153 lines
22 KiB
XML
153 lines
22 KiB
XML
<document id="84491FC228AB41DF1924638D9419140A" ID-DOI="10.11646/zootaxa.3704.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="6507447" ID-ZooBank="341ED5F6-2DCA-4E73-83D6-389DF0DD347F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629898207969" checkinUser="felipe" docAuthor="KITS, JOEL H. & MARSHALL, STEPHEN A." docDate="2013" docId="03A18780C614FFDF0DD3D98AFD4CF90C" docLanguage="en" docName="zt03704p113.pdf" docOrigin="Zootaxa 3704 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3704.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Photoantrops echinus KITS & MARSHALL 2013, sp. nov." docType="treatment" docVersion="4" lastPageNumber="93" masterDocId="FF98FFF8C648FF830D44D91DFF80FFB2" masterDocTitle="<p class = " HeadingRunIn " align = " left "> <strong> Generic classification of the Archiborborinae (Diptera: Sphaeroceridae), with a revision of <em> Antrops </ em> Enderlein, <em> Coloantrops </ em> gen. nov., <em> Maculantrops </ em> gen. nov., <em> Photoantrops </ em> gen. nov., and <em> Poecilantrops </ em> gen. nov. </ strong> </ p>" masterLastPageNumber="113" masterPageNumber="1" pageNumber="93" updateTime="1699126277967" updateUser="plazi">
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<mods:title id="352B2F261F0BACED4601FB9A1CBC6F3C"><p class = " HeadingRunIn " align = " left "> <strong> Generic classification of the Archiborborinae (Diptera: Sphaeroceridae), with a revision of <em> Antrops </ em> Enderlein, <em> Coloantrops </ em> gen. nov., <em> Maculantrops </ em> gen. nov., <em> Photoantrops </ em> gen. nov., and <em> Poecilantrops </ em> gen. nov. </ strong> </ p></mods:title>
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<mods:namePart id="678829BF421F56F95F3CC9D547B1C0E3">KITS, JOEL H.</mods:namePart>
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<mods:namePart id="2F5EA6D2B7B0D7778676AD4E28F5EEE8">MARSHALL, STEPHEN A.</mods:namePart>
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<mods:title id="920F4440C39C938B1E930EEE69FC0580">Zootaxa</mods:title>
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<mods:part id="4703B3AB7D0363EB5E7A0277C472B53C">
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<mods:date id="063AF1305C5BD81C5EAF3B4179FC8905">2013</mods:date>
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<mods:number id="01FF87D207EB92F32F469C8AE53D4331">2013-09-02</mods:number>
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<mods:number id="368849900D7BA2A4DBF879B759908B9F">3704</mods:number>
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<mods:url id="15287D23A447CE8B487C8D9ECA2D33DD">http://dx.doi.org/10.11646/zootaxa.3704.1.1</mods:url>
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<mods:classification id="12084894B9758A243E0A441764B802F7">journal article</mods:classification>
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<mods:identifier id="879D32B0C2298A094C8789FBCBC78C38" type="DOI">10.11646/zootaxa.3704.1.1</mods:identifier>
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<treatment id="03A18780C614FFDF0DD3D98AFD4CF90C" ID-DOI="http://doi.org/10.5281/zenodo.6509242" ID-Zenodo-Dep="6509242" LSID="urn:lsid:plazi:treatment:03A18780C614FFDF0DD3D98AFD4CF90C" httpUri="http://treatment.plazi.org/id/03A18780C614FFDF0DD3D98AFD4CF90C" lastPageNumber="93" pageId="92" pageNumber="93">
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<subSubSection id="C312651DC614FFDF0DD3D98AFD84FF00" box="[151,516,151,178]" pageId="92" pageNumber="93" type="nomenclature">
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<paragraph id="8BB73696C614FFDF0DD3D98AFD84FF00" blockId="92.[151,516,151,211]" box="[151,516,151,178]" pageId="92" pageNumber="93">
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<heading id="D0FF81FAC614FFDF0DD3D98AFD84FF00" bold="true" box="[151,516,151,178]" fontSize="11" level="2" pageId="92" pageNumber="93" reason="4">
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<emphasis id="B97CEA84C614FFDF0DD3D98AFD84FF00" bold="true" box="[151,516,151,178]" pageId="92" pageNumber="93">
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<taxonomicName id="4C084D15C614FFDF0DD3D98AFE1FFF03" ID-CoL="8DBXT" authority="KITS & MARSHALL, 2013" authorityName="KITS & MARSHALL" authorityYear="2013" box="[151,415,151,177]" class="Insecta" family="Sphaeroceridae" genus="Photoantrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="echinus" status="sp. nov.">
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<emphasis id="B97CEA84C614FFDF0DD3D98AFE1FFF03" bold="true" box="[151,415,151,177]" italics="true" pageId="92" pageNumber="93">Photoantrops echinus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24F57FFC614FFDF0CE3D985FD84FF00" box="[423,516,152,178]" pageId="92" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C312651DC614FFDF0DD3D9A7FDFDF9B8" pageId="92" pageNumber="93" type="description">
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<paragraph id="8BB73696C614FFDF0DD3D9A7FE2CFF61" blockId="92.[151,516,151,211]" box="[151,428,186,211]" pageId="92" pageNumber="93">
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(
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<figureCitation id="13332A13C614FFDF0DDBD9A7FF7CFF61" box="[159,252,186,211]" captionStart="FIGURES 7–12" captionStartId="10.[151,264,1695,1718]" captionTargetBox="[151,1436,226,1646]" captionTargetId="figure-79@10.[151,1436,193,1646]" captionTargetPageId="10" captionText="FIGURES 7–12. Archiborborine adults, habitus photos: (7) Antrops truncipennis, female, Bird Island, South Georgia Islands; (8) A. siberia, male holotype, Yungas de la Siberia, Bolivia; (9) A. tetrastichus, male paratype, Quito–Baeza pass; (10) Photoantrops echinus, male holotype, SierrAzul Lodge, Ecuador; (11) Coloantrops daedalus, female paratype, Laraquete, Chile; (12) Poecilantrops psilosternum, female paratype, Nova Friburgo, Brazil." figureDoi="http://doi.org/10.5281/zenodo.6507451" httpUri="https://zenodo.org/record/6507451/files/figure.png" pageId="92" pageNumber="93">Figs. 10</figureCitation>
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,
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<figureCitation id="13332A13C614FFDF0C4CD9A7FEECFF61" box="[264,364,186,211]" captionStart="FIGURES 173–180" captionStartId="93.[151,264,1862,1885]" captionTargetBox="[158,1432,213,1829]" captionTargetId="figure-87@93.[151,1436,195,1841]" captionTargetPageId="93" captionText="FIGURES 173–180. Photoantrops echinus male terminalia. (173) phallus, postgonite, and ejaculatory apodeme, lateral view, (174) epandrium, cerci, and surstyli, posterior view, (175) sternite 5. Poecilantrops male terminalia. (176) P. baorucensis, phallus, postgonite, and phallapodeme, lateral view; (177) P. boraceiensis, phallus, postgonite, and phallapodeme, lateral view; (178) P. crocidosternum, phallus, postgonite, and phallapodeme, lateral view; (179) P. crocidosternum, sternite 5; (180) P. boraceiensis, sternite 5." figureDoi="http://doi.org/10.5281/zenodo.6507505" httpUri="https://zenodo.org/record/6507505/files/figure.png" pageId="92" pageNumber="93">173–175</figureCitation>
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,
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<figureCitation id="13332A13C614FFDF0C3CD9A7FE23FF61" box="[376,419,186,211]" captionStart="FIGURES 231–239" captionStartId="109.[151,264,1710,1733]" captionTargetBox="[151,1436,193,1690]" captionTargetId="figure-68@109.[150,1436,193,1690]" captionTargetPageId="109" captionText="FIGURES 231–239. Archiborborine distribution maps. (231) Maculantrops hirtipes; (232) Photoantrops echinus; (233) Poecilantrops baorucensis (stars), P. dominicus (circles); (234) P. stellans; (235) P. boraceiensis (circles), P. plaumanni (stars); (236) P. crocidosternum; (237) P. psilosternum; (238) P. flavifemur; (239) P.marensis (circles), P. vittifrons (stars)." figureDoi="http://doi.org/10.5281/zenodo.6507519" httpUri="https://zenodo.org/record/6507519/files/figure.png" pageId="92" pageNumber="93">232</figureCitation>
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)
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</paragraph>
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<paragraph id="8BB73696C614FFDF0DD3D81FFB03FE35" blockId="92.[151,1437,258,1726]" pageId="92" pageNumber="93">
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<emphasis id="B97CEA84C614FFDF0DD3D81FFEA9FEA9" bold="true" box="[151,297,258,283]" pageId="92" pageNumber="93">Description.</emphasis>
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Head yellow, interfrontal plates and a small spot at ventral margin of eye brown. Mostly covered with microtomentum, frons with thin shiny lines lateral to ocelli. Ocellar bristles absent, only 1 orbital bristle. Subvibrissal bristle about 0.5X length of vibrissa, anterior genal bristle about 0.3X length of vibrissa. Gena about 0.3X height of eye in male, about 0.3x height of eye in female. Occiput strongly rounded.
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</paragraph>
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<paragraph id="8BB73696C614FFDF0D83D88FFB53FDA5" blockId="92.[151,1437,258,1726]" pageId="92" pageNumber="93">Thorax mostly brown, yellow on postpronotum, in a spot anterior to presutural dorsocentral, proepisternum, in a median spot on anepisternum, and ventral half of anepimeron, black on scutal suture. Mostly shiny; scutum, ventral margin of katepisternum, metapleuron, and laterotergite with microtomentum. Bristles greatly enlarged, other setae on scutum greatly reduced, barely visible in male. Halter whitish, brown below knob.</paragraph>
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<paragraph id="8BB73696C614FFDF0D83DB3EFE01FD15" blockId="92.[151,1437,258,1726]" pageId="92" pageNumber="93">Legs mostly yellow, base of fore femur and all of mid and hind coxae and femora dark brown. Hind femur greatly swollen in male, about 7X width of tibia. Mid tibia with row of anterodorsal, row of posterodorsal, 1 anteroventral, and 4 subapical bristles. Hind tibia with row of anterodorsal, row of posterodorsal, and 2 ventroapical bristles.</paragraph>
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<paragraph id="8BB73696C614FFDF0D83DBAFFCE3FD78" blockId="92.[151,1437,258,1726]" box="[199,867,690,719]" pageId="92" pageNumber="93">
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Wing yellowish, with a small dark smudge at tip of vein R
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<subScript id="178C34D3C614FFDF0E10DBDCFCDCFD7D" attach="left" box="[852,860,705,719]" fontSize="6" pageId="92" pageNumber="93">2</subScript>
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.
