152 lines
18 KiB
XML
152 lines
18 KiB
XML
<document id="0376C3A82A8EC4938FB851EC4717A92E" ID-DOI="10.11646/zootaxa.4994.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5043261" ID-ZooBank="107FCA64-345F-40A4-99D3-5C1441EEAD93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624993528766" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="03A187DEFFE0FFE36194FEBA05EBF2C6" docLanguage="en" docName="zootaxa.4994.1.1.pdf" docOrigin="Zootaxa 4994 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus liquidus Perkins 2021, new species" docType="treatment" docVersion="4" lastPageNumber="29" masterDocId="FF98FFA6FFFCFFFF6103FFF0045FF707" masterDocTitle="Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)" masterLastPageNumber="93" masterPageNumber="1" pageNumber="29" updateTime="1698978858223" updateUser="plazi">
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<mods:titleInfo id="7F01B94EFA83DF40DA77285B25A5880A">
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<mods:title id="38EF6B685167189D75A8BFB991001801">Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="0D1DF9A9B742B9BCA081272B62419314" type="personal">
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<mods:roleTerm id="8224BC06A088C63AF9C451865E84B1DF">Author</mods:roleTerm>
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<mods:namePart id="380550123D884FBB4537457EE659B1E6">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="09DBB638C4E9E1D3BFC29E15D2948BBA">text</mods:typeOfResource>
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<mods:title id="019E5C1C9A4CD1E6E5FB6C051F57AD0C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="1BDCA865043A85CBF113E28BE2437E97">
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<mods:date id="A0520523280AB6C444C5A67BAC1ED702">2021</mods:date>
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<mods:detail id="8B3B60F5367BC9A3998E86BE1C7A1CDC" type="pubDate">
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<mods:number id="8E2A4F003444F30E05F3D95006F27EB4">2021-06-28</mods:number>
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<mods:detail id="C57F05A7F3E8CCF7274376BD72554FD0" type="volume">
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<mods:number id="71A557C36035EEF311AF080445DFE9B8">4994</mods:number>
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</mods:detail>
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<mods:detail id="B81D5CA123870C6C027C1D32183E74C5" type="issue">
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<mods:number id="27AA014C612CAC07BA2A2D71915B4E2A">1</mods:number>
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<mods:start id="158AB01EBA9BA626C97B525F3ABB9530">1</mods:start>
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<mods:end id="FE0EF1C0B8B512E3CB2035DE4907F3C1">93</mods:end>
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<mods:classification id="4035B9666283F21BF3DFA16F8E33F371">journal article</mods:classification>
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<mods:identifier id="1BC92C84C33EF3F14AE5C068ADC2E5D5" type="DOI">10.11646/zootaxa.4994.1.1</mods:identifier>
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<mods:identifier id="DDA9911D8D289307B38199716D71E09E" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="26C5435400A07E4C05A36EF9DEB31B4C" type="Zenodo-Dep">5043261</mods:identifier>
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<mods:identifier id="558809501D273F96D2ACBF21FFC736F6" type="ZooBank">107FCA64-345F-40A4-99D3-5C1441EEAD93</mods:identifier>
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<treatment id="03A187DEFFE0FFE36194FEBA05EBF2C6" ID-DOI="http://doi.org/10.5281/zenodo.5043966" ID-Zenodo-Dep="5043966" LSID="urn:lsid:plazi:treatment:03A187DEFFE0FFE36194FEBA05EBF2C6" httpUri="http://treatment.plazi.org/id/03A187DEFFE0FFE36194FEBA05EBF2C6" lastPageNumber="29" pageId="28" pageNumber="29">
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<subSubSection id="C3126543FFE0FFE36194FEBA067BF663" box="[151,548,330,357]" pageId="28" pageNumber="29" type="nomenclature">
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<paragraph id="8BB736C8FFE0FFE36194FEBA067BF663" blockId="28.[151,548,330,393]" box="[151,548,330,357]" pageId="28" pageNumber="29">
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<heading id="D0FF81A4FFE0FFE36194FEBA067BF663" bold="true" box="[151,548,330,357]" fontSize="11" level="2" pageId="28" pageNumber="29" reason="4">
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<emphasis id="B97CEADAFFE0FFE36194FEBA067BF663" bold="true" box="[151,548,330,357]" pageId="28" pageNumber="29">
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||
