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<document id="852967D546D3B584CC397BAE41F0CDC9" ID-CLB-Dataset="26131" ID-DOI="10.11646/zootaxa.4646.2.4" ID-GBIF-Dataset="e1a3cc15-4c28-478d-a067-52c7f9224b6a" ID-ISSN="1175-5326" ID-Zenodo-Dep="3349449" ID-ZooBank="E031D297-36C5-46D3-A7B0-7355719AAD64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1563947505712" checkinUser="plazi" docAuthor="Santanna, Manoela, Rodrigues, Everton Nei Lopes, Cizauskas, Igor &amp; Brescovit, Antonio Domingos" docDate="2019" docId="03D80F63FFB5FFA5FF0F7E45FD0CF810" docLanguage="en" docName="zootaxa.4646.2.4.pdf" docOrigin="Zootaxa 4646 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cryptachaea Archer 1946" docType="treatment" docVersion="12" lastPageNumber="272" masterDocId="FFE1771BFFB4FFA4FF987B78FFECFF82" masterDocTitle="On the spider genus Cryptachaea from Peru, Bolivia and cave environments in Brazil: a new species, additional descriptions and new records (Araneae, Theridiidae)" masterLastPageNumber="292" masterPageNumber="271" pageNumber="272" updateTime="1698747171170" updateUser="ExternalLinkService">
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<mods:title id="6DEC7FE15AF88D8E9AFA7EC888E389EE">On the spider genus Cryptachaea from Peru, Bolivia and cave environments in Brazil: a new species, additional descriptions and new records (Araneae, Theridiidae)</mods:title>
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<mods:namePart id="80622D1CB02C6D7982880B7EF047324A">Santanna, Manoela</mods:namePart>
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<mods:namePart id="D1110B38C9364B2618420831A4A0DE33">Rodrigues, Everton Nei Lopes</mods:namePart>
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<mods:namePart id="D8A3E6BAC8AF03C6FF464575357075DC">Cizauskas, Igor</mods:namePart>
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<mods:namePart id="DAF51825C7275112DAB302C6D4DDB820">Brescovit, Antonio Domingos</mods:namePart>
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<mods:title id="3FB6998D2E50563455E88B30FB7C7387">Zootaxa</mods:title>
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<mods:date id="335B2BE03655E6C5BFAA7CD7F4362A94">2019</mods:date>
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<mods:number id="3FFD4D28F12245A5D1B91308B91F684B">2019-07-24</mods:number>
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<mods:identifier id="14048C571447A94C9E1783C78AAB90A3" type="DOI">10.11646/zootaxa.4646.2.4</mods:identifier>
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<treatment id="03D80F63FFB5FFA5FF0F7E45FD0CF810" ID-DOI="http://doi.org/10.5281/zenodo.5944246" ID-GBIF-Taxon="158852376" ID-Zenodo-Dep="5944246" LSID="urn:lsid:plazi:treatment:03D80F63FFB5FFA5FF0F7E45FD0CF810" httpUri="http://treatment.plazi.org/id/03D80F63FFB5FFA5FF0F7E45FD0CF810" lastPageNumber="272" pageId="1" pageNumber="272">
<subSubSection id="C36BEDFEFFB5FFA5FF0F7E45FE3AFAD5" box="[151,470,1341,1367]" pageId="1" pageNumber="272" type="nomenclature">
<paragraph id="8BCEBE75FFB5FFA5FF0F7E45FE3AFAD5" blockId="1.[151,470,1341,1367]" box="[151,470,1341,1367]" pageId="1" pageNumber="272">
<heading id="D0860919FFB5FFA5FF0F7E45FE3AFAD5" bold="true" box="[151,470,1341,1367]" fontSize="11" level="1" pageId="1" pageNumber="272" reason="1">
<taxonomicName id="4C71C5F6FFB5FFA5FF0F7E45FE3AFAD5" ID-CoL="62P4W" ID-ENA="653443" authority="Archer, 1946" authorityName="Archer" authorityYear="1946" box="[151,470,1341,1367]" class="Arachnida" family="Theridiidae" genus="Cryptachaea" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="B9056267FFB5FFA5FF0F7E45FE3AFAD5" bold="true" box="[151,470,1341,1367]" pageId="1" pageNumber="272">
<emphasis id="B9056267FFB5FFA5FF0F7E45FEDCFAD5" bold="true" box="[151,304,1341,1367]" italics="true" pageId="1" pageNumber="272">Cryptachaea</emphasis>
Archer, 1946
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36BEDFEFFB5FFA5FF0F7EF9FD0CF810" pageId="1" pageNumber="272" type="materials_examined">
<paragraph id="8BCEBE75FFB5FFA5FF0F7EF9FD63FA18" blockId="1.[151,655,1409,1434]" box="[151,655,1409,1434]" pageId="1" pageNumber="272">
<emphasis id="B9056267FFB5FFA5FF0F7EF9FECFFA18" bold="true" box="[151,291,1409,1434]" pageId="1" pageNumber="272">
<typeStatus id="54CA00D7FFB5FFA5FF0F7EF9FF20FA1B" box="[151,204,1409,1433]" pageId="1" pageNumber="272">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C71C5F6FFB5FFA5FEB27EF9FD63FA18" authority="(Banks, 1896)" baseAuthorityName="Banks" baseAuthorityYear="1896" box="[298,655,1409,1434]" class="Arachnida" family="Theridiidae" genus="Cryptachaea" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="272" phylum="Arthropoda" rank="species" species="porteri">
<emphasis id="B9056267FFB5FFA5FEB27EF9FE15FA1B" box="[298,505,1409,1433]" italics="true" pageId="1" pageNumber="272">Cryptachaea porteri</emphasis>
(Banks, 1896)
</taxonomicName>
</paragraph>
<paragraph id="8BCEBE75FFB5FFA5FF0F7EBFFCB4F95C" blockId="1.[151,1436,1479,1938]" pageId="1" pageNumber="272">
<emphasis id="B9056267FFB5FFA5FF0F7EBFFEFEFA60" bold="true" box="[151,274,1479,1506]" pageId="1" pageNumber="272">Diagnosis.</emphasis>
<taxonomicName id="4C71C5F6FFB5FFA5FE817EB0FE46FA60" authorityName="Archer" authorityYear="1946" box="[281,426,1480,1506]" class="Arachnida" family="Theridiidae" genus="Cryptachaea" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="B9056267FFB5FFA5FE817EB0FE46FA60" box="[281,426,1480,1506]" italics="true" pageId="1" pageNumber="272">Cryptachaea</emphasis>
</taxonomicName>
species lack the theridiid tegular apophysis, and as other members of the subfamily, also lack a colulus (
<bibRefCitation id="EFE0C384FFB5FFA5FEA67E94FDEDF984" author="Agnarsson, I." box="[318,513,1516,1542]" pageId="1" pageNumber="272" pagination="447 - 626" refId="ref17149" refString="Agnarsson, I. (2004) Morphological phylogeny of combweb spiders and their relatives (Araneae, Theridiidae). Zoological Journal of the Linnean Society, 141, 447 - 626. https: // doi. org / 10.1111 / j. 1096 - 3642.2004.00120. x" type="journal article" year="2004">Agnarsson, 2004</bibRefCitation>
;
<bibRefCitation id="EFE0C384FFB5FFA5FD937E94FD05F984" author="Buckup, E. H. &amp; Marques, M. A. L. &amp; Rodrigues, E. N. L." box="[523,745,1516,1542]" pageId="1" pageNumber="272" pagination="206 - 211" refId="ref17642" refString="Buckup, E. H., Marques, M. A. L. &amp; Rodrigues, E. N. L. (2012) Tres novas especies sul-americanas de Cryptachaea e acrescimos taxonomicos em Achaearanea (Araneae, Theridiidae). Iheringia, Serie Zoologia, 102, 206 - 211. https: // doi. org / 10.1590 / S 0073 - 47212012000200013" type="journal article" year="2012">
Buckup
<emphasis id="B9056267FFB5FFA5FDF07E94FD75F984" box="[616,665,1516,1542]" italics="true" pageId="1" pageNumber="272">et al</emphasis>
., 2012
</bibRefCitation>
;
<bibRefCitation id="EFE0C384FFB5FFA5FD6B7E94FBFEF984" author="Rodrigues, E. N. L. &amp; Poeta, M. R. M." box="[755,1042,1515,1542]" pageId="1" pageNumber="272" pagination="26 - 33" refId="ref18157" refString="Rodrigues, E. N. L. &amp; Poeta, M. R. M. (2015) Twelve new Neotropical species of the spider genus Cryptachaea (Araneae: Theridiidae). Journal of Arachnology, 43, 26 - 33. https: // doi. org / 10.1636 / K 14 - 47.1" type="journal article" year="2015">
<collectingRegion id="49B57097FFB5FFA5FD6B7E94FC8AF984" box="[755,870,1516,1542]" country="Mauritius" name="Rodrigues" pageId="1" pageNumber="272">Rodrigues</collectingRegion>
&amp; Poeta, 2015
</bibRefCitation>
). The genus is similar to
<taxonomicName id="4C71C5F6FFB5FFA5FABE7E94FEB6F9A8" authority="Strand, 1929" authorityName="Strand" authorityYear="1929" class="Arachnida" family="Theridiidae" genus="Achaearanea" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="B9056267FFB5FFA5FABE7E94FF53F9AB" italics="true" pageId="1" pageNumber="272">Achaearanea</emphasis>
Strand, 1929
</taxonomicName>
,
<taxonomicName id="4C71C5F6FFB5FFA5FEFD7D68FD77F9AB" authority="Archer, 1946" authorityName="Archer" authorityYear="1946" box="[357,667,1551,1578]" class="Arachnida" family="Theridiidae" genus="Hentziectypus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="B9056267FFB5FFA5FEFD7D68FDEEF9AB" box="[357,514,1552,1577]" italics="true" pageId="1" pageNumber="272">Hentziectypus</emphasis>
Archer, 1946
</taxonomicName>
and
<taxonomicName id="4C71C5F6FFB5FFA5FD4B7D68FBE9F9AB" authority="Archer, 1946" authorityName="Archer" authorityYear="1946" box="[723,1029,1551,1578]" class="Arachnida" family="Theridiidae" genus="Parasteatoda" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="B9056267FFB5FFA5FD4B7D68FC86F9A8" box="[723,874,1552,1578]" italics="true" pageId="1" pageNumber="272">Parasteatoda</emphasis>
Archer, 1946
</taxonomicName>
, however can be distinguished from these genera by the male palp with a medium to short embolus, which is not articulated and not supported on the back of the cymbium; the base is fused to the median apophysis lodged in the cymbial hood (
<bibRefCitation id="EFE0C384FFB5FFA5FB4B7D20FA74F9F0" author="Agnarsson, I." box="[1235,1432,1624,1650]" pageId="1" pageNumber="272" pagination="447 - 626" refId="ref17149" refString="Agnarsson, I. (2004) Morphological phylogeny of combweb spiders and their relatives (Araneae, Theridiidae). Zoological Journal of the Linnean Society, 141, 447 - 626. https: // doi. org / 10.1111 / j. 1096 - 3642.2004.00120. x" type="journal article" year="2004">Agnarsson, 2004</bibRefCitation>
;
<bibRefCitation id="EFE0C384FFB5FFA5FF0F7D04FED6F914" author="Yoshida, H." box="[151,314,1660,1686]" pageId="1" pageNumber="272" pagination="37 - 40" refId="ref18378" refString="Yoshida, H. (2008) A revision of the genus Achaearanea (Araneae: Theridiidae). Acta Arachnologica, 1, 37 - 40. https: // doi. org / 10.2476 / asjaa. 57.37" type="journal article" year="2008">Yoshida, 2008</bibRefCitation>
;
<bibRefCitation id="EFE0C384FFB5FFA5FEDD7D04FDCFF914" author="Buckup, E. H. &amp; Marques, M. A. L. &amp; Rodrigues, E. N. L." box="[325,547,1660,1686]" pageId="1" pageNumber="272" pagination="206 - 211" refId="ref17642" refString="Buckup, E. H., Marques, M. A. L. &amp; Rodrigues, E. N. L. (2012) Tres novas especies sul-americanas de Cryptachaea e acrescimos taxonomicos em Achaearanea (Araneae, Theridiidae). Iheringia, Serie Zoologia, 102, 206 - 211. https: // doi. org / 10.1590 / S 0073 - 47212012000200013" type="journal article" year="2012">
Buckup
<emphasis id="B9056267FFB5FFA5FE3A7D04FE36F914" box="[418,474,1660,1686]" italics="true" pageId="1" pageNumber="272">et al.</emphasis>
, 2012
</bibRefCitation>
;
<bibRefCitation id="EFE0C384FFB5FFA5FDB57D04FCA7F914" author="Rodrigues, E. N. L. &amp; Poeta, M. R. M." box="[557,843,1659,1686]" pageId="1" pageNumber="272" pagination="26 - 33" refId="ref18157" refString="Rodrigues, E. N. L. &amp; Poeta, M. R. M. (2015) Twelve new Neotropical species of the spider genus Cryptachaea (Araneae: Theridiidae). Journal of Arachnology, 43, 26 - 33. https: // doi. org / 10.1636 / K 14 - 47.1" type="journal article" year="2015">
<collectingRegion id="49B57097FFB5FFA5FDB57D04FD4CF914" box="[557,672,1660,1686]" country="Mauritius" name="Rodrigues" pageId="1" pageNumber="272">Rodrigues</collectingRegion>
&amp; Poeta, 2015
</bibRefCitation>
). The epigynum of
<taxonomicName id="4C71C5F6FFB5FFA5FBB97D04FB5EF914" authorityName="Archer" authorityYear="1946" box="[1057,1202,1660,1686]" class="Arachnida" family="Theridiidae" genus="Cryptachaea" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="B9056267FFB5FFA5FBB97D04FB5EF914" box="[1057,1202,1660,1686]" italics="true" pageId="1" pageNumber="272">Cryptachaea</emphasis>
</taxonomicName>
has simple, uncoiled copulatory ducts leading to a pair of simple subcircular spermathecae and a slightly sclerotized epigynal plate (
<bibRefCitation id="EFE0C384FFB5FFA5FAF77DD8FEDBF95C" author="Agnarsson, I." pageId="1" pageNumber="272" pagination="447 - 626" refId="ref17149" refString="Agnarsson, I. (2004) Morphological phylogeny of combweb spiders and their relatives (Araneae, Theridiidae). Zoological Journal of the Linnean Society, 141, 447 - 626. https: // doi. org / 10.1111 / j. 1096 - 3642.2004.00120. x" type="journal article" year="2004">Agnarsson, 2004</bibRefCitation>
;
<bibRefCitation id="EFE0C384FFB5FFA5FEDA7DBCFDCEF95C" author="Buckup, E. H. &amp; Marques, M. A. L. &amp; Rodrigues, E. N. L." box="[322,546,1732,1758]" pageId="1" pageNumber="272" pagination="206 - 211" refId="ref17642" refString="Buckup, E. H., Marques, M. A. L. &amp; Rodrigues, E. N. L. (2012) Tres novas especies sul-americanas de Cryptachaea e acrescimos taxonomicos em Achaearanea (Araneae, Theridiidae). Iheringia, Serie Zoologia, 102, 206 - 211. https: // doi. org / 10.1590 / S 0073 - 47212012000200013" type="journal article" year="2012">
Buckup
<emphasis id="B9056267FFB5FFA5FE387DBCFE3DF95C" box="[416,465,1732,1758]" italics="true" pageId="1" pageNumber="272">et al</emphasis>
., 2012
</bibRefCitation>
;
<bibRefCitation id="EFE0C384FFB5FFA5FDB47DBCFCA0F95C" author="Rodrigues, E. N. L. &amp; Poeta, M. R. M." box="[556,844,1731,1758]" pageId="1" pageNumber="272" pagination="26 - 33" refId="ref18157" refString="Rodrigues, E. N. L. &amp; Poeta, M. R. M. (2015) Twelve new Neotropical species of the spider genus Cryptachaea (Araneae: Theridiidae). Journal of Arachnology, 43, 26 - 33. https: // doi. org / 10.1636 / K 14 - 47.1" type="journal article" year="2015">
<collectingRegion id="49B57097FFB5FFA5FDB47DBCFD73F95C" box="[556,671,1732,1758]" country="Mauritius" name="Rodrigues" pageId="1" pageNumber="272">Rodrigues</collectingRegion>
&amp; Poeta, 2015
</bibRefCitation>
).
</paragraph>
<paragraph id="8BCEBE75FFB5FFA5FF5F7D9FFD0CF810" blockId="1.[151,1436,1479,1938]" pageId="1" pageNumber="272">
<emphasis id="B9056267FFB5FFA5FF5F7D9FFDC2F880" bold="true" box="[199,558,1767,1794]" pageId="1" pageNumber="272">Composition and distribution.</emphasis>
90 species, including the new species described here.
<taxonomicName id="4C71C5F6FFB5FFA5FB1E7D90FAFBF880" authorityName="Archer" authorityYear="1946" box="[1158,1303,1768,1794]" class="Arachnida" family="Theridiidae" genus="Cryptachaea" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="B9056267FFB5FFA5FB1E7D90FAFBF880" box="[1158,1303,1768,1794]" italics="true" pageId="1" pageNumber="272">Cryptachaea</emphasis>
</taxonomicName>
includes 73 species exclusive from Neotropical Region, seven species exclusive to the Nearctic Region, five species exclusive to the Palearctic Region, three species exclusive to the Australian Region, one species shared between Neotropical Region and Nearctic Region, and one species shared between Neotropical Region, Nearctic Region, Palearctic Region, Afrotropical Region and Australian Region.
</paragraph>
</subSubSection>
</treatment>
</document>
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