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</paragraph>
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<paragraph id="8BB73696C614FFDF0D83DBC4FEF1FCEC" blockId="92.[151,1437,258,1726]" pageId="92" pageNumber="93">Abdomen with tergites brown, shiny. Syntergite 1+2 twice the width of tergite 3–5. Tergites 4 and 5 narrow anteriorly, with a shallow notch posteriorly. Setae along posterior margins of tergites and some pleural setae enlarged, longer in male than female. Sternites 2–3 narrow, weakly sclerotized, sternite 4 wider, lateral edges heavily sclerotized.</paragraph>
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<paragraph id="8BB73696C614FFDF0D83DA74FE8EFABC" blockId="92.[151,1437,258,1726]" pageId="92" pageNumber="93">
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Male postabdomen: Sternite 5 with curved lateral margins, narrow anteriorly, without anterior apodeme (
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<figureCitation id="13332A13C614FFDF082ADA74FF45FC14" captionStart="FIGURES 173–180" captionStartId="93.[151,264,1862,1885]" captionTargetBox="[158,1432,213,1829]" captionTargetId="figure-87@93.[151,1436,195,1841]" captionTargetPageId="93" captionText="FIGURES 173–180. Photoantrops echinus male terminalia. (173) phallus, postgonite, and ejaculatory apodeme, lateral view, (174) epandrium, cerci, and surstyli, posterior view, (175) sternite 5. Poecilantrops male terminalia. (176) P. baorucensis, phallus, postgonite, and phallapodeme, lateral view; (177) P. boraceiensis, phallus, postgonite, and phallapodeme, lateral view; (178) P. crocidosternum, phallus, postgonite, and phallapodeme, lateral view; (179) P. crocidosternum, sternite 5; (180) P. boraceiensis, sternite 5." figureDoi="http://doi.org/10.5281/zenodo.6507505" httpUri="https://zenodo.org/record/6507505/files/figure.png" pageId="92" pageNumber="93">Fig. 175</figureCitation>
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). Synsternite 6+7 asymmetrical, narrow, one section detached and forming a fan-shaped accessory sclerite flanking distiphallus in genital pouch; dorsal corner fused to sternite 8. Sternite 8 fused to epandrium along right side. Epandrium more or less symmetrical, with a cleft above anterior edge of surstylus, anterior setae greatly enlarged (
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<figureCitation id="13332A13C614FFDF0C4DDAE4FEEEFBA0" box="[265,366,1017,1042]" captionStart="FIGURES 173–180" captionStartId="93.[151,264,1862,1885]" captionTargetBox="[158,1432,213,1829]" captionTargetId="figure-87@93.[151,1436,195,1841]" captionTargetPageId="93" captionText="FIGURES 173–180. Photoantrops echinus male terminalia. (173) phallus, postgonite, and ejaculatory apodeme, lateral view, (174) epandrium, cerci, and surstyli, posterior view, (175) sternite 5. Poecilantrops male terminalia. (176) P. baorucensis, phallus, postgonite, and phallapodeme, lateral view; (177) P. boraceiensis, phallus, postgonite, and phallapodeme, lateral view; (178) P. crocidosternum, phallus, postgonite, and phallapodeme, lateral view; (179) P. crocidosternum, sternite 5; (180) P. boraceiensis, sternite 5." figureDoi="http://doi.org/10.5281/zenodo.6507505" httpUri="https://zenodo.org/record/6507505/files/figure.png" pageId="92" pageNumber="93">Fig. 174</figureCitation>
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). Cerci small and medially fused. Subepandrial sclerite Y -shaped, articulated with cerci and surstyli. Surstylus broad, subrectangular, articulated with epandrium and subepandrial sclerite. Hypandrial arms weakly fused with ventral edge of epandrium; hypandrial apodeme well-developed, rod-like, weakly fused with arms. Phallapodeme well-developed, rod-like. Pregonite small, fully fused with postgonite. Postgonite long, bilobed apically, anterior lobe rounded, posterior lobe narrow, curved, pointed, articulated with phallapodeme, hypandrium, and basiphallus. Ejaculatory apodeme large, fan-shaped apically. Basiphallus long, with a long thin epiphallus with a spur near base, no distinct preepiphallus. Distiphallus complex, without a distinct dorsal tube (
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<figureCitation id="13332A13C614FFDF0DDBDDE8FE82FABC" box="[159,258,1269,1294]" captionStart="FIGURES 173–180" captionStartId="93.[151,264,1862,1885]" captionTargetBox="[158,1432,213,1829]" captionTargetId="figure-87@93.[151,1436,195,1841]" captionTargetPageId="93" captionText="FIGURES 173–180. Photoantrops echinus male terminalia. (173) phallus, postgonite, and ejaculatory apodeme, lateral view, (174) epandrium, cerci, and surstyli, posterior view, (175) sternite 5. Poecilantrops male terminalia. (176) P. baorucensis, phallus, postgonite, and phallapodeme, lateral view; (177) P. boraceiensis, phallus, postgonite, and phallapodeme, lateral view; (178) P. crocidosternum, phallus, postgonite, and phallapodeme, lateral view; (179) P. crocidosternum, sternite 5; (180) P. boraceiensis, sternite 5." figureDoi="http://doi.org/10.5281/zenodo.6507505" httpUri="https://zenodo.org/record/6507505/files/figure.png" pageId="92" pageNumber="93">Fig. 173</figureCitation>
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).
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</paragraph>
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<paragraph id="8BB73696C614FFDF0D83DC04FDFDF9B8" blockId="92.[151,1437,258,1726]" pageId="92" pageNumber="93">Female postabdomen: Tergite 5 long and narrow, emarginated anteriorly. Tergites 6–7 and sternites 6–7 narrow, hourglass-shaped, anterior and lateral margins heavily sclerotized, weakly sclerotized medially. Tergites with 3 posterior strips, sternites with 2 posterior strips. Tergite 8 shiny, narrow, emarginated anteriorly, with a rounded lobe projecting laterally. Epiproct shiny, lozenge-shaped, anterior arms narrow, with 2 pairs of setae. Cerci long. Sclerites of sternite 8 shiny, broad, slightly narrower anteriorly, pinched medially. Hypoproct large, square, covered with microtomentum and setae. Spermathecae barrel-shaped, slightly longer than wide, with basal and apical invaginations, sclerotized duct short.</paragraph>
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</subSubSection>
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<subSubSection id="C312651DC614FFDF0D83DF08FE97F9C4" pageId="92" pageNumber="93" type="materials_examined">
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<paragraph id="8BB73696C614FFDF0D83DF08FE97F9C4" blockId="92.[151,1437,258,1726]" pageId="92" pageNumber="93">
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<emphasis id="B97CEA84C614FFDF0D83DF08FEF4F99C" bold="true" box="[199,372,1557,1582]" pageId="92" pageNumber="93">Type material.</emphasis>
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<materialsCitation id="3B603CCBC614FFDF0C3FDF08FD04F9E0" collectingDate="2002-05-08" collectionCode="QCAZ" collectorName="S. M. Paiero" country="Ecuador" elevation="2200" location="SierrAzul Lodge" pageId="92" pageNumber="93" specimenCount="1" specimenCount-male="1" stateProvince="Napo" typeStatus="holotype">
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<typeStatus id="54B38834C614FFDF0C3FDF08FE63F99C" box="[379,483,1557,1582]" pageId="92" pageNumber="93" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D0EFD1FC614FFDF0CADDF0BFE7EF99F" box="[489,510,1558,1581]" pageId="92" pageNumber="93" type="male">♂</specimenCount>
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:
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<collectingCountry id="F31F7606C614FFDF0F49DF08FD19F99C" box="[525,665,1557,1582]" name="Ecuador" pageId="92" pageNumber="93">ECUADOR</collectingCountry>
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:
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<emphasis id="B97CEA84C614FFDF0FE7DF08FD63F99C" bold="true" box="[675,739,1557,1582]" pageId="92" pageNumber="93">
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<collectingRegion id="49CCF874C614FFDF0FE7DF08FD63F99C" box="[675,739,1557,1582]" country="Ecuador" name="Napo" pageId="92" pageNumber="93">Napo</collectingRegion>
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</emphasis>
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:
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<location id="8ED7604DC614FFDF0FB5DF08FC2FF99C" LSID="urn:lsid:plazi:treatment:03A18780C614FFDF0DD3D98AFD4CF90C:8ED7604DC614FFDF0FB5DF08FC2FF99C" box="[753,943,1557,1582]" country="Ecuador" name="SierrAzul Lodge" pageId="92" pageNumber="93" stateProvince="Napo">SierrAzul Lodge</location>
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,
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<locationDeviation id="9A052FB4C614FFDF0EFEDF08FB0BF99C" box="[954,1163,1557,1582]" location="Cosanga" pageId="92" pageNumber="93">
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<quantity id="4CF09B73C614FFDF0EFEDF08FB81F99C" box="[954,1025,1557,1582]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="92" pageNumber="93" unit="km" value="14.0">14 km</quantity>
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W Cosanga
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</locationDeviation>
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,
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<quantity id="4CF09B73C614FFDF09D2DF08FB6DF99F" box="[1174,1261,1557,1582]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="92" pageNumber="93" unit="m" value="2200.0">
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<elevation id="0025D1A5C614FFDF09D2DF08FB6DF99F" box="[1174,1261,1557,1582]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="92" pageNumber="93" unit="m" value="2200.0">2200 m</elevation>
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</quantity>
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, at light,
|
||
<date id="FFB61056C614FFDF0810DF08FF52F9E0" pageId="92" pageNumber="93" value="2002-05-08">
|
||
<collectingDate id="EFF2E9BEC614FFDF0810DF08FF52F9E0" pageId="92" pageNumber="93" value="2002-05-08">8 May 2002</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26FD5340C614FFDF0D9ADF24FEE8F9E0" box="[222,360,1593,1618]" pageId="92" pageNumber="93">S.M. Paiero</collectorName>
|
||
, debu00186468 (
|
||
<collectionCode id="ED19AE53C614FFDF0F6EDF24FDFDF9E0" box="[554,637,1593,1618]" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="92" pageNumber="93" type="Museum">QCAZ</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B603CCBC614FFDF0FD4DF27FE93F9C4" collectingDate="2002-05-08" collectingDateMax="2002-05-10" collectingDateMin="2002-05-08" collectionCode="DEBU" collectorName="O. Lonsdale" country="Ecuador" elevation="2200" location="SierrAzul Lodge" pageId="92" pageNumber="93" specimenCount="1" specimenCount-female="1" stateProvince="Napo" typeStatus="paratype">
|
||
<typeStatus id="54B38834C614FFDF0FD4DF27FD77F9E0" box="[656,759,1594,1618]" pageId="92" pageNumber="93" type="paratype">Paratype</typeStatus>
|
||
: same data as holotype but
|
||
<date id="FFB61056C614FFDF0974DF24FB66F9E0" box="[1072,1254,1593,1618]" pageId="92" pageNumber="93" value="2002-05-08" valueMax="2002-05-10" valueMin="2002-05-08">
|
||
<collectingDate id="EFF2E9BEC614FFDF0974DF24FB66F9E0" box="[1072,1254,1593,1618]" pageId="92" pageNumber="93" value="2002-05-08" valueMax="2002-05-10" valueMin="2002-05-08">8–10 May 2002</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26FD5340C614FFDF09B7DF24FAFDF9E0" box="[1267,1405,1593,1618]" pageId="92" pageNumber="93">O. Lonsdale</collectorName>
|
||
(
|
||
<specimenCount id="9D0EFD1FC614FFDF08D4DF24FF2CF9C7" pageId="92" pageNumber="93" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED19AE53C614FFDF0DFEDF40FE8BF9C4" box="[186,267,1629,1654]" country="Canada" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="92" pageNumber="93">DEBU</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C312651DC614FFDF0D83DF9CFD4CF90C" pageId="92" pageNumber="93" type="discussion">
|
||
<paragraph id="8BB73696C614FFDF0D83DF9CFD4CF90C" blockId="92.[151,1437,258,1726]" pageId="92" pageNumber="93">
|
||
<emphasis id="B97CEA84C614FFDF0D83DF9CFECFF928" bold="true" box="[199,335,1665,1690]" pageId="92" pageNumber="93">Comments.</emphasis>
|
||
The species name is the Latin word for a sea urchin and refers to the spiny appearance of this fly which has enlarged bristles on the thorax and legs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |