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<document id="421BDB4313A50083C2F6FA3EDB34BD61" ID="10.11646/zootaxa.4157.1.1" ID-DOI="10.11646/zootaxa.4157.1.1" ID-GBIF-Dataset="2389f078-1812-4b4e-826a-561cd7e46654" ID-ISSN="1175-5326" ID-Zenodo-Dep="262413" ID-ZooBank="6BF4514A-892F-499F-BC1E-B7920C7A00B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1472198526050" checkinUser="plazi" docAuthor="Jałoszyński, Paweł &amp; Perkovsky, Evgeny" docDate="2016" docId="03A187A9297EFFB6FF03C06D981AF8A2" docLanguage="en" docName="zootaxa.4157.1.1.pdf" docOrigin="Zootaxa 4157 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Glandulariini" docType="treatment" docVersion="8" lastPageNumber="72" masterDocId="FF98FFD12939FFF1FF94C52D9D54FFD1" masterDocTitle="Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber" masterLastPageNumber="85" masterPageNumber="1" pageNumber="72" updateTime="1698382676775" updateUser="plazi">
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<mods:title id="1717288706A9ED3392861A08FDBF990F">Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber</mods:title>
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<mods:namePart id="764417225292F7DED8DB5E8BA1F2B3C4">Jałoszyński, Paweł</mods:namePart>
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<mods:namePart id="3ACA459C6E0CAA34FA1F0AABFA2BC4DF">Perkovsky, Evgeny</mods:namePart>
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<mods:date id="DF868E3655D77BC5DEFD46D05437771C">2016</mods:date>
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<mods:number id="E48065EC68EF75FA3CBCE773AF4A9B0B">4157</mods:number>
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<mods:identifier id="975165ABC11FD63F7765F24DA14AFD14" type="DOI">10.11646/zootaxa.4157.1.1</mods:identifier>
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<treatment id="03A187A9297EFFB6FF03C06D981AF8A2" ID-DOI="http://doi.org/10.5281/zenodo.5681863" ID-GBIF-Taxon="124407831" ID-Zenodo-Dep="5681863" LSID="urn:lsid:plazi:treatment:03A187A9297EFFB6FF03C06D981AF8A2" httpUri="http://treatment.plazi.org/id/03A187A9297EFFB6FF03C06D981AF8A2" lastPageNumber="72" pageId="71" pageNumber="72">
<subSubSection id="C3126534297EFFB6FF03C06D9C39FAAD" pageId="71" pageNumber="72" type="nomenclature">
<paragraph id="8BB736BF297EFFB6FF03C06D9F27FA8B" blockId="71.[151,627,1344,1404]" box="[151,627,1344,1370]" pageId="71" pageNumber="72">
<heading id="D0FF81D3297EFFB6FF03C06D9F27FA8B" bold="true" box="[151,627,1344,1370]" fontSize="11" level="1" pageId="71" pageNumber="72" reason="1">
<emphasis id="B97CEAAD297EFFB6FF03C06D9F27FA8B" bold="true" box="[151,627,1344,1370]" pageId="71" pageNumber="72">
<taxonomicName id="4C084D3C297EFFB6FF03C06D9C16FA8B" box="[151,322,1344,1370]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="tribe" tribe="Glandulariini">Glandulariini</taxonomicName>
genus &amp; species indet. 1
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB736BF297EFFB6FF03C04E9C39FAAD" blockId="71.[151,627,1344,1404]" box="[151,365,1379,1404]" pageId="71" pageNumber="72">
(
<figureCitation id="13332A3A297EFFB6FF0BC04E9DA1FAAD" box="[159,245,1379,1404]" captionStart="FIGURES 21 – 34" captionStartId="8.[151,264,1436,1459]" captionTargetBox="[163,1421,202,1404]" captionTargetId="figure@8.[151,1436,193,1415]" captionTargetPageId="8" captionText="FIGURES 21 – 34. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 8972, Rovnoscydmus sp. specimen 8 (21); K- 3322, Rovnoscydmus sp. specimen 9 (22); K- 15548,? Rovnoscydmus sp. specimen 10 (23); K- 8971,? Rovnoscydmus sp. specimen 11 (24); K- 6045, Euconnus palaeogenus sp. n. (25); K- 9126, Euconnus sp. 1 (26); K- 24859,? Euconnus sp. 2 (27); K- 24302, Stenichnus proavus sp. n. (28); UA- 2216, Microscydmus sp. 1 (29); K- 6943,? Microscydmus sp. 2 (30); K- 3657 a,? Microscydmus sp. 3 (31); K- 6764,? Scydmoraphes sp. 1 (32); K- 6274, Glandulariini gen. &amp; sp. indet. 1 (33); K- 7089, Glandulariini gen. &amp; sp. indet. 2 (34)." httpUri="https://zenodo.org/record/262415/files/figure.png" pageId="71" pageNumber="72">Figs 33</figureCitation>
,
<figureCitation id="13332A3A297EFFB6FE95C0499C31FAAD" box="[257,357,1380,1404]" captionStart="FIGURES 217 – 220" captionStartId="72.[151,264,1930,1953]" captionTargetBox="[151,1436,193,1909]" captionTargetId="figure@72.[151,1436,193,1909]" captionTargetPageId="72" captionText="FIGURES 217 – 220. Glandulariini gen. &amp; sp. indet. 1 (K- 6274). Habitus in ventrolateral (217, 218) and dorsolateral (219) views, right profemur with a foretic deutonymph of? Acaridae (220)." httpUri="https://zenodo.org/record/262462/files/figure.png" pageId="71" pageNumber="72">217–220</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3126534297EFFB6FF03C0869F73F982" pageId="71" pageNumber="72" type="materials_examined">
<paragraph id="8BB736BF297EFFB6FF03C0869EDAFA36" blockId="71.[151,1437,1451,1908]" pageId="71" pageNumber="72">
<materialsCitation id="3B603CE2297EFFB6FF03C0869EDAFA36" ID-GBIF-Occurrence="1300093510" collectionCode="SIZK" county="Material" location="Europe" municipality="Late Eocene" pageId="71" pageNumber="72" specimenCount="1" stateProvince="Rovno">
<emphasis id="B97CEAAD297EFFB6FF03C0869E6AFA15" bold="true" box="[151,830,1451,1476]" pageId="71" pageNumber="72">
<collectingCounty id="62D64E33297EFFB6FF03C0869DABFA15" box="[151,255,1451,1476]" pageId="71" pageNumber="72">Material</collectingCounty>
studied.
<collectingMunicipality id="6BD3ACC5297EFFB6FEFAC0869F55FA15" box="[366,513,1451,1476]" pageId="71" pageNumber="72">Late Eocene</collectingMunicipality>
of
<location id="8ED76064297EFFB6FDBEC0869FD0FA15" LSID="urn:lsid:plazi:treatment:03A187A9297EFFB6FF03C06D981AF8A2:8ED76064297EFFB6FDBEC0869FD0FA15" box="[554,644,1451,1476]" county="Material" municipality="Late Eocene" name="Europe" pageId="71" pageNumber="72" stateProvince="Rovno">Europe</location>
,
<collectingRegion id="49CCF85D297EFFB6FD05C0869F8BFA15" box="[657,735,1451,1476]" country="Ukraine" name="Rivne" pageId="71" pageNumber="72">Rovno</collectingRegion>
amber:
</emphasis>
sex unknown; inclusion in slightly irregular pyramid of amber
<quantity id="4CF09B5A297EFFB6FE96C0E29C0CFA36" box="[258,344,1487,1512]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="71" pageNumber="72" unit="mm" value="9.5">9.5 mm</quantity>
long (
<figureCitation id="13332A3A297EFFB6FE34C0E29CA7FA36" box="[416,499,1487,1512]" captionStart="FIGURES 21 – 34" captionStartId="8.[151,264,1436,1459]" captionTargetBox="[163,1421,202,1404]" captionTargetId="figure@8.[151,1436,193,1415]" captionTargetPageId="8" captionText="FIGURES 21 – 34. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 8972, Rovnoscydmus sp. specimen 8 (21); K- 3322, Rovnoscydmus sp. specimen 9 (22); K- 15548,? Rovnoscydmus sp. specimen 10 (23); K- 8971,? Rovnoscydmus sp. specimen 11 (24); K- 6045, Euconnus palaeogenus sp. n. (25); K- 9126, Euconnus sp. 1 (26); K- 24859,? Euconnus sp. 2 (27); K- 24302, Stenichnus proavus sp. n. (28); UA- 2216, Microscydmus sp. 1 (29); K- 6943,? Microscydmus sp. 2 (30); K- 3657 a,? Microscydmus sp. 3 (31); K- 6764,? Scydmoraphes sp. 1 (32); K- 6274, Glandulariini gen. &amp; sp. indet. 1 (33); K- 7089, Glandulariini gen. &amp; sp. indet. 2 (34)." httpUri="https://zenodo.org/record/262415/files/figure.png" pageId="71" pageNumber="72">Fig. 33</figureCitation>
), collection number K-6274 (
<collectionCode id="ED19AE7A297EFFB6FCAAC0E29ED6FA36" box="[830,898,1487,1511]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="71" pageNumber="72">SIZK</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BB736BF297EFFB6FF53C0DE9F73F982" blockId="71.[151,1437,1451,1908]" pageId="71" pageNumber="72">
This specimen is very poorly visible inside particularly turbid piece of amber, and proper description is not possible. The beetle has BL about
<quantity id="4CF09B5A297EFFB6FDB3C33A9FC4F9FE" box="[551,656,1559,1583]" metricMagnitude="-3" metricUnit="m" metricValue="1.73" pageId="71" pageNumber="72" unit="mm" value="1.73">1.73 mm</quantity>
; HL about
<quantity id="4CF09B5A297EFFB6FC80C33A9E29F9FE" box="[788,893,1559,1583]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="71" pageNumber="72" unit="mm" value="0.3">0.30 mm</quantity>
, HW about
<quantity id="4CF09B5A297EFFB6FB9EC33A9926F9FE" box="[1034,1138,1559,1583]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="71" pageNumber="72" unit="mm" value="0.3">0.30 mm</quantity>
, AnL about
<quantity id="4CF09B5A297EFFB6FA91C33A983AF9FE" box="[1285,1390,1559,1583]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="71" pageNumber="72" unit="mm" value="0.34">0.34 mm</quantity>
; PL about
<quantity id="4CF09B5A297EFFB6FF48C3119C16F982" box="[220,322,1596,1620]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="71" pageNumber="72" unit="mm" value="0.28">0.28 mm</quantity>
, EL about
<quantity id="4CF09B5A297EFFB6FE28C3169F77F982" box="[444,547,1595,1620]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="71" pageNumber="72" unit="mm" value="0.55">0.55 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3126534297EFFB6FF53C372981AF8A2" pageId="71" pageNumber="72" type="discussion">
<paragraph id="8BB736BF297EFFB6FF53C372981AF8A2" blockId="71.[151,1437,1451,1908]" pageId="71" pageNumber="72">
<emphasis id="B97CEAAD297EFFB6FF53C3729C6FF9A9" bold="true" box="[199,315,1631,1656]" pageId="71" pageNumber="72">Remarks.</emphasis>
This specimen can only be assigned to the tribe (based on the structure of maxillary palps). It is exceptionally interesting because of a hypopus (phoretic deutonymph) of a mite belonging most likely to the family
<taxonomicName id="4C084D3C297EFFB6FF03C38A9DA8F911" box="[151,252,1703,1728]" class="Arachnida" family="Acaridae" kingdom="Animalia" order="Astigmata" pageId="71" pageNumber="72" phylum="Arthropoda" rank="family">Acaridae</taxonomicName>
attached to the clavate portion of the left profemur (
<figureCitation id="13332A3A297EFFB6FCF8C38A9944F96E" box="[876,1040,1703,1728]" captionStart="FIGURES 217 – 220" captionStartId="72.[151,264,1930,1953]" captionTargetBox="[151,1436,193,1909]" captionTargetId="figure@72.[151,1436,193,1909]" captionTargetPageId="72" captionText="FIGURES 217 – 220. Glandulariini gen. &amp; sp. indet. 1 (K- 6274). Habitus in ventrolateral (217, 218) and dorsolateral (219) views, right profemur with a foretic deutonymph of? Acaridae (220)." httpUri="https://zenodo.org/record/262462/files/figure.png" pageId="71" pageNumber="72">Figs 219–220</figureCitation>
). The first author frequently finds hypopi of astigmatid (acarid and histiostomatid) mites attached to various body parts of extant species of
<taxonomicName id="4C084D3C297EFFB6FF03C3C29C69F8D9" box="[151,317,1775,1800]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="subFamily" subFamily="Scydmaeninae">Scydmaeninae</taxonomicName>
, especially those associated with habitats rich in decaying plant debris, although this association remains unrecorded in literature. For the first time we demonstrate that astigmatid mites have been using
<taxonomicName id="4C084D3C297EFFB6FF03C21A9C6EF881" box="[151,314,1847,1872]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="subFamily" subFamily="Scydmaeninae">Scydmaeninae</taxonomicName>
(of which many are known as predators of acaridid, oribatid and/or uropodine mites, see
<bibRefCitation id="EF994B4E297EFFB6FA8FC21A9DA6F8A5" author="Jaloszynski" pageId="71" pageNumber="72" refString="Jaloszynski, P. (2012 d) Adults of European ant-like stone beetles (Coleoptera: Staphylinidae: Scydmaeninae) Scydmaenus tarsatus Muller &amp; Kunze and S. hellwigii (Herbst) prey on soft-bodied arthropods. Entomological Science, 15, 35 - 41. http: // dx. doi. org / 10.1111 / j. 1479 - 8298.2011.00479. x" type="journal article" year="2012" yearSuffix="d">Jałoszyński (2012d)</bibRefCitation>
,
<bibRefCitation id="EF994B4E297EFFB6FF69C2769F2AF8A5" author="Jaloszynski" box="[253,638,1883,1908]" pageId="71" pageNumber="72" refString="Jaloszynski, P. &amp; Olszanowski, Z. (2013) Specialized feeding of Euconnus pubicollis (Coleoptera: Staphylinidae, Scydmaeninae) on oribatid mites: prey preferences and hunting behaviour. European Journal of Entomology, 110 (2), 339 - 353. http: // dx. doi. org / 10.14411 / eje. 2013.047" type="journal article" year="2013">Jałoszyński &amp; Olszanowski (2013</bibRefCitation>
, 2015, 2016) as means of dispersal at least since Lower Eocene.
</paragraph>
</subSubSection>
</treatment>
</document>
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