204 lines
40 KiB
XML
204 lines
40 KiB
XML
<document id="DDB2F82A0A30649C0720CDDF91AFC924" ID="10.11646/zootaxa.4157.1.1" ID-DOI="10.11646/zootaxa.4157.1.1" ID-GBIF-Dataset="2389f078-1812-4b4e-826a-561cd7e46654" ID-ISSN="1175-5326" ID-Zenodo-Dep="262413" ID-ZooBank="6BF4514A-892F-499F-BC1E-B7920C7A00B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1472198526050" checkinUser="plazi" docAuthor="Jałoszyński, Paweł & Perkovsky, Evgeny" docDate="2016" docId="03A187A92902FFCAFF03C6E49EA2F825" docLanguage="en" docName="zootaxa.4157.1.1.pdf" docOrigin="Zootaxa 4157 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Euconnus" docType="treatment" docVersion="9" lastPageNumber="60" masterDocId="FF98FFD12939FFF1FF94C52D9D54FFD1" masterDocTitle="Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber" masterLastPageNumber="85" masterPageNumber="1" pageNumber="60" updateTime="1698382676775" updateUser="plazi">
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<mods:title id="10C625972E96DE3FB57E180036DFB147">Diversity of Scydmaeninae (Coleoptera: Staphylinidae) in Upper Eocene Rovno amber</mods:title>
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<mods:namePart id="565D14F172271B35683610B7942356A0">Jałoszyński, Paweł</mods:namePart>
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<mods:namePart id="DA4DE83FC248C3EE4FB586F9823EFB43">Perkovsky, Evgeny</mods:namePart>
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<mods:typeOfResource id="9A19A64951252306931306548A1192DE">text</mods:typeOfResource>
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<mods:title id="BE4311F8B0F7C6ABDDE1034705E7B7C4">Zootaxa</mods:title>
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<mods:part id="60B7B9B275DC3B9A2195453D9C84DD58">
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<mods:date id="0763029ECBAEDB7E438660292D6BBBE7">2016</mods:date>
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<mods:number id="7AFFBEC7A3F3062C9C58DFE79988E5DE">4157</mods:number>
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<mods:identifier id="497ED11C20DAA132C83C51E657AED064" type="DOI">10.11646/zootaxa.4157.1.1</mods:identifier>
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<treatment id="03A187A92902FFCAFF03C6E49EA2F825" ID-DOI="http://doi.org/10.5281/zenodo.5681822" ID-GBIF-Taxon="124407866" ID-Zenodo-Dep="5681822" LSID="urn:lsid:plazi:treatment:03A187A92902FFCAFF03C6E49EA2F825" httpUri="http://treatment.plazi.org/id/03A187A92902FFCAFF03C6E49EA2F825" lastPageNumber="60" pageId="59" pageNumber="60">
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<subSubSection id="C31265342902FFCAFF03C6E49C39FBD5" pageId="59" pageNumber="60" type="nomenclature">
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<paragraph id="8BB736BF2902FFCAFF03C6E49F45FC32" blockId="59.[151,529,969,1029]" box="[151,529,969,995]" pageId="59" pageNumber="60">
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<heading id="D0FF81D32902FFCAFF03C6E49F45FC32" bold="true" box="[151,529,969,995]" fontSize="11" level="1" pageId="59" pageNumber="60" reason="1">
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<emphasis id="B97CEAAD2902FFCAFF03C6E49F45FC32" bold="true" box="[151,529,969,995]" pageId="59" pageNumber="60">
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<taxonomicName id="4C084D3C2902FFCAFF03C6E49C47FC32" box="[151,275,969,995]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="genus">
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<emphasis id="B97CEAAD2902FFCAFF03C6E49C47FC32" bold="true" box="[151,275,969,995]" italics="true" pageId="59" pageNumber="60">Euconnus</emphasis>
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</taxonomicName>
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(
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<emphasis id="B97CEAAD2902FFCAFEB0C6E49C93FC32" bold="true" box="[292,455,969,995]" italics="true" pageId="59" pageNumber="60">incertae sedis</emphasis>
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) sp. 1
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB736BF2902FFCAFF03C6C19C39FBD5" blockId="59.[151,529,969,1029]" box="[151,365,1004,1029]" pageId="59" pageNumber="60">
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(
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<figureCitation id="13332A3A2902FFCAFF0BC6C19DA1FBD4" box="[159,245,1004,1029]" captionStart="FIGURES 21 – 34" captionStartId="8.[151,264,1436,1459]" captionTargetBox="[163,1421,202,1404]" captionTargetId="figure@8.[151,1436,193,1415]" captionTargetPageId="8" captionText="FIGURES 21 – 34. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 8972, Rovnoscydmus sp. specimen 8 (21); K- 3322, Rovnoscydmus sp. specimen 9 (22); K- 15548,? Rovnoscydmus sp. specimen 10 (23); K- 8971,? Rovnoscydmus sp. specimen 11 (24); K- 6045, Euconnus palaeogenus sp. n. (25); K- 9126, Euconnus sp. 1 (26); K- 24859,? Euconnus sp. 2 (27); K- 24302, Stenichnus proavus sp. n. (28); UA- 2216, Microscydmus sp. 1 (29); K- 6943,? Microscydmus sp. 2 (30); K- 3657 a,? Microscydmus sp. 3 (31); K- 6764,? Scydmoraphes sp. 1 (32); K- 6274, Glandulariini gen. & sp. indet. 1 (33); K- 7089, Glandulariini gen. & sp. indet. 2 (34)." httpUri="https://zenodo.org/record/262415/files/figure.png" pageId="59" pageNumber="60">Figs 26</figureCitation>
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,
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<figureCitation id="13332A3A2902FFCAFE95C6C19C30FBD5" box="[257,356,1004,1028]" captionStart-0="FIGURES 182 – 185" captionStart-1="FIGURES 186 – 188" captionStartId-0="60.[151,264,1745,1768]" captionStartId-1="61.[151,264,1190,1213]" captionTargetBox-0="[151,1436,195,1724]" captionTargetBox-1="[161,1428,201,1163]" captionTargetId-0="figure@60.[151,1436,195,1724]" captionTargetId-1="figure@61.[151,1436,193,1169]" captionTargetPageId-0="60" captionTargetPageId-1="61" captionText-0="FIGURES 182 – 185. Euconnus sp. 1 (K- 9126). Dorsal (182), ventral (183), right lateral (184) and left lateral (185) views." captionText-1="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri-0="https://zenodo.org/record/262452/files/figure.png" httpUri-1="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">182–188</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C31265342902FFCAFF03C1199ECFFBA1" pageId="59" pageNumber="60" type="materials_examined">
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<paragraph id="8BB736BF2902FFCAFF03C1199ECFFBA1" blockId="59.[151,1437,1076,2037]" pageId="59" pageNumber="60">
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<materialsCitation id="3B603CE22902FFCAFF03C1199ECFFBA1" ID-GBIF-Occurrence="1300093492" collectionCode="SIZK" county="Material" location="Europe" municipality="Late Eocene" pageId="59" pageNumber="60" specimenCount="1" stateProvince="Rovno">
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<emphasis id="B97CEAAD2902FFCAFF03C1199E61FB9C" bold="true" box="[151,821,1076,1101]" pageId="59" pageNumber="60">
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<collectingCounty id="62D64E332902FFCAFF03C1199DABFB9C" box="[151,255,1076,1101]" pageId="59" pageNumber="60">Material</collectingCounty>
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studied.
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<collectingMunicipality id="6BD3ACC52902FFCAFEFFC1199CAAFB9C" box="[363,510,1076,1101]" pageId="59" pageNumber="60">Late Eocene</collectingMunicipality>
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of
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<location id="8ED760642902FFCAFDB0C1199F2AFB9C" LSID="urn:lsid:plazi:treatment:03A187A92902FFCAFF03C6E49EA2F825:8ED760642902FFCAFDB0C1199F2AFB9C" box="[548,638,1076,1101]" county="Material" municipality="Late Eocene" name="Europe" pageId="59" pageNumber="60" stateProvince="Rovno">Europe</location>
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,
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<collectingRegion id="49CCF85D2902FFCAFD1EC1199F8CFB9C" box="[650,728,1076,1101]" country="Ukraine" name="Rivne" pageId="59" pageNumber="60">Rovno</collectingRegion>
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amber:
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</emphasis>
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sex unknown; inclusion in elongate, rectangular prism of amber
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<quantity id="4CF09B5A2902FFCAFE96C17A9C31FBBE" box="[258,357,1111,1136]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="59" pageNumber="60" unit="mm" value="11.5">11.5 mm</quantity>
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long (
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<figureCitation id="13332A3A2902FFCAFE39C17A9F54FBA1" box="[429,512,1111,1136]" captionStart="FIGURES 21 – 34" captionStartId="8.[151,264,1436,1459]" captionTargetBox="[163,1421,202,1404]" captionTargetId="figure@8.[151,1436,193,1415]" captionTargetPageId="8" captionText="FIGURES 21 – 34. Specimens of Rovno amber with inclusions of Scydmaeninae. K- 8972, Rovnoscydmus sp. specimen 8 (21); K- 3322, Rovnoscydmus sp. specimen 9 (22); K- 15548,? Rovnoscydmus sp. specimen 10 (23); K- 8971,? Rovnoscydmus sp. specimen 11 (24); K- 6045, Euconnus palaeogenus sp. n. (25); K- 9126, Euconnus sp. 1 (26); K- 24859,? Euconnus sp. 2 (27); K- 24302, Stenichnus proavus sp. n. (28); UA- 2216, Microscydmus sp. 1 (29); K- 6943,? Microscydmus sp. 2 (30); K- 3657 a,? Microscydmus sp. 3 (31); K- 6764,? Scydmoraphes sp. 1 (32); K- 6274, Glandulariini gen. & sp. indet. 1 (33); K- 7089, Glandulariini gen. & sp. indet. 2 (34)." httpUri="https://zenodo.org/record/262415/files/figure.png" pageId="59" pageNumber="60">Fig. 26</figureCitation>
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), collection number K-9126 (
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||
<collectionCode id="ED19AE7A2902FFCAFCDFC1759EDBFBA1" box="[843,911,1112,1136]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="59" pageNumber="60">SIZK</collectionCode>
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).
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||
</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31265342902FFCAFF53C1519ECFF961" pageId="59" pageNumber="60" type="description">
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<paragraph id="8BB736BF2902FFCAFF53C15198D3FB45" blockId="59.[151,1437,1076,2037]" box="[199,1415,1148,1173]" pageId="59" pageNumber="60">
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<emphasis id="B97CEAAD2902FFCAFF53C1519C0DFB44" bold="true" box="[199,345,1148,1173]" pageId="59" pageNumber="60">Description.</emphasis>
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Body (
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<figureCitation id="13332A3A2902FFCAFE3EC1519F12FB45" box="[426,582,1148,1173]" captionStart-0="FIGURES 182 – 185" captionStart-1="FIGURES 186 – 188" captionStartId-0="60.[151,264,1745,1768]" captionStartId-1="61.[151,264,1190,1213]" captionTargetBox-0="[151,1436,195,1724]" captionTargetBox-1="[161,1428,201,1163]" captionTargetId-0="figure@60.[151,1436,195,1724]" captionTargetId-1="figure@61.[151,1436,193,1169]" captionTargetPageId-0="60" captionTargetPageId-1="61" captionText-0="FIGURES 182 – 185. Euconnus sp. 1 (K- 9126). Dorsal (182), ventral (183), right lateral (184) and left lateral (185) views." captionText-1="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri-0="https://zenodo.org/record/262452/files/figure.png" httpUri-1="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">Figs 182–188</figureCitation>
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) elongate, relatively stout and strongly convex, dark brown; BL
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<quantity id="4CF09B5A2902FFCAFA8FC15198D7FB45" box="[1307,1411,1148,1173]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="59" pageNumber="60" unit="mm" value="1.5">1.50 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BB736BF2902FFCAFF53C1B299CDFA96" blockId="59.[151,1437,1076,2037]" pageId="59" pageNumber="60">
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Head (
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<figureCitation id="13332A3A2902FFCAFE80C1B29CE0FB69" box="[276,436,1183,1208]" captionStart-0="FIGURES 182 – 185" captionStart-1="FIGURES 186 – 188" captionStartId-0="60.[151,264,1745,1768]" captionStartId-1="61.[151,264,1190,1213]" captionTargetBox-0="[151,1436,195,1724]" captionTargetBox-1="[161,1428,201,1163]" captionTargetId-0="figure@60.[151,1436,195,1724]" captionTargetId-1="figure@61.[151,1436,193,1169]" captionTargetPageId-0="60" captionTargetPageId-1="61" captionText-0="FIGURES 182 – 185. Euconnus sp. 1 (K- 9126). Dorsal (182), ventral (183), right lateral (184) and left lateral (185) views." captionText-1="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri-0="https://zenodo.org/record/262452/files/figure.png" httpUri-1="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">Figs 182–188</figureCitation>
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) broadest at eyes, HL
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||
<quantity id="4CF09B5A2902FFCAFD56C18D9E78FB66" box="[706,812,1184,1208]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="59" pageNumber="60" unit="mm" value="0.33">0.33 mm</quantity>
|
||
, HW
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||
<quantity id="4CF09B5A2902FFCAFCE0C1B29E8BFB66" box="[884,991,1183,1208]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="59" pageNumber="60" unit="mm" value="0.35">0.35 mm</quantity>
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; shape of vertex and frons difficult to assess; tempora seem shorter than eyes. Punctures on frons and vertex (
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<figureCitation id="13332A3A2902FFCAFC23C1E99900FB0C" box="[951,1108,1220,1245]" captionStart="FIGURES 186 – 188" captionStartId="61.[151,264,1190,1213]" captionTargetBox="[161,1428,201,1163]" captionTargetId="figure@61.[151,1436,193,1169]" captionTargetPageId="61" captionText="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">Figs 186–187</figureCitation>
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||
) fine, separated by spaces 3– 4 × as wide as diameters of punctures; setae (
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<figureCitation id="13332A3A2902FFCAFD19C1CA9E7DFAD1" box="[653,809,1255,1280]" captionStart="FIGURES 186 – 188" captionStartId="61.[151,264,1190,1213]" captionTargetBox="[161,1428,201,1163]" captionTargetId="figure@61.[151,1436,193,1169]" captionTargetPageId="61" captionText="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">Figs 186–187</figureCitation>
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) sparse, short, suberect. Antennae (
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<figureCitation id="13332A3A2902FFCAFB20C1CA981BFAD1" box="[1204,1359,1255,1280]" captionStart-0="FIGURES 182 – 185" captionStart-1="FIGURES 186 – 188" captionStartId-0="60.[151,264,1745,1768]" captionStartId-1="61.[151,264,1190,1213]" captionTargetBox-0="[151,1436,195,1724]" captionTargetBox-1="[161,1428,201,1163]" captionTargetId-0="figure@60.[151,1436,195,1724]" captionTargetId-1="figure@61.[151,1436,193,1169]" captionTargetPageId-0="60" captionTargetPageId-1="61" captionText-0="FIGURES 182 – 185. Euconnus sp. 1 (K- 9126). Dorsal (182), ventral (183), right lateral (184) and left lateral (185) views." captionText-1="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri-0="https://zenodo.org/record/262452/files/figure.png" httpUri-1="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">Figs 182–188</figureCitation>
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||
) much shorter than half BL, AnL
|
||
<quantity id="4CF09B5A2902FFCAFE50C0219F7EFAF5" box="[452,554,1292,1317]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="59" pageNumber="60" unit="mm" value="0.55">0.55 mm</quantity>
|
||
, shapes of antennomeres difficult to assess, but antennae seem to be gradually thickened distally and compactly assembled, with at least some of flagellomeres transverse.
|
||
</paragraph>
|
||
<paragraph id="8BB736BF2902FFCAFF53C0789FF8F9F1" blockId="59.[151,1437,1076,2037]" pageId="59" pageNumber="60">
|
||
Pronotum (
|
||
<figureCitation id="13332A3A2902FFCAFED0C0799CFCFABD" box="[324,424,1364,1389]" captionStart="FIGURES 182 – 185" captionStartId="60.[151,264,1745,1768]" captionTargetBox="[151,1436,195,1724]" captionTargetId="figure@60.[151,1436,195,1724]" captionTargetPageId="60" captionText="FIGURES 182 – 185. Euconnus sp. 1 (K- 9126). Dorsal (182), ventral (183), right lateral (184) and left lateral (185) views." httpUri="https://zenodo.org/record/262452/files/figure.png" pageId="59" pageNumber="60">Figs 182</figureCitation>
|
||
,
|
||
<figureCitation id="13332A3A2902FFCAFE20C0799C8BFABC" box="[436,479,1364,1389]" captionStart="FIGURES 186 – 188" captionStartId="61.[151,264,1190,1213]" captionTargetBox="[161,1428,201,1163]" captionTargetId="figure@61.[151,1436,193,1169]" captionTargetPageId="61" captionText="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">186</figureCitation>
|
||
) slightly broader than long, broadest site difficult to define, but most likely in front of middle; PL
|
||
<quantity id="4CF09B5A2902FFCAFEB1C05A9CC4FA5E" box="[293,400,1399,1424]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="59" pageNumber="60" unit="mm" value="0.35">0.35 mm</quantity>
|
||
, PW
|
||
<quantity id="4CF09B5A2902FFCAFE42C0559F16FA5E" box="[470,578,1400,1424]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="59" pageNumber="60" unit="mm" value="0.38">0.38 mm</quantity>
|
||
; anterior margin arcuate; posterior margin shallowly bisinuate, shapes of pronotal corners and lateral margins difficult to assess; base of pronotum with short but distinct sublateral carinae (
|
||
<figureCitation id="13332A3A2902FFCAFF0BC0929C54FA09" box="[159,256,1471,1496]" captionStart="FIGURES 186 – 188" captionStartId="61.[151,264,1190,1213]" captionTargetBox="[161,1428,201,1163]" captionTargetId="figure@61.[151,1436,193,1169]" captionTargetPageId="61" captionText="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">Fig. 186</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2902FFCAFE98C0929C7FFA09" box="[268,299,1471,1496]" italics="true" pageId="59" pageNumber="60">slc</emphasis>
|
||
) and one (inner) pair of antebasal pits (
|
||
<figureCitation id="13332A3A2902FFCAFD76C0929E17FA09" box="[738,835,1471,1496]" captionStart="FIGURES 186 – 188" captionStartId="61.[151,264,1190,1213]" captionTargetBox="[161,1428,201,1163]" captionTargetId="figure@61.[151,1436,193,1169]" captionTargetPageId="61" captionText="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">Fig. 186</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2902FFCAFCDAC0929E2CFA09" box="[846,888,1471,1496]" italics="true" pageId="59" pageNumber="60">abp</emphasis>
|
||
). Punctures on pronotal disc (
|
||
<figureCitation id="13332A3A2902FFCAFB53C0929872FA09" box="[1223,1318,1471,1496]" captionStart="FIGURES 186 – 188" captionStartId="61.[151,264,1190,1213]" captionTargetBox="[161,1428,201,1163]" captionTargetId="figure@61.[151,1436,193,1169]" captionTargetPageId="61" captionText="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">Fig. 186</figureCitation>
|
||
) small but distinct, separated by spaces 3–4 × as wide as diameters of punctures; setae (
|
||
<figureCitation id="13332A3A2902FFCAFC64C0C9991BFA2C" box="[1008,1103,1508,1533]" captionStart="FIGURES 186 – 188" captionStartId="61.[151,264,1190,1213]" captionTargetBox="[161,1428,201,1163]" captionTargetId="figure@61.[151,1436,193,1169]" captionTargetPageId="61" captionText="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">Fig. 186</figureCitation>
|
||
) short, sparse, suberect; sides of pronotum with thick bristles (
|
||
<figureCitation id="13332A3A2902FFCAFD95C32A9FF4F9F1" box="[513,672,1543,1568]" captionStart="FIGURES 186 – 188" captionStartId="61.[151,264,1190,1213]" captionTargetBox="[161,1428,201,1163]" captionTargetId="figure@61.[151,1436,193,1169]" captionTargetPageId="61" captionText="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">Figs 186–188</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB736BF2902FFCAFF53C3019CB4F95C" blockId="59.[151,1437,1076,2037]" pageId="59" pageNumber="60">
|
||
Elytra (
|
||
<figureCitation id="13332A3A2902FFCAFE8EC3019F71F995" box="[282,549,1580,1605]" captionStart-0="FIGURES 182 – 185" captionStart-1="FIGURES 186 – 188" captionStartId-0="60.[151,264,1745,1768]" captionStartId-1="61.[151,264,1190,1213]" captionTargetBox-0="[151,1436,195,1724]" captionTargetBox-1="[161,1428,201,1163]" captionTargetId-0="figure@60.[151,1436,195,1724]" captionTargetId-1="figure@61.[151,1436,193,1169]" captionTargetPageId-0="60" captionTargetPageId-1="61" captionText-0="FIGURES 182 – 185. Euconnus sp. 1 (K- 9126). Dorsal (182), ventral (183), right lateral (184) and left lateral (185) views." captionText-1="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri-0="https://zenodo.org/record/262452/files/figure.png" httpUri-1="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">Figs 182, 184–186, 188</figureCitation>
|
||
) elongate oval, strongly convex, broadest anterior to middle, EL
|
||
<quantity id="4CF09B5A2902FFCAFB6BC3019831F995" box="[1279,1381,1580,1604]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="59" pageNumber="60" unit="mm" value="0.83">0.83 mm</quantity>
|
||
, EW
|
||
<quantity id="4CF09B5A2902FFCAFF03C3629DAAF9B6" box="[151,254,1615,1640]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="59" pageNumber="60" unit="mm" value="0.6">0.60 mm</quantity>
|
||
, EI 1.38; punctures and setae similar to those on pronotum but slightly sparser (
|
||
<figureCitation id="13332A3A2902FFCAFB1CC36299BFF9B9" box="[1160,1259,1615,1640]" captionStart="FIGURES 186 – 188" captionStartId="61.[151,264,1190,1213]" captionTargetBox="[161,1428,201,1163]" captionTargetId="figure@61.[151,1436,193,1169]" captionTargetPageId="61" captionText="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">Fig. 186</figureCitation>
|
||
). Scutellum not visible between elytral bases.
|
||
</paragraph>
|
||
<paragraph id="8BB736BF2902FFCAFF53C3B59ECFF961" blockId="59.[151,1437,1076,2037]" box="[199,923,1687,1712]" pageId="59" pageNumber="60">
|
||
Legs (
|
||
<figureCitation id="13332A3A2902FFCAFE98C3BA9C25F961" box="[268,369,1687,1712]" captionStart="FIGURES 182 – 185" captionStartId="60.[151,264,1745,1768]" captionTargetBox="[151,1436,195,1724]" captionTargetId="figure@60.[151,1436,195,1724]" captionTargetPageId="60" captionText="FIGURES 182 – 185. Euconnus sp. 1 (K- 9126). Dorsal (182), ventral (183), right lateral (184) and left lateral (185) views." httpUri="https://zenodo.org/record/262452/files/figure.png" pageId="59" pageNumber="60">Figs 183</figureCitation>
|
||
,
|
||
<figureCitation id="13332A3A2902FFCAFEE9C3B59CB4F961" box="[381,480,1688,1712]" captionStart="FIGURES 186 – 188" captionStartId="61.[151,264,1190,1213]" captionTargetBox="[161,1428,201,1163]" captionTargetId="figure@61.[151,1436,193,1169]" captionTargetPageId="61" captionText="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">187–188</figureCitation>
|
||
) moderately long, slender, unmodified.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31265342902FFCAFF53C3919EA2F825" pageId="59" pageNumber="60" type="discussion">
|
||
<paragraph id="8BB736BF2902FFCAFF53C3919EA2F825" blockId="59.[151,1437,1076,2037]" pageId="59" pageNumber="60">
|
||
<emphasis id="B97CEAAD2902FFCAFF53C3919C6FF904" bold="true" box="[199,315,1724,1749]" pageId="59" pageNumber="60">Remarks.</emphasis>
|
||
This species can be placed in
|
||
<taxonomicName id="4C084D3C2902FFCAFD3AC3909E48F905" box="[686,796,1725,1748]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2902FFCAFD3AC3909E48F905" box="[686,796,1725,1748]" italics="true" pageId="59" pageNumber="60">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
on the basis of the carinate and elevated mesoventral intercoxal process (
|
||
<figureCitation id="13332A3A2902FFCAFEEEC3F29CB4F929" box="[378,480,1759,1784]" captionStart="FIGURES 186 – 188" captionStartId="61.[151,264,1190,1213]" captionTargetBox="[161,1428,201,1163]" captionTargetId="figure@61.[151,1436,193,1169]" captionTargetPageId="61" captionText="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">Fig. 187</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2902FFCAFE64C3CC9F7EF929" box="[496,554,1761,1784]" italics="true" pageId="59" pageNumber="60">msvp</emphasis>
|
||
) the metaventral intercoxal process (
|
||
<figureCitation id="13332A3A2902FFCAFC4BC3F29911F929" box="[991,1093,1759,1784]" captionStart="FIGURES 186 – 188" captionStartId="61.[151,264,1190,1213]" captionTargetBox="[161,1428,201,1163]" captionTargetId="figure@61.[151,1436,193,1169]" captionTargetPageId="61" captionText="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">Fig. 187</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2902FFCAFBC1C3CC99DFF929" box="[1109,1163,1761,1784]" italics="true" pageId="59" pageNumber="60">mtvp</emphasis>
|
||
) short and very broad, broadly separating metacoxae, the scutellum not exposed between elytral bases (
|
||
<figureCitation id="13332A3A2902FFCAFBB1C22999DCF8CC" box="[1061,1160,1796,1821]" captionStart="FIGURES 186 – 188" captionStartId="61.[151,264,1190,1213]" captionTargetBox="[161,1428,201,1163]" captionTargetId="figure@61.[151,1436,193,1169]" captionTargetPageId="61" captionText="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">Fig. 186</figureCitation>
|
||
), the subtriangular inner (adcoxal) part of prothoracic hypomeron (
|
||
<figureCitation id="13332A3A2902FFCAFDEFC20A9F8BF891" box="[635,735,1831,1856]" captionStart="FIGURES 186 – 188" captionStartId="61.[151,264,1190,1213]" captionTargetBox="[161,1428,201,1163]" captionTargetId="figure@61.[151,1436,193,1169]" captionTargetPageId="61" captionText="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">Fig. 187</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2902FFCAFD7AC20A9E5DF891" box="[750,777,1831,1856]" italics="true" pageId="59" pageNumber="60">hy</emphasis>
|
||
) demarcated laterally by complete hypomeral ridge (
|
||
<figureCitation id="13332A3A2902FFCAFAFBC20A9D90F8B5" captionStart="FIGURES 186 – 188" captionStartId="61.[151,264,1190,1213]" captionTargetBox="[161,1428,201,1163]" captionTargetId="figure@61.[151,1436,193,1169]" captionTargetPageId="61" captionText="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">Fig. 187</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2902FFCAFF47C2619DACF8B4" box="[211,248,1868,1893]" italics="true" pageId="59" pageNumber="60">hyr</emphasis>
|
||
), and two basal elytral foveae (
|
||
<figureCitation id="13332A3A2902FFCAFDFAC2619F87F8B4" box="[622,723,1868,1893]" captionStart="FIGURES 186 – 188" captionStartId="61.[151,264,1190,1213]" captionTargetBox="[161,1428,201,1163]" captionTargetId="figure@61.[151,1436,193,1169]" captionTargetPageId="61" captionText="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">Fig. 186</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2902FFCAFD76C2619E50F8B4" box="[738,772,1868,1893]" italics="true" pageId="59" pageNumber="60">bef</emphasis>
|
||
) on each elytron. Additionally, the lateral bristles on the pronotum and sublateral pronotal carinae (
|
||
<figureCitation id="13332A3A2902FFCAFDFBC2429F84F859" box="[623,720,1903,1928]" captionStart="FIGURES 186 – 188" captionStartId="61.[151,264,1190,1213]" captionTargetBox="[161,1428,201,1163]" captionTargetId="figure@61.[151,1436,193,1169]" captionTargetPageId="61" captionText="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">Fig. 186</figureCitation>
|
||
;
|
||
<emphasis id="B97CEAAD2902FFCAFD4FC2429FAEF859" box="[731,762,1903,1928]" italics="true" pageId="59" pageNumber="60">slc</emphasis>
|
||
) accompanied by small antebasal pits (
|
||
<figureCitation id="13332A3A2902FFCAFB3BC2429844F859" box="[1199,1296,1903,1928]" captionStart="FIGURES 186 – 188" captionStartId="61.[151,264,1190,1213]" captionTargetBox="[161,1428,201,1163]" captionTargetId="figure@61.[151,1436,193,1169]" captionTargetPageId="61" captionText="FIGURES 186 – 188. Euconnus sp. 1 (K- 9126). Nearly dorsal (186), ventral (187) and lateral (188) views. Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; cl, clypeus; fr, frons; hy, hypomeron; hyr, hypomeral ridge; lbr, labrum; lp, labial palp; md, mandible; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; mxp 3 – 4, maxillary palpomere III – IV; pg, pygidium; slc, sublateral carina; st 3 – 8, abdominal sternite III – VIII; v 3, metaventrite." httpUri="https://zenodo.org/record/262453/files/figure.png" pageId="59" pageNumber="60">Fig. 186</figureCitation>
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;
|
||
<emphasis id="B97CEAAD2902FFCAFA8FC2429811F859" box="[1307,1349,1903,1928]" italics="true" pageId="59" pageNumber="60">abp</emphasis>
|
||
) can be found in many extant species of
|
||
<taxonomicName id="4C084D3C2902FFCAFD97C2B89F25F87D" box="[515,625,1941,1964]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97CEAAD2902FFCAFD97C2B89F25F87D" box="[515,625,1941,1964]" italics="true" pageId="59" pageNumber="60">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
, but these characters are not restricted to this genus. This fossil does not show any conspicuous characters that could be used to reliably define a new species; also it was not possible to determine whether this species belongs to any of the extant subgenera or not.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |