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<document id="DC4ABD90BD735BAE0C248A0E62027811" ID-DOI="10.11646/zootaxa.5040.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5531372" ID-ZooBank="5E9EF3EC-6179-44B9-96A3-A788BA3C1DE9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632748248110" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2021" docId="03A187C1087B2448FF06FA62FCA51690" docLanguage="en" docName="zootaxa.5040.3.1.pdf" docOrigin="Zootaxa 5040 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mannerheimia afghanica Coiffait 1982" docType="treatment" docVersion="3" lastPageNumber="307" masterDocId="FF98FFB9087D244EFF91FFE7FFB6104E" masterDocTitle="On the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), with taxonomic and faunistic notes on some species" masterLastPageNumber="333" masterPageNumber="301" pageNumber="307" updateTime="1699415462997" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="11341B1A3C44C1EA6CA9C1F62FA0667E">On the genus Mannerheimia Mäklin, 1880 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), with taxonomic and faunistic notes on some species</mods:title>
</mods:titleInfo>
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<mods:namePart id="802ACB987401F3D90072BE53E400C6F5">Shavrin, Alexey V.</mods:namePart>
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<mods:typeOfResource id="6A2C63078BE04A77BD0F68FC7827D383">text</mods:typeOfResource>
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<mods:title id="75FD09E5F5C89BCBE5A1BE596DE65D23">Zootaxa</mods:title>
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<mods:date id="0C73B0006D57E1F030BABDD95407EC80">2021</mods:date>
<mods:detail id="DA95291BD2D8C012EFEBF89A9D91E45E" type="pubDate">
<mods:number id="77622FCC67E124551C584F4D8B31DFB6">2021-09-22</mods:number>
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<mods:number id="926488B3455E71C4D24292FA335AB2EE">5040</mods:number>
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<mods:number id="E8F40071D067573491312792380EC0EB">3</mods:number>
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<mods:classification id="AAB7AAB67196D83AAEE2B4AEBE662E1B">journal article</mods:classification>
<mods:identifier id="7157226CC75E135681EB7003AC48EE90" type="DOI">10.11646/zootaxa.5040.3.1</mods:identifier>
<mods:identifier id="3A711D46EAEEB0B092B409F89D0A0568" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E3F565B85654262D9A215EAB79E722B0" type="Zenodo-Dep">5531372</mods:identifier>
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<treatment id="03A187C1087B2448FF06FA62FCA51690" LSID="urn:lsid:plazi:treatment:03A187C1087B2448FF06FA62FCA51690" httpUri="http://treatment.plazi.org/id/03A187C1087B2448FF06FA62FCA51690" lastPageNumber="307" pageId="6" pageNumber="307">
<subSubSection id="C312655C087B2448FF06FA62FDC915D1" box="[151,639,1413,1440]" pageId="6" pageNumber="307" type="nomenclature">
<paragraph id="8BB736D7087B2448FF06FA62FDC915D1" blockId="6.[151,639,1413,1475]" box="[151,639,1413,1440]" pageId="6" pageNumber="307">
<heading id="D0FF81BB087B2448FF06FA62FDC915D1" bold="true" box="[151,639,1413,1440]" fontSize="11" level="1" pageId="6" pageNumber="307" reason="1">
<taxonomicName id="4C084D54087B2448FF06FA62FDC915D1" ID-CoL="3XVM5" authority="Coiffait, 1982" authorityName="Coiffait" authorityYear="1982" box="[151,639,1413,1440]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="afghanica">
<emphasis id="B97CEAC5087B2448FF06FA62FDC915D1" bold="true" box="[151,639,1413,1440]" pageId="6" pageNumber="307">
<emphasis id="B97CEAC5087B2448FF06FA62FE7B15EE" bold="true" box="[151,461,1413,1440]" italics="true" pageId="6" pageNumber="307">Mannerheimia afghanica</emphasis>
<bibRefCitation id="EF994B26087B2448FE44FA62FDC915D1" author="Coiffait, H." box="[469,639,1413,1439]" pageId="6" pageNumber="307" pagination="21 - 179" refId="ref24135" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179." type="journal article" year="1982">Coiffait, 1982</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C312655C087B2448FF06FA4EFEFF158D" box="[151,329,1449,1475]" pageId="6" pageNumber="307" type="description">
<paragraph id="8BB736D7087B2448FF06FA4EFEFF158D" blockId="6.[151,639,1413,1475]" box="[151,329,1449,1475]" pageId="6" pageNumber="307">
(
<figureCitation id="13332A52087B2448FF0EFA4EFF5B158D" box="[159,237,1449,1475]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1955,1979]" captionTargetBox="[187,1400,190,1920]" captionTargetId="figure-16@3.[173,1414,181,1931]" captionTargetPageId="3" captionText="FIGURES 1–9. Habitus of Mannerheimia: 1—M. afghanica (paratype), 2—M. arctica, 3—M. asiatica, 4—M. brevipennis, 5—M. japonica, 6—M. obscura, 7—M. pleshanovi (paratype), 8—M. yasudai, 9—M. zanettii (paratype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5531374" httpUri="https://zenodo.org/record/5531374/files/figure.png" pageId="6" pageNumber="307">Figs. 1</figureCitation>
,
<figureCitation id="13332A52087B2448FF6AFA4EFEF6158D" box="[251,320,1449,1475]" captionStart="FIGURES 10–11" captionStartId="7.[151,264,1477,1501]" captionTargetBox="[161,1421,192,1444]" captionTargetId="figure-16@7.[151,1436,181,1453]" captionTargetPageId="7" captionText="FIGURES 10–11. Aedeagus of Mannerheimia afganica (paratype): 10—parameral view, 11—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531376" httpUri="https://zenodo.org/record/5531376/files/figure.png" pageId="6" pageNumber="307">10–11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C312655C087B2448FF06FA09FDE41648" box="[151,594,1518,1542]" pageId="6" pageNumber="307" type="reference_group">
<paragraph id="8BB736D7087B2448FF06FA09FDE41648" blockId="6.[151,594,1518,1542]" box="[151,594,1518,1542]" pageId="6" pageNumber="307">
<treatmentCitationGroup id="AB1811F9087B2448FF06FA09FDE41648" box="[151,594,1518,1542]" pageId="6" pageNumber="307">
<treatmentCitation id="0AA910C6087B2448FF06FA09FDE41648" author="Coiffait, H." box="[151,594,1518,1542]" page="82" pageId="6" pageNumber="307" year="1982">
<taxonomicName id="4C084D54087B2448FF06FA09FDE41648" ID-CoL="3XVM5" authority="Coiffait, 1982: 82" authorityName="Coiffait" authorityPageNumber="82" authorityYear="1982" box="[151,594,1518,1542]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="afghanica">
<emphasis id="B97CEAC5087B2448FF06FA09FE211648" box="[151,407,1518,1542]" italics="true" pageId="6" pageNumber="307">Mannerheimia afghanica</emphasis>
<bibRefCitation id="EF994B26087B2448FE0FFA09FDE41648" author="Coiffait, H." box="[414,594,1518,1542]" pageId="6" pageNumber="307" pagination="21 - 179" refId="ref24135" refString="Coiffait, H. (1982) Contribution a la connaissance des Staphylinides de l'Himalaya (Nepal, Ladakh, Cachemire) (Insecta: Coleoptera: Staphylinidae). Senckenbergiana Biologica, 62, 21 - 179." type="journal article" year="1982">Coiffait, 1982: 82</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C312655C087B2448FF06F9D3FB0D16D8" pageId="6" pageNumber="307" type="materials_examined">
<paragraph id="8BB736D7087B2448FF06F9D3FB0D16D8" blockId="6.[151,1436,1588,1758]" pageId="6" pageNumber="307">
<emphasis id="B97CEAC5087B2448FF06F9D3FE741600" bold="true" box="[151,450,1588,1614]" pageId="6" pageNumber="307">Type material examined:</emphasis>
<typeStatus id="54B38875087B2448FE5BF9D3FD8D1600" box="[458,571,1588,1614]" pageId="6" pageNumber="307" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D0EFD5E087B2448FDD7F9D3FDC21600" box="[582,628,1588,1614]" pageId="6" pageNumber="307" type="male">1 ♂</specimenCount>
: ‘Paratype’ &lt;red, printed&gt;, ‘COTYPUS |
<taxonomicName id="4C084D54087B2448FBD6F9D3FB5C1600" authorityName="Maklin" authorityYear="1880" box="[1095,1258,1588,1614]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">Mannerheimia</taxonomicName>
| afghanica | O. Scheerpeltz’ &lt;pink, printed&gt;, ‘
<materialsCitation id="3B603C8A087B2448FE7CF9BFFD8916D8" collectionCode="ZFMK" collectorName="J. Klapperich &amp; Kamdesch" elevation="2200" location="Nuristan" pageId="6" pageNumber="307" specimenCount="1" specimenCount-male="1" stateProvince="Nuristan" typeStatus="paratype">
<specimenCount id="9D0EFD5E087B2448FE7CF9BFFDB1163C" box="[493,519,1624,1650]" pageId="6" pageNumber="307" type="male">♂</specimenCount>
’ &lt;handwritten&gt;, ‘
<collectorName id="26FD5301087B2448FD5EF9BFFCDE163C" box="[719,872,1624,1650]" pageId="6" pageNumber="307">J. Klapperich</collectorName>
|
<collectorName id="26FD5301087B2448FCEFF9BFFC42163C" box="[894,1012,1624,1650]" pageId="6" pageNumber="307">Kamdesch</collectorName>
<quantity id="4CF09B32087B2448FC6DF9BFFBE4163C" box="[1020,1106,1624,1650]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="307" unit="m" value="2200.0">
