514 lines
62 KiB
XML
514 lines
62 KiB
XML
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<mods:title id="FD809FA799F4488E71068D08A0B62B22">Contribution to the cockroach genus Ctenoneura Hanitsch, 1925 (Blattodea: Corydioidea: Corydiidae) with descriptions of seven new species from China</mods:title>
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<mods:namePart id="AE1D566F00DDA3E6521B7928D87FEADE">Qiu, Lu</mods:namePart>
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<mods:namePart id="3CDFC48098EEFE062DC82334AFEAA3BE">Che, Yan-Li</mods:namePart>
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<mods:namePart id="0EC8C623DFE06146CBD5B968951280FE">Wang, Zong-Qing</mods:namePart>
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<treatment id="03D2F661FFC8FF96658EFA45FE0E1DAD" ID-DOI="http://doi.org/10.5281/zenodo.6053077" ID-GBIF-Taxon="127710821" ID-Zenodo-Dep="6053077" LSID="urn:lsid:plazi:treatment:03D2F661FFC8FF96658EFA45FE0E1DAD" httpUri="http://treatment.plazi.org/id/03D2F661FFC8FF96658EFA45FE0E1DAD" lastPageId="6" lastPageNumber="271" pageId="3" pageNumber="268">
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<paragraph id="8BC44777FFC8FF93658EFA45FE231DEE" blockId="3.[151,488,1439,1466]" box="[151,488,1439,1466]" pageId="3" pageNumber="268">
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<taxonomicName id="4C7B3CF4FFC8FF93658EFA45FE231DEE" ID-CoL="8MRDJ" authority="Hanitsch, 1925" authorityName="Hanitsch" authorityYear="1925" box="[151,488,1439,1466]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="268" phylum="Arthropoda" rank="genus">
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<emphasis id="B90F9B65FFC8FF93658EFA45FE231DEE" bold="true" box="[151,488,1439,1466]" pageId="3" pageNumber="268">
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<emphasis id="B90F9B65FFC8FF93658EFA45FEED1DED" bold="true" box="[151,294,1439,1465]" italics="true" pageId="3" pageNumber="268">Ctenoneura</emphasis>
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<bibRefCitation id="EFEA3A86FFC8FF936434FA7AFE231DEE" author="Hanitsch" box="[301,488,1440,1466]" pageId="3" pageNumber="268" refString="Hanitsch, R. (1925) On a collection of Blattidae from northern Sarawak, chiefly Mt. Murud and Mt. Dulit. Sarawak Museum Journal, 3, 75 - 106." type="journal article" year="1925">Hanitsch, 1925</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C36114FCFFC8FF93658EFA3DFCEB1E4F" pageId="3" pageNumber="268" type="reference_group">
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<paragraph id="8BC44777FFC8FF93658EFA3DFCEB1E4F" blockId="3.[151,1436,1510,1594]" pageId="3" pageNumber="268">
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<treatmentCitationGroup id="AB6B6059FFC8FF93658EFA3DFCEB1E4F" pageId="3" pageNumber="268">
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<taxonomicName id="4C7B3CF4FFC8FF93658EFA3DFE3D1DA8" ID-CoL="8MRDJ" authority="Hanitsch, 1925: 100" authorityName="Hanitsch" authorityPageNumber="100" authorityYear="1925" box="[151,502,1510,1532]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="268" phylum="Arthropoda" rank="genus">
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<emphasis id="B90F9B65FFC8FF93658EFA3DFEC41DA8" box="[151,271,1511,1532]" italics="true" pageId="3" pageNumber="268">Ctenoneura</emphasis>
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<treatmentCitation id="0ADA6166FFC8FF936402FA3CFE3D1DA8" author="Hanitsch" box="[283,502,1510,1532]" page="100" pageId="3" pageNumber="268" year="1925">
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<bibRefCitation id="EFEA3A86FFC8FF936402FA3CFE761DA8" author="Hanitsch" box="[283,445,1510,1532]" pageId="3" pageNumber="268" refString="Hanitsch, R. (1925) On a collection of Blattidae from northern Sarawak, chiefly Mt. Murud and Mt. Dulit. Sarawak Museum Journal, 3, 75 - 106." type="journal article" year="1925">Hanitsch, 1925</bibRefCitation>
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: 100
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</treatmentCitation>
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</taxonomicName>
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;
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<treatmentCitation id="0ADA6166FFC8FF93671FFA3CFD031DA8" author="Princis" box="[518,712,1510,1532]" page="204" pageId="3" pageNumber="268" year="1950">
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<bibRefCitation id="EFEA3A86FFC8FF93671FFA3CFD441DA8" author="Princis" box="[518,655,1510,1532]" pageId="3" pageNumber="268" refString="Princis, K. (1950) Entomological results from the Swedish Expedition 1934 to Burma and British India, Blattaria. Arkiv for Zoologi, 1, 203 - 222." type="journal article" year="1950">Princis 1950</bibRefCitation>
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: 204
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||
</treatmentCitation>
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;
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||
<treatmentCitation id="0ADA6166FFC8FF9367C1FA3CFC521DA8" author="Princis" box="[728,921,1510,1532]" page="207" pageId="3" pageNumber="268" year="1953">
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<bibRefCitation id="EFEA3A86FFC8FF9367C1FA3CFCAA1DA8" author="Princis" box="[728,865,1510,1532]" pageId="3" pageNumber="268" refString="Princis, K. (1953) Kleine Beitrage zur Kenntnis der Blattarien und ihrer Verbreitung. VI. Entomologisk Tidskrift, 74, 203 - 213." type="journal article" year="1953">Princis 1953</bibRefCitation>
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: 207
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||
</treatmentCitation>
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;
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<treatmentCitation id="0ADA6166FFC8FF9366B3FA3CFBA31DA8" author="Princis" box="[938,1128,1510,1532]" page="101" pageId="3" pageNumber="268" year="1963">
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<bibRefCitation id="EFEA3A86FFC8FF9366B3FA3CFBF81DA8" author="Princis" box="[938,1075,1510,1532]" pageId="3" pageNumber="268" refString="Princis, K. (1963) Blattariae: Suborde Polyphagoidea: Fam.: Homoeogamiidae, Euthyrrhaphidae, Latindiidae, Anacompsidae, Atticolidae, Attaphilidae; Subordo Blaberoidea: Fam. Blaberidae. In: Beier, M. (Ed.), Orthopterorum Catalogus. Pars 4. Uitgeverij Dr. W. Junk, ' s-Gravenhage, pp. 77 - 172." type="journal article" year="1963">Princis 1963</bibRefCitation>
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: 101
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||
</treatmentCitation>
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(Catalogue);
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<treatmentCitation id="0ADA6166FFC8FF9361E6FA3CFA571DA8" author="Roth" box="[1279,1436,1510,1532]" page="83" pageId="3" pageNumber="268" year="1993">
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<bibRefCitation id="EFEA3A86FFC8FF9361E6FA3CFAB81DA8" author="Roth" box="[1279,1395,1510,1532]" pageId="3" pageNumber="268" refString="Roth, L. M. (1993) Revision of the cockroach genus Ctenoneura Hanitsch (Blattaria, Polyphagidae). Tijdschrift voor Entomologie, 136, 83 - 109." type="journal article" year="1993">Roth 1993</bibRefCitation>
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: 83
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</treatmentCitation>
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(Revision);
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||
<treatmentCitation id="0ADA6166FFC8FF936426F9DFFE231E4F" author="Roth" box="[319,488,1541,1563]" page="117" pageId="3" pageNumber="268" year="1995">
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<bibRefCitation id="EFEA3A86FFC8FF936426F9DFFE711E4F" author="Roth" box="[319,442,1541,1563]" pageId="3" pageNumber="268" refString="Roth, L. M. (1995 b) Description of a new species of Ctenoneura Hanitsch from Sabah (Blattaria, Polyphagidae). Tijdschrift voor Entomologie, 138, 117 - 119." type="journal article" year="1995" yearSuffix="b">Roth 1995b</bibRefCitation>
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: 117
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||
</treatmentCitation>
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||
(Rediagnosis);
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||
<treatmentCitation id="0ADA6166FFC8FF936790F9DFFCD71E4F" author="Roth" box="[649,796,1541,1563]" page="34" pageId="3" pageNumber="268" year="2003">
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<bibRefCitation id="EFEA3A86FFC8FF936790F9DFFD3D1E4F" author="Roth" box="[649,758,1541,1563]" pageId="3" pageNumber="268" refString="Roth, L. M. (2003) Systematics and phylogeny of cockroaches (Dictyoptera: Blattaria). Oriental Insects, 37, 1 - 186. https: // doi. org / 10.1080 / 00305316.2003.10417344" type="journal article" year="2003">Roth 2003</bibRefCitation>
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: 34
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||
</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36114FCFFC8FF93658EF9FFFC791E6E" box="[151,946,1572,1594]" pageId="3" pageNumber="268" type="materials_examined">
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<paragraph id="8BC44777FFC8FF93658EF9FFFC791E6E" blockId="3.[151,1436,1510,1594]" box="[151,946,1572,1594]" pageId="3" pageNumber="268">
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Type Species:
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<taxonomicName id="4C7B3CF4FFC8FF936433F9FEFD4F1E6E" authority="Hanitsch, 1925" authorityName="Hanitsch" authorityYear="1925" box="[298,644,1572,1594]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="268" phylum="Arthropoda" rank="species" species="major">
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<emphasis id="B90F9B65FFC8FF936433F9FEFE2F1E6E" box="[298,484,1572,1594]" italics="true" pageId="3" pageNumber="268">Ctenoneura major</emphasis>
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||
<bibRefCitation id="EFEA3A86FFC8FF9364F3F9FEFD4F1E6E" author="Hanitsch" box="[490,644,1572,1594]" pageId="3" pageNumber="268" refString="Hanitsch, R. (1925) On a collection of Blattidae from northern Sarawak, chiefly Mt. Murud and Mt. Dulit. Sarawak Museum Journal, 3, 75 - 106." type="journal article" year="1925">Hanitsch, 1925</bibRefCitation>
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</taxonomicName>
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||
(Designated by
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||
<bibRefCitation id="EFEA3A86FFC8FF936633F9FEFC601E6E" author="Princis" box="[810,939,1572,1594]" pageId="3" pageNumber="268" refString="Princis, K. (1950) Entomological results from the Swedish Expedition 1934 to Burma and British India, Blattaria. Arkiv for Zoologi, 1, 203 - 222." type="journal article" year="1950">Princis 1950</bibRefCitation>
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||
)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36114FCFFC8FF93658EF9BCFC961FCA" pageId="3" pageNumber="268" type="diagnosis">
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<paragraph id="8BC44777FFC8FF93658EF9BCFC961FCA" blockId="3.[151,1436,1638,2023]" pageId="3" pageNumber="268">
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||
<emphasis id="B90F9B65FFC8FF93658EF9BCFEDA1E2B" bold="true" box="[151,273,1638,1663]" pageId="3" pageNumber="268">Diagnosis.</emphasis>
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This genus is peculiar among the
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<taxonomicName id="4C7B3CF4FFC8FF9367D6F9BCFC841E2B" box="[719,847,1638,1663]" class="Insecta" family="Corydiidae" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="268" phylum="Arthropoda" rank="family">Corydiidae</taxonomicName>
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. The small, narrow and smooth body distinctly differentiates it from many other
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<taxonomicName id="4C7B3CF4FFC8FF93673BF950FD551EF7" box="[546,670,1674,1699]" class="Insecta" family="Corydiidae" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="268" phylum="Arthropoda" rank="family">Corydiidae</taxonomicName>
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genera whose body is pubescent and oval shaped.
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<bibRefCitation id="EFEA3A86FFC8FF936018F950FA571EF7" author="Roth" box="[1281,1436,1674,1699]" pageId="3" pageNumber="268" refString="Roth, L. M. (1995 b) Description of a new species of Ctenoneura Hanitsch from Sabah (Blattaria, Polyphagidae). Tijdschrift voor Entomologie, 138, 117 - 119." type="journal article" year="1995" yearSuffix="b">Roth (1995b)</bibRefCitation>
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||
considered this genus most resembles genus
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||
<taxonomicName id="4C7B3CF4FFC8FF93678DF974FC881E93" box="[660,835,1710,1735]" class="Insecta" family="Corydiidae" genus="Homopteroidea" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="268" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90F9B65FFC8FF93678DF974FC881E93" box="[660,835,1710,1735]" italics="true" pageId="3" pageNumber="268">Homopteroidea</emphasis>
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||
</taxonomicName>
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, a detailed generic comparison has been provided in
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<bibRefCitation id="EFEA3A86FFC8FF93658EF908FEF91EBF" author="Roth" box="[151,306,1746,1771]" pageId="3" pageNumber="268" refString="Roth, L. M. (1995 b) Description of a new species of Ctenoneura Hanitsch from Sabah (Blattaria, Polyphagidae). Tijdschrift voor Entomologie, 138, 117 - 119." type="journal article" year="1995" yearSuffix="b">Roth (1995b)</bibRefCitation>
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, we add the following comparison as supplement: 1) body smooth, not pubescent in
|
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<taxonomicName id="4C7B3CF4FFC8FF936008F909FA5E1EBE" box="[1297,1429,1747,1770]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="268" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B90F9B65FFC8FF936008F909FA5E1EBE" box="[1297,1429,1747,1770]" italics="true" pageId="3" pageNumber="268">Ctenoneura</emphasis>
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||
</taxonomicName>
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, while body smooth, with a few setae in
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<taxonomicName id="4C7B3CF4FFC8FF936740F92CFCC31F5B" box="[601,776,1782,1807]" class="Insecta" family="Corydiidae" genus="Homopteroidea" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="268" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90F9B65FFC8FF936740F92CFCC31F5B" box="[601,776,1782,1807]" italics="true" pageId="3" pageNumber="268">Homopteroidea</emphasis>
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||
</taxonomicName>
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||
; 2) tegmen with well-developed M, but simplified CuA in
|
||
<taxonomicName id="4C7B3CF4FFC8FF93658EF8C1FED71F66" box="[151,284,1819,1842]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="268" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90F9B65FFC8FF93658EF8C1FED71F66" box="[151,284,1819,1842]" italics="true" pageId="3" pageNumber="268">Ctenoneura</emphasis>
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||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13405BF2FFC8FF936432F8C0FEBB1F67" box="[299,368,1818,1843]" captionStart="FIGURE 2" captionStartId="3.[151,250,1268,1290]" captionTargetBox="[185,1430,874,1218]" captionTargetId="figure@3.[168,1453,802,1254]" captionTargetPageId="3" captionText="FIGURE 2. Tegmen and wing. Venation abbreviations see text." httpUri="https://zenodo.org/record/343798/files/figure.png" pageId="3" pageNumber="268">Fig. 2</figureCitation>
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||
), while M simple, CuA developed into presutural vein in
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<taxonomicName id="4C7B3CF4FFC8FF9366E5F8C0FB611F67" box="[1020,1194,1818,1843]" class="Insecta" family="Corydiidae" genus="Homopteroidea" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="268" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90F9B65FFC8FF9366E5F8C0FB611F67" box="[1020,1194,1818,1843]" italics="true" pageId="3" pageNumber="268">Homopteroidea</emphasis>
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||
</taxonomicName>
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||
; 3) both tegmina and wings with an intercalary vein between R and M in
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||
<taxonomicName id="4C7B3CF4FFC8FF936618F8E5FC4E1F02" box="[769,901,1855,1878]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="268" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90F9B65FFC8FF936618F8E5FC4E1F02" box="[769,901,1855,1878]" italics="true" pageId="3" pageNumber="268">Ctenoneura</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13405BF2FFC8FF936680F8E4FC2A1F03" box="[921,993,1854,1879]" captionStart="FIGURE 2" captionStartId="3.[151,250,1268,1290]" captionTargetBox="[185,1430,874,1218]" captionTargetId="figure@3.[168,1453,802,1254]" captionTargetPageId="3" captionText="FIGURE 2. Tegmen and wing. Venation abbreviations see text." httpUri="https://zenodo.org/record/343798/files/figure.png" pageId="3" pageNumber="268">Fig. 2</figureCitation>
|
||
), while without an intercalary vein in
|
||
<taxonomicName id="4C7B3CF4FFC8FF93658EF8B8FE8D1F2F" box="[151,326,1890,1915]" class="Insecta" family="Corydiidae" genus="Homopteroidea" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="268" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90F9B65FFC8FF93658EF8B8FE8D1F2F" box="[151,326,1890,1915]" italics="true" pageId="3" pageNumber="268">Homopteroidea</emphasis>
|
||
</taxonomicName>
|
||
; 4) male genitalia simple, with reduced left phallomere, genital hook absent in
|
||
<taxonomicName id="4C7B3CF4FFC8FF9361DCF8B9FA821F2E" box="[1221,1353,1891,1914]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="268" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90F9B65FFC8FF9361DCF8B9FA821F2E" box="[1221,1353,1891,1914]" italics="true" pageId="3" pageNumber="268">Ctenoneura</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13405BF2FFC8FF936040F8B8FF651FCB" captionStart="FIGURE 1" captionStartId="2.[151,250,1982,2004]" captionTargetBox="[168,1412,1043,1926]" captionTargetId="figure@2.[155,1440,1023,1934]" captionTargetPageId="2" captionText="FIGURE 1. A – D. Genitalia of Ctenoneura and its terms: A. The original positions, dorsal view; B. stretched phallomeres, dorsal view; C – D. stretched phallomeres, ventral view." httpUri="https://zenodo.org/record/343797/files/figure.png" pageId="3" pageNumber="268">Fig. 1 D</figureCitation>
|
||
), while male genitalia extremely complex in
|
||
<taxonomicName id="4C7B3CF4FFC8FF9367BEF85CFC9D1FCB" box="[679,854,1926,1951]" class="Insecta" family="Corydiidae" genus="Homopteroidea" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="268" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90F9B65FFC8FF9367BEF85CFC9D1FCB" box="[679,854,1926,1951]" italics="true" pageId="3" pageNumber="268">Homopteroidea</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36114FCFFC8FF9465DEF870FE161C47" lastPageId="4" lastPageNumber="269" pageId="3" pageNumber="268" type="description">
|
||
<paragraph id="8BC44777FFC8FF9365DEF870FA5D1FB3" blockId="3.[151,1436,1638,2023]" pageId="3" pageNumber="268">
|
||
<emphasis id="B90F9B65FFC8FF9365DEF870FE0B1F97" bold="true" box="[199,448,1962,1987]" pageId="3" pageNumber="268">Redescription. Male.</emphasis>
|
||
Small size, resembles small
|
||
<taxonomicName id="4C7B3CF4FFC8FF936610F870FC4A1F97" box="[777,897,1962,1987]" class="Insecta" family="Ectobiidae" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="268" phylum="Arthropoda" rank="family">Ectobiidae</taxonomicName>
|
||
cockroaches, body length (without wings) 5.0–
|
||
<quantity id="4C83EA92FFC8FF93658EF815FF241FB2" box="[151,239,1999,2023]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="3" pageNumber="268" unit="mm" value="9.7">9.7 mm</quantity>
|
||
. Body smooth and shining, not pubescent, usually coloration ranging from brownish yellow to dark brown.
|
||
</paragraph>
|
||
<paragraph id="8BC44777FFCFFF9465DEFF4DFAC31AFF" blockId="4.[151,1437,151,2016]" pageId="4" pageNumber="269">
|
||
<emphasis id="B90F9B65FFCFFF9465DEFF4DFEDB18E4" bold="true" box="[199,272,151,176]" pageId="4" pageNumber="269">Head:</emphasis>
|
||
hidden or not, vertex round, eyes wide apart, ocelli very small, nearly degenerate, antennal sockets round, large.
|
||
<emphasis id="B90F9B65FFCFFF946429FF66FE791881" bold="true" box="[304,434,188,213]" pageId="4" pageNumber="269">Pronotum:</emphasis>
|
||
generally subcircular or oval.
|
||
<emphasis id="B90F9B65FFCFFF94660BFF66FBCD1881" bold="true" box="[786,1030,188,213]" pageId="4" pageNumber="269">Tegmina and wings:</emphasis>
|
||
fully developed.
|
||
<taxonomicName id="4C7B3CF4FFCFFF9461C8FF66FADE1881" box="[1233,1301,188,213]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="299" phylum="Arthropoda" rank="species" species="major">Major</taxonomicName>
|
||
veins often densely reticulate with numerous cross veins. Tegmina horny, Sc and branches of R usually oblique, M welldeveloped, usually with 3–6 branches, an intercalary vein present between R and M, sometimes incomplete, CuA usually simple, sometimes bifurcate; wings with a mass of spots between RA and RP, usually with an intercalary vein between R and M, sometimes incomplete, CuA usually with 3 to 8 branches that generally are parallel, and with numerous small cross veins connecting them, CuP usually slender, anal area folded over the rest of the wing in resting position as in other
|
||
<taxonomicName id="4C7B3CF4FFCFFF9464DFFE4EFD8819F9" box="[454,579,404,429]" class="Insecta" family="Corydiidae" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="family">Corydiidae</taxonomicName>
|
||
genera, AA thick, AP branches fan-like (
|
||
<figureCitation id="13405BF2FFCFFF94610AFE4EFB9219F8" box="[1043,1113,404,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1268,1290]" captionTargetBox="[185,1430,874,1218]" captionTargetId="figure@3.[168,1453,802,1254]" captionTargetPageId="3" captionText="FIGURE 2. Tegmen and wing. Venation abbreviations see text." httpUri="https://zenodo.org/record/343798/files/figure.png" pageId="4" pageNumber="269">Fig. 2</figureCitation>
|
||
).
|
||
<emphasis id="B90F9B65FFCFFF946175FE4EFB6519F9" bold="true" box="[1132,1198,404,429]" pageId="4" pageNumber="269">Legs:</emphasis>
|
||
front femur
|
||
<typeStatus id="54C0F9D5FFCFFF946025FE4FFAB819F9" box="[1340,1395,405,429]" pageId="4" pageNumber="269">Type</typeStatus>
|
||
C1, sometimes with small processes on hind margin basally and setae on the surface (
|
||
<figureCitation id="13405BF2FFCFFF94613BFE60FBB61987" box="[1058,1149,442,467]" captionStart="FIGURE 23" captionStartId="29.[151,250,1868,1890]" captionTargetBox="[247,1345,221,1812]" captionTargetId="figure@29.[230,1357,205,1822]" captionTargetPageId="29" captionText="FIGURE 23. Ctenoneura helicata sp. nov., male from Yunnan, China: A. pronotum with exposed head; B. tegmen; C. wing; D. front femur; E. supra-anal plate with T 9, dorsal view; F. subgenital plate, ventral view; G. subgenital plate, dorsal view; H. subgenital sclerite (sgs); I. genitalia, ventral view." httpUri="https://zenodo.org/record/343819/files/figure.png" pageId="4" pageNumber="269">Figs. 23</figureCitation>
|
||
D; 24 D). Pulvilli absent, tarsal claws symmetrical, simple, arolia small or absent.
|
||
<emphasis id="B90F9B65FFCFFF946634FE04FC6019A3" bold="true" box="[813,939,478,503]" pageId="4" pageNumber="269">Abdomen:</emphasis>
|
||
abdominal terga unspecialized. Supra-anal plate transverse, with many different shapes. Cerci long, pubescent, sometimes the basal segments specialized (round or slightly protruded and directed medially, e.g.
|
||
<figureCitation id="13405BF2FFCFFF94660BFDFCFCA81A6B" box="[786,867,550,575]" captionStart="FIGURE 20" captionStartId="26.[151,250,1881,1903]" captionTargetBox="[178,1425,228,1825]" captionTargetId="figure@26.[160,1445,193,1834]" captionTargetPageId="26" captionText="FIGURE 20. Ctenoneura papillaris sp. nov., male from Hainan, China: A. pronotum with exposed head; B. tegmen; C. wing; D. front femur; E. supra-anal plate with T 9, dorsal view; F. subgenital plate, ventral view; G. subgenital plate, dorsal view; H. genitalia, ventral view." httpUri="https://zenodo.org/record/343816/files/figure.png" pageId="4" pageNumber="269">Fig. 20</figureCitation>
|
||
E). Subgenital plate asymmetrical, different from one species to another, usually with one stylus or none (
|
||
<bibRefCitation id="EFEA3A86FFCFFF94663BFD90FC651A37" author="Roth" box="[802,942,586,611]" pageId="4" pageNumber="269" refString="Roth, L. M. (1993) Revision of the cockroach genus Ctenoneura Hanitsch (Blattaria, Polyphagidae). Tijdschrift voor Entomologie, 136, 83 - 109." type="journal article" year="1993">Roth (1993)</bibRefCitation>
|
||
stated some species without stylus). Some species with one slender and curved subgenital sclerite (sgs) present above the subgenital plate, below the genitalia, or integrated with subgenital plate, and an extending structure (eds) to hold the sgs (
|
||
<figureCitation id="13405BF2FFCFFF94615DFD48FB5C1AFF" box="[1092,1175,658,683]" captionStart="FIGURE 13" captionStartId="19.[151,250,1694,1716]" captionTargetBox="[201,1419,1273,1641]" captionTargetId="figure@19.[163,1448,1228,1661]" captionTargetPageId="19" captionText="FIGURE 13. The subgenital plate with subgenital sclerite (sgs): A. C. helicata sp. nov.; B. C. simulans; C. C. qiuae sp. nov.." httpUri="https://zenodo.org/record/343809/files/figure.png" pageId="4" pageNumber="269">Fig. 13</figureCitation>
|
||
; 22 F–G).
|
||
</paragraph>
|
||
<paragraph id="8BC44777FFCFFF9465DEFD6CFAEC1BD7" blockId="4.[151,1437,151,2016]" pageId="4" pageNumber="269">
|
||
<emphasis id="B90F9B65FFCFFF9465DEFD6CFEB01A9B" bold="true" box="[199,379,694,719]" pageId="4" pageNumber="269">Male genitalia:</emphasis>
|
||
consists of three parts, viz. left phallomere, right phallomere and transverse sclerite (tvs). Left phallomere strongly reduced, two parts present, viz. left dorsal phallomere (ldp) and left ventral phallomere (lvp); lvp small, usually with irregular processes posteriorly, anterior usually acute, like a parrot peak; ldp connects lvp by the whole anterior margin, posterior with a
|
||
<emphasis id="B90F9B65FFCFFF946787FCF8FCFB1B6F" bold="true" box="[670,816,802,827]" pageId="4" pageNumber="269">curved part</emphasis>
|
||
(
|
||
<emphasis id="B90F9B65FFCFFF94665DFCF8FCA51B6F" bold="true" box="[836,878,802,827]" pageId="4" pageNumber="269">cvp</emphasis>
|
||
), small, towards the anterior. Right phallomere consists of three parts, viz. R1M, R2 and R3, R1M irregular, R2 with three parts: a robust process (rop), a slender process (slp) and an elongate process (elp), R3 concave. Transverse sclerite (tvs) long, irregular (
|
||
<figureCitation id="13405BF2FFCFFF9461CCFCB0FADD1BD7" box="[1237,1302,874,899]" captionStart="FIGURE 1" captionStartId="2.[151,250,1982,2004]" captionTargetBox="[168,1412,1043,1926]" captionTargetId="figure@2.[155,1440,1023,1934]" captionTargetPageId="2" captionText="FIGURE 1. A – D. Genitalia of Ctenoneura and its terms: A. The original positions, dorsal view; B. stretched phallomeres, dorsal view; C – D. stretched phallomeres, ventral view." httpUri="https://zenodo.org/record/343797/files/figure.png" pageId="4" pageNumber="269">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC44777FFCFFF9465DEFC54FB4F1B9F" blockId="4.[151,1437,151,2016]" pageId="4" pageNumber="269">
|
||
<emphasis id="B90F9B65FFCFFF9465DEFC54FEEE1BF3" bold="true" box="[199,293,910,935]" pageId="4" pageNumber="269">Female.</emphasis>
|
||
Apterous, body smooth, eyes reduced, arolia absent, supra-anal plate transverse, suboval, apex slightly truncated, cerci robust, apex with a spine. Subgenital plate valvular (
|
||
<figureCitation id="13405BF2FFCFFF9466EBFC68FBB31B9F" box="[1010,1144,946,971]" captionStart="FIGURE 5" captionStartId="9.[151,250,1737,1759]" captionTargetBox="[165,1427,420,1691]" captionTargetId="figure@9.[160,1445,409,1717]" captionTargetPageId="9" captionText="FIGURE 5. A – L. Ctenoneura heixuanfeng sp. nov.: A – B. male, paratype: A. dorsal view; B. ventral view. C – D. male supraanal plate: C. with T 9, dorsal view; D. close-up, dorsal view. E. male stylus, ventral view. F. male subgenital plate, ventral view. G – H. female, paratype: G. dorsal view; H. ventral view. I. female supra-anal plate, dorsal view; J. female subgenital plate, ventral view. K – L. ootheca: K. lateral view; L. dorsal view." httpUri="https://zenodo.org/record/343801/files/figure.png" pageId="4" pageNumber="269">Figs. 5 G–J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC44777FFCFFF9465DEFC0CFE161C47" blockId="4.[151,1437,151,2016]" pageId="4" pageNumber="269">
|
||
<emphasis id="B90F9B65FFCFFF9465DEFC0CFEFA1BBB" bold="true" box="[199,305,982,1007]" pageId="4" pageNumber="269">Ootheca.</emphasis>
|
||
Oval, with small serrations on the keel. The surface with many ribbed lines, eggs filled, can be seen in dorsal view (
|
||
<figureCitation id="13405BF2FFCFFF94645CFC20FE191C47" box="[325,466,1018,1043]" captionStart="FIGURE 5" captionStartId="9.[151,250,1737,1759]" captionTargetBox="[165,1427,420,1691]" captionTargetId="figure@9.[160,1445,409,1717]" captionTargetPageId="9" captionText="FIGURE 5. A – L. Ctenoneura heixuanfeng sp. nov.: A – B. male, paratype: A. dorsal view; B. ventral view. C – D. male supraanal plate: C. with T 9, dorsal view; D. close-up, dorsal view. E. male stylus, ventral view. F. male subgenital plate, ventral view. G – H. female, paratype: G. dorsal view; H. ventral view. I. female supra-anal plate, dorsal view; J. female subgenital plate, ventral view. K – L. ootheca: K. lateral view; L. dorsal view." httpUri="https://zenodo.org/record/343801/files/figure.png" pageId="4" pageNumber="269">Figs. 5 K–L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36114FCFFCFFF9465DEFBC4FD6A1C2B" pageId="4" pageNumber="269" type="distribution">
|
||
<paragraph id="8BC44777FFCFFF9465DEFBC4FDE91C63" blockId="4.[151,1437,151,2016]" box="[199,546,1054,1079]" pageId="4" pageNumber="269">
|
||
<emphasis id="B90F9B65FFCFFF9465DEFBC4FEAB1C63" bold="true" box="[199,352,1054,1079]" pageId="4" pageNumber="269">Distribution.</emphasis>
|
||
Oriental Region.
|
||
</paragraph>
|
||
<paragraph id="8BC44777FFCFFF9465DEFB98FD6A1C2B" blockId="4.[151,1437,151,2016]" pageId="4" pageNumber="269">
|
||
<emphasis id="B90F9B65FFCFFF9465DEFB98FE4C1C0F" bold="true" box="[199,391,1090,1115]" pageId="4" pageNumber="269">Natural history.</emphasis>
|
||
<taxonomicName id="4C7B3CF4FFCFFF946488FB99FDDD1C0E" box="[401,534,1091,1114]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90F9B65FFCFFF946488FB99FDDD1C0E" box="[401,534,1091,1114]" italics="true" pageId="4" pageNumber="269">Ctenoneura</emphasis>
|
||
</taxonomicName>
|
||
should be one kind of wood-feeding cockroach. For more details see “natural history” under
|
||
<taxonomicName id="4C7B3CF4FFCFFF946426FBBDFE231C2B" box="[319,488,1126,1151]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="heixuanfeng">
|
||
<emphasis id="B90F9B65FFCFFF946426FBBDFE231C2B" box="[319,488,1126,1151]" italics="true" pageId="4" pageNumber="269">C. heixuanfeng</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7B3CF4FFCFFF946706FBBDFD521C2B" box="[543,665,1126,1151]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="delicata">
|
||
<emphasis id="B90F9B65FFCFFF946706FBBDFD521C2B" box="[543,665,1126,1151]" italics="true" pageId="4" pageNumber="269">C. delicata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36114FCFFCFFF9465DEFB50FA571F8A" pageId="4" pageNumber="269" type="discussion">
|
||
<paragraph id="8BC44777FFCFFF9465DEFB50FA561D67" blockId="4.[151,1437,151,2016]" pageId="4" pageNumber="269">
|
||
<materialsCitation id="3B134D2AFFCFFF9465DEFB50FA521D67" ID-GBIF-Occurrence="1437751814" country="China" location="Yunnan" pageId="4" pageNumber="269" specimenCount="1" stateProvince="Yunnan">
|
||
<emphasis id="B90F9B65FFCFFF9465DEFB50FCD01CF7" bold="true" box="[199,795,1162,1187]" pageId="4" pageNumber="269">Discussion. The estimate of species abundance.</emphasis>
|
||
<taxonomicName id="4C7B3CF4FFCFFF946633FB51FC641CF6" box="[810,943,1163,1186]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90F9B65FFCFFF946633FB51FC641CF6" box="[810,943,1163,1186]" italics="true" pageId="4" pageNumber="269">Ctenoneura</emphasis>
|
||
</taxonomicName>
|
||
must be a genus rich in species in the
|
||
<taxonomicName id="4C7B3CF4FFCFFF94658EFB74FEDC1C93" box="[151,279,1198,1223]" class="Insecta" family="Corydiidae" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="family">Corydiidae</taxonomicName>
|
||
. Members of this rarely-collected genus are mostly recorded from Southeast Asia. Former studies only found four species from
|
||
<collectingCountry id="F36C07E7FFCFFF9464B2FB08FE251CBF" box="[427,494,1234,1259]" name="China" pageId="4" pageNumber="269">China</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFEA3A86FFCFFF9464E4FB08FD031CBF" author="Bey-Bienko" box="[509,712,1234,1259]" pageId="4" pageNumber="269" refString="Bey-Bienko, G. Y. (1957) Blattoidea of Szechuan and Yunnan. Communication I. The results of the Chinese-Soviet zoologicobotanical expeditions to southwestern China 1955 - 1956. Entomologicheskoe Obozrenie, 36, 895 - 915." type="journal article" year="1957">Bey-Bienko 1957</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFEA3A86FFCFFF9467CDFB08FCDB1CBF" author="Bey-Bienko" box="[724,784,1234,1259]" pageId="4" pageNumber="269" refString="Bey-Bienko, G. Y. (1969) New genera and species of cockroaches (Blattoptera) from tropical and subtropical Asia. Entomologicheskoe Obozrenie, 48, 832 - 862." type="journal article" year="1969">1969</bibRefCitation>
|
||
), one species from
|
||
<collectingCountry id="F36C07E7FFCFFF9466F0FB09FB9D1CBF" box="[1001,1110,1235,1259]" name="Myanmar" pageId="4" pageNumber="269">Myanmar</collectingCountry>
|
||
and one undescribed species from
|
||
<collectingCountry id="F36C07E7FFCFFF9465CFFB2CFEFE1D5B" box="[214,309,1270,1295]" name="Vietnam" pageId="4" pageNumber="269">Vietnam</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFEA3A86FFCFFF94645CFB2CFE741D5B" author="Roth" box="[325,447,1270,1295]" pageId="4" pageNumber="269" refString="Roth, L. M. (1993) Revision of the cockroach genus Ctenoneura Hanitsch (Blattaria, Polyphagidae). Tijdschrift voor Entomologie, 136, 83 - 109." type="journal article" year="1993">Roth 1993</bibRefCitation>
|
||
).This paper adds three species from
|
||
<collectingRegion id="49BF8995FFCFFF94667FFB22FC751D5B" box="[870,958,1272,1295]" country="China" name="Yunnan" pageId="4" pageNumber="269">Yunnan</collectingRegion>
|
||
, four species from
|
||
<collectingRegion id="49BF8995FFCFFF946182FB2CFB271D5B" box="[1179,1260,1270,1295]" country="China" name="Hainan" pageId="4" pageNumber="269">Hainan</collectingRegion>
|
||
Island and one unknown female from
|
||
<collectingRegion id="49BF8995FFCFFF94648FFAC0FE311D67" box="[406,506,1306,1331]" country="China" name="Guangxi" pageId="4" pageNumber="269">Guangxi</collectingRegion>
|
||
. We suspect that there must be more new species in
|
||
<collectingCountry id="F36C07E7FFCFFF94615BFAC0FA521D67" box="[1090,1433,1306,1331]" isEnumeration="true" pageId="4" pageNumber="269">China, Myanmar and Vietnam</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC44777FFCFFF9465DEFAE4FE301F53" blockId="4.[151,1437,151,2016]" pageId="4" pageNumber="269">
|
||
<emphasis id="B90F9B65FFCFFF9465DEFAE4FDCA1D03" bold="true" box="[199,513,1342,1367]" pageId="4" pageNumber="269">Species groups suggestion.</emphasis>
|
||
After dissecting the specimens, we found
|
||
<taxonomicName id="4C7B3CF4FFCFFF9466C7FAE5FBA91D03" box="[990,1122,1342,1367]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="simulans">
|
||
<emphasis id="B90F9B65FFCFFF9466C7FAE5FBA91D03" box="[990,1122,1342,1367]" italics="true" pageId="4" pageNumber="269">C. simulans</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7B3CF4FFCFFF946169FAE5FB201D03" box="[1136,1259,1342,1367]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="helicata" status="sp. nov.">
|
||
<emphasis id="B90F9B65FFCFFF946169FAE5FB201D03" box="[1136,1259,1342,1367]" italics="true" pageId="4" pageNumber="269">C. helicata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23C261EFFCFFF9461EBFAE4FA871D03" box="[1266,1356,1342,1367]" pageId="4" pageNumber="269" rank="species">
|
||
<emphasis id="B90F9B65FFCFFF9461EBFAE4FA871D03" bold="true" box="[1266,1356,1342,1367]" pageId="4" pageNumber="269">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C7B3CF4FFCFFF94609BFAE5FF1E1D2F" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="qiuae" status="sp. nov.">
|
||
<emphasis id="B90F9B65FFCFFF94609BFAE5FF1E1D2F" italics="true" pageId="4" pageNumber="269">C. qiuae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23C261EFFCFFF9465C5FAB8FEFE1D2F" box="[220,309,1378,1403]" pageId="4" pageNumber="269" rank="species">
|
||
<emphasis id="B90F9B65FFCFFF9465C5FAB8FEFE1D2F" bold="true" box="[220,309,1378,1403]" pageId="4" pageNumber="269">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
share some similar characters: 1) supra-anal plate simple, unspecialized; 2) subgenital plate with eds forming a groove, and separated sgs partly resting in the groove; 3) hind margin of subgenital plate usually with an incision or excavation, one stylus originating from the left of the incision or excavation. Here we build a
|
||
<emphasis id="B90F9B65FFCFFF946033FA70FF151DB3" bold="true" pageId="4" pageNumber="269">
|
||
<taxonomicName id="4C7B3CF4FFCFFF946033FA70FA591D97" box="[1322,1426,1450,1475]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="simulans">
|
||
<emphasis id="B90F9B65FFCFFF946033FA70FA591D97" bold="true" box="[1322,1426,1450,1475]" italics="true" pageId="4" pageNumber="269">simulans</emphasis>
|
||
</taxonomicName>
|
||
- group
|
||
</emphasis>
|
||
to contain
|
||
<taxonomicName id="4C7B3CF4FFCFFF946471FA15FD1A1DB3" authority="Bey-Bienko, 1969" authorityName="Bey-Bienko" authorityYear="1969" box="[360,721,1486,1511]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="simulans">
|
||
<emphasis id="B90F9B65FFCFFF946471FA15FE3A1DB3" box="[360,497,1486,1511]" italics="true" pageId="4" pageNumber="269">C. simulans</emphasis>
|
||
<bibRefCitation id="EFEA3A86FFCFFF9464E5FA14FD1A1DB3" author="Bey-Bienko" box="[508,721,1486,1511]" pageId="4" pageNumber="269" refString="Bey-Bienko, G. Y. (1969) New genera and species of cockroaches (Blattoptera) from tropical and subtropical Asia. Entomologicheskoe Obozrenie, 48, 832 - 862." type="journal article" year="1969">Bey-Bienko, 1969</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7B3CF4FFCFFF9467F8FA15FCAB1DB3" box="[737,864,1486,1511]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="helicata" status="sp. nov.">
|
||
<emphasis id="B90F9B65FFCFFF9467F8FA15FCAB1DB3" box="[737,864,1486,1511]" italics="true" pageId="4" pageNumber="269">C. helicata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23C261EFFCFFF946675FA14FC011DB3" box="[876,970,1486,1511]" pageId="4" pageNumber="269" rank="species">
|
||
<emphasis id="B90F9B65FFCFFF946675FA14FC011DB3" bold="true" box="[876,970,1486,1511]" pageId="4" pageNumber="269">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C7B3CF4FFCFFF946110FA15FBA71DB3" box="[1033,1132,1487,1511]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="qiuae" status="sp. nov.">
|
||
<emphasis id="B90F9B65FFCFFF946110FA15FBA71DB3" box="[1033,1132,1487,1511]" italics="true" pageId="4" pageNumber="269">C. qiuae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23C261EFFCFFF946161FA14FB1D1DB3" box="[1144,1238,1486,1511]" pageId="4" pageNumber="269" rank="species">
|
||
<emphasis id="B90F9B65FFCFFF946161FA14FB1D1DB3" bold="true" box="[1144,1238,1486,1511]" pageId="4" pageNumber="269">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
that all own the characters listed above.
|
||
<taxonomicName id="4C7B3CF4FFCFFF9464BAFA29FDED1E5F" box="[419,550,1522,1547]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="elongata" status="sp. nov.">
|
||
<emphasis id="B90F9B65FFCFFF9464BAFA29FDED1E5F" box="[419,550,1522,1547]" italics="true" pageId="4" pageNumber="269">C. elongata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23C261EFFCFFF946735FA28FD4E1E5F" box="[556,645,1522,1547]" pageId="4" pageNumber="269" rank="species">
|
||
<emphasis id="B90F9B65FFCFFF946735FA28FD4E1E5F" bold="true" box="[556,645,1522,1547]" pageId="4" pageNumber="269">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is different from the typical
|
||
<taxonomicName id="4C7B3CF4FFCFFF9466DCFA28FBE21E5F" box="[965,1065,1522,1547]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="simulans">
|
||
<emphasis id="B90F9B65FFCFFF9466DCFA28FBE21E5F" box="[965,1065,1522,1547]" italics="true" pageId="4" pageNumber="269">simulans</emphasis>
|
||
</taxonomicName>
|
||
-group, it is distinct in owning an unseparated sgs and an extremely large eds; we include it in this group as an “aberrant” species.
|
||
<taxonomicName id="4C7B3CF4FFCFFF94601CF9CDFF231E07" authority="Princis" authorityName="Princis" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="birmanica">
|
||
<emphasis id="B90F9B65FFCFFF94601CF9CDFA571E7B" box="[1285,1436,1558,1583]" italics="true" pageId="4" pageNumber="269">C. birmanica</emphasis>
|
||
Princis
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7B3CF4FFCFFF9465EDF9E1FE3A1E07" authority="Bey-Bienko" authorityName="Bey-Bienko" box="[244,497,1594,1619]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="misera">
|
||
<emphasis id="B90F9B65FFCFFF9465EDF9E1FEA91E07" box="[244,354,1595,1619]" italics="true" pageId="4" pageNumber="269">C. misera</emphasis>
|
||
Bey-Bienko
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7B3CF4FFCFFF946733F9E1FC831E07" authority="Bey-Bienko" authorityName="Bey-Bienko" box="[554,840,1594,1619]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="yunnanea">
|
||
<emphasis id="B90F9B65FFCFFF946733F9E1FD721E07" box="[554,697,1595,1619]" italics="true" pageId="4" pageNumber="269">C. yunnanea</emphasis>
|
||
Bey-Bienko
|
||
</taxonomicName>
|
||
may belong to this group, but we lacked material to examine. The species from
|
||
<collectingRegion id="49BF8995FFCFFF9464C7F984FDE41E23" box="[478,559,1630,1655]" country="China" name="Hainan" pageId="4" pageNumber="269">Hainan</collectingRegion>
|
||
<collectingCountry id="F36C07E7FFCFFF946723F984FDB51E23" box="[570,638,1630,1655]" name="Iceland" pageId="4" pageNumber="269">Island</collectingCountry>
|
||
are quite distinguished from each other in the supra-anal plate and subgenital plate, but they all share some other identical characters: 1) supra-anal plate all specialized into different shapes; 2) subgenital plate protrudes toward right; and 3) body dark brown to brownish black, usually with small yellow areas in lateral fore borders of the pronotum. We suggest building a
|
||
<emphasis id="B90F9B65FFCFFF946111F910FB211EB7" bold="true" box="[1032,1258,1738,1763]" pageId="4" pageNumber="269">
|
||
<taxonomicName id="4C7B3CF4FFCFFF946111F910FB511EB7" box="[1032,1178,1738,1763]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="heixuanfeng">
|
||
<emphasis id="B90F9B65FFCFFF946111F910FB511EB7" bold="true" box="[1032,1178,1738,1763]" italics="true" pageId="4" pageNumber="269">heixuanfeng</emphasis>
|
||
</taxonomicName>
|
||
-group
|
||
</emphasis>
|
||
for the
|
||
<collectingRegion id="49BF8995FFCFFF946052F910FA571EB7" box="[1355,1436,1738,1763]" country="China" name="Hainan" pageId="4" pageNumber="269">Hainan</collectingRegion>
|
||
species described in this article.
|
||
</paragraph>
|
||
<paragraph id="8BC44777FFCFFF9465DEF8C8FA571F8A" blockId="4.[151,1437,151,2016]" pageId="4" pageNumber="269">
|
||
<emphasis id="B90F9B65FFCFFF9465DEF8C8FE001F7E" bold="true" box="[199,459,1810,1835]" pageId="4" pageNumber="269">Discussion of females.</emphasis>
|
||
The apterous female reported in this article is one important discovery on
|
||
<emphasis id="B90F9B65FFCFFF946008F8C9FA571F7E" box="[1297,1436,1811,1834]" italics="true" pageId="4" pageNumber="269">
|
||
<taxonomicName id="4C7B3CF4FFCFFF946008F8C9FA521F7E" box="[1297,1433,1811,1834]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="genus">Ctenoneura</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The previous records of the females (
|
||
<bibRefCitation id="EFEA3A86FFCFFF94674CF8ECFCCA1F1B" author="Hanitsch" box="[597,769,1846,1871]" pageId="4" pageNumber="269" refString="Hanitsch, R. (1925) On a collection of Blattidae from northern Sarawak, chiefly Mt. Murud and Mt. Dulit. Sarawak Museum Journal, 3, 75 - 106." type="journal article" year="1925">Hanitsch 1925</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEA3A86FFCFFF94660BF8ECFC141F1B" author="Bey-Bienko" box="[786,991,1846,1871]" pageId="4" pageNumber="269" refString="Bey-Bienko, G. Y. (1957) Blattoidea of Szechuan and Yunnan. Communication I. The results of the Chinese-Soviet zoologicobotanical expeditions to southwestern China 1955 - 1956. Entomologicheskoe Obozrenie, 36, 895 - 915." type="journal article" year="1957">Bey-Bienko 1957</bibRefCitation>
|
||
) indicates macropterous, but they are doubtful.
|
||
<bibRefCitation id="EFEA3A86FFCFFF94641AF880FE731F27" author="Hanitsch" box="[259,440,1882,1907]" pageId="4" pageNumber="269" refString="Hanitsch, R. (1925) On a collection of Blattidae from northern Sarawak, chiefly Mt. Murud and Mt. Dulit. Sarawak Museum Journal, 3, 75 - 106." type="journal article" year="1925">Hanitsch (1925)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="4C7B3CF4FFCFFF946729F881FD431F27" box="[560,648,1882,1907]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="fulva">
|
||
<emphasis id="B90F9B65FFCFFF946729F881FD431F27" box="[560,648,1882,1907]" italics="true" pageId="4" pageNumber="269">C. fulva</emphasis>
|
||
</taxonomicName>
|
||
based on six specimens, and
|
||
<taxonomicName id="4C7B3CF4FFCFFF9466D6F881FBF91F27" box="[975,1074,1883,1907]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="species" species="major">
|
||
<emphasis id="B90F9B65FFCFFF9466D6F881FBF91F27" box="[975,1074,1883,1907]" italics="true" pageId="4" pageNumber="269">C. major</emphasis>
|
||
</taxonomicName>
|
||
based on two specimens; within these were three macropterous females determined by Hanitsch himself. But there were two significant concerns about Hanitsch’s knowledge of
|
||
<emphasis id="B90F9B65FFCFFF946719F879FD401FEE" box="[512,651,1955,1978]" italics="true" pageId="4" pageNumber="269">
|
||
<taxonomicName id="4C7B3CF4FFCFFF946719F879FD431FEE" box="[512,648,1955,1978]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="269" phylum="Arthropoda" rank="genus">Ctenoneura</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
One is the generic diagnosis of this genus having the male with two styli which later proved to have the male with one stylus or none. The other concern is about the
|
||
<typeStatus id="54C0F9D5FFCFFF9461FDF812FADF1FB4" box="[1252,1300,1992,2016]" pageId="4" pageNumber="269">type</typeStatus>
|
||
series of
|
||
<emphasis id="B90F9B65FFCFFF94609BF81DFA571F8A" box="[1410,1436,1991,2014]" italics="true" pageId="4" pageNumber="269">C.</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36114FCFFCEFF96658EFF4DFE0E1DAD" lastPageId="6" lastPageNumber="271" pageId="5" pageNumber="270" type="reference_group">
|
||
<paragraph id="8BC44777FFCEFF95658EFF4DFAE419F9" blockId="5.[151,1436,151,429]" pageId="5" pageNumber="270">
|
||
<emphasis id="B90F9B65FFCEFF95658EFF4DFF1C18E4" box="[151,215,151,176]" italics="true" pageId="5" pageNumber="270">
|
||
<taxonomicName id="4C7B3CF4FFCEFF95658EFF4DFF1818E4" ID-CoL="3266Q" box="[151,211,151,176]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="fulva">fulva</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<bibRefCitation id="EFEA3A86FFCEFF9565C4FF4DFEAD18E4" author="Roth" box="[221,358,151,176]" pageId="5" pageNumber="270" refString="Roth, L. M. (1993) Revision of the cockroach genus Ctenoneura Hanitsch (Blattaria, Polyphagidae). Tijdschrift voor Entomologie, 136, 83 - 109." type="journal article" year="1993">Roth (1993)</bibRefCitation>
|
||
has examined the
|
||
<typeStatus id="54C0F9D5FFCEFF956723FF42FDA118E4" box="[570,618,152,176]" pageId="5" pageNumber="270">type</typeStatus>
|
||
specimens of
|
||
<taxonomicName id="4C7B3CF4FFCEFF956614FF42FCA318E4" ID-CoL="3266Q" box="[781,872,151,176]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="fulva">
|
||
<emphasis id="B90F9B65FFCEFF956614FF42FCA318E4" box="[781,872,151,176]" italics="true" pageId="5" pageNumber="270">C. fulva</emphasis>
|
||
</taxonomicName>
|
||
in the Museum of Oxford University. No females were found, but he discovered two new species within the
|
||
<typeStatus id="54C0F9D5FFCEFF956642FF67FC401881" box="[859,907,189,213]" pageId="5" pageNumber="270">type</typeStatus>
|
||
series of
|
||
<taxonomicName id="4C7B3CF4FFCEFF95611EFF67FBAE1881" ID-CoL="3266Q" box="[1031,1125,188,213]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="fulva">
|
||
<emphasis id="B90F9B65FFCEFF95611EFF67FBAE1881" box="[1031,1125,188,213]" italics="true" pageId="5" pageNumber="270">C. fulva</emphasis>
|
||
</taxonomicName>
|
||
(one was described as
|
||
<taxonomicName id="4C7B3CF4FFCEFF95609BFF67FED318AC" ID-CoL="32672" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="murudensis">
|
||
<emphasis id="B90F9B65FFCEFF95609BFF67FED318AC" italics="true" pageId="5" pageNumber="270">C. murudensis</emphasis>
|
||
</taxonomicName>
|
||
, the other was not described due to its bad condition). Such limited knowledge may lead to mistaken identification. At that time, the male subgenital plate of
|
||
<taxonomicName id="4C7B3CF4FFCEFF956609FEDFFC5F1948" ID-CoL="8MRDJ" box="[784,916,261,284]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="270" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90F9B65FFCEFF956609FEDFFC5F1948" box="[784,916,261,284]" italics="true" pageId="5" pageNumber="270">Ctenoneura</emphasis>
|
||
</taxonomicName>
|
||
was poorly studied. Hanitsch probably mixed in other male species as the females of
|
||
<taxonomicName id="4C7B3CF4FFCEFF956755FEF2FD6F1914" ID-CoL="3266Q" box="[588,676,295,320]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="fulva">
|
||
<emphasis id="B90F9B65FFCEFF956755FEF2FD6F1914" box="[588,676,295,320]" italics="true" pageId="5" pageNumber="270">C. fulva</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7B3CF4FFCEFF9567C2FEF2FCF61914" ID-CoL="3266W" box="[731,829,296,320]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="major">
|
||
<emphasis id="B90F9B65FFCEFF9567C2FEF2FCF61914" box="[731,829,296,320]" italics="true" pageId="5" pageNumber="270">C. major</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFEA3A86FFCEFF956652FEFDFBE81914" author="Bey-Bienko" box="[843,1059,295,320]" pageId="5" pageNumber="270" refString="Bey-Bienko, G. Y. (1957) Blattoidea of Szechuan and Yunnan. Communication I. The results of the Chinese-Soviet zoologicobotanical expeditions to southwestern China 1955 - 1956. Entomologicheskoe Obozrenie, 36, 895 - 915." type="journal article" year="1957">Bey-Bienko (1957)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="4C7B3CF4FFCEFF956185FEF2FAF91914" ID-CoL="3266J" box="[1180,1330,296,320]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="270" phylum="Arthropoda" rank="species" species="acuticerca">
|
||
<emphasis id="B90F9B65FFCEFF956185FEF2FAF91914" box="[1180,1330,296,320]" italics="true" pageId="5" pageNumber="270">C. acuticerca</emphasis>
|
||
</taxonomicName>
|
||
based on two macropterous females, but this species may not belong to
|
||
<taxonomicName id="4C7B3CF4FFCEFF956657FE97FC181930" ID-CoL="8MRDJ" box="[846,979,333,356]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="270" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90F9B65FFCEFF956657FE97FC181930" box="[846,979,333,356]" italics="true" pageId="5" pageNumber="270">Ctenoneura</emphasis>
|
||
</taxonomicName>
|
||
due to the absence of an intercalary vein which is an important character for
|
||
<taxonomicName id="4C7B3CF4FFCEFF95673AFEAAFD6C19D3" ID-CoL="8MRDJ" box="[547,679,368,391]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="270" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90F9B65FFCEFF95673AFEAAFD6C19D3" box="[547,679,368,391]" italics="true" pageId="5" pageNumber="270">Ctenoneura</emphasis>
|
||
</taxonomicName>
|
||
. The discovery of the apteral female in this article may explain why the previous studies are absence of females; the sexual dimorphism may lead to overlooking the females.
|
||
</paragraph>
|
||
<caption id="DF0417FFFFCEFF95658EF8A0FBDE1FC4" box="[151,1045,1914,1936]" httpUri="https://zenodo.org/record/343799/files/figure.png" pageId="5" pageNumber="270" targetBox="[211,1345,495,1880]" targetPageId="5">
|
||
<paragraph id="8BC44777FFCEFF95658EF8A0FBDE1FC4" blockId="5.[151,1045,1914,1936]" box="[151,1045,1914,1936]" pageId="5" pageNumber="270">
|
||
<emphasis id="B90F9B65FFCEFF95658EF8A0FED91FDB" bold="true" box="[151,274,1914,1936]" pageId="5" pageNumber="270">FIGURE 3.</emphasis>
|
||
Distribution map of
|
||
<taxonomicName id="4C7B3CF4FFCEFF9564FEF8A1FD941FC4" ID-CoL="8MRDJ" box="[487,607,1915,1936]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="270" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90F9B65FFCEFF9564FEF8A1FD941FC4" box="[487,607,1915,1936]" italics="true" pageId="5" pageNumber="270">Ctenoneura</emphasis>
|
||
</taxonomicName>
|
||
species in Yunnan Province and Myanmar.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0417FFFFCDFF96658EFB7DFCA81CE9" box="[151,867,1191,1213]" httpUri="https://zenodo.org/record/343800/files/figure.png" pageId="6" pageNumber="271" targetBox="[217,1351,193,1167]" targetPageId="6">
|
||
<paragraph id="8BC44777FFCDFF96658EFB7DFCA81CE9" blockId="6.[151,867,1191,1213]" box="[151,867,1191,1213]" pageId="6" pageNumber="271">
|
||
<emphasis id="B90F9B65FFCDFF96658EFB7DFED91CE8" bold="true" box="[151,274,1191,1213]" pageId="6" pageNumber="271">FIGURE 4.</emphasis>
|
||
Distribution map of
|
||
<taxonomicName id="4C7B3CF4FFCDFF9664FEFB72FD941CE9" ID-CoL="8MRDJ" box="[487,607,1192,1213]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90F9B65FFCDFF9664FEFB72FD941CE9" box="[487,607,1192,1213]" italics="true" pageId="6" pageNumber="271">Ctenoneura</emphasis>
|
||
</taxonomicName>
|
||
species in Hainan Island.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC44777FFCDFF9665DEFB3EFE0E1DAD" blockId="6.[151,1436,1252,1529]" pageId="6" pageNumber="271">
|
||
<emphasis id="B90F9B65FFCDFF9665DEFB3EFE551CA9" bold="true" box="[199,414,1252,1277]" pageId="6" pageNumber="271">Taxonomic status.</emphasis>
|
||
So far the subfamily status of
|
||
<taxonomicName id="4C7B3CF4FFCDFF9667ECFB3CFCB21CA9" ID-CoL="8MRDJ" box="[757,889,1254,1277]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90F9B65FFCDFF9667ECFB3CFCB21CA9" box="[757,889,1254,1277]" italics="true" pageId="6" pageNumber="271">Ctenoneura</emphasis>
|
||
</taxonomicName>
|
||
is still unclear.
|
||
<bibRefCitation id="EFEA3A86FFCDFF966133FB3EFB081CA9" author="Princis" box="[1066,1219,1252,1277]" pageId="6" pageNumber="271" refString="Princis, K. (1950) Entomological results from the Swedish Expedition 1934 to Burma and British India, Blattaria. Arkiv for Zoologi, 1, 203 - 222." type="journal article" year="1950">Princis (1950</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFEA3A86FFCDFF9661D7FB3EFAC21CA9" author="Princis" box="[1230,1289,1252,1277]" pageId="6" pageNumber="271" refString="Princis, K. (1953) Kleine Beitrage zur Kenntnis der Blattarien und ihrer Verbreitung. VI. Entomologisk Tidskrift, 74, 203 - 213." type="journal article" year="1953">1953</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFEA3A86FFCDFF96600CFB3EFA9B1CA9" author="Princis" box="[1301,1360,1252,1277]" pageId="6" pageNumber="271" refString="Princis, K. (1963) Blattariae: Suborde Polyphagoidea: Fam.: Homoeogamiidae, Euthyrrhaphidae, Latindiidae, Anacompsidae, Atticolidae, Attaphilidae; Subordo Blaberoidea: Fam. Blaberidae. In: Beier, M. (Ed.), Orthopterorum Catalogus. Pars 4. Uitgeverij Dr. W. Junk, ' s-Gravenhage, pp. 77 - 172." type="journal article" year="1963">1963</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFEA3A86FFCDFF966042FB3EFA5F1CA9" author="Princis" box="[1371,1428,1252,1277]" pageId="6" pageNumber="271" refString="Princis, K. (1967) Kleine Beitrage zur Kenntnis der Blattarien und ihrer Verbreitung. X. Opuscula Entomologica Lund, 32, 141 - 151." type="journal article" year="1967">1967</bibRefCitation>
|
||
) put this genus in
|
||
<taxonomicName id="4C7B3CF4FFCDFF96647BFAD2FE2A1D75" ID-CoL="8N7YB" box="[354,481,1288,1313]" class="Insecta" family="Corydiidae" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="271" phylum="Arthropoda" rank="subFamily" subFamily="Latindiinae">Latindiinae</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C7B3CF4FFCDFF9664EAFAD2FDB91D75" box="[499,626,1288,1313]" class="Insecta" family="Latindiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="271" phylum="Arthropoda" rank="family">Latindiidae</taxonomicName>
|
||
without giving any reasons,
|
||
<bibRefCitation id="EFEA3A86FFCDFF9666DCFAD2FB821D75" author="Roth" box="[965,1097,1288,1313]" pageId="6" pageNumber="271" refString="Roth, L. M. (1993) Revision of the cockroach genus Ctenoneura Hanitsch (Blattaria, Polyphagidae). Tijdschrift voor Entomologie, 136, 83 - 109." type="journal article" year="1993">Roth (1993</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFEA3A86FFCDFF966141FAD2FB6A1D75" author="Roth" box="[1112,1185,1288,1313]" pageId="6" pageNumber="271" refString="Roth, L. M. (1995 b) Description of a new species of Ctenoneura Hanitsch from Sabah (Blattaria, Polyphagidae). Tijdschrift voor Entomologie, 138, 117 - 119." type="journal article" year="1995" yearSuffix="b">1995b</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFEA3A86FFCDFF9661B6FAD3FB221D75" author="Roth" box="[1199,1257,1289,1313]" pageId="6" pageNumber="271" refString="Roth, L. M. (2003) Systematics and phylogeny of cockroaches (Dictyoptera: Blattaria). Oriental Insects, 37, 1 - 186. https: // doi. org / 10.1080 / 00305316.2003.10417344" type="journal article" year="2003">2003</bibRefCitation>
|
||
) and
|
||
<bibRefCitation id="EFEA3A86FFCDFF966035FAD2FF2A1D11" author="Beccaloni" pageId="6" pageNumber="271" refString="Beccaloni, G. W. (2014) Cockroach Species File Online. Version 5.0 / 5.0. World Wide Web electronic publication. Available from: http: // Cockroach. SpeciesFile. org (accessed 11 January 2016)" type="journal article" year="2014">Beccaloni (2014)</bibRefCitation>
|
||
did not place it in any subfamily. The many distinctive characters (the much reduced left phallomere, the absence of genital hook, specialized and asymmetrical subgenital plate, single stylus, smooth body, intercalary vein presenting in tegmina and wings, well developed M and simplified CuA in tegmen) make it unique in the
|
||
<taxonomicName id="4C7B3CF4FFCDFF96658EFA42FEDC1DE5" ID-CoL="8MPDM" box="[151,279,1432,1457]" class="Insecta" family="Corydiidae" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="271" phylum="Arthropoda" rank="family">Corydiidae</taxonomicName>
|
||
. It may belong to a new subfamily named Ctenoneuriinae. However, more
|
||
<taxonomicName id="4C7B3CF4FFCDFF966170FA43FB251DE4" ID-CoL="8MRDJ" box="[1129,1262,1433,1456]" class="Insecta" family="Corydiidae" genus="Ctenoneura" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="271" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90F9B65FFCDFF966170FA43FB251DE4" box="[1129,1262,1433,1456]" italics="true" pageId="6" pageNumber="271">Ctenoneura</emphasis>
|
||
</taxonomicName>
|
||
examples need to studied, more comparison studies within Corydioidea genera need to be done, and molecular methods need to be used to solve this problem.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |