242 lines
37 KiB
XML
242 lines
37 KiB
XML
<document id="4E2CDB24E1FAB304DBB309D6F075A76A" ID="10.11646/zootaxa.4157.1.1" ID-DOI="10.11646/zootaxa.4157.1.1" ID-GBIF-Dataset="2389f078-1812-4b4e-826a-561cd7e46654" ID-ISSN="1175-5326" ID-Zenodo-Dep="262413" ID-ZooBank="6BF4514A-892F-499F-BC1E-B7920C7A00B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1472198526050" checkinUser="plazi" docAuthor="Jałoszyński, Paweł & Perkovsky, Evgeny" docDate="2016" docId="03A187A92978FFB2FF03C21B9C34FA85" docLanguage="en" docName="zootaxa.4157.1.1.pdf" docOrigin="Zootaxa 4157 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Microscydmus" docType="treatment" docVersion="9" lastPageNumber="68" masterDocId="FF98FFD12939FFF1FF94C52D9D54FFD1" masterDocTitle="Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber" masterLastPageNumber="85" masterPageNumber="1" pageNumber="66" updateTime="1698382676775" updateUser="plazi">
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<mods:titleInfo id="55DBD97691A8429EE0DD1345598BDF24">
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<mods:title id="1CC0A041849283A1E8C09841F2DECD9C">Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber</mods:title>
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<mods:name id="539039C9C43C7FFC7777DD6DA4E3B0A7" type="personal">
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<mods:namePart id="2FEFA15C538AC636E719A8A562F80946">Jałoszyński, Paweł</mods:namePart>
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<mods:roleTerm id="8E9B61307587D00326737FB58BDC9BAD">Author</mods:roleTerm>
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<mods:namePart id="229DD376D0B8EB922CECD8C21768D87A">Perkovsky, Evgeny</mods:namePart>
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<mods:typeOfResource id="2FC548404B406B06EA5DF51BF3F880D8">text</mods:typeOfResource>
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<mods:title id="82ACA78A6D7DEF3B5CF56E440B600C8E">Zootaxa</mods:title>
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<mods:part id="4131C48B118B0896A9B36BF4D29C1194">
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<mods:date id="30F2AEACDB57E4B983187E995E7EE92A">2016</mods:date>
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<mods:number id="6784C906BD69C6A8558206B9BE8CE651">4157</mods:number>
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<mods:identifier id="A6BDAEEE93287981E5328348AE6B9E16" type="DOI">10.11646/zootaxa.4157.1.1</mods:identifier>
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<treatment id="03A187A92978FFB2FF03C21B9C34FA85" ID-DOI="http://doi.org/10.5281/zenodo.5681841" ID-GBIF-Taxon="124407829" ID-Zenodo-Dep="5681841" LSID="urn:lsid:plazi:treatment:03A187A92978FFB2FF03C21B9C34FA85" httpUri="http://treatment.plazi.org/id/03A187A92978FFB2FF03C21B9C34FA85" lastPageId="67" lastPageNumber="68" pageId="65" pageNumber="66">
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<subSubSection id="C31265342978FFB0FF03C21B9C39F8A3" pageId="65" pageNumber="66" type="nomenclature">
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<paragraph id="8BB736BF2978FFB0FF03C21B9CD3F881" blockId="65.[151,391,1846,1906]" box="[151,391,1846,1872]" pageId="65" pageNumber="66">
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<heading id="D0FF81D32978FFB0FF03C21B9CD3F881" bold="true" box="[151,391,1846,1872]" fontSize="11" level="1" pageId="65" pageNumber="66" reason="1">
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<emphasis id="B97CEAAD2978FFB0FF03C21B9CD3F881" bold="true" box="[151,391,1846,1872]" pageId="65" pageNumber="66">
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<taxonomicName id="4C084D3C2978FFB0FF03C21B9C13F881" ID-CoL="8VRV9" box="[151,327,1846,1872]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="genus">
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<emphasis id="B97CEAAD2978FFB0FF03C21B9C13F881" bold="true" box="[151,327,1846,1872]" italics="true" pageId="65" pageNumber="66">Microscydmus</emphasis>
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</taxonomicName>
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sp. 1
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB736BF2978FFB0FF03C2749C39F8A3" blockId="65.[151,391,1846,1906]" box="[151,365,1881,1906]" pageId="65" pageNumber="66">
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(
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<figureCitation id="13332A3A2978FFB0FF0BC2749DA1F8A3" box="[159,245,1881,1906]" captionStart="FIGURES 21 – 34" captionStartId="8.[151,264,1436,1459]" captionTargetBox="[163,1421,202,1404]" captionTargetId="figure@8.[151,1436,193,1415]" captionTargetPageId="8" captionText="FIGURES 21 – 34. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 8972, Rovnoscydmus sp. specimen 8 (21); K- 3322, Rovnoscydmus sp. specimen 9 (22); K- 15548,? Rovnoscydmus sp. specimen 10 (23); K- 8971,? Rovnoscydmus sp. specimen 11 (24); K- 6045, Euconnus palaeogenus sp. n. (25); K- 9126, Euconnus sp. 1 (26); K- 24859,? Euconnus sp. 2 (27); K- 24302, Stenichnus proavus sp. n. (28); UA- 2216, Microscydmus sp. 1 (29); K- 6943,? Microscydmus sp. 2 (30); K- 3657 a,? Microscydmus sp. 3 (31); K- 6764,? Scydmoraphes sp. 1 (32); K- 6274, Glandulariini gen. & sp. indet. 1 (33); K- 7089, Glandulariini gen. & sp. indet. 2 (34)." httpUri="https://zenodo.org/record/262415/files/figure.png" pageId="65" pageNumber="66">Figs 29</figureCitation>
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,
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<figureCitation id="13332A3A2978FFB0FE95C2779C31F8A3" box="[257,357,1882,1906]" captionStart="FIGURES 199 – 203" captionStartId="66.[151,264,1758,1781]" captionTargetBox="[151,1436,193,1728]" captionTargetId="figure@66.[151,1436,193,1738]" captionTargetPageId="66" captionText="FIGURES 199 – 203. Microscydmus sp. 1 (UA- 2216). Dorsal (199, 202), right lateral (200, 203) and left lateral (201) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; fr, frons; hyr, hypomeral ridge; mcp, mesocoxal projection; nr, ' neck' region; vt, vertex." httpUri="https://zenodo.org/record/262458/files/figure.png" pageId="65" pageNumber="66">199–203</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31265342978FFB0FF03C28C9EA2F80F" pageId="65" pageNumber="66" type="materials_examined">
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<paragraph id="8BB736BF2978FFB0FF03C28C9EA2F80F" blockId="65.[151,1436,1953,2014]" pageId="65" pageNumber="66">
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<materialsCitation id="3B603CE22978FFB0FF03C28C9EA2F80F" ID-GBIF-Occurrence="1300093518" collectionCode="SIZK" county="Material" location="Europe" municipality="Late Eocene" pageId="65" pageNumber="66" specimenCount="1" stateProvince="Rovno">
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<emphasis id="B97CEAAD2978FFB0FF03C28C9E13F86B" bold="true" box="[151,839,1953,1978]" pageId="65" pageNumber="66">
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<collectingCounty id="62D64E332978FFB0FF03C28C9DABF86B" box="[151,255,1953,1978]" pageId="65" pageNumber="66">Material</collectingCounty>
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studied.
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<collectingMunicipality id="6BD3ACC52978FFB0FEE4C28C9F51F86B" box="[368,517,1953,1978]" pageId="65" pageNumber="66">Late Eocene</collectingMunicipality>
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of
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<location id="8ED760642978FFB0FDA4C28C9FDEF86B" LSID="urn:lsid:plazi:treatment:03A187A92978FFB2FF03C21B9C34FA85:8ED760642978FFB0FDA4C28C9FDEF86B" box="[560,650,1953,1978]" county="Material" municipality="Late Eocene" name="Europe" pageId="65" pageNumber="66" stateProvince="Rovno">Europe</location>
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,
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<collectingRegion id="49CCF85D2978FFB0FD0DC28C9FB3F86B" box="[665,743,1953,1978]" country="Ukraine" name="Rivne" pageId="65" pageNumber="66">Rovno</collectingRegion>
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amber:
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</emphasis>
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sex unknown; inclusion in elongate, subtrapezoidal prism of amber
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<quantity id="4CF09B5A2978FFB0FEDDC2E89CF8F80C" box="[329,428,1989,2014]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="65" pageNumber="66" unit="mm" value="11.5">11.5 mm</quantity>
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long (
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<figureCitation id="13332A3A2978FFB0FE60C2E89F13F80F" box="[500,583,1989,2014]" captionStart="FIGURES 21 – 34" captionStartId="8.[151,264,1436,1459]" captionTargetBox="[163,1421,202,1404]" captionTargetId="figure@8.[151,1436,193,1415]" captionTargetPageId="8" captionText="FIGURES 21 – 34. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 8972, Rovnoscydmus sp. specimen 8 (21); K- 3322, Rovnoscydmus sp. specimen 9 (22); K- 15548,? Rovnoscydmus sp. specimen 10 (23); K- 8971,? Rovnoscydmus sp. specimen 11 (24); K- 6045, Euconnus palaeogenus sp. n. (25); K- 9126, Euconnus sp. 1 (26); K- 24859,? Euconnus sp. 2 (27); K- 24302, Stenichnus proavus sp. n. (28); UA- 2216, Microscydmus sp. 1 (29); K- 6943,? Microscydmus sp. 2 (30); K- 3657 a,? Microscydmus sp. 3 (31); K- 6764,? Scydmoraphes sp. 1 (32); K- 6274, Glandulariini gen. & sp. indet. 1 (33); K- 7089, Glandulariini gen. & sp. indet. 2 (34)." httpUri="https://zenodo.org/record/262415/files/figure.png" pageId="65" pageNumber="66">Fig. 29</figureCitation>
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), collection number UA-2216 (
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<collectionCode id="ED19AE7A2978FFB0FC32C2EB9EBEF80F" box="[934,1002,1990,2014]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="65" pageNumber="66">SIZK</collectionCode>
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).