<elevation id="0025D1E4087B2448FC6DF9BFFBE4163C" box="[1020,1106,1624,1650]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="307" unit="m" value="2200.0">2200 m</elevation>
</quantity>
|
<collectingRegion id="49CCF835087B2448FBF6F9BFFB71163C" box="[1127,1223,1624,1650]" country="Afghanistan" name="Nuristan" pageId="6" pageNumber="307">Nuristan</collectingRegion>
28. 4. [19]53 | Af- ghanistan’ &lt;printed in blue&gt; (
<collectionCode id="ED19AE12087B2448FE73F99BFD8016D8" box="[482,566,1660,1686]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="6" pageNumber="307" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B603C8A087B2448FDDBF99BFCC316D8" box="[586,885,1660,1686]" collectionCode="ZFMK" collectorName="J. Klapperich &amp; Kamdesch" elevation="2200" location="Nuristan" pageId="6" pageNumber="307" specimenCount="1" specimenCount-female="1" stateProvince="Nuristan" typeStatus="paratype">
<specimenCount id="9D0EFD5E087B2448FDDBF99BFDC016D8" box="[586,630,1660,1686]" pageId="6" pageNumber="307" type="female">1 ♀</specimenCount>
: same data, except for
</materialsCitation>
: ‘
<materialsCitation id="3B603C8A087B2448FC15F99BFB0E16D8" box="[900,1208,1660,1686]" collectionCode="ZFMK" pageId="6" pageNumber="307" specimenCount="1" specimenCount-female="1" stateProvince="Nuristan" typeStatus="paratype">
<specimenCount id="9D0EFD5E087B2448FC15F99BFC2416D8" box="[900,914,1660,1686]" pageId="6" pageNumber="307" type="female">♀</specimenCount>
’&lt;handwritten&gt; (
<collectionCode id="ED19AE12087B2448FBCAF99BFB1916D8" box="[1115,1199,1660,1686]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="6" pageNumber="307" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C312655C087B2448FF56F947FCA51690" pageId="6" pageNumber="307" type="discussion">
<paragraph id="8BB736D7087B2448FF56F947FCA51690" blockId="6.[151,1436,1588,1758]" pageId="6" pageNumber="307">
<emphasis id="B97CEAC5087B2448FF56F947FE8D16F4" bold="true" box="[199,315,1696,1722]" pageId="6" pageNumber="307">Remarks.</emphasis>
<taxonomicName id="4C084D54087B2448FED5F947FDD416F4" authorityName="Coiffait" authorityYear="1982" box="[324,610,1696,1722]" class="Insecta" family="Staphylinidae" genus="Mannerheimia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="afghanica">
<emphasis id="B97CEAC5087B2448FED5F947FDD416F4" box="[324,610,1696,1722]" italics="true" pageId="6" pageNumber="307">Mannerheimia afghanica</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="54B38875087B2448FC12F947FC0516F4" box="[899,947,1696,1722]" pageId="6" pageNumber="307">type</typeStatus>
locality in “
<collectingCountry id="F31F7647087B2448FBD3F947FB6716F4" box="[1090,1233,1696,1722]" name="Afghanistan" pageId="6" pageNumber="307">Afghanistan</collectingCountry>
,
<collectingRegion id="49CCF835087B2448FB4FF947FAF916F4" box="[1246,1359,1696,1722]" country="Afghanistan" name="Nuristan" pageId="6" pageNumber="307">Nouristan</collectingRegion>
, Kamdesch”. Habitus as in
<figureCitation id="13332A52087B2448FE1AF923FE7B1690" box="[395,461,1732,1758]" captionStart="FIGURES 1–9" captionStartId="3.[151,264,1955,1979]" captionTargetBox="[187,1400,190,1920]" captionTargetId="figure-16@3.[173,1414,181,1931]" captionTargetPageId="3" captionText="FIGURES 1–9. Habitus of Mannerheimia: 1—M. afghanica (paratype), 2—M. arctica, 3—M. asiatica, 4—M. brevipennis, 5—M. japonica, 6—M. obscura, 7—M. pleshanovi (paratype), 8—M. yasudai, 9—M. zanettii (paratype). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5531374" httpUri="https://zenodo.org/record/5531374/files/figure.png" pageId="6" pageNumber="307">Fig. 1</figureCitation>
. Aedeagus as in
<figureCitation id="13332A52087B2448FD18F923FCB91690" box="[649,783,1732,1758]" captionStart="FIGURES 10–11" captionStartId="7.[151,264,1477,1501]" captionTargetBox="[161,1421,192,1444]" captionTargetId="figure-16@7.[151,1436,181,1453]" captionTargetPageId="7" captionText="FIGURES 10–11. Aedeagus of Mannerheimia afganica (paratype): 10—parameral view, 11—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5531376" httpUri="https://zenodo.org/record/5531376/files/figure.png" pageId="6" pageNumber="307">Figs. 10–11</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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