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<document id="AB87050344345BCC869FB0997FB36B40" ID-DOI="10.11646/phytotaxa.598.4.3" ID-ISSN="1179-3163" ID-Zenodo-Dep="7983752" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1685426207320" checkinUser="plazi" docAuthor="Morales, Nicolás Gutiérrez, Moreno, Juan Sebastián, Karremans, Adam P. &amp; Gil-Amaya, Karen" docDate="2023" docId="03CE1958DF636B4CFF1DFA81841DDE68" docLanguage="en" docName="phytotaxa.598.4.3.pdf" docOrigin="Phytotaxa 598 (4)" docSource="http://dx.doi.org/10.11646/phytotaxa.598.4.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.6:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="6" docTitle="Restrepia santanderensis N. Gut. &amp; K. Gil-Amaya" docType="treatment" docVersion="2" lastPageNumber="299" masterDocId="FFF76120DF626B4AFF95FF758117DF21" masterDocTitle="Restrepia santanderensis (Orchidaceae: Pleurothallidinae), a new species from the western slope of the eastern Andes in Colombia" masterLastPageNumber="300" masterPageNumber="293" pageNumber="294" updateTime="1685426709795" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="F1A34DD2F7AA5C424A3CAB8AD576812C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="A97497F8621C2AF49553937B9FB1AD54">Restrepia santanderensis (Orchidaceae: Pleurothallidinae), a new species from the western slope of the eastern Andes in Colombia</mods:title>
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<mods:namePart id="CE02154B21825029D1367FC81DA7A402">Morales, Nicolás Gutiérrez</mods:namePart>
<mods:nameIdentifier id="7C246C49889FE51A13AA7A9B27A8C187" type="ORCID">0000-0002-6592-9326</mods:nameIdentifier>
<mods:affiliation id="65989918E89208EA4491D4FADEEA50DE">Instituto de Biociências de Rio Claro, Departamento de Botânica, Universidade Estadual Paulista - UNESP, Av. 24 A 1515, Bela Vista, 13506 - 900, Caixa Postal 199, Rio Claro, São Paulo, Brazil. &amp; Grupo de Investigación Schultes, Fundación Ecotonos, Valle del Cauca, Cali, Colombia &amp; nicolaiequal @ gmail. com; https: // orcid. org / 0000 - 0002 - 6592 - 9326</mods:affiliation>
<mods:nameIdentifier id="F8C90835848F6385D73BC064A156EEE2" type="email">nicolaiequal@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B10B9B6D78CFA2427F7EEF01E0F6FFAE">Moreno, Juan Sebastián</mods:namePart>
<mods:nameIdentifier id="1F541F77EDB976EC5958AEAE7EC4349A" type="ORCID">0000-0002-0875-9498</mods:nameIdentifier>
<mods:affiliation id="9F7C4FA122681556BDC5D31E7744F8CF">Grupo de Investigación Schultes, Fundación Ecotonos, Valle del Cauca, Cali, Colombia &amp; Departamento de Biología, Universidad del Valle, Calle 13 # 100 - 00, Cali, Colombia &amp; semoreno 113 @ gmail. com; https: // orcid. org / 0000 - 0002 - 0875 - 9498</mods:affiliation>
<mods:nameIdentifier id="098A05A95FCFA736CCE79F21A4C293DB" type="email">semoreno113@gmail.com</mods:nameIdentifier>
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<mods:role id="07AD56B9E328FD6C28D672C6BA4EED98">
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<mods:namePart id="4DA94DC917F6A4BA52FED6D956BCB699">Karremans, Adam P.</mods:namePart>
<mods:nameIdentifier id="69F3E367FDF69664D9354EED8A5AF6F8" type="ORCID">0000-0001-5987-7710</mods:nameIdentifier>
<mods:affiliation id="B0D0DC4378B4761CCFDDCB40610596D6">Lankester Botanical Garden, University of Costa Rica, P. O. Box 302 - 7050 Cartago, Costa Rica &amp; Naturalis Biodiversity Center, Evolutionary Ecology Group, Sylviusweg 72, 2333 BE, Leiden, The Netherlands &amp; adam. karremans @ ucr. ac. cr; https: // orcid. org / 0000 - 0001 - 5987 - 7710</mods:affiliation>
<mods:nameIdentifier id="3060C263FB6487E024CF40CFEECDE29E" type="email">adam.karremans@ucr.ac.cr</mods:nameIdentifier>
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<mods:namePart id="2451C75D9F597F19E3A4A6EEB6CD8279">Gil-Amaya, Karen</mods:namePart>
<mods:nameIdentifier id="3832C8431A5A36D86B70255197AA0CD4" type="ORCID">0000-0002-2088-1541</mods:nameIdentifier>
<mods:affiliation id="E544433EC12A0AEE9332E64EE46F49D4">Grupo de Investigación Schultes, Fundación Ecotonos, Valle del Cauca, Cali, Colombia &amp; Lankester Botanical Garden, University of Costa Rica, P. O. Box 302 - 7050 Cartago, Costa Rica &amp; kgilecologa @ gmail. com; https: // orcid. org / 0000 - 0002 - 2088 - 1541</mods:affiliation>
<mods:nameIdentifier id="B54BD988B41A58B7415B5F45F84FD358" type="email">kgilecologa@gmail.com</mods:nameIdentifier>
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<mods:title id="14B1B6C7B24E1F5DA273790F0EF9FCE9">Phytotaxa</mods:title>
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<mods:date id="2D5689234F90E88A5E4F7C17AE09D299">2023</mods:date>
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<mods:number id="2E2E1879F7A4EB6E58CC5D1D648FF55A">2023-05-29</mods:number>
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<mods:number id="6B9A05DBFBCF825B0C6D1BD424CA447F">598</mods:number>
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<mods:url id="4B622016E2FDE6ADAC0871D209ADDEE8">http://dx.doi.org/10.11646/phytotaxa.598.4.3</mods:url>
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<mods:classification id="2D78899E3E5B6F2A530E856DC4CE7F8D">journal article</mods:classification>
<mods:identifier id="9A12FC7BE66EE78ACBDF4BFBEB3B801A" type="DOI">10.11646/phytotaxa.598.4.3</mods:identifier>
<mods:identifier id="157DD874BD13CB0B6FBDFC1D08D695D7" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="0441BBCE3AA8AAD3B6AF751A5C31238B" type="Zenodo-Dep">7983752</mods:identifier>
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<treatment id="03CE1958DF636B4CFF1DFA81841DDE68" ID-DOI="http://doi.org/10.5281/zenodo.7983754" ID-Zenodo-Dep="7983754" LSID="urn:lsid:plazi:treatment:03CE1958DF636B4CFF1DFA81841DDE68" httpUri="http://treatment.plazi.org/id/03CE1958DF636B4CFF1DFA81841DDE68" lastPageId="6" lastPageNumber="299" pageId="1" pageNumber="294">
<subSubSection id="C37DFBC5DF636B4BFF1DFA818531D92F" box="[136,1062,1524,1551]" pageId="1" pageNumber="294" type="nomenclature">
<paragraph id="8BD8A84EDF636B4BFF1DFA818531D92F" blockId="1.[136,1062,1524,1551]" box="[136,1062,1524,1551]" pageId="1" pageNumber="294">
<heading id="D0901F22DF636B4BFF1DFA818531D92F" box="[136,1062,1524,1551]" fontSize="11" level="2" pageId="1" pageNumber="294" reason="3">
<taxonomicName id="4C67D3CDDF636B4BFF1DFA8183DDD92F" authority="N. Gut. &amp; K. Gil-Amaya" authorityName="N. Gut. &amp; K. Gil-Amaya" box="[136,714,1524,1551]" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="294" phylum="Tracheophyta" rank="species" species="santanderensis">
<emphasis id="B913745CDF636B4BFF1DFA8180BED92E" bold="true" box="[136,425,1524,1551]" italics="true" pageId="1" pageNumber="294">Restrepia santanderensis</emphasis>
N. Gut. &amp; K. Gil-Amaya
</taxonomicName>
,
<emphasis id="B913745CDF636B4BFD40FA808234D92F" box="[725,803,1525,1550]" italics="true" pageId="1" pageNumber="294">sp nov.</emphasis>
(
<figureCitation id="135CB4CBDF636B4BFCA4FA8182B5D92F" box="[817,930,1524,1550]" captionStart="FIGURE 1" captionStartId="2.[136,229,1877,1899]" captionTargetBox="[151,1421,188,1843]" captionTargetId="figure-23@2.[151,1436,187,1850]" captionTargetPageId="2" captionText="FIGURE 1. Illustration of Restrepia santanderensis, A. Habit. B. Flower. C. Dissected perianth with magnified trichomes of the synsepal. D. Lip, column, ovary, floral bract in lateral view. E. Lip, lateral and adaxial view. F. Column in ventral view, without the anther G. Anther cap and pollinaria. Illustration by Nicolás Gutiérrez Morales and Sebastián Moreno from the holotype." figureDoi="http://doi.org/10.5281/zenodo.7983758" httpUri="https://zenodo.org/record/7983758/files/figure.png" pageId="1" pageNumber="294">Figures. 1</figureCitation>
,
<figureCitation id="135CB4CBDF636B4BFC25FA8182D7D92F" box="[944,960,1524,1550]" captionStart="FIGURE 2" captionStartId="3.[136,229,1877,1899]" captionTargetBox="[151,1436,190,1853]" captionTargetId="figure-21@3.[151,1436,190,1853]" captionTargetPageId="3" captionText="FIGURE 2. Lankester composite dissection plate (LCDP) of Restrepia santanderensis. A. Habit. B. Flower. C. Dissected perianth with magnified trichomes of the synsepal. D. Lip and column, lateral view. E. Lip, adaxial view. F. Column in ventral view. Photographs by Nicolás Gutiérrez Morales, from the holotype." figureDoi="http://doi.org/10.5281/zenodo.7983760" httpUri="https://zenodo.org/record/7983760/files/figure.png" pageId="1" pageNumber="294">2</figureCitation>
,
<figureCitation id="135CB4CBDF636B4BFC59FA818509D92F" box="[972,1054,1524,1550]" captionStart="FIGURE 3" captionStartId="4.[136,229,1741,1763]" captionTargetBox="[262,1326,190,1716]" captionTargetId="figure-23@4.[262,1326,190,1716]" captionTargetPageId="4" captionText="FIGURE 3. Restrepia santanderensis N. Gutiérrez &amp; K. Gil-Amaya, and similar morphological species. A. R. flosculata. B. R. mendozae. C. R. santanderensis. D. Capsules of R. santanderensis. Photographs byAndreas Key (A), Joost Riksen (B) from cultivated specimens and Nicolás Gutiérrez Morales (C, D) from the paratype." figureDoi="http://doi.org/10.5281/zenodo.7983764" httpUri="https://zenodo.org/record/7983764/files/figure.png" pageId="1" pageNumber="294">3C, 3D</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37DFBC5DF636B4BFF1DF96981C5D873" pageId="1" pageNumber="294" type="materials_examined">
<paragraph id="8BD8A84EDF636B4BFF1DF9698330D974" blockId="1.[136,1452,1564,1622]" pageId="1" pageNumber="294">
<emphasis id="B913745CDF636B4BFF1DF96981D5D910" bold="true" box="[136,194,1564,1585]" pageId="1" pageNumber="294">Type:</emphasis>
—
<materialsCitation id="3B0FA213DF636B4BFF4EF9698334D974" collectingDate="2016-01" collectingDateMax="2020-01" collectingDateMin="2016-01" collectorName="La Belleza &amp; M. Calderon" country="Colombia" elevation="1950" location="Vereda Berlin" municipality="La Belleza" pageId="1" pageNumber="294" specimenCount="1" stateProvince="Santander" typeStatus="holotype">
<collectingCountry id="F370E8DEDF636B4BFF4EF969804AD910" box="[219,349,1564,1585]" name="Colombia" pageId="1" pageNumber="294">COLOMBIA</collectingCountry>
.
<collectingRegion id="49A366ACDF636B4BFEFCF96980DAD913" box="[361,461,1564,1586]" country="Colombia" name="Santander" pageId="1" pageNumber="294">Santander</collectingRegion>
:
<collectorName id="2692CD98DF636B4BFE4DF9698353D913" box="[472,580,1564,1586]" pageId="1" pageNumber="294">
<collectingMunicipality id="6BBC3234DF636B4BFE4DF9698353D913" box="[472,580,1564,1586]" pageId="1" pageNumber="294">La Belleza</collectingMunicipality>
</collectorName>
,
<location id="8EB8FE95DF636B4BFDC5F96983CED913" LSID="urn:lsid:plazi:treatment:03CE1958DF636B4CFF1DFA81841DDE68:8EB8FE95DF636B4BFDC5F96983CED913" box="[592,729,1564,1586]" country="Colombia" municipality="La Belleza" name="Vereda Berlin" pageId="1" pageNumber="294" stateProvince="Santander">Vereda Berlin</location>
,
<quantity id="4C9F05ABDF636B4BFD70F9698224D910" box="[741,819,1564,1586]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="1" pageNumber="294" unit="m" value="1950.0">
<elevation id="004A4F7DDF636B4BFD70F9698224D910" box="[741,819,1564,1586]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="1" pageNumber="294" unit="m" value="1950.0">1950 m</elevation>
</quantity>
,
<date id="FFD98E8EDF636B4BFCAAF96982D3D913" box="[831,964,1564,1586]" pageId="1" pageNumber="294" value="2016-01">
<collectingDate id="EF9D7766DF636B4BFCAAF96982D3D913" box="[831,964,1564,1586]" pageId="1" pageNumber="294" value="2016-01">January 2016</collectingDate>
</date>
, flowering under cultivation in
<date id="FFD98E8EDF636B4BFA95F9698492D910" box="[1280,1413,1564,1585]" pageId="1" pageNumber="294" value="2020-01">
<collectingDate id="EF9D7766DF636B4BFA95F9698492D910" box="[1280,1413,1564,1585]" pageId="1" pageNumber="294" value="2020-01">January 2020</collectingDate>
</date>
,
<emphasis id="B913745CDF636B4BFA04F969806FD974" italics="true" pageId="1" pageNumber="294">
<collectorName id="2692CD98DF636B4BFA04F969800FD974" pageId="1" pageNumber="294">M. Calderón</collectorName>
et al. 191
</emphasis>
(
<typeStatus id="54DC16ECDF636B4BFE10F93580C9D977" box="[389,478,1600,1622]" pageId="1" pageNumber="294">holotype</typeStatus>
: JBB!)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD8A84EDF636B4BFF1DF9F081C5D873" blockId="1.[136,1452,1668,2091]" pageId="1" pageNumber="294">
<taxonomicName id="4C67D3CDDF636B4BFF1DF9F080B5D9BF" box="[136,418,1668,1694]" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="294" phylum="Tracheophyta" rank="species" species="santanderensis">
<emphasis id="B913745CDF636B4BFF1DF9F080B5D9BF" box="[136,418,1668,1694]" italics="true" pageId="1" pageNumber="294">Restrepia santanderensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C67D3CDDF636B4BFDACF9F08280D9BE" authority="Luer (1996: 157)" authorityName="Luer" authorityPageNumber="157" authorityYear="1996" box="[569,919,1668,1695]" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="294" phylum="Tracheophyta" rank="species" species="mendozae">
<emphasis id="B913745CDF636B4BFDACF9F083DDD9BF" box="[569,714,1668,1694]" italics="true" pageId="1" pageNumber="294">R. mendozae</emphasis>
<bibRefCitation id="EFF6D5BFDF636B4BFD47F9F08280D9BE" author="Luer, C. A." box="[722,919,1668,1695]" pageId="1" pageNumber="294" pagination="1 - 168" refId="ref4089" refString="Luer, C. A. (1996) Icones pleurothallidinarum XIII. Systematics of the genus Restrepia (Orchidaceae). Monographs in Systematic Botany from the Missouri Botanical Garden 59: 1 - 168." type="journal article" year="1996">Luer (1996: 157)</bibRefCitation>
</taxonomicName>
, but can be distinguished by the sheaths of the ramicauls completely brown-dotted (
<emphasis id="B913745CDF636B4BFDBAF9DC8351D9E3" box="[559,582,1705,1730]" italics="true" pageId="1" pageNumber="294">vs</emphasis>
. only lowermost sheath dotted), the sepals partially spreading, yellow with vinaceous stripes in the synsepal (
<emphasis id="B913745CDF636B4BFD97F9B8830ED9C7" box="[514,537,1741,1766]" italics="true" pageId="1" pageNumber="294">vs</emphasis>
. sepals completely spreading, yellow with purple spots), the dorsal sepal prostrate (
<emphasis id="B913745CDF636B4BFF04F98481BFD82B" box="[145,168,1777,1802]" italics="true" pageId="1" pageNumber="294">vs</emphasis>
. dorsal sepal erect), the synsepal with trichomes along the veins on the adaxial surface (
<emphasis id="B913745CDF636B4BFB3CF98485D7D82B" box="[1193,1216,1777,1802]" italics="true" pageId="1" pageNumber="294">vs</emphasis>
. synsepal glabrous), the straight petals (
<emphasis id="B913745CDF636B4BFEF7F8608097D80F" box="[354,384,1813,1838]" italics="true" pageId="1" pageNumber="294">vs.</emphasis>
decurved petals), and the lip yellow with intense rose at base (
<emphasis id="B913745CDF636B4BFBC5F8608578D80F" box="[1104,1135,1813,1838]" italics="true" pageId="1" pageNumber="294">vs.</emphasis>
entirely yellow with purple spots).
</paragraph>
</subSubSection>
<subSubSection id="C37DFBC5DF636B4FFF28F8298307DC45" lastPageId="5" lastPageNumber="298" pageId="1" pageNumber="294" type="description">
<paragraph id="8BD8A84EDF636B4FFF28F8298307DC45" blockId="1.[136,1452,1668,2091]" lastBlockId="5.[136,1452,158,2093]" lastPageId="5" lastPageNumber="298" pageId="1" pageNumber="294">
<emphasis id="B913745CDF636B4BFF28F829807AD857" bold="true" box="[189,365,1884,1910]" pageId="1" pageNumber="294">Description:—</emphasis>
<emphasis id="B913745CDF636B4BFEF8F82980BDD857" box="[365,426,1884,1910]" italics="true" pageId="1" pageNumber="294">Plant</emphasis>
epiphytic, caespitose, occasionally prolific, erect to sub-erect, up to
<quantity id="4C9F05ABDF636B4BFB4BF829843FD856" box="[1246,1320,1884,1911]" metricMagnitude="-1" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="294" unit="cm" value="24.0">24 cm</quantity>
tall.
<emphasis id="B913745CDF636B4BFAF9F82884BBD857" box="[1388,1452,1885,1910]" italics="true" pageId="1" pageNumber="294">Roots</emphasis>
slender, flexuous,
<emphasis id="B913745CDF636B4BFECCF8F4806DD8BB" box="[345,378,1921,1946]" italics="true" pageId="1" pageNumber="294">ca.</emphasis>
<quantity id="4C9F05ABDF636B4BFE11F8F580D0D8BA" box="[388,455,1920,1947]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="294" unit="mm" value="1.0">1 mm</quantity>
in diameter.
<emphasis id="B913745CDF636B4BFDF5F8F583C1D8BB" box="[608,726,1920,1946]" italics="true" pageId="1" pageNumber="294">Ramicauls</emphasis>
erect,
<quantity id="4C9F05ABDF636B4BFCB3F8F5829ED8BA" box="[806,905,1920,1947]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="13.0" metricValueMin="2.0" pageId="1" pageNumber="294" unit="cm" value="7.5" valueMax="13.0" valueMin="2.0">2–13 cm</quantity>
long, enclosed by 5–6, whitish, brown-spotted, papery, loose, imbricating, oblique, laterally compressed, acute sheaths, the uppermost prominent, extended as long as the pedicel, acuminate.
<emphasis id="B913745CDF636B4BFE2FF8BC831FD8C3" box="[442,520,1993,2018]" italics="true" pageId="1" pageNumber="294">Leaves</emphasis>
erect, elliptical-ovate, coriaceous, sub-acute, minutely tri-denticulate, 4.5–11.2 ×
<quantity id="4C9F05ABDF636B4BFF1DF899801FD726" box="[136,264,2028,2055]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="4.3" metricValueMin="2.2" pageId="1" pageNumber="294" unit="cm" value="3.25" valueMax="4.3" valueMin="2.2">2.2–4.3 cm</quantity>
, the base cuneate contracted into a short, twisted petiole.
<emphasis id="B913745CDF636B4BFC18F8998534D727" box="[909,1059,2028,2054]" italics="true" pageId="1" pageNumber="294">Inflorescence</emphasis>
up to
<quantity id="4C9F05ABDF636B4BFBFCF89985AED726" box="[1129,1209,2028,2055]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="1" pageNumber="294" unit="cm" value="3.9">3.9 cm</quantity>
, emerging without an annulus
<quantity id="4C9F05ABDF636B4BFF7DF7658050D70A" box="[232,327,2064,2091]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="1" pageNumber="294" unit="mm" value="7.0" valueMax="9.0" valueMin="5.0">5–9 mm</quantity>
below the apex of the ramicaul, with a solitary yellow flower, produced successively in a fascicle behind the leaf, in a slender, sub-erect, 2.0–
<quantity id="4C9F05ABDF676B4FFDFBFFEB83AEDF98" box="[622,697,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="5" pageNumber="298" unit="cm" value="2.7">2.7 cm</quantity>
long peduncle, subtended by a tubular, oblique spathe,
<quantity id="4C9F05ABDF676B4FFAB0FFEB84BBDF98" box="[1317,1452,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="9.55" metricValueMax="9.9" metricValueMin="9.2" pageId="5" pageNumber="298" unit="mm" value="9.55" valueMax="9.9" valueMin="9.2">9.2–9.9 mm</quantity>
long, floral bract translucent, oblique, slightly inflated, apiculate, to
<quantity id="4C9F05ABDF676B4FFC1AFFB782F1DFFC" box="[911,998,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="5" pageNumber="298" unit="mm" value="6.9">6.9 mm</quantity>
long, enclosing the short, stout pedicel, 1.0–
<quantity id="4C9F05ABDF676B4FFF2EFF938019DE20" box="[187,270,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="298" unit="mm" value="1.3">1.3 mm</quantity>
long, with a filament
<quantity id="4C9F05ABDF676B4FFD95FF938391DE20" box="[512,646,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.5" metricValueMin="2.8" pageId="5" pageNumber="298" unit="mm" value="3.15" valueMax="3.5" valueMin="2.8">2.8–3.5 mm</quantity>
long.
<emphasis id="B913745CDF676B4FFD5FFF928207DE20" box="[714,784,231,257]" italics="true" pageId="5" pageNumber="298">Ovary</emphasis>
slightly arcuate, lightly costate,
<quantity id="4C9F05ABDF676B4FFBE2FF93841CDE20" box="[1143,1291,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="5.435" metricValueMax="6.17" metricValueMin="4.7" pageId="5" pageNumber="298" unit="mm" value="5.435" valueMax="6.17" valueMin="4.7">4.7–6.17 mm</quantity>
long.
<emphasis id="B913745CDF676B4FFAC5FF9384BBDE21" box="[1360,1452,230,256]" italics="true" pageId="5" pageNumber="298">Flowers</emphasis>
resupinate, intermediate-sized, to
<quantity id="4C9F05ABDF676B4FFD96FE7F8358DE04" box="[515,591,266,293]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="298" unit="cm" value="2.5">2.5 cm</quantity>
long.
<emphasis id="B913745CDF676B4FFD02FE7F83F7DE05" box="[663,736,266,292]" italics="true" pageId="5" pageNumber="298">Sepals</emphasis>
pale yellow, membranous, not completely spreading, the dorsal sepal free, narrowly ovate and slightly concave below the lower third, 5-veined, contracted into a long tail above the lower third, with the apex clavate-thickened,
<quantity id="4C9F05ABDF676B4FFD10FE2783D0DE4C" box="[645,711,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="298" unit="mm" value="1.0">1 mm</quantity>
thick at the apex, 17.0–
<quantity id="4C9F05ABDF676B4FFC41FE27852EDE4C" box="[980,1081,338,365]" metricMagnitude="-2" metricUnit="m" metricValue="2.26" pageId="5" pageNumber="298" unit="mm" value="22.6">22.6 mm</quantity>
long,
<quantity id="4C9F05ABDF676B4FFB15FE278433DE4C" box="[1152,1316,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="2.235" metricValueMax="2.56" metricValueMin="1.91" pageId="5" pageNumber="298" unit="mm" value="2.235" valueMax="2.56" valueMin="1.91">1.91–2.56 mm</quantity>
wide above the base, the lateral sepals somewhat striped in vinaceous near the margins of the basal third, each 7-veined, connate ca. 19.0–
<quantity id="4C9F05ABDF676B4FFF78FEEF8047DE94" box="[237,336,410,437]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" pageId="5" pageNumber="298" unit="mm" value="21.8">21.8 mm</quantity>
into a narrowly oblong, slightly concave at base, bifid, subacute lamina, with glandular trichomes in the middle region and in the apex of the adaxial surface, along the veins,
<quantity id="4C9F05ABDF676B4FFC56FECB852BDEF8" box="[963,1084,446,473]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="5" pageNumber="298" unit="mm" value="23.5" valueMax="25.0" valueMin="22.0">22–25 mm</quantity>
long,
<quantity id="4C9F05ABDF676B4FFB11FECB840EDEF8" box="[1156,1305,446,473]" metricMagnitude="-3" metricUnit="m" metricValue="7.574999999999999" metricValueMax="7.8" metricValueMin="7.35" pageId="5" pageNumber="298" unit="mm" value="7.574999999999999" valueMax="7.8" valueMin="7.35">7.35–7.8 mm</quantity>
wide.
<emphasis id="B913745CDF676B4FFAF0FECB84BCDEF9" box="[1381,1451,446,472]" italics="true" pageId="5" pageNumber="298">Petals</emphasis>
translucent yellow, slender, narrowly linear-triangular, membranous, 3-veined, the margins minutely serrate near the base, attenuated above the middle, the apex clavate-thickened, 11.0–
<quantity id="4C9F05ABDF676B4FFC1FFD7382FBDD00" box="[906,1004,518,545]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="5" pageNumber="298" unit="mm" value="11.6">11.6 mm</quantity>
long, 1.0–
<quantity id="4C9F05ABDF676B4FFBF0FD7385ACDD00" box="[1125,1211,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="298" unit="mm" value="1.2">1.2 mm</quantity>
wide above the base.
<emphasis id="B913745CDF676B4FFF1DFD5E81BADD65" box="[136,173,555,580]" italics="true" pageId="5" pageNumber="298">Lip</emphasis>
yellow, vinaceous along the veins, the base intense rose and white, rectangular, 3-veined, truncated, 3-veined,
<quantity id="4C9F05ABDF676B4FFAEFFD5F81FCDD48" metricMagnitude="-3" metricUnit="m" metricValue="9.25" metricValueMax="10.9" metricValueMin="7.6" pageId="5" pageNumber="298" unit="mm" value="9.25" valueMax="10.9" valueMin="7.6">7.6– 10.9 mm</quantity>
long,
<quantity id="4C9F05ABDF676B4FFEA5FD3B80A0DD48" box="[304,439,590,617]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.6" metricValueMin="2.3" pageId="5" pageNumber="298" unit="mm" value="2.95" valueMax="3.6" valueMin="2.3">2.3–3.6 mm</quantity>
wide, the base a slightly concave, with erect margins, the sides with a translucent, uncinate process or cirrhi,
<emphasis id="B913745CDF676B4FFED8FD068078DDAD" box="[333,367,627,652]" italics="true" pageId="5" pageNumber="298">ca.</emphasis>
<quantity id="4C9F05ABDF676B4FFEE3FD0780EADDAC" box="[374,509,626,653]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.8" metricValueMin="1.4" pageId="5" pageNumber="298" unit="mm" value="1.6" valueMax="1.8" valueMin="1.4">1.4–1.8 mm</quantity>
long, each with a minute tooth at the base, the lip with a pair of low calli extending forward from the base of near the middle, the base connected to the column-foot by a white, rigid, cylindrical neck, the apical portion microscopically verrucose.
<emphasis id="B913745CDF676B4FFDF0FDCF83A9DDF5" box="[613,702,698,724]" italics="true" pageId="5" pageNumber="298">Column</emphasis>
greenish-white, the basal half slender, clavate, the margins of the apical half irregular, marked with intense yellow,
<quantity id="4C9F05ABDF676B4FFD22FDAB8229DDD8" box="[695,830,734,761]" metricMagnitude="-3" metricUnit="m" metricValue="5.35" metricValueMax="6.1" metricValueMin="4.6" pageId="5" pageNumber="298" unit="mm" value="5.35" valueMax="6.1" valueMin="4.6">4.6–6.1 mm</quantity>
long, the base pedestal-like with a pair of longitudinal, minutely papillose calli, ending in a pair of light orange, obtuse points, the anther and stigma ventral, the stigma subcuneiform.
<emphasis id="B913745CDF676B4FFEA7FC5380B8DC61" box="[306,431,806,832]" italics="true" pageId="5" pageNumber="298">Anther cap</emphasis>
incumbent, deciduous, whitish.
<taxonomicName id="4C67D3CDDF676B4FFCB6FC53826ADC61" box="[803,893,806,832]" class="Liliopsida" family="Poaceae" genus="Pollinia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Poales" pageId="5" pageNumber="298" phylum="Tracheophyta" rank="genus">
<emphasis id="B913745CDF676B4FFCB6FC53826ADC61" box="[803,893,806,832]" italics="true" pageId="5" pageNumber="298">Pollinia</emphasis>
</taxonomicName>
four, ovoid, in two pairs, united by a viscidium.
<emphasis id="B913745CDF676B4FFF1DFC3F81F3DC45" box="[136,228,842,868]" italics="true" pageId="5" pageNumber="298">Capsule</emphasis>
dehiscent,
<emphasis id="B913745CDF676B4FFEF6FC3E806ADC45" box="[355,381,843,868]" italics="true" pageId="5" pageNumber="298">ca</emphasis>
.
<quantity id="4C9F05ABDF676B4FFE1EFC3F80C0DC44" box="[395,471,842,869]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="298" unit="cm" value="1.5">1.5 cm</quantity>
long
</paragraph>
</subSubSection>
<caption id="DF18F8C6DF606B48FF1DF8208575D892" ID-DOI="http://doi.org/10.5281/zenodo.7983758" ID-Zenodo-Dep="7983758" httpUri="https://zenodo.org/record/7983758/files/figure.png" pageId="2" pageNumber="295" startId="2.[136,229,1877,1899]" targetBox="[151,1421,188,1843]" targetPageId="2" targetType="figure">
<paragraph id="8BD8A84EDF606B48FF1DF8208575D892" blockId="2.[136,1452,1877,1971]" pageId="2" pageNumber="295">
<emphasis id="B913745CDF606B48FF1DF82081EAD84B" bold="true" box="[136,253,1877,1899]" pageId="2" pageNumber="295">FIGURE 1.</emphasis>
Illustration of
<taxonomicName id="4C67D3CDDF606B48FE1CF82382D3D84A" authority=", A. Habit. B. Flower. C. Dissected" box="[393,964,1877,1899]" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="295" phylum="Tracheophyta" rank="species" species="santanderensis">
<emphasis id="B913745CDF606B48FE1CF823836ED84B" box="[393,633,1877,1899]" italics="true" pageId="2" pageNumber="295">Restrepia santanderensis</emphasis>
,
<emphasis id="B913745CDF606B48FD11F820838BD84B" bold="true" box="[644,668,1877,1898]" pageId="2" pageNumber="295">A.</emphasis>
Habit.
<emphasis id="B913745CDF606B48FD74F82383E0D84A" bold="true" box="[737,759,1878,1899]" pageId="2" pageNumber="295">B.</emphasis>
Flower.
<emphasis id="B913745CDF606B48FCDFF8208275D84A" bold="true" box="[842,866,1877,1899]" pageId="2" pageNumber="295">C.</emphasis>
Dissected
</taxonomicName>
perianth with magnified trichomes of the synsepal.
<emphasis id="B913745CDF606B48FF1DF80F81B7D8AE" bold="true" box="[136,160,1914,1935]" pageId="2" pageNumber="295">D.</emphasis>
Lip, column, ovary, floral bract in lateral view.
<emphasis id="B913745CDF606B48FDF0F80F836BD8AE" bold="true" box="[613,636,1914,1935]" pageId="2" pageNumber="295">E.</emphasis>
Lip, lateral and adaxial view.
<emphasis id="B913745CDF606B48FC0DF80F82BCD8AE" bold="true" box="[920,939,1914,1935]" pageId="2" pageNumber="295">F.</emphasis>
Column in ventral view, without the anther
<emphasis id="B913745CDF606B48FAD9F80C8472D8AE" bold="true" box="[1356,1381,1913,1935]" pageId="2" pageNumber="295">G.</emphasis>
Anther cap and pollinaria. Illustration by Nicolás Gutiérrez Morales and Sebastián Moreno from the holotype.
</paragraph>
</caption>
<caption id="DF18F8C6DF616B49FF1DF8208351D892" ID-DOI="http://doi.org/10.5281/zenodo.7983760" ID-Zenodo-Dep="7983760" httpUri="https://zenodo.org/record/7983760/files/figure.png" pageId="3" pageNumber="296" startId="3.[136,229,1877,1899]" targetBox="[151,1436,190,1853]" targetPageId="3" targetType="figure">
<paragraph id="8BD8A84EDF616B49FF1DF8208351D892" blockId="3.[136,1452,1877,1971]" pageId="3" pageNumber="296">
<emphasis id="B913745CDF616B49FF1DF82081E9D84B" bold="true" box="[136,254,1877,1899]" pageId="3" pageNumber="296">FIGURE 2.</emphasis>
Lankester composite dissection plate (LCDP) of
<taxonomicName id="4C67D3CDDF616B49FD4AF82382C7D84B" box="[735,976,1877,1899]" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="296" phylum="Tracheophyta" rank="species" species="santanderensis">
<emphasis id="B913745CDF616B49FD4AF82382C7D84B" box="[735,976,1877,1899]" italics="true" pageId="3" pageNumber="296">Restrepia santanderensis</emphasis>
</taxonomicName>
.
<emphasis id="B913745CDF616B49FC48F82082E3D84B" bold="true" box="[989,1012,1877,1898]" pageId="3" pageNumber="296">A.</emphasis>
Habit.
<emphasis id="B913745CDF616B49FBA8F8238544D84A" bold="true" box="[1085,1107,1878,1899]" pageId="3" pageNumber="296">B.</emphasis>
Flower.
<emphasis id="B913745CDF616B49FB3CF82085D6D84A" bold="true" box="[1193,1217,1877,1899]" pageId="3" pageNumber="296">C.</emphasis>
Dissected perianth with magnified trichomes of the synsepal.
<emphasis id="B913745CDF616B49FE60F80F831AD8AE" bold="true" box="[501,525,1914,1935]" pageId="3" pageNumber="296">D.</emphasis>
Lip and column, lateral view.
<emphasis id="B913745CDF616B49FCA2F80F8259D8AE" bold="true" box="[823,846,1914,1935]" pageId="3" pageNumber="296">E.</emphasis>
Lip, adaxial view.
<emphasis id="B913745CDF616B49FB9CF80F850CD8AE" bold="true" box="[1033,1051,1914,1935]" pageId="3" pageNumber="296">F.</emphasis>
Column in ventral view. Photographs by Nicolás Gutiérrez Morales, from the holotype.
</paragraph>
</caption>
<caption id="DF18F8C6DF666B4EFF1DF9B88395D80B" ID-DOI="http://doi.org/10.5281/zenodo.7983764" ID-Zenodo-Dep="7983764" httpUri="https://zenodo.org/record/7983764/files/figure.png" pageId="4" pageNumber="297" startId="4.[136,229,1741,1763]" targetBox="[262,1326,190,1716]" targetPageId="4" targetType="figure">
<paragraph id="8BD8A84EDF666B4EFF1DF9B88395D80B" blockId="4.[136,1452,1741,1835]" pageId="4" pageNumber="297">
<emphasis id="B913745CDF666B4EFF1DF9B881EAD9C3" bold="true" box="[136,253,1741,1763]" pageId="4" pageNumber="297">FIGURE 3.</emphasis>
<taxonomicName id="4C67D3CDDF666B4EFE97F9B88202D9C2" authority="N. Gutierrez &amp; K. Gil-Amaya" authorityName="N. Gutierrez &amp; K. Gil-Amaya" box="[258,789,1741,1763]" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="297" phylum="Tracheophyta" rank="species" species="santanderensis">
<emphasis id="B913745CDF666B4EFE97F9B880E6D9C3" box="[258,497,1741,1762]" italics="true" pageId="4" pageNumber="297">Restrepia santanderensis</emphasis>
N. Gutiérrez &amp; K. Gil-Amaya
</taxonomicName>
, and similar morphological species.
<emphasis id="B913745CDF666B4EFBE5F9B8859FD9C3" bold="true" box="[1136,1160,1741,1762]" pageId="4" pageNumber="297">A.</emphasis>
<taxonomicName id="4C67D3CDDF666B4EFB19F9B88412D9C3" authorityName="Luer" authorityYear="1982" box="[1164,1285,1741,1762]" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="297" phylum="Tracheophyta" rank="species" species="flosculata">
<emphasis id="B913745CDF666B4EFB19F9B88412D9C3" box="[1164,1285,1741,1762]" italics="true" pageId="4" pageNumber="297">R. flosculata</emphasis>
</taxonomicName>
.
<emphasis id="B913745CDF666B4EFA85F9B88431D9C3" bold="true" box="[1296,1318,1741,1762]" pageId="4" pageNumber="297">B.</emphasis>
<taxonomicName id="4C67D3CDDF666B4EFABEF9B884B2D9C3" authorityName="Luer" authorityYear="1996" box="[1323,1445,1741,1762]" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="297" phylum="Tracheophyta" rank="species" species="mendozae">
<emphasis id="B913745CDF666B4EFABEF9B884B2D9C3" box="[1323,1445,1741,1762]" italics="true" pageId="4" pageNumber="297">R. mendozae</emphasis>
</taxonomicName>
.
<emphasis id="B913745CDF666B4EFF1DF98481B7D826" bold="true" box="[136,160,1777,1799]" pageId="4" pageNumber="297">C.</emphasis>
<taxonomicName id="4C67D3CDDF666B4EFF31F984805AD827" box="[164,333,1777,1798]" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="297" phylum="Tracheophyta" rank="species" species="santanderensis">
<emphasis id="B913745CDF666B4EFF31F984805AD827" box="[164,333,1777,1798]" italics="true" pageId="4" pageNumber="297">R. santanderensis</emphasis>
</taxonomicName>
.
<emphasis id="B913745CDF666B4EFECDF9848067D827" bold="true" box="[344,368,1777,1798]" pageId="4" pageNumber="297">D.</emphasis>
Capsules of
<taxonomicName id="4C67D3CDDF666B4EFE7FF9848385D827" box="[490,658,1777,1798]" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="297" phylum="Tracheophyta" rank="species" species="santanderensis">
<emphasis id="B913745CDF666B4EFE7FF9848385D827" box="[490,658,1777,1798]" italics="true" pageId="4" pageNumber="297">R. santanderensis</emphasis>
</taxonomicName>
. Photographs by Andreas Key (A), Joost Riksen (B) from cultivated specimens and Nicolás Gutiérrez Morales (C, D) from the paratype.
</paragraph>
</caption>
<subSubSection id="C37DFBC5DF676B4FFF28FC1B85F6DBD0" pageId="5" pageNumber="298" type="distribution">
<paragraph id="8BD8A84EDF676B4FFF28FC1B85F6DBD0" blockId="5.[136,1452,158,2093]" pageId="5" pageNumber="298">
<emphasis id="B913745CDF676B4FFF28FC1B80E6DCA9" bold="true" box="[189,497,878,904]" pageId="5" pageNumber="298">Distribution and Ecology:</emphasis>
—
<taxonomicName id="4C67D3CDDF676B4FFD98FC1A8233DCA9" box="[525,804,878,904]" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="298" phylum="Tracheophyta" rank="species" species="santanderensis">
<emphasis id="B913745CDF676B4FFD98FC1A8233DCA9" box="[525,804,878,904]" italics="true" pageId="5" pageNumber="298">Restrepia santanderensis</emphasis>
</taxonomicName>
occurs in a small area the eastern Andes of
<collectingCountry id="F370E8DEDF676B4FFA90FC1B8460DCA9" box="[1285,1399,878,904]" name="Colombia" pageId="5" pageNumber="298">Colombia</collectingCountry>
. The specimen that served as
<typeStatus id="54DC16ECDF676B4FFE0BFCE680D9DC8C" box="[414,462,915,941]" pageId="5" pageNumber="298">type</typeStatus>
was found in
<quantity id="4C9F05ABDF676B4FFDE6FCE783DDDC8D" box="[627,714,914,940]" metricMagnitude="1" metricUnit="m" metricValue="5.12064" pageId="5" pageNumber="298" unit="in" value="2016.0">2016 in</quantity>
the municipality of La Belleza,
<collectingRegion id="49A366ACDF676B4FFBA8FCE785B8DC8D" box="[1085,1199,914,940]" country="Colombia" name="Santander" pageId="5" pageNumber="298">Santander</collectingRegion>
, near to the top of the waterfall called “El chorro de la Humareda” (
<figureCitation id="135CB4CBDF676B4FFD10FCC383FFDCF1" box="[645,744,950,976]" captionStart="FIGURE 4" captionStartId="6.[136,229,1080,1102]" captionTargetBox="[151,1436,353,1056]" captionTargetId="figure-117@6.[151,1436,353,1056]" captionTargetPageId="6" captionText="FIGURE 4. View of the Restrepia santanderensis habitat in the western slope of the eastern Andes in Colombia, near to the top of “La Humareda” waterfall, in La Belleza (Santander) municipality." figureDoi="http://doi.org/10.5281/zenodo.7983770" httpUri="https://zenodo.org/record/7983770/files/figure.png" pageId="5" pageNumber="298">Figure 4</figureCitation>
). The plant was found growing epiphytically, partially exposed to direct sunlight, at around four meters above the ground, on an isolated tree among pastures, not so far from the edge of the forest. Other orchid species in the area were
<taxonomicName id="4C67D3CDDF676B4FFC94FC8B85D3DB39" authority="Luer (1976: 108)" authorityName="Luer" authorityPageNumber="108" authorityYear="1976" box="[769,1220,1022,1049]" class="Liliopsida" family="Orchidaceae" genus="Pleurothallis" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="298" phylum="Tracheophyta" rank="species" species="furcifera">
<emphasis id="B913745CDF676B4FFC94FC8B82EDDB39" box="[769,1018,1022,1048]" italics="true" pageId="5" pageNumber="298">Pleurothallis furcifera</emphasis>
<bibRefCitation id="EFF6D5BFDF676B4FFB94FC8A85D3DB39" author="Luer, C. A." box="[1025,1220,1022,1049]" pageId="5" pageNumber="298" pagination="38 - 201" refId="ref4068" refString="Luer, C. A. (1976) Miscellaneous species of Pleurothallis. Selbyana 3: 38 - 201." type="journal article" year="1976">Luer (1976: 108)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C67D3CDDF676B4FFB5AFC8B808DDB1D" authority="Christenson (2013: 174)" authorityName="Christenson" authorityPageNumber="174" authorityYear="2013" class="Liliopsida" family="Orchidaceae" genus="Maxillaria" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="298" phylum="Tracheophyta" rank="species" species="carrilloi">
<emphasis id="B913745CDF676B4FFB5AFC8B84BCDB39" box="[1231,1451,1022,1048]" italics="true" pageId="5" pageNumber="298">Maxillaria carrilloi</emphasis>
<bibRefCitation id="EFF6D5BFDF676B4FFF1DFB57808DDB1D" author="Christenson, E. A." box="[136,410,1058,1084]" pageId="5" pageNumber="298" pagination="174 - 175" refId="ref3552" refString="Christenson, E. A. (2013) Maxillaria carrilloi. In: Christenson, E. A. Maxillaria, an unfinished monograph, Volume 1. published for Robert Christenson by Patricia Ann Harding, Lebanon, pp. 174 - 175." type="book chapter" year="2013">Christenson (2013: 174)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C67D3CDDF676B4FFE33FB578524DB1D" authority="F. Lehmann. &amp; Kraenzl (1899: 476)" authorityName="F. Lehmann. &amp; Kraenzl" authorityPageNumber="476" authorityYear="1899" box="[422,1075,1058,1085]" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="298" phylum="Tracheophyta" rank="species" species="cupreum">
<emphasis id="B913745CDF676B4FFE33FB57838EDB1D" box="[422,665,1058,1084]" italics="true" pageId="5" pageNumber="298">Epidendrum cupreum</emphasis>
F. Lehmann. &amp; Kraenzl (1899: 476)
</taxonomicName>
and
<taxonomicName id="4C67D3CDDF676B4FFBFCFB578360DB41" authority="E. Parra, Hagsater &amp; L. Sanchez (2013: 69)" authorityName="E. Parra, Hagsater &amp; L. Sanchez" authorityPageNumber="69" authorityYear="2013" class="Liliopsida" family="Orchidaceae" genus="Epidendrum" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="298" phylum="Tracheophyta" rank="species" species="fusagasugaense">
<emphasis id="B913745CDF676B4FFBFCFB5784BCDB1D" box="[1129,1451,1058,1084]" italics="true" pageId="5" pageNumber="298">Epidendrum fusagasugaense</emphasis>
E. Parra, Hágsater &amp; L. Sánchez (2013: 69)
</taxonomicName>
. A short expedition in May of 2022 failed to recover more individuals of the new species, plantations of
<taxonomicName id="4C67D3CDDF676B4FFE7BFB1F8296DBA5" authority="Lam (1794: 16)" authorityName="Lam" authorityPageNumber="16" authorityYear="1794" box="[494,897,1130,1157]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="5" pageNumber="298" phylum="Tracheophyta" rank="species" species="quitoense">
<emphasis id="B913745CDF676B4FFE7BFB1F83D4DBA5" box="[494,707,1130,1156]" italics="true" pageId="5" pageNumber="298">Solanum quitoense</emphasis>
Lam (1794: 16)
</taxonomicName>
, “lulo”, were introduced in recent years, and the tree where the original specimen was collected was not found. However, the steep walls of the canyon that receives the “Peña Bonita” river, which falls for more than 220 meters into a waterfall, has a well-preserved forest remnant in a gradient of elevation of
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, and may host a population of
<taxonomicName id="4C67D3CDDF676B4FFC57FBA285CDDBD1" box="[962,1242,1238,1264]" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="298" phylum="Tracheophyta" rank="species" species="santanderensis">
<emphasis id="B913745CDF676B4FFC57FBA285CDDBD1" box="[962,1242,1238,1264]" italics="true" pageId="5" pageNumber="298">Restrepia santanderensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C37DFBC5DF676B4CFF28FB8F841DDE68" lastPageId="6" lastPageNumber="299" pageId="5" pageNumber="298" type="etymology">
<paragraph id="8BD8A84EDF676B4FFF28FB8F8438DA35" blockId="5.[136,1452,158,2093]" box="[189,1327,1274,1301]" pageId="5" pageNumber="298">
<emphasis id="B913745CDF676B4FFF28FB8F805FDA35" bold="true" box="[189,328,1274,1300]" pageId="5" pageNumber="298">Etymology:</emphasis>
—Named for the Department of
<collectingRegion id="49A366ACDF676B4FFD23FB8F823FDA35" box="[694,808,1274,1300]" country="Colombia" name="Santander" pageId="5" pageNumber="298">Santander</collectingRegion>
,
<collectingCountry id="F370E8DEDF676B4FFCA6FB8F82B3DA35" box="[819,932,1274,1300]" name="Colombia" pageId="5" pageNumber="298">Colombia</collectingCountry>
, where the species was discovered.
</paragraph>
<paragraph id="8BD8A84EDF676B4FFF28FA6B85D8DA7D" blockId="5.[136,1452,158,2093]" pageId="5" pageNumber="298">
<materialsCitation id="3B0FA213DF676B4FFF28FA6B85DCDA7D" collectingDate="2016-01" collectingDateMax="2023-01" collectingDateMin="2016-01" collectorName="La Belleza &amp; N. Gutierrez &amp; J. de Jesus" country="Colombia" elevation="1950" location="Vereda Berlin" municipality="La Belleza" pageId="5" pageNumber="298" specimenCount="2" stateProvince="Santander">
<emphasis id="B913745CDF676B4FFF28FA6B83D7DA19" bold="true" box="[189,704,1310,1336]" pageId="5" pageNumber="298">
<typeStatus id="54DC16ECDF676B4FFF28FA6B833ADA19" box="[189,557,1310,1336]" isOtherTypeStatus="true" pageId="5" pageNumber="298">Additional material examined</typeStatus>
(
<typeStatus id="54DC16ECDF676B4FFDD4FA6B83A4DA19" box="[577,691,1310,1336]" pageId="5" pageNumber="298">paratype</typeStatus>
):
</emphasis>
—
<collectingCountry id="F370E8DEDF676B4FFD48FA6B8262DA19" box="[733,885,1310,1336]" name="Colombia" pageId="5" pageNumber="298">COLOMBIA</collectingCountry>
.
<collectingRegion id="49A366ACDF676B4FFC10FA6B82EEDA19" box="[901,1017,1310,1336]" country="Colombia" name="Santander" pageId="5" pageNumber="298">Santander</collectingRegion>
:
<collectorName id="2692CD98DF676B4FFB9DFA6A859EDA19" box="[1032,1161,1310,1337]" pageId="5" pageNumber="298">
<collectingMunicipality id="6BBC3234DF676B4FFB9DFA6A859EDA19" box="[1032,1161,1310,1337]" pageId="5" pageNumber="298">La Belleza</collectingMunicipality>
</collectorName>
,
<location id="8EB8FE95DF676B4FFB0DFA6B842CDA19" LSID="urn:lsid:plazi:treatment:03CE1958DF636B4CFF1DFA81841DDE68:8EB8FE95DF676B4FFB0DFA6B842CDA19" box="[1176,1339,1310,1336]" country="Colombia" municipality="La Belleza" name="Vereda Berlin" pageId="5" pageNumber="298" stateProvince="Santander">Vereda Berlin</location>
,
<quantity id="4C9F05ABDF676B4FFADEFA6B84B0DA18" box="[1355,1447,1310,1337]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="5" pageNumber="298" unit="m" value="1950.0">
<elevation id="004A4F7DDF676B4FFADEFA6B84B0DA18" box="[1355,1447,1310,1337]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="5" pageNumber="298" unit="m" value="1950.0">1950 m</elevation>
</quantity>
,
<date id="FFD98E8EDF676B4FFF1DFA368036DA7D" box="[136,289,1346,1373]" pageId="5" pageNumber="298" value="2016-01">
<collectingDate id="EF9D7766DF676B4FFF1DFA368036DA7D" box="[136,289,1346,1373]" pageId="5" pageNumber="298" value="2016-01">January 2016</collectingDate>
</date>
, flowered under cultivation in
<date id="FFD98E8EDF676B4FFDECFA368205DA7D" box="[633,786,1346,1373]" pageId="5" pageNumber="298" value="2023-01">
<collectingDate id="EF9D7766DF676B4FFDECFA368205DA7D" box="[633,786,1346,1373]" pageId="5" pageNumber="298" value="2023-01">January 2023</collectingDate>
</date>
,
<emphasis id="B913745CDF676B4FFC88FA368562DA7D" box="[797,1141,1346,1373]" italics="true" pageId="5" pageNumber="298">
<collectorName id="2692CD98DF676B4FFC88FA3682BFDA7C" box="[797,936,1347,1373]" pageId="5" pageNumber="298">N. Gutiérrez</collectorName>
&amp;
<collectorName id="2692CD98DF676B4FFC59FA368552DA7D" box="[972,1093,1346,1372]" pageId="5" pageNumber="298">J. de Jesus</collectorName>
241
</emphasis>
(JBB!)
</materialsCitation>
.
</paragraph>
<paragraph id="8BD8A84EDF676B4FFF28FA1382F0D930" blockId="5.[136,1452,158,2093]" pageId="5" pageNumber="298">
<emphasis id="B913745CDF676B4FFF28FA1380DADAA1" bold="true" box="[189,461,1382,1408]" pageId="5" pageNumber="298">Taxonomic Discussion:</emphasis>
—
<taxonomicName id="4C67D3CDDF676B4FFE7CFA1283E8DAA1" box="[489,767,1382,1408]" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="298" phylum="Tracheophyta" rank="species" species="santanderensis">
<emphasis id="B913745CDF676B4FFE7CFA1283E8DAA1" box="[489,767,1382,1408]" italics="true" pageId="5" pageNumber="298">Restrepia santanderensis</emphasis>
</taxonomicName>
(
<figureCitation id="135CB4CBDF676B4FFC9EFA13826DDAA1" box="[779,890,1382,1408]" captionStart="FIGURE 1" captionStartId="2.[136,229,1877,1899]" captionTargetBox="[151,1421,188,1843]" captionTargetId="figure-23@2.[151,1436,187,1850]" captionTargetPageId="2" captionText="FIGURE 1. Illustration of Restrepia santanderensis, A. Habit. B. Flower. C. Dissected perianth with magnified trichomes of the synsepal. D. Lip, column, ovary, floral bract in lateral view. E. Lip, lateral and adaxial view. F. Column in ventral view, without the anther G. Anther cap and pollinaria. Illustration by Nicolás Gutiérrez Morales and Sebastián Moreno from the holotype." figureDoi="http://doi.org/10.5281/zenodo.7983758" httpUri="https://zenodo.org/record/7983758/files/figure.png" pageId="5" pageNumber="298">Figures. 1</figureCitation>
,
<figureCitation id="135CB4CBDF676B4FFC13FA138281DAA1" box="[902,918,1382,1408]" captionStart="FIGURE 2" captionStartId="3.[136,229,1877,1899]" captionTargetBox="[151,1436,190,1853]" captionTargetId="figure-21@3.[151,1436,190,1853]" captionTargetPageId="3" captionText="FIGURE 2. Lankester composite dissection plate (LCDP) of Restrepia santanderensis. A. Habit. B. Flower. C. Dissected perianth with magnified trichomes of the synsepal. D. Lip and column, lateral view. E. Lip, adaxial view. F. Column in ventral view. Photographs by Nicolás Gutiérrez Morales, from the holotype." figureDoi="http://doi.org/10.5281/zenodo.7983760" httpUri="https://zenodo.org/record/7983760/files/figure.png" pageId="5" pageNumber="298">2</figureCitation>
,
<figureCitation id="135CB4CBDF676B4FFC35FA1382F8DAA1" box="[928,1007,1382,1408]" captionStart="FIGURE 3" captionStartId="4.[136,229,1741,1763]" captionTargetBox="[262,1326,190,1716]" captionTargetId="figure-23@4.[262,1326,190,1716]" captionTargetPageId="4" captionText="FIGURE 3. Restrepia santanderensis N. Gutiérrez &amp; K. Gil-Amaya, and similar morphological species. A. R. flosculata. B. R. mendozae. C. R. santanderensis. D. Capsules of R. santanderensis. Photographs byAndreas Key (A), Joost Riksen (B) from cultivated specimens and Nicolás Gutiérrez Morales (C, D) from the paratype." figureDoi="http://doi.org/10.5281/zenodo.7983764" httpUri="https://zenodo.org/record/7983764/files/figure.png" pageId="5" pageNumber="298">3C, 3D</figureCitation>
) can be distinguished from other species in the genus by the ramicauls covered by densely brown-spotted sheaths, the elongate uppermost sheath extending as long as the pedicel, the short inflorescences with intermediate-sized, partially spreading, yellow flowers, the elongate, prostrate dorsal sepal, the synsepal with trichomes along the veins on the adaxial surface, and the rectangular lip, truncate at the apex, with intense rose at base and microscopically verrucose.
</paragraph>
<paragraph id="8BD8A84EDF676B4FFF28F96F82B3D9E5" blockId="5.[136,1452,158,2093]" pageId="5" pageNumber="298">
The most similar species morphologically are
<taxonomicName id="4C67D3CDDF676B4FFD5DF96E8508D915" authority="Luer (1982: 127)" authorityName="Luer" authorityPageNumber="127" authorityYear="1982" box="[712,1055,1562,1589]" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="298" phylum="Tracheophyta" rank="species" species="flosculata">
<emphasis id="B913745CDF676B4FFD5DF96E8241D915" box="[712,854,1562,1588]" italics="true" pageId="5" pageNumber="298">R. flosculata</emphasis>
<bibRefCitation id="EFF6D5BFDF676B4FFCCAF96E8508D915" author="Luer, C. A." box="[863,1055,1562,1589]" pageId="5" pageNumber="298" pagination="127" refId="ref4042" refString="Luer, C. A. (1982) Miscellaneous new species and combinations in the Pleurothallidinae (Orchidaceae). Selbyana 7: 127." type="journal article" year="1982">Luer (1982: 127)</bibRefCitation>
</taxonomicName>
(
<figureCitation id="135CB4CBDF676B4FFBBAF96F85B0D915" box="[1071,1191,1562,1588]" captionStart="FIGURE 3" captionStartId="4.[136,229,1741,1763]" captionTargetBox="[262,1326,190,1716]" captionTargetId="figure-23@4.[262,1326,190,1716]" captionTargetPageId="4" captionText="FIGURE 3. Restrepia santanderensis N. Gutiérrez &amp; K. Gil-Amaya, and similar morphological species. A. R. flosculata. B. R. mendozae. C. R. santanderensis. D. Capsules of R. santanderensis. Photographs byAndreas Key (A), Joost Riksen (B) from cultivated specimens and Nicolás Gutiérrez Morales (C, D) from the paratype." figureDoi="http://doi.org/10.5281/zenodo.7983764" httpUri="https://zenodo.org/record/7983764/files/figure.png" pageId="5" pageNumber="298">Figure 3A</figureCitation>
), from
<collectingRegion id="49A366ACDF676B4FFB6FF96F8011D979" country="Colombia" name="Valle del Cauca" pageId="5" pageNumber="298">Valle del Cauca department</collectingRegion>
in
<collectingCountry id="F370E8DEDF676B4FFEB8F94B808CD979" box="[301,411,1598,1624]" name="Colombia" pageId="5" pageNumber="298">Colombia</collectingCountry>
and northwestern
<collectingCountry id="F370E8DEDF676B4FFDE5F94B83D8D979" box="[624,719,1598,1624]" name="Ecuador" pageId="5" pageNumber="298">Ecuador</collectingCountry>
, a species that present various color forms, dotted or stripped in purple to orange (
<bibRefCitation id="EFF6D5BFDF676B4FFED8F91780D0D95D" author="Luer, C. A." box="[333,455,1634,1660]" pageId="5" pageNumber="298" pagination="1 - 168" refId="ref4089" refString="Luer, C. A. (1996) Icones pleurothallidinarum XIII. Systematics of the genus Restrepia (Orchidaceae). Monographs in Systematic Botany from the Missouri Botanical Garden 59: 1 - 168." type="journal article" year="1996">Luer 1996</bibRefCitation>
), and
<taxonomicName id="4C67D3CDDF676B4FFD9CF916838FD95D" authorityName="Luer" authorityYear="1996" box="[521,664,1634,1660]" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="298" phylum="Tracheophyta" rank="species" species="mendozae">
<emphasis id="B913745CDF676B4FFD9CF916838FD95D" box="[521,664,1634,1660]" italics="true" pageId="5" pageNumber="298">R. mendozae</emphasis>
</taxonomicName>
(
<figureCitation id="135CB4CBDF676B4FFD33F917820DD95D" box="[678,794,1634,1660]" captionStart="FIGURE 3" captionStartId="4.[136,229,1741,1763]" captionTargetBox="[262,1326,190,1716]" captionTargetId="figure-23@4.[262,1326,190,1716]" captionTargetPageId="4" captionText="FIGURE 3. Restrepia santanderensis N. Gutiérrez &amp; K. Gil-Amaya, and similar morphological species. A. R. flosculata. B. R. mendozae. C. R. santanderensis. D. Capsules of R. santanderensis. Photographs byAndreas Key (A), Joost Riksen (B) from cultivated specimens and Nicolás Gutiérrez Morales (C, D) from the paratype." figureDoi="http://doi.org/10.5281/zenodo.7983764" httpUri="https://zenodo.org/record/7983764/files/figure.png" pageId="5" pageNumber="298">Figure 3B</figureCitation>
) known from southeastern
<collectingCountry id="F370E8DEDF676B4FFBD9F91785BED95D" box="[1100,1193,1634,1660]" name="Ecuador" pageId="5" pageNumber="298">Ecuador</collectingCountry>
(
<bibRefCitation id="EFF6D5BFDF676B4FFB2DF9178426D95D" author="Luer, C. A." box="[1208,1329,1634,1660]" pageId="5" pageNumber="298" pagination="1 - 168" refId="ref4089" refString="Luer, C. A. (1996) Icones pleurothallidinarum XIII. Systematics of the genus Restrepia (Orchidaceae). Monographs in Systematic Botany from the Missouri Botanical Garden 59: 1 - 168." type="journal article" year="1996">Luer 1996</bibRefCitation>
), with pale yellow flowers dotted in dark purple. They all share the overall similar size, shape of the leaves, the short-pedunculate inflorescence, and the pale yellow, intermediate to small-sized flowers.
</paragraph>
<paragraph id="8BD8A84EDF676B4FFF28F9BB807FD8E1" blockId="5.[136,1452,158,2093]" pageId="5" pageNumber="298">
The flowers of
<taxonomicName id="4C67D3CDDF676B4FFEE2F9BA831ED9C9" authorityName="Luer" authorityYear="1982" box="[375,521,1742,1768]" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="298" phylum="Tracheophyta" rank="species" species="flosculata">
<emphasis id="B913745CDF676B4FFEE2F9BA831ED9C9" box="[375,521,1742,1768]" italics="true" pageId="5" pageNumber="298">R. flosculata</emphasis>
</taxonomicName>
may have a similar color pattern to that of
<taxonomicName id="4C67D3CDDF676B4FFB89F9BA85F0D9C9" box="[1052,1255,1742,1768]" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="298" phylum="Tracheophyta" rank="species" species="santanderensis">
<emphasis id="B913745CDF676B4FFB89F9BA85F0D9C9" box="[1052,1255,1742,1768]" italics="true" pageId="5" pageNumber="298">R. santanderensis</emphasis>
</taxonomicName>
. However, in
<taxonomicName id="4C67D3CDDF676B4FFA06F9BA81E1D82D" authorityName="Luer" authorityYear="1982" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="298" phylum="Tracheophyta" rank="species" species="flosculata">
<emphasis id="B913745CDF676B4FFA06F9BA81E1D82D" italics="true" pageId="5" pageNumber="298">R. flosculata</emphasis>
</taxonomicName>
, only the basal sheaths of the ramicauls are brown-dotted (
<emphasis id="B913745CDF676B4FFC39F98682DCD82D" box="[940,971,1779,1804]" italics="true" pageId="5" pageNumber="298">vs.</emphasis>
all sheaths brown-dotted), the uppermost sheath is oblique, acute (
<emphasis id="B913745CDF676B4FFE31F86280D5D811" box="[420,450,1815,1840]" italics="true" pageId="5" pageNumber="298">vs.</emphasis>
elongate, acuminate), the sepals are shorter, to
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long (
<emphasis id="B913745CDF676B4FFBEBF862858AD811" box="[1150,1181,1815,1840]" italics="true" pageId="5" pageNumber="298">vs.</emphasis>
sepals to
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long), the dorsal sepal is narrowly triangular below the middle, attenuate above (
<emphasis id="B913745CDF676B4FFC40F84E82E4D875" box="[981,1011,1851,1876]" italics="true" pageId="5" pageNumber="298">vs.</emphasis>
dorsal sepal narrowly ovate below the lower third, contracted into a tail above), with a notoriously glabrous, obovate synsepal (
<emphasis id="B913745CDF676B4FFB19F82A85B4D859" box="[1164,1187,1887,1912]" italics="true" pageId="5" pageNumber="298">vs</emphasis>
. synsepal with evident trichomes, oblong), decurved petals (
<emphasis id="B913745CDF676B4FFDBCF8F68357D8BD" box="[553,576,1923,1948]" italics="true" pageId="5" pageNumber="298">vs</emphasis>
. straight petals), and a broadly oblong lip with retuse apex (
<emphasis id="B913745CDF676B4FFB76F8F68415D8BD" box="[1251,1282,1923,1948]" italics="true" pageId="5" pageNumber="298">vs.</emphasis>
rectangular lip with truncate apex).
</paragraph>
<paragraph id="8BD8A84EDF676B4CFF28F8BE85BEDFFD" blockId="5.[136,1452,158,2093]" lastBlockId="6.[136,1452,158,329]" lastPageId="6" lastPageNumber="299" pageId="5" pageNumber="298">
In
<taxonomicName id="4C67D3CDDF676B4FFF4BF8BE8078D8C5" authorityName="Luer" authorityYear="1996" box="[222,367,1994,2020]" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="298" phylum="Tracheophyta" rank="species" species="mendozae">
<emphasis id="B913745CDF676B4FFF4BF8BE8078D8C5" box="[222,367,1994,2020]" italics="true" pageId="5" pageNumber="298">R. mendozae</emphasis>
</taxonomicName>
the shapes and size of sepals and petals are very similar to that of
<taxonomicName id="4C67D3CDDF676B4FFBE0F8BE842AD8C5" box="[1141,1341,1994,2020]" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="298" phylum="Tracheophyta" rank="species" species="santanderensis">
<emphasis id="B913745CDF676B4FFBE0F8BE842AD8C5" box="[1141,1341,1994,2020]" italics="true" pageId="5" pageNumber="298">R. santanderensis</emphasis>
</taxonomicName>
, but only the lowermost sheath is brown-dotted. In addition, the sepals are spreading, yellow with purple dots (
<emphasis id="B913745CDF676B4FFAB3F89A842AD729" box="[1318,1341,2031,2056]" italics="true" pageId="5" pageNumber="298">vs</emphasis>
. partially spreading sepals, yellow with vinaceous strips in the synsepal), the dorsal sepal is erect (
<emphasis id="B913745CDF676B4FFBFCF7668597D70D" box="[1129,1152,2067,2092]" italics="true" pageId="5" pageNumber="298">vs</emphasis>
. prostrate), the synsepal is glabrous (
<emphasis id="B913745CDF646B4CFF6DFFEA8018DF99" box="[248,271,159,184]" italics="true" pageId="6" pageNumber="299">vs</emphasis>
. synsepal with evident trichomes), the petals are decurved (
<emphasis id="B913745CDF646B4CFC30FFEA82ABDF99" box="[933,956,159,184]" italics="true" pageId="6" pageNumber="299">vs</emphasis>
. straight), and the lip is entirely yellow, with dark purple spots (
<emphasis id="B913745CDF646B4CFECCFFB6806FDFFD" box="[345,376,195,220]" italics="true" pageId="6" pageNumber="299">vs.</emphasis>
yellow with intense rose at the base and vinaceous, longitudinal stripes).
</paragraph>
<paragraph id="8BD8A84EDF646B4CFF28FF93841DDE68" blockId="6.[136,1452,158,329]" pageId="6" pageNumber="299">
The arrangement of the trichomes along the veins of the synsepal in
<taxonomicName id="4C67D3CDDF646B4CFC67FF9285ABDE21" box="[1010,1212,230,256]" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="299" phylum="Tracheophyta" rank="species" species="santanderensis">
<emphasis id="B913745CDF646B4CFC67FF9285ABDE21" box="[1010,1212,230,256]" italics="true" pageId="6" pageNumber="299">R. santanderensis</emphasis>
</taxonomicName>
, matches
<bibRefCitation id="EFF6D5BFDF646B4CFAA3FF93805DDE05" author="Millner, H. J. &amp; Baldwin, T. C." pageId="6" pageNumber="299" pagination="10 - 19" refId="ref4179" refString="Millner, H. J. &amp; Baldwin, T. C. (2016) Floral micromorphology of the genus Restrepia (Orchidaceae) and the potential consequences for pollination. Flora-Morphology, Distribution, Functional Ecology of Plants 225: 10 - 19. https: // doi. org / 10.1016 / j. flora. 2016.09.007" type="journal article" year="2016">Millner &amp; Baldwin’s (2016)</bibRefCitation>
description of the papillae in
<taxonomicName id="4C67D3CDDF646B4CFD07FE7E8232DE05" box="[658,805,266,292]" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="299" phylum="Tracheophyta" rank="species" species="brachypus">
<emphasis id="B913745CDF646B4CFD07FE7E8232DE05" box="[658,805,266,292]" italics="true" pageId="6" pageNumber="299">R. brachypus</emphasis>
</taxonomicName>
, following the stripes of the synsepal. This as ‘false nectar guides’, part of the non-rewarding pollination strategy that is supposed to occur in the genus
<taxonomicName id="4C67D3CDDF646B4CFB0DFE5A8415DE69" authorityName="Kunth" authorityYear="1815" box="[1176,1282,303,328]" class="Liliopsida" family="Orchidaceae" genus="Restrepia" kingdom="Plantae" order="Asparagales" pageId="6" pageNumber="299" phylum="Tracheophyta" rank="genus">
<emphasis id="B913745CDF646B4CFB0DFE5A8415DE69" box="[1176,1282,303,328]" italics="true" pageId="6" pageNumber="299">Restrepia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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