<taxonomicName id="4C084D4BFFE0FFE36194FEBA05D9F662" ID-CoL="8Y4QK" authority="Perkins, 2021" authorityName="Perkins" authorityYear="2021" box="[151,390,330,357]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="liquidus" status="sp. nov.">
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<emphasis id="B97CEADAFFE0FFE36194FEBA05D9F662" bold="true" box="[151,390,330,357]" italics="true" pageId="28" pageNumber="29">Hydrochus liquidus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24F57A1FFE0FFE36096FEBB067BF663" box="[405,548,330,357]" pageId="28" pageNumber="29" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3126543FFE0FFE36194FE9F0568F68E" box="[151,311,366,393]" pageId="28" pageNumber="29" type="description">
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||
<paragraph id="8BB736C8FFE0FFE36194FE9F0568F68E" blockId="28.[151,548,330,393]" box="[151,311,366,393]" pageId="28" pageNumber="29">
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<figureCitation id="13332A4DFFE0FFE36194FE9F04BBF68E" box="[151,228,367,393]" captionStart="FIGURE 3" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[169,1397,189,1912]" captionTargetId="figure-28@8.[154,1433,181,1919]" captionTargetPageId="8" captionText="FIGURE 3. Dorsal habitus of Hydrochus species in study area." figureDoi="http://doi.org/10.5281/zenodo.5043269" httpUri="https://zenodo.org/record/5043269/files/figure.png" pageId="28" pageNumber="29">Figs. 3</figureCitation>
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,
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<figureCitation id="13332A4DFFE0FFE361F2FE9F0550F68E" box="[241,271,367,393]" captionStart="FIGURES 38–39" captionStartId="71.[151,264,1944,1968]" captionTargetBox="[157,1430,203,1847]" captionTargetId="figure-15@71.[151,1435,181,1920]" captionTargetPageId="71" captionText="FIGURES 38–39. Habitus and male genitalia. 38. Hydrochus liquidus n. sp., holotype. 39. Hydrochus jenniferiduae Makhan." figureDoi="http://doi.org/10.5281/zenodo.5043324" httpUri="https://zenodo.org/record/5043324/files/figure.png" pageId="28" pageNumber="29">38</figureCitation>
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,
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<figureCitation id="13332A4DFFE0FFE36018FE9E0568F68E" box="[283,311,366,393]" captionStart="FIGURES 55–56" captionStartId="80.[151,264,1943,1967]" captionTargetBox="[165,1422,181,1920]" captionTargetId="figure-28@80.[165,1422,181,1920]" captionTargetPageId="80" captionText="FIGURES 55–56. Geographical distributions. 55. Endemics and near endemics. 56. H. bicarinatus and H. merkli." figureDoi="http://doi.org/10.5281/zenodo.5043352" httpUri="https://zenodo.org/record/5043352/files/figure.png" pageId="28" pageNumber="29">55</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE0FFE36194FE470687F51E" pageId="28" pageNumber="29" type="materials_examined">
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||
<paragraph id="8BB736C8FFE0FFE36194FE470687F51E" blockId="28.[151,1437,438,1473]" pageId="28" pageNumber="29">
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<emphasis id="B97CEADAFFE0FFE36194FE470514F6D7" bold="true" box="[151,331,438,465]" pageId="28" pageNumber="29">Type Material.</emphasis>
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<materialsCitation id="3B603C95FFE0FFE36056FE4701C6F6F2" collectingDate="2005-08-05" collectionCode="INPA" collectorName="Hamada, N. & M. Landeiro" country="Brazil" county="Piata" latitude="-13.269166" location="Agnaldo" municipality="Corrego" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Bahia" typeStatus="holotype">
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<emphasis id="B97CEADAFFE0FFE36056FE47067DF6D7" bold="true" box="[341,546,438,465]" pageId="28" pageNumber="29">
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<typeStatus id="54B3886AFFE0FFE36056FE47059DF6D6" box="[341,450,439,465]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
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(male):
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</emphasis>
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“
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<collectingCountry id="F31F7658FFE0FFE36335FE4706FDF6D6" box="[566,674,439,465]" name="Brazil" pageId="28" pageNumber="29">BRAZIL</collectingCountry>
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:
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<collectingRegion id="49CCF82AFFE0FFE363B3FE4706ABF6D6" box="[688,756,439,465]" country="Brazil" name="Bahia" pageId="28" pageNumber="29">Bahia</collectingRegion>
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,
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<collectingCounty id="62D64E44FFE0FFE36201FE470762F6D6" box="[770,829,439,465]" pageId="28" pageNumber="29">Piatã</collectingCounty>
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,
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<collectingMunicipality id="6BD3ACB2FFE0FFE36248FE4707F8F6D6" box="[843,935,439,465]" pageId="28" pageNumber="29">Córrego</collectingMunicipality>
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na trilha para a casa do
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<location id="8ED76013FFE0FFE365C8FE470171F6D6" LSID="urn:lsid:plazi:treatment:03A187DEFFE0FFE36194FEBA05EBF2C6:8ED76013FFE0FFE365C8FE470171F6D6" box="[1227,1326,439,465]" country="Brazil" county="Piata" latitude="-13.269166" municipality="Corrego" name="Agnaldo" pageId="28" pageNumber="29" stateProvince="Bahia">Agnaldo</location>
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,
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<location id="8ED76013FFE0FFE3643FFE4704ADF6F2" LSID="urn:lsid:plazi:treatment:03A187DEFFE0FFE36194FEBA05EBF2C6:8ED76013FFE0FFE3643FFE4704ADF6F2" country="Brazil" county="Piata" latitude="-13.269166" municipality="Corrego" name="Serra da Tromba" pageId="28" pageNumber="29" stateProvince="Bahia">Serra da Tromba</location>
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, (
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<geoCoordinate id="EE3C500FFFE0FFE3600AFE2B05DAF6F2" box="[265,389,475,501]" degrees="13" direction="south" minutes="16" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="9" value="-13.269166">13°16’9’’S</geoCoordinate>
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, 41°46’18’’),
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<date id="FFB61008FFE0FFE36324FE2A06FEF6F2" box="[551,673,474,501]" pageId="28" pageNumber="29" value="2005-08-05">
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<collectingDate id="EFF2E9E0FFE0FFE36324FE2A06FEF6F2" box="[551,673,474,501]" pageId="28" pageNumber="29" value="2005-08-05">5.viii.2005</collectingDate>
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</date>
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// stream, leg.
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<collectorName id="26FD531EFFE0FFE36252FE2B0783F6F2" box="[849,988,475,501]" pageId="28" pageNumber="29">Hamada, N.</collectorName>
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, Pepinelli,
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<collectorName id="26FD531EFFE0FFE3655CFE2B00AEF6F2" box="[1119,1265,475,501]" pageId="28" pageNumber="29">M. Landeiro</collectorName>
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, V.L.” (
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<collectionCode id="ED19AE0DFFE0FFE3644FFE2B01CDF6F2" box="[1356,1426,475,501]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="28" pageNumber="29">INPA</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B603C95FFE0FFE36194FE0F068BF51E" box="[151,724,511,537]" collectingDate="2005-08-05" collectionCode="INPA" collectorName="Hamada, N. & M. Landeiro" country="Brazil" county="Piata" latitude="-13.269166" location="Agnaldo" municipality="Corrego" pageId="28" pageNumber="29" specimenCount="1" specimenCount-female="1" stateProvince="Bahia" typeStatus="paratype">
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<emphasis id="B97CEADAFFE0FFE36194FE0F0552F51E" bold="true" box="[151,269,511,537]" pageId="28" pageNumber="29">
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<typeStatus id="54B3886AFFE0FFE36194FE0F0557F51E" box="[151,264,511,537]" pageId="28" pageNumber="29" type="paratype">Paratype</typeStatus>
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:
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</emphasis>
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Same data as holotype (
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<specimenCount id="9D0EFD41FFE0FFE36321FE0F06DEF51E" box="[546,641,511,537]" pageId="28" pageNumber="29" type="female">1 female</specimenCount>
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<collectionCode id="ED19AE0DFFE0FFE3638BFE0F0693F51E" box="[648,716,511,537]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="28" pageNumber="29">INPA</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3126543FFE0FFE361C4FDD20156F5AE" pageId="28" pageNumber="29" type="diagnosis">
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<paragraph id="8BB736C8FFE0FFE361C4FDD20156F5AE" blockId="28.[151,1437,438,1473]" pageId="28" pageNumber="29">
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<emphasis id="B97CEADAFFE0FFE361C4FDD2058CF53B" bold="true" box="[199,467,546,572]" pageId="28" pageNumber="29">Differential Diagnosis.</emphasis>
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Among Brazilian species, recognized by the combination of small size (ca.
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<quantity id="4CF09B2DFFE0FFE3642BFDD201CFF53A" box="[1320,1424,546,573]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="28" pageNumber="29" unit="mm" value="2.5">2.50 mm</quantity>
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), the black dorsum, the elytral odd numbered interstriae subcostate to costate, the smooth rounded pronotal reliefs, the deep and coarsely densely punctate pronotal depressions, and the male genitalia (
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<figureCitation id="13332A4DFFE0FFE36514FD9B0036F582" box="[1047,1129,619,645]" captionStart="FIGURES 38–39" captionStartId="71.[151,264,1944,1968]" captionTargetBox="[157,1430,203,1847]" captionTargetId="figure-15@71.[151,1435,181,1920]" captionTargetPageId="71" captionText="FIGURES 38–39. Habitus and male genitalia. 38. Hydrochus liquidus n. sp., holotype. 39. Hydrochus jenniferiduae Makhan." figureDoi="http://doi.org/10.5281/zenodo.5043324" httpUri="https://zenodo.org/record/5043324/files/figure.png" pageId="28" pageNumber="29">Fig. 38</figureCitation>
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; described below). Reliable determinations of similarly sized black specimens with costate elytra will require dissection of males.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE0FFE361C4FD4200B3F27E" pageId="28" pageNumber="29" type="description">
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<paragraph id="8BB736C8FFE0FFE361C4FD420016F45A" blockId="28.[151,1437,438,1473]" pageId="28" pageNumber="29">
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<emphasis id="B97CEADAFFE0FFE361C4FD42050EF5CB" bold="true" box="[199,337,690,716]" pageId="28" pageNumber="29">Description</emphasis>
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. Size:
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<typeStatus id="54B3886AFFE0FFE360A7FD430659F5CA" box="[420,518,691,717]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 2.50/1.02; head width 0.66; pronotum l/w 0.57/0.67; PA 0.67; PB 0.50; elytra 1.66/1.02. Habitus as illustrated (
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<figureCitation id="13332A4DFFE0FFE36528FD270020F5F6" box="[1067,1151,727,753]" captionStart="FIGURES 38–39" captionStartId="71.[151,264,1944,1968]" captionTargetBox="[157,1430,203,1847]" captionTargetId="figure-15@71.[151,1435,181,1920]" captionTargetPageId="71" captionText="FIGURES 38–39. Habitus and male genitalia. 38. Hydrochus liquidus n. sp., holotype. 39. Hydrochus jenniferiduae Makhan." figureDoi="http://doi.org/10.5281/zenodo.5043324" httpUri="https://zenodo.org/record/5043324/files/figure.png" pageId="28" pageNumber="29">Fig. 38</figureCitation>
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). Body size moderate (ca.
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||
<quantity id="4CF09B2DFFE0FFE36194FD0A04A1F412" box="[151,254,762,789]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="28" pageNumber="29" unit="mm" value="2.5">2.50 mm</quantity>
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). Dorsum black, with very slight or no iridescence, elytra without black spots. Legs brown, with tibiofemoral joints darker. Punctation on elytra ca. 3–5x that of pronotum. Elytra interstriae ca. 0.5x strial puncture diameter; odd numbered interstriae subcostate to costate. Usual area of callus on 5
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<superScript id="7C7D9B80FFE0FFE362C3FCB20792F457" attach="left" box="[960,973,834,848]" fontSize="6" pageId="28" pageNumber="29">th</superScript>
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not raised.
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</paragraph>
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<paragraph id="8BB736C8FFE0FFE361C4FC97012DF4CE" blockId="28.[151,1437,438,1473]" pageId="28" pageNumber="29">Pronotum slightly wider than long (ratio ca. 36/31), widest at anterior margin, narrowed at base, sides weakly sinuate, smooth to microdenticulate; anterior margin sinuate; depressions deep; pronotum coarsely densely punctate, more densely punctate in depressions than on reliefs. Lateral and anterior margins with row of short setae.</paragraph>
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<paragraph id="8BB736C8FFE0FFE361C4FC230023F316" blockId="28.[151,1437,438,1473]" pageId="28" pageNumber="29">Elytra with spaces between strial punctures ca. 0.5x their diameter; apices conjointly rounded in dorsal view, in lateral view outer margin with slight angulation; lateral most row of punctures in groove.</paragraph>
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<paragraph id="8BB736C8FFE0FFE361C4FBEB0095F332" blockId="28.[151,1437,438,1473]" box="[199,1226,1051,1077]" pageId="28" pageNumber="29">Ventral characters: Mentum punctate, with central depression; submentum with two foveae.</paragraph>
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<paragraph id="8BB736C8FFE0FFE361C4FBCF0744F20A" blockId="28.[151,1437,438,1473]" pageId="28" pageNumber="29">
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Male genitalia (
|
||
<figureCitation id="13332A4DFFE0FFE3607BFBCF0595F35E" box="[376,458,1087,1113]" captionStart="FIGURES 38–39" captionStartId="71.[151,264,1944,1968]" captionTargetBox="[157,1430,203,1847]" captionTargetId="figure-15@71.[151,1435,181,1920]" captionTargetPageId="71" captionText="FIGURES 38–39. Habitus and male genitalia. 38. Hydrochus liquidus n. sp., holotype. 39. Hydrochus jenniferiduae Makhan." figureDoi="http://doi.org/10.5281/zenodo.5043324" httpUri="https://zenodo.org/record/5043324/files/figure.png" pageId="28" pageNumber="29">Fig. 38</figureCitation>
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) general characters: genitalia very narrow, elongate, length ca. 7x greatest width, basal piece shorter than parameres (ratio ca. 4/7); paramere tips extending very slightly beyond aedeagus tip, medial margins beyond aedeagus tip parallel, plsa absent; parameres in dorsal/ventral views very narrow; in lateral view parameres straight, very sharply pointed apically; aedeagus very narrow, wider apically and sub-basally than at midlength; basal piece in dorsal/ventral views widening very slightly from orifice to distal end, in lateral view very asymmetrical, constricted just before lobe-shaped orifice.
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</paragraph>
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<paragraph id="8BB736C8FFE0FFE361C4FAE7060AF252" blockId="28.[151,1437,438,1473]" pageId="28" pageNumber="29">Dorsal surface: adtl near base, just distal to adbl; pdmm approximating alm; adbl moderately large, basal end transverse, width slightly less than agw.</paragraph>
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<paragraph id="8BB736C8FFE0FFE361C4FAAF00B3F27E" blockId="28.[151,1437,438,1473]" box="[199,1260,1375,1401]" pageId="28" pageNumber="29">Ventral surface: avtl not apparent; pvmm separation similar to pdmm, neither overlapping alm.</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE0FFE361C4FA7305EBF2C6" pageId="28" pageNumber="29" type="etymology">
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<paragraph id="8BB736C8FFE0FFE361C4FA7305EBF2C6" blockId="28.[151,1437,438,1473]" pageId="28" pageNumber="29">
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<emphasis id="B97CEADAFFE0FFE361C4FA730517F29A" bold="true" box="[199,328,1411,1437]" pageId="28" pageNumber="29">Etymology</emphasis>
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. Named in reference to the pronotal sculpture, with very smooth reliefs, which give the impression of being melted, flowing.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |