" refString="Pietsch, T. W. 2009. Oceanic Anglerfishes: Extraordinary Diversity in the Deep-sea. University of California Press, Berkeley and Los Angeles .." title="Oceanic Anglerfishes: Extraordinary Diversity in the Deep-sea" type="book" year="2009">
Pietsch,
<number box="[689,748,1881,1903]" pageId="3" value="2009.0">2009</number>
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="3" type="reference_group">
<paragraph blockId="3.[145,815,1942,2024]" lastBlockId="3.[862,1532,136,248]" pageId="3">
<emphasis box="[145,646,1942,1965]" italics="true" pageId="3">
<emphasis bold="true" box="[145,425,1943,1965]" italics="true" pageId="3">Geographic distribution.—</emphasis>
<taxonomicName box="[425,646,1942,1964]" class="Actinopterygii" family="Thaumatichthyidae" genus="Lasiognathus" kingdom="Animalia" order="Lophiiformes" pageId="3" phylum="Chordata" rank="species" species="dinema">Lasiognathus dinema</taxonomicName>
</emphasis>
is known from three female specimens collected in the northern Gulf of
<collectingCountry box="[145,230,2001,2024]" name="Mexico" pageId="3">Mexico</collectingCountry>
, over the Mississippi Fan bathyal accumulation (
<number box="[870,938,136,157]" pageId="3" value="27.5" valueMax="28.0" valueMin="27.0">2728</number>
°N,
<number box="[984,1051,136,157]" pageId="3" value="88.5" valueMax="90.0" valueMin="87.0">8790</number>
°W; bottom depths
<quantity box="[1284,1443,136,158]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="3" unit="m" value="2500.0" valueMax="3000.0" valueMin="2000.0">20003000 m</quantity>
). Specimens were collected in closing trawls fished between
<number pageId="3" value="900.0" valueMax="1000.0" valueMin="800.0">800 1000</number>
and
<quantity box="[979,1135,196,217]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="3" unit="m" value="1100.0" valueMax="1200.0" valueMin="1000.0">10001200 m</quantity>
, and in a non-closing trawl fished between
<quantity box="[964,1078,225,247]" metricMagnitude="2" metricUnit="m" metricValue="6.3549999999999995" metricValueMax="12.709999999999999" metricValueMin="0.0" pageId="3" unit="m" value="635.5" valueMax="1271.0" valueMin="0.0">01271 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection pageId="3" type="etymology">
<paragraph blockId="3.[862,1532,287,429]" pageId="3">
<emphasis bold="true" box="[862,1003,287,309]" italics="true" pageId="3">Etymology.—</emphasis>
The name
<taxonomicName box="[1120,1197,287,309]" class="Actinopterygii" family="Thaumatichthyidae" genus="Lasiognathus" kingdom="Animalia" order="Lophiiformes" pageId="3" phylum="Chordata" rank="species" species="dinema">
<emphasis box="[1120,1197,287,309]" italics="true" pageId="3">dinema</emphasis>
</taxonomicName>
, is derived from the Greek,
<emphasis box="[1505,1525,287,309]" italics="true" pageId="3">di</emphasis>
, a prefix meaning two, and
<emphasis box="[1195,1252,318,339]" italics="true" pageId="3">nema</emphasis>
, thread, in allusion to the two elongate, thread-like prolongations emerging anteriorly from the bases of the escal hooks of this species (
<figureCitation box="[870,970,406,429]" captionStart-0="Fig. 1" captionStart-1="Fig. 2" captionStart-2="Fig. 3" captionStartId-0="2.[97,124,1130,1150]" captionStartId-1="2.[815,842,1897,1917]" captionStartId-2="3.[145,172,1207,1227]" captionTargetBox-0="[97,1486,133,1093]" captionTargetBox-1="[815,1484,1302,1860]" captionTargetBox-2="[145,814,133,1171]" captionTargetId-0="figure@2.[97,1486,133,1095]" captionTargetId-1="figure@2.[815,1485,1302,1862]" captionTargetId-2="figure@3.[145,815,133,1171]" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Fig. 1. Lasiognathus dinema Pietsch and Sutton, new species, paratype, UW 152604, female, 30 mm SL." captionText-1="Fig. 2. Escae of Lasiognathus dinema Pietsch and Sutton, new species, in preservation: (A) Holotype, UW 152603, female, 95 mm SL, left lateral view; (B) Paratype, UW 152604, female, 30 mm SL, right lateral view (prolongation of distal escal appendage rotated to show anterior face)." captionText-2="Fig. 3. Esca of Lasiognathus dinema Pietsch and Sutton, new species, paratype, UW 154626, female, 36 mm SL, left lateral view, in preservation. Image courtesy of L. Malarky (NSUOC)." httpUri-0="https://zenodo.org/record/269573/files/figure.png" httpUri-1="https://zenodo.org/record/269574/files/figure.png" httpUri-2="https://zenodo.org/record/269575/files/figure.png" pageId="3">
Figs.
<number box="[928,970,406,428]" pageId="3" value="2.0" valueMax="3.0" valueMin="1.0">13</number>
</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="3" type="discussion">
<paragraph blockId="3.[862,1005,476,498]" box="[862,1005,476,498]" pageId="3">
<heading allCaps="true" bold="true" box="[862,1005,476,498]" fontSize="9" level="4" pageId="3" reason="2">
<emphasis bold="true" box="[862,1005,476,498]" pageId="3">DISCUSSION</emphasis>
</heading>
</paragraph>
<paragraph blockId="3.[862,1533,520,1880]" pageId="3">
The specimens for this study were collected as part of an ongoing Natural Resource Damage Assessment Process conducted by the National Oceanic and Atmospheric Administration (for more information, see http://www.darrp.noaa.gov/about/nrda.html) pursuant to the
<emphasis box="[910,1113,670,691]" italics="true" pageId="3">Deepwater Horizon</emphasis>
Oil Spill (
<collectionCode box="[1237,1338,669,691]" pageId="3">DWHOS</collectionCode>
). The stations at which the
<typeStatus box="[984,1033,699,720]" pageId="3">type</typeStatus>
specimens were collected were within a
<quantity metricMagnitude="5" metricUnit="m" metricValue="2.5" pageId="3" unit="km" value="250.0">250 km</quantity>
radius of the Macondo wellhead, at depths between
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="3" unit="m" value="1050.0" valueMax="1300.0" valueMin="800.0">800 1300 m</quantity>
. The depths coincided with the depths of massive horizontal intrusions of hydrocarbons during and after
<collectionCode box="[862,956,818,839]" pageId="3">DWHOS</collectionCode>
(~
<quantity box="[995,1138,818,839]" metricMagnitude="3" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="3" unit="m" value="1000.0" valueMax="1200.0" valueMin="800.0">8001200 m</quantity>
;
<bibRefCitation author="Camilli, R. &amp; C. M. Reddy &amp; D. R. Yoerger &amp; B. A. S. Van Mooy &amp; M. V. Jakuba &amp; J. C. Kinsey &amp; C. P. McIntyre &amp; S. P. Sylva &amp; J. V. Maloney" box="[1153,1378,817,840]" journalOrPublisher="Science" pageId="3" pagination="201 - 204" part="330" refString="Camilli, R., C. M. Reddy, D. R. Yoerger, B. A. S. Van Mooy, M. V. Jakuba, J. C. Kinsey, C. P. McIntyre, S. P. Sylva, and J. V. Maloney. 2010. Tracking hydrocarbon plume transport and biodegradation at Deepwater Horizon. Science 330: 201 - 204." title="Tracking hydrocarbon plume transport and biodegradation at Deepwater Horizon" type="journal article" year="2010">
Camilli et al.,
<number box="[1320,1378,818,839]" pageId="3" value="2010.0">2010</number>
</bibRefCitation>
;
<bibRefCitation author="Valentine, D. L. &amp; J. D. Kessler &amp; M. C. Redmond &amp; S. D. Mendes &amp; M. B. Heintz &amp; C. Farwell &amp; L. Hu &amp; F. S. Kinnaman &amp; S. Yvon-Lewis &amp; M. Du &amp; E. W. Chan &amp; F. Garcia Tigreros &amp; C. J. Villanueva" journalOrPublisher="Science" pageId="3" pagination="208 - 211" part="330" refString="Valentine, D. L., J. D. Kessler, M. C. Redmond, S. D. Mendes, M. B. Heintz, C. Farwell, L. Hu, F. S. Kinnaman, S. Yvon-Lewis, M. Du, E. W. Chan, F. Garcia Tigreros, and C. J. Villanueva. 2010. Propane respiration jumpstarts microbial response to a deep oil spill. Science 330: 208 - 211." title="Propane respiration jumpstarts microbial response to a deep oil spill" type="journal article" year="2010">
Valentine et al.,
<number box="[913,972,848,869]" pageId="3" value="2010.0">2010</number>
</bibRefCitation>
;
<bibRefCitation author="Kessler, J. D. &amp; D. L. Valentine &amp; M. C. Redmond &amp; M. R. Du &amp; E. W. Chan &amp; S. D. Mendes &amp; E. W. Quiroz &amp; C. J. Villanueva &amp; S. S. Shusta &amp; L. M. Werra &amp; S. A. Yvon-Lewis &amp; T. C. Weber" box="[992,1233,847,870]" journalOrPublisher="Science" pageId="3" pagination="312 - 315" part="331" refString="Kessler, J. D., D. L. Valentine, M. C. Redmond, M. R. Du, E. W. Chan, S. D. Mendes, E. W. Quiroz, C. J. Villanueva, S. S. Shusta, L. M. Werra, S. A. Yvon-Lewis, and T. C. Weber. 2011. A persistent oxygen anomaly reveals the fate of spilled methane in the deep Gulf of Mexico. Science 331: 312 - 315." title="A persistent oxygen anomaly reveals the fate of spilled methane in the deep Gulf of Mexico" type="journal article" year="2011">
Kessler et al.,
<number box="[1174,1233,848,870]" pageId="3" value="2011.0">2011</number>
</bibRefCitation>
), though the specimens described herein were not necessarily exposed. The finding of a new species in this region adds to our knowledge of the deep-pelagic Gulf in particular, and to the bathypelagic zone in general, which post-Census of Marine Life surveys have identified as the most chronically under-sampled environment on earth (
<bibRefCitation author="Webb, T. J. &amp; E. V. Berghe &amp; R. O'Dor" box="[1037,1237,1025,1048]" journalOrPublisher="PLOS ONE 5" pageId="3" pagination="10223" part="8" refString="Webb, T. J., E. V. Berghe, and R. O'Dor. 2010. Biodiversity's big wet secret: the global distribution of marine biological records reveals chronic under-exploration of the deep pelagic ocean. PLOS ONE 5 (8): e 10223." title="Biodiversity's big wet secret: the global distribution of marine biological records reveals chronic under-exploration of the deep pelagic ocean" type="journal article" year="2010">
Webb et al.,
<number box="[1178,1237,1026,1048]" pageId="3" value="2010.0">2010</number>
</bibRefCitation>
). In addition to describing a new species, this report is the first record of the genus
<taxonomicName box="[862,999,1084,1106]" class="Actinopterygii" family="Thaumatichthyidae" genus="Lasiognathus" kingdom="Animalia" order="Lophiiformes" pageId="3" phylum="Chordata" rank="genus">
<emphasis box="[862,999,1084,1106]" italics="true" pageId="3">Lasiognathus</emphasis>
</taxonomicName>
in the Gulf.
</paragraph>
<paragraph blockId="3.[862,1533,520,1880]" pageId="3">
Like the majority of ceratioid anglerfishes, examination of all available material of
<taxonomicName box="[1176,1318,1144,1166]" class="Actinopterygii" family="Thaumatichthyidae" genus="Lasiognathus" kingdom="Animalia" order="Lophiiformes" pageId="3" phylum="Chordata" rank="genus">
<emphasis box="[1176,1318,1144,1166]" italics="true" pageId="3">Lasiognathus</emphasis>
</taxonomicName>
has revealed no characters that allow separation of the species other than those of the esca and its appendages (
<bibRefCitation author="Bertelsen, E. &amp; T. W. Pietsch" journalOrPublisher="Copeia" pageId="3" pagination="401 - 409" part="1996" refString="Bertelsen, E., and T. W. Pietsch. 1996. A revision of the deep-sea anglerfish genus Lasiognathus (Lophiiformes: Thaumatichthyidae), with the description of a new species. Copeia 1996: 401 - 409." title="A revision of the deep-sea anglerfish genus Lasiognathus (Lophiiformes: Thaumatichthyidae), with the description of a new species" type="journal article" year="1996">
Bertelsen and Pietsch,
<number box="[862,919,1234,1257]" pageId="3" value="1996.0">1996</number>
</bibRefCitation>
;
<bibRefCitation author="Pietsch, T. W." box="[936,1092,1234,1257]" journalOrPublisher="Copeia" pageId="3" pagination="77 - 81" part="2005" refString="Pietsch, T. W. 2005. A new species of the ceratioid anglerfish genus Lasiognathus Regan (Lophiiformes: Thaumatichthyidae) from the Eastern North Atlantic off Madeira. Copeia 2005: 77 - 81." title="A new species of the ceratioid anglerfish genus Lasiognathus Regan (Lophiiformes: Thaumatichthyidae) from the Eastern North Atlantic off Madeira" type="journal article" year="2005">Pietsch, 2005</bibRefCitation>
,
<bibRefCitation author="Pietsch, T. W." box="[1108,1167,1234,1256]" journalOrPublisher="University of California Press, Berkeley and Los Angeles .." pageId="3" refString="Pietsch, T. W. 2009. Oceanic Anglerfishes: Extraordinary Diversity in the Deep-sea. University of California Press, Berkeley and Los Angeles .." title="Oceanic Anglerfishes: Extraordinary Diversity in the Deep-sea" type="book" year="2009">2009</bibRefCitation>
). The differences in escal morphology, however, now strongly indicate the existence of six species.
<taxonomicName box="[964,1197,1293,1315]" class="Actinopterygii" family="Thaumatichthyidae" genus="Lasiognathus" kingdom="Animalia" order="Lophiiformes" pageId="3" phylum="Chordata" rank="species" species="dinema">
<emphasis box="[964,1197,1293,1315]" italics="true" pageId="3">Lasiognathus dinema</emphasis>
</taxonomicName>
differs from all previously recognized species of the genus in having an anterior escal appendage and a pair of elongate, filamentous prolongations of the distal escal appendage, the former having no apparent homolog in any of its congeners; the latter represented by a single prolongation in
<taxonomicName box="[1341,1491,1443,1464]" class="Actinopterygii" family="Thaumatichthyidae" genus="Lasiognathus" kingdom="Animalia" order="Lophiiformes" pageId="3" phylum="Chordata" rank="species" species="saccostoma">
<emphasis box="[1341,1491,1443,1464]" italics="true" pageId="3">L. saccostoma</emphasis>
</taxonomicName>
,
<taxonomicName class="Actinopterygii" family="Thaumatichthyidae" genus="Lasiognathus" kingdom="Animalia" order="Lophiiformes" pageId="3" phylum="Chordata" rank="species" species="intermedius">
<emphasis italics="true" pageId="3">L. intermedius</emphasis>
</taxonomicName>
, and
<taxonomicName box="[1048,1247,1471,1493]" class="Actinopterygii" family="Thaumatichthyidae" genus="Lasiognathus" kingdom="Animalia" order="Lophiiformes" pageId="3" phylum="Chordata" rank="species" species="amphirhamphus">
<emphasis box="[1048,1247,1471,1493]" italics="true" pageId="3">L. amphirhamphus</emphasis>
</taxonomicName>
, but absent altogether in
<taxonomicName box="[862,953,1501,1523]" class="Actinopterygii" family="Thaumatichthyidae" genus="Lasiognathus" kingdom="Animalia" order="Lophiiformes" pageId="3" phylum="Chordata" rank="species" species="beebei">
<emphasis box="[862,953,1501,1523]" italics="true" pageId="3">L. beebei</emphasis>
</taxonomicName>
and
<taxonomicName box="[1010,1116,1501,1523]" class="Actinopterygii" family="Thaumatichthyidae" genus="Lasiognathus" kingdom="Animalia" order="Lophiiformes" pageId="3" phylum="Chordata" rank="species" species="waltoni">
<emphasis box="[1010,1116,1501,1523]" italics="true" pageId="3">L. waltoni</emphasis>
</taxonomicName>
. The new species differs further from
<taxonomicName box="[862,955,1530,1553]" class="Actinopterygii" family="Thaumatichthyidae" genus="Lasiognathus" kingdom="Animalia" order="Lophiiformes" pageId="3" phylum="Chordata" rank="species" species="beebei">
<emphasis box="[862,955,1530,1553]" italics="true" pageId="3">L. beebei</emphasis>
</taxonomicName>
in having the escal hooks emerging from a cylindrical distal appendage, rather than from a transverse, fanshaped distal appendage (
<bibRefCitation author="Bertelsen, E. &amp; T. W. Pietsch" journalOrPublisher="Copeia" pageId="3" pagination="401 - 409" part="1996" refString="Bertelsen, E., and T. W. Pietsch. 1996. A revision of the deep-sea anglerfish genus Lasiognathus (Lophiiformes: Thaumatichthyidae), with the description of a new species. Copeia 1996: 401 - 409." title="A revision of the deep-sea anglerfish genus Lasiognathus (Lophiiformes: Thaumatichthyidae), with the description of a new species" type="journal article" year="1996">
Bertelsen and Pietsch,
<number box="[1422,1479,1590,1613]" pageId="3" value="1996.0">1996</number>
:
<number box="[1483,1528,1590,1613]" pageId="3" value="404.0">404</number>
, fig.
<number box="[906,921,1620,1642]" pageId="3" value="2.0">2</number>
</bibRefCitation>
); from
<taxonomicName box="[1006,1156,1620,1642]" class="Actinopterygii" family="Thaumatichthyidae" genus="Lasiognathus" kingdom="Animalia" order="Lophiiformes" pageId="3" phylum="Chordata" rank="species" species="intermedius">
<emphasis box="[1006,1156,1620,1642]" italics="true" pageId="3">L. intermedius</emphasis>
</taxonomicName>
in having a broad-based, anteroposteriorly compressed posterior escal appendage instead of a cylindrical appendage (
<bibRefCitation author="Bertelsen, E. &amp; T. W. Pietsch" journalOrPublisher="Copeia" pageId="3" pagination="401 - 409" part="1996" refString="Bertelsen, E., and T. W. Pietsch. 1996. A revision of the deep-sea anglerfish genus Lasiognathus (Lophiiformes: Thaumatichthyidae), with the description of a new species. Copeia 1996: 401 - 409." title="A revision of the deep-sea anglerfish genus Lasiognathus (Lophiiformes: Thaumatichthyidae), with the description of a new species" type="journal article" year="1996">
Bertelsen and Pietsch,
<number box="[1422,1479,1679,1702]" pageId="3" value="1996.0">1996</number>
:
<number box="[1483,1528,1679,1702]" pageId="3" value="406.0">406</number>
, fig.
<number box="[906,922,1709,1731]" pageId="3" value="4.0">4</number>
</bibRefCitation>
); from
<taxonomicName box="[1005,1150,1710,1731]" class="Actinopterygii" family="Thaumatichthyidae" genus="Lasiognathus" kingdom="Animalia" order="Lophiiformes" pageId="3" phylum="Chordata" rank="species" species="saccostoma">
<emphasis box="[1005,1150,1710,1731]" italics="true" pageId="3">L. saccostoma</emphasis>
</taxonomicName>
in having simple distal prolongations of the distal escal appendage instead of bearing numerous lateral serrations or filaments (
<bibRefCitation author="Bertelsen, E. &amp; T. W. Pietsch" journalOrPublisher="Copeia" pageId="3" pagination="401 - 409" part="1996" refString="Bertelsen, E., and T. W. Pietsch. 1996. A revision of the deep-sea anglerfish genus Lasiognathus (Lophiiformes: Thaumatichthyidae), with the description of a new species. Copeia 1996: 401 - 409." title="A revision of the deep-sea anglerfish genus Lasiognathus (Lophiiformes: Thaumatichthyidae), with the description of a new species" type="journal article" year="1996">
Bertelsen and Pietsch,
<number box="[961,1018,1798,1821]" pageId="3" value="1996.0">1996</number>
:
<number box="[1022,1066,1798,1821]" pageId="3" value="407.0">407</number>
, fig.
<number box="[1131,1146,1798,1821]" pageId="3" value="5.0">5</number>
</bibRefCitation>
); and from
<taxonomicName box="[1293,1497,1798,1820]" class="Actinopterygii" family="Thaumatichthyidae" genus="Lasiognathus" kingdom="Animalia" order="Lophiiformes" pageId="3" phylum="Chordata" rank="species" species="amphirhamphus">
<emphasis box="[1293,1497,1798,1820]" italics="true" pageId="3">L. amphirhamphus</emphasis>
</taxonomicName>
in having three escal hooks instead of only two (
<bibRefCitation author="Pietsch, T. W." journalOrPublisher="Copeia" pageId="3" pagination="77 - 81" part="2005" refString="Pietsch, T. W. 2005. A new species of the ceratioid anglerfish genus Lasiognathus Regan (Lophiiformes: Thaumatichthyidae) from the Eastern North Atlantic off Madeira. Copeia 2005: 77 - 81." title="A new species of the ceratioid anglerfish genus Lasiognathus Regan (Lophiiformes: Thaumatichthyidae) from the Eastern North Atlantic off Madeira" type="journal article" year="2005">
Pietsch,
<number box="[862,921,1857,1880]" pageId="3" value="2005.0">2005</number>
:
<number box="[925,954,1857,1880]" pageId="3" value="78.0">78</number>
, fig.
<number box="[1010,1026,1858,1880]" pageId="3" value="2.0">2</number>
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>