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</materialsCitation>
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||
</paragraph>
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</subSubSection>
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<caption id="DF776637297BFFB3FF03C3F39FC6F8E3" httpUri="https://zenodo.org/record/262458/files/figure.png" pageId="66" pageNumber="67" targetBox="[151,1436,193,1728]" targetPageId="66">
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<paragraph id="8BB736BF297BFFB3FF03C3F39FC6F8E3" blockId="66.[151,1437,1758,1842]" pageId="66" pageNumber="67">
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<emphasis id="B97CEAAD297BFFB3FF03C3F39C3BF925" bold="true" box="[151,367,1758,1781]" pageId="66" pageNumber="67">FIGURES 199–203.</emphasis>
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<taxonomicName id="4C084D3C297BFFB3FEE2C3F29F5DF924" box="[374,521,1759,1781]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="genus">
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<emphasis id="B97CEAAD297BFFB3FEE2C3F29F5DF924" box="[374,521,1759,1781]" italics="true" pageId="66" pageNumber="67">Microscydmus</emphasis>
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</taxonomicName>
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sp. 1 (UA-2216). Dorsal (199, 202), right lateral (200, 203) and left lateral (201) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; fr, frons; hyr, hypomeral ridge; mcp, mesocoxal projection; nr, 'neck' region; vt, vertex.
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</paragraph>
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</caption>
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||
<subSubSection id="C3126534297BFFB2FF53C2779ECFFD54" lastPageId="67" lastPageNumber="68" pageId="66" pageNumber="67" type="description">
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||
<paragraph id="8BB736BF297BFFB3FF53C2779C79F847" blockId="66.[151,1436,1882,2015]" pageId="66" pageNumber="67">
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<emphasis id="B97CEAAD297BFFB3FF53C2779C0DF8A2" bold="true" box="[199,345,1882,1907]" pageId="66" pageNumber="67">Description.</emphasis>
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Body (
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<figureCitation id="13332A3A297BFFB3FE38C2779F1DF8A3" box="[428,585,1882,1907]" captionStart="FIGURES 199 – 203" captionStartId="66.[151,264,1758,1781]" captionTargetBox="[151,1436,193,1728]" captionTargetId="figure@66.[151,1436,193,1738]" captionTargetPageId="66" captionText="FIGURES 199 – 203. Microscydmus sp. 1 (UA- 2216). Dorsal (199, 202), right lateral (200, 203) and left lateral (201) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; fr, frons; hyr, hypomeral ridge; mcp, mesocoxal projection; nr, ' neck' region; vt, vertex." httpUri="https://zenodo.org/record/262458/files/figure.png" pageId="66" pageNumber="67">Figs 199–203</figureCitation>
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) elongate, relatively stout and moderately convex, moderately light brown; BL
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<quantity id="4CF09B5A297BFFB3FF56C2509C7DF847" box="[194,297,1917,1942]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="66" pageNumber="67" unit="mm" value="0.58">0.58 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BB736BF297BFFB2FF53C28F9FCEFEB5" blockId="66.[151,1436,1882,2015]" lastBlockId="67.[151,1437,151,1365]" lastPageId="67" lastPageNumber="68" pageId="66" pageNumber="67">
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Head (
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<figureCitation id="13332A3A297BFFB3FE85C28F9CF9F86B" box="[273,429,1954,1979]" captionStart="FIGURES 199 – 203" captionStartId="66.[151,264,1758,1781]" captionTargetBox="[151,1436,193,1728]" captionTargetId="figure@66.[151,1436,193,1738]" captionTargetPageId="66" captionText="FIGURES 199 – 203. Microscydmus sp. 1 (UA- 2216). Dorsal (199, 202), right lateral (200, 203) and left lateral (201) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; fr, frons; hyr, hypomeral ridge; mcp, mesocoxal projection; nr, ' neck' region; vt, vertex." httpUri="https://zenodo.org/record/262458/files/figure.png" pageId="66" pageNumber="67">Figs 199–203</figureCitation>
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) short and subtrapezoidal, broadest at eyes, HL
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<quantity id="4CF09B5A297BFFB3FC5CC28F997AF86B" box="[968,1070,1954,1978]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="66" pageNumber="67" unit="mm" value="0.13">0.13 mm</quantity>
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, HW
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||
<quantity id="4CF09B5A297BFFB3FBFBC28F9982F86B" box="[1135,1238,1954,1978]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="66" pageNumber="67" unit="mm" value="0.13">0.13 mm</quantity>
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||
; vertex (
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||
<figureCitation id="13332A3A297BFFB3FAA2C28F98CCF86B" box="[1334,1432,1954,1979]" captionStart="FIGURES 199 – 203" captionStartId="66.[151,264,1758,1781]" captionTargetBox="[151,1436,193,1728]" captionTargetId="figure@66.[151,1436,193,1738]" captionTargetPageId="66" captionText="FIGURES 199 – 203. Microscydmus sp. 1 (UA- 2216). Dorsal (199, 202), right lateral (200, 203) and left lateral (201) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; fr, frons; hyr, hypomeral ridge; mcp, mesocoxal projection; nr, ' neck' region; vt, vertex." httpUri="https://zenodo.org/record/262458/files/figure.png" pageId="66" pageNumber="67">Fig. 202</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD297BFFB3FF03C2EA9DFFF80F" box="[151,171,1991,2014]" italics="true" pageId="66" pageNumber="67">vt</emphasis>
|
||
) and frons (
|
||
<figureCitation id="13332A3A297BFFB3FEA3C2E89CCEF80F" box="[311,410,1989,2014]" captionStart="FIGURES 199 – 203" captionStartId="66.[151,264,1758,1781]" captionTargetBox="[151,1436,193,1728]" captionTargetId="figure@66.[151,1436,193,1738]" captionTargetPageId="66" captionText="FIGURES 199 – 203. Microscydmus sp. 1 (UA- 2216). Dorsal (199, 202), right lateral (200, 203) and left lateral (201) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; fr, frons; hyr, hypomeral ridge; mcp, mesocoxal projection; nr, ' neck' region; vt, vertex." httpUri="https://zenodo.org/record/262458/files/figure.png" pageId="66" pageNumber="67">Fig. 202</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD297BFFB3FE33C2EB9CEEF80F" box="[423,442,1990,2014]" italics="true" pageId="66" pageNumber="67">fr</emphasis>
|
||
) confluent, weakly convex and together subtrapezoidal, strongly transverse; eyes large, semioval, strongly convex; tempora distinctly shorter than eyes. Punctures (
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||
<figureCitation id="13332A3A297AFFB2FC7DC5BA991DFF61" box="[1001,1097,151,176]" captionStart="FIGURES 199 – 203" captionStartId="66.[151,264,1758,1781]" captionTargetBox="[151,1436,193,1728]" captionTargetId="figure@66.[151,1436,193,1738]" captionTargetPageId="66" captionText="FIGURES 199 – 203. Microscydmus sp. 1 (UA- 2216). Dorsal (199, 202), right lateral (200, 203) and left lateral (201) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; fr, frons; hyr, hypomeral ridge; mcp, mesocoxal projection; nr, ' neck' region; vt, vertex." httpUri="https://zenodo.org/record/262458/files/figure.png" pageId="67" pageNumber="68">Fig. 202</figureCitation>
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||
) on frons and vertex very fine but discernible, separated by spaces 2–3 × as wide as diameters of punctures; setae (
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||
<figureCitation id="13332A3A297AFFB2FBD8C59199F8FF05" box="[1100,1196,188,213]" captionStart="FIGURES 199 – 203" captionStartId="66.[151,264,1758,1781]" captionTargetBox="[151,1436,193,1728]" captionTargetId="figure@66.[151,1436,193,1738]" captionTargetPageId="66" captionText="FIGURES 199 – 203. Microscydmus sp. 1 (UA- 2216). Dorsal (199, 202), right lateral (200, 203) and left lateral (201) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; fr, frons; hyr, hypomeral ridge; mcp, mesocoxal projection; nr, ' neck' region; vt, vertex." httpUri="https://zenodo.org/record/262458/files/figure.png" pageId="67" pageNumber="68">Fig. 202</figureCitation>
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) sparse, short, nearly recumbent. Antennae (
|
||
<figureCitation id="13332A3A297AFFB2FE0AC5F29F6BFF29" box="[414,575,223,248]" captionStart="FIGURES 199 – 203" captionStartId="66.[151,264,1758,1781]" captionTargetBox="[151,1436,193,1728]" captionTargetId="figure@66.[151,1436,193,1738]" captionTargetPageId="66" captionText="FIGURES 199 – 203. Microscydmus sp. 1 (UA- 2216). Dorsal (199, 202), right lateral (200, 203) and left lateral (201) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; fr, frons; hyr, hypomeral ridge; mcp, mesocoxal projection; nr, ' neck' region; vt, vertex." httpUri="https://zenodo.org/record/262458/files/figure.png" pageId="67" pageNumber="68">Figs 199–203</figureCitation>
|
||
) distinctly shorter than half BL, AnL about
|
||
<quantity id="4CF09B5A297AFFB2FBC6C5CD99E9FF26" box="[1106,1213,224,248]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="67" pageNumber="68" unit="mm" value="0.2">0.20 mm</quantity>
|
||
, scape and pedicel strongly elongate, shapes of most of flagellomeres difficult to assess, antennomeres IX–XI distinctly broader than VIII (
|
||
<figureCitation id="13332A3A297AFFB2FF43C40A9C6EFE91" box="[215,314,295,320]" captionStart="FIGURES 199 – 203" captionStartId="66.[151,264,1758,1781]" captionTargetBox="[151,1436,193,1728]" captionTargetId="figure@66.[151,1436,193,1738]" captionTargetPageId="66" captionText="FIGURES 199 – 203. Microscydmus sp. 1 (UA- 2216). Dorsal (199, 202), right lateral (200, 203) and left lateral (201) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; fr, frons; hyr, hypomeral ridge; mcp, mesocoxal projection; nr, ' neck' region; vt, vertex." httpUri="https://zenodo.org/record/262458/files/figure.png" pageId="67" pageNumber="68">Fig. 202</figureCitation>
|
||
), IX and X slightly transverse, XI much shorter than IX–X combined, about 1.2 × as long as broad, slightly narrower than X and rounded at apex.
|
||
</paragraph>
|
||
<paragraph id="8BB736BF297AFFB2FF53C45D98DAFDC9" blockId="67.[151,1437,151,1365]" pageId="67" pageNumber="68">
|
||
Pronotum (
|
||
<figureCitation id="13332A3A297AFFB2FEDCC4429CBEFE59" box="[328,490,367,392]" captionStart="FIGURES 199 – 203" captionStartId="66.[151,264,1758,1781]" captionTargetBox="[151,1436,193,1728]" captionTargetId="figure@66.[151,1436,193,1738]" captionTargetPageId="66" captionText="FIGURES 199 – 203. Microscydmus sp. 1 (UA- 2216). Dorsal (199, 202), right lateral (200, 203) and left lateral (201) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; fr, frons; hyr, hypomeral ridge; mcp, mesocoxal projection; nr, ' neck' region; vt, vertex." httpUri="https://zenodo.org/record/262458/files/figure.png" pageId="67" pageNumber="68">Figs 199, 202</figureCitation>
|
||
) round, broadest near middle; PL
|
||
<quantity id="4CF09B5A297AFFB2FCEAC4429EBCFE56" box="[894,1000,367,392]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="67" pageNumber="68" unit="mm" value="0.15">0.15 mm</quantity>
|
||
, PW
|
||
<quantity id="4CF09B5A297AFFB2FBBFC45D99C2FE56" box="[1067,1174,368,392]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="67" pageNumber="68" unit="mm" value="0.18">0.18 mm</quantity>
|
||
; anterior and posterior margins nearly straight, anterior and posterior pronotal corners broadly obtuse-angled and blunt; sides strongly rounded; base with short but distinct transverse antebasal groove (
|
||
<figureCitation id="13332A3A297AFFB2FC14C49A9EB6FE01" box="[896,994,439,464]" captionStart="FIGURES 199 – 203" captionStartId="66.[151,264,1758,1781]" captionTargetBox="[151,1436,193,1728]" captionTargetId="figure@66.[151,1436,193,1738]" captionTargetPageId="66" captionText="FIGURES 199 – 203. Microscydmus sp. 1 (UA- 2216). Dorsal (199, 202), right lateral (200, 203) and left lateral (201) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; fr, frons; hyr, hypomeral ridge; mcp, mesocoxal projection; nr, ' neck' region; vt, vertex." httpUri="https://zenodo.org/record/262458/files/figure.png" pageId="67" pageNumber="68">Fig. 202</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD297AFFB2FC79C49A9943FE01" box="[1005,1047,439,464]" italics="true" pageId="67" pageNumber="68">abg</emphasis>
|
||
) and one large and shallow lateral antebasal pit (
|
||
<figureCitation id="13332A3A297AFFB2FEAAC4F19CF2FE25" box="[318,422,476,501]" captionStart="FIGURES 199 – 203" captionStartId="66.[151,264,1758,1781]" captionTargetBox="[151,1436,193,1728]" captionTargetId="figure@66.[151,1436,193,1738]" captionTargetPageId="66" captionText="FIGURES 199 – 203. Microscydmus sp. 1 (UA- 2216). Dorsal (199, 202), right lateral (200, 203) and left lateral (201) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; fr, frons; hyr, hypomeral ridge; mcp, mesocoxal projection; nr, ' neck' region; vt, vertex." httpUri="https://zenodo.org/record/262458/files/figure.png" pageId="67" pageNumber="68">Fig. 202</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD297AFFB2FE22C4F19CB4FE24" box="[438,480,476,501]" italics="true" pageId="67" pageNumber="68">abp</emphasis>
|
||
) at each side. Punctures on pronotal disc (
|
||
<figureCitation id="13332A3A297AFFB2FC7CC4F19919FE25" box="[1000,1101,476,501]" captionStart="FIGURES 199 – 203" captionStartId="66.[151,264,1758,1781]" captionTargetBox="[151,1436,193,1728]" captionTargetId="figure@66.[151,1436,193,1738]" captionTargetPageId="66" captionText="FIGURES 199 – 203. Microscydmus sp. 1 (UA- 2216). Dorsal (199, 202), right lateral (200, 203) and left lateral (201) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; fr, frons; hyr, hypomeral ridge; mcp, mesocoxal projection; nr, ' neck' region; vt, vertex." httpUri="https://zenodo.org/record/262458/files/figure.png" pageId="67" pageNumber="68">Fig. 202</figureCitation>
|
||
) very small but discernible, separated by spaces 3–4 × as wide as diameters of punctures; setae (
|
||
<figureCitation id="13332A3A297AFFB2FC06C4D29EA6FDC9" box="[914,1010,511,536]" captionStart="FIGURES 199 – 203" captionStartId="66.[151,264,1758,1781]" captionTargetBox="[151,1436,193,1728]" captionTargetId="figure@66.[151,1436,193,1738]" captionTargetPageId="66" captionText="FIGURES 199 – 203. Microscydmus sp. 1 (UA- 2216). Dorsal (199, 202), right lateral (200, 203) and left lateral (201) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; fr, frons; hyr, hypomeral ridge; mcp, mesocoxal projection; nr, ' neck' region; vt, vertex." httpUri="https://zenodo.org/record/262458/files/figure.png" pageId="67" pageNumber="68">Fig. 202</figureCitation>
|
||
) short and sparse, nearly recumbent.
|
||
</paragraph>
|
||
<paragraph id="8BB736BF297AFFB2FF53C7099E38FDB1" blockId="67.[151,1437,151,1365]" pageId="67" pageNumber="68">
|
||
Elytra (
|
||
<figureCitation id="13332A3A297AFFB2FE8FC7099CEDFDED" box="[283,441,548,573]" captionStart="FIGURES 199 – 203" captionStartId="66.[151,264,1758,1781]" captionTargetBox="[151,1436,193,1728]" captionTargetId="figure@66.[151,1436,193,1738]" captionTargetPageId="66" captionText="FIGURES 199 – 203. Microscydmus sp. 1 (UA- 2216). Dorsal (199, 202), right lateral (200, 203) and left lateral (201) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; fr, frons; hyr, hypomeral ridge; mcp, mesocoxal projection; nr, ' neck' region; vt, vertex." httpUri="https://zenodo.org/record/262458/files/figure.png" pageId="67" pageNumber="68">Figs 199–203</figureCitation>
|
||
) elongate oval, moderately convex, EL
|
||
<quantity id="4CF09B5A297AFFB2FC14C7099EB3FDED" box="[896,999,548,572]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="67" pageNumber="68" unit="mm" value="0.3">0.30 mm</quantity>
|
||
, EW
|
||
<quantity id="4CF09B5A297AFFB2FBB3C70999DBFDED" box="[1063,1167,548,572]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="67" pageNumber="68" unit="mm" value="0.2">0.20 mm</quantity>
|
||
, EI 1.50; punctures and setae similar to those on pronotum (
|
||
<figureCitation id="13332A3A297AFFB2FDBDC76A9FDDFDB1" box="[553,649,583,608]" captionStart="FIGURES 199 – 203" captionStartId="66.[151,264,1758,1781]" captionTargetBox="[151,1436,193,1728]" captionTargetId="figure@66.[151,1436,193,1738]" captionTargetPageId="66" captionText="FIGURES 199 – 203. Microscydmus sp. 1 (UA- 2216). Dorsal (199, 202), right lateral (200, 203) and left lateral (201) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; fr, frons; hyr, hypomeral ridge; mcp, mesocoxal projection; nr, ' neck' region; vt, vertex." httpUri="https://zenodo.org/record/262458/files/figure.png" pageId="67" pageNumber="68">Fig. 202</figureCitation>
|
||
) but slightly denser.
|
||
</paragraph>
|
||
<paragraph id="8BB736BF297AFFB2FF53C7409ECFFD54" blockId="67.[151,1437,151,1365]" box="[199,923,620,645]" pageId="67" pageNumber="68">
|
||
Legs (
|
||
<figureCitation id="13332A3A297AFFB2FE98C7419CB4FD55" box="[268,480,620,645]" captionStart="FIGURES 199 – 203" captionStartId="66.[151,264,1758,1781]" captionTargetBox="[151,1436,193,1728]" captionTargetId="figure@66.[151,1436,193,1738]" captionTargetPageId="66" captionText="FIGURES 199 – 203. Microscydmus sp. 1 (UA- 2216). Dorsal (199, 202), right lateral (200, 203) and left lateral (201) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; fr, frons; hyr, hypomeral ridge; mcp, mesocoxal projection; nr, ' neck' region; vt, vertex." httpUri="https://zenodo.org/record/262458/files/figure.png" pageId="67" pageNumber="68">Figs 200–201, 203</figureCitation>
|
||
) moderately long, slender, unmodified.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3126534297AFFB2FF53C7A29C34FA85" pageId="67" pageNumber="68" type="discussion">
|
||
<paragraph id="8BB736BF297AFFB2FF53C7A29F0BFC3D" blockId="67.[151,1437,151,1365]" pageId="67" pageNumber="68">
|
||
<emphasis id="B97CEAAD297AFFB2FF53C7A29C6FFD79" bold="true" box="[199,315,655,680]" pageId="67" pageNumber="68">Remarks.</emphasis>
|
||
Diagnostic characters of
|
||
<taxonomicName id="4C084D3C297AFFB2FDF9C7A29E5BFD79" box="[621,783,655,680]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD297AFFB2FDF9C7A29E5BFD79" box="[621,783,655,680]" italics="true" pageId="67" pageNumber="68">Microscydmus</emphasis>
|
||
</taxonomicName>
|
||
were illustrated and discussed by
|
||
<bibRefCitation id="EF994B4E297AFFB2FB3BC7A298CCFD79" author="Jaloszynski" box="[1199,1432,655,680]" pageId="67" pageNumber="68" refString="Jaloszynski, P. (2014 d) The smallest Cyrtoscydmini of Australia: revision of Microscydmus Saulcy & Croissandeau and Penicillidmus gen. n. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 3774 (1), 1 - 30. http: // dx. doi. org / 10.11646 / zootaxa. 3774.1.1" type="journal article" year="2014" yearSuffix="d">Jałoszyński (2014d)</bibRefCitation>
|
||
. These are typically very small beetles, rarely exceeding
|
||
<quantity id="4CF09B5A297AFFB2FC9CC7999E1FFD1D" box="[776,843,692,716]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="67" pageNumber="68" unit="mm" value="1.0">1 mm</quantity>
|
||
, and often below
|
||
<quantity id="4CF09B5A297AFFB2FB84C799993CFD1D" box="[1040,1128,692,716]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="67" pageNumber="68" unit="mm" value="0.8">0.8 mm</quantity>
|
||
. The specimen from
|
||
<collectingRegion id="49CCF85D297AFFB2FAC6C79898C9FD1D" box="[1362,1437,693,716]" country="Ukraine" name="Rivne" pageId="67" pageNumber="68">Rovno</collectingRegion>
|
||
amber is extremely small (BL merely
|
||
<quantity id="4CF09B5A297AFFB2FDAAC7FA9FF7FD21" box="[574,675,727,752]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="67" pageNumber="68" unit="mm" value="0.58">0.58 mm</quantity>
|
||
) and has most of the important ventral structures obscured by turbid or darkened amber. However, besides the small body, the specimen has the general shape typical of
|
||
<taxonomicName id="4C084D3C297AFFB2FB67C7D198C1FCC4" box="[1267,1429,764,789]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD297AFFB2FB67C7D198C1FCC4" box="[1267,1429,764,789]" italics="true" pageId="67" pageNumber="68">Microscydmus</emphasis>
|
||
</taxonomicName>
|
||
, with a short head and round pronotum; its antennae have clubs composed of three distal antennomeres; the prothoracic hypomera have hypomeral ridges (
|
||
<figureCitation id="13332A3A297AFFB2FD24C6699E40FC8D" box="[688,788,836,861]" captionStart="FIGURES 199 – 203" captionStartId="66.[151,264,1758,1781]" captionTargetBox="[151,1436,193,1728]" captionTargetId="figure@66.[151,1436,193,1738]" captionTargetPageId="66" captionText="FIGURES 199 – 203. Microscydmus sp. 1 (UA- 2216). Dorsal (199, 202), right lateral (200, 203) and left lateral (201) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; fr, frons; hyr, hypomeral ridge; mcp, mesocoxal projection; nr, ' neck' region; vt, vertex." httpUri="https://zenodo.org/record/262458/files/figure.png" pageId="67" pageNumber="68">Fig. 203</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD297AFFB2FCB7C6699E1CFC8C" box="[803,840,836,861]" italics="true" pageId="67" pageNumber="68">hyr</emphasis>
|
||
) and, above all, the pronotal base bears a short but deep median transverse antebasal groove (
|
||
<figureCitation id="13332A3A297AFFB2FDE4C64A9F86FC51" box="[624,722,871,896]" captionStart="FIGURES 199 – 203" captionStartId="66.[151,264,1758,1781]" captionTargetBox="[151,1436,193,1728]" captionTargetId="figure@66.[151,1436,193,1738]" captionTargetPageId="66" captionText="FIGURES 199 – 203. Microscydmus sp. 1 (UA- 2216). Dorsal (199, 202), right lateral (200, 203) and left lateral (201) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; fr, frons; hyr, hypomeral ridge; mcp, mesocoxal projection; nr, ' neck' region; vt, vertex." httpUri="https://zenodo.org/record/262458/files/figure.png" pageId="67" pageNumber="68">Fig. 202</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD297AFFB2FD49C64A9E53FC51" box="[733,775,871,896]" italics="true" pageId="67" pageNumber="68">abg</emphasis>
|
||
) flanked at each side by one lateral antebasal pit (
|
||
<figureCitation id="13332A3A297AFFB2FAA2C64A98C3FC51" box="[1334,1431,871,896]" captionStart="FIGURES 199 – 203" captionStartId="66.[151,264,1758,1781]" captionTargetBox="[151,1436,193,1728]" captionTargetId="figure@66.[151,1436,193,1738]" captionTargetPageId="66" captionText="FIGURES 199 – 203. Microscydmus sp. 1 (UA- 2216). Dorsal (199, 202), right lateral (200, 203) and left lateral (201) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; fr, frons; hyr, hypomeral ridge; mcp, mesocoxal projection; nr, ' neck' region; vt, vertex." httpUri="https://zenodo.org/record/262458/files/figure.png" pageId="67" pageNumber="68">Fig. 202</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD297AFFB2FF03C6A19D95FC74" box="[151,193,908,933]" italics="true" pageId="67" pageNumber="68">abp</emphasis>
|
||
), and each elytron has one small basal fovea (
|
||
<figureCitation id="13332A3A297AFFB2FD51C6A19E72FC75" box="[709,806,908,933]" captionStart="FIGURES 199 – 203" captionStartId="66.[151,264,1758,1781]" captionTargetBox="[151,1436,193,1728]" captionTargetId="figure@66.[151,1436,193,1738]" captionTargetPageId="66" captionText="FIGURES 199 – 203. Microscydmus sp. 1 (UA- 2216). Dorsal (199, 202), right lateral (200, 203) and left lateral (201) views. Abbreviations: abg, antebasal groove; abp, antebasal pit; bef, basal elytral fovea; fr, frons; hyr, hypomeral ridge; mcp, mesocoxal projection; nr, ' neck' region; vt, vertex." httpUri="https://zenodo.org/record/262458/files/figure.png" pageId="67" pageNumber="68">Fig. 202</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD297AFFB2FCA5C6A19E07FC74" box="[817,851,908,933]" italics="true" pageId="67" pageNumber="68">bef</emphasis>
|
||
) located in a strikingly large and deep, nearly round basal impression. Especially the latter character is a hallmark of most species of
|
||
<taxonomicName id="4C084D3C297AFFB2FBB8C682999AFC19" box="[1068,1230,943,968]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD297AFFB2FBB8C682999AFC19" box="[1068,1230,943,968]" italics="true" pageId="67" pageNumber="68">Microscydmus</emphasis>
|
||
</taxonomicName>
|
||
. This specimen is consequently assigned to
|
||
<taxonomicName id="4C084D3C297AFFB2FE22C6F99F0CFC3C" box="[438,600,980,1005]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD297AFFB2FE22C6F99F0CFC3C" box="[438,600,980,1005]" italics="true" pageId="67" pageNumber="68">Microscydmus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB736BF297AFFB2FF53C6D59C34FA85" blockId="67.[151,1437,151,1365]" pageId="67" pageNumber="68">
|
||
Even in extant species generic diagnostic characters are poorly visible and the scanning electron microscopy or examination of specimens in transparent mounts are unavoidable to confirm their generic placement. Among about 170 currently known species of
|
||
<taxonomicName id="4C084D3C297AFFB2FE69C1129FCBFB89" box="[509,671,1087,1112]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD297AFFB2FE69C1129FCBFB89" box="[509,671,1087,1112]" italics="true" pageId="67" pageNumber="68">Microscydmus</emphasis>
|
||
</taxonomicName>
|
||
, many have been misplaced, diagnoses having been based solely on the tiny body and general appearance (Jałoszyński, unpublished obs.). Besides the monstrous genus
|
||
<taxonomicName id="4C084D3C297AFFB2FAB2C14898C0FBAD" box="[1318,1428,1125,1148]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD297AFFB2FAB2C14898C0FBAD" box="[1318,1428,1125,1148]" italics="true" pageId="67" pageNumber="68">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C084D3C297AFFB2FF03C1AA9C6DFB71" box="[151,313,1159,1184]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD297AFFB2FF03C1AA9C6DFB71" box="[151,313,1159,1184]" italics="true" pageId="67" pageNumber="68">Microscydmus</emphasis>
|
||
</taxonomicName>
|
||
is the most challenging taxon among all
|
||
<taxonomicName id="4C084D3C297AFFB2FCDBC1AA9EB3FB71" box="[847,999,1159,1184]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="tribe" tribe="Glandulariini">Glandulariini</taxonomicName>
|
||
, and a complex and heterogeneous conglomerate of several distinct genera hides behind this name. Aedeagi, extremely small, thin-walled and prone to distortion during preparation, additionally complicate the situation. External features can be used to define new species only in exceptional cases, e.g., when conspicuous modifications of exoskeletal structures have been found (see e.g.,
|
||
<bibRefCitation id="EF994B4E297AFFB2FE86C03A9F53FAE1" author="Jaloszynski" box="[274,519,1303,1328]" pageId="67" pageNumber="68" refString="Jaloszynski, P. (2009 c) First record of Microscydmus Saulcy et Croissandeau from China, with descriptions of two new species (Coleoptera, Scydmaenidae). Zootaxa, 2078, 63 - 68." type="journal article" year="2009" yearSuffix="c">Jałoszyński (2009c))</bibRefCitation>
|
||
. The fossil described above is rather unremarkable and it must remain as
|
||
<taxonomicName id="4C084D3C297AFFB2FF03C0119C6DFA84" box="[151,313,1340,1365]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD297AFFB2FF03C0119C6DFA84" box="[151,313,1340,1365]" italics="true" pageId="67" pageNumber="68">Microscydmus</emphasis>
|
||
</taxonomicName>
|
||
sp.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |