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<document id="06D3B630FBE58AAD21958DFA5FA8A979" ID-DOI="10.6620/ZS.2017.56-06" ID-ISSN="1810-522X" ID-PMC="PMC6517739" ID-PubMed="31966205" ID-Zenodo-Dep="12824488" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721913255345" checkinUser="felipe" docAuthor="Balakirev, Alexander Evgenievich, Abramov, Alexei Vladimirovich &amp; Rozhnov, Viatcheslav Vladimirovich" docDate="2017" docId="0394BA61FFC0A705B7536C71E94E1C76" docLanguage="en" docName="ZoolStud.56.6.1-19.pdf.imf" docOrigin="Zoological Studies 56 (6)" docSource="http://dx.doi.org/10.5281/zenodo.8060414" docStyle="DocumentStyle:CB122E4603BA284FA287D3847868CAF6.2:ZoolStud.2016-2018.journal_article" docStyleId="CB122E4603BA284FA287D3847868CAF6" docStyleName="ZoolStud.2016-2018.journal_article" docStyleVersion="2" docTitle="Maxomys surifer subsp. tonkinensis Balakirev 2017" docType="treatment" docVersion="3" lastPageNumber="17" masterDocId="FFADC219FFCEA715B40A6A6CEB7B1E20" masterDocTitle="Zavreliella inawaheia Sasa, Kitami &amp; Suzuki 2001" masterLastPageNumber="19" masterPageNumber="1" pageNumber="15" updateTime="1721929600552" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="124717E36884DC78FE1F0B92DE8FA521" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F2C20A114634C265073D3B097E7D28D0">
<mods:title id="57687161D38BD9360ED50AE2A059BA59">Zavreliella inawaheia Sasa, Kitami &amp; Suzuki 2001</mods:title>
</mods:titleInfo>
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</mods:role>
<mods:namePart id="950E8EC02D99A51EEA8A624D40FE4A03">Balakirev, Alexander Evgenievich</mods:namePart>
<mods:affiliation id="57C96ACABE3721C3FF6644F9C724AE16">Joint Russian-Vietnamese Tropical Research and Technological Centre, Nguyen Van Huyen, Nghia Do, Cau Giay, Hanoi, Vietnam &amp; A. N. Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Leninskii pr. 33, Moscow 119071, Russia.</mods:affiliation>
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</mods:role>
<mods:namePart id="E9E4E1C372931090E0EFBDD1D2EA9357">Abramov, Alexei Vladimirovich</mods:namePart>
<mods:affiliation id="22186B416A9B2E81FA0934D8D209FEF1">Joint Russian-Vietnamese Tropical Research and Technological Centre, Nguyen Van Huyen, Nghia Do, Cau Giay, Hanoi, Vietnam &amp; Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, Saint Petersburg 199034, Russia.</mods:affiliation>
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<mods:namePart id="BF870EE9F251428E26AAE776DBD2C81B">Rozhnov, Viatcheslav Vladimirovich</mods:namePart>
<mods:affiliation id="2BE9ACAD4205E6B8A4B5D597DF417E87">Joint Russian-Vietnamese Tropical Research and Technological Centre, Nguyen Van Huyen, Nghia Do, Cau Giay, Hanoi, Vietnam &amp; A. N. Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Leninskii pr. 33, Moscow 119071, Russia.</mods:affiliation>
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<mods:title id="DB2802FEF685063ED4202E3C0D80A87F">Zoological Studies</mods:title>
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<mods:date id="9ED1984F161DEDD094ED18F3E30E2358">2017</mods:date>
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<mods:number id="C126E2937BE230DF6CDB4873B7C899DE">2017-04-19</mods:number>
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<mods:number id="FE9EC952AD0DF8992508B60707833948">56</mods:number>
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<mods:identifier id="AD29B7D16B772A360EBF23EE21D72835" type="ISSN">1810-522X</mods:identifier>
<mods:identifier id="6F522117F415C498FF718074B8270716" type="PMC">PMC6517739</mods:identifier>
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<treatment id="0394BA61FFC0A705B7536C71E94E1C76" LSID="urn:lsid:plazi:treatment:0394BA61FFC0A705B7536C71E94E1C76" httpUri="http://treatment.plazi.org/id/0394BA61FFC0A705B7536C71E94E1C76" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
<subSubSection id="C32758FCFFC0A71BB7536C71EFE21876" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8B820B77FFC0A71BB7536C71EEDF1816" blockId="14.[857,1444,1565,1679]" box="[857,1444,1565,1590]" pageId="14" pageNumber="15">
<heading id="D0CABC1BFFC0A71BB7536C71EEDF1816" bold="true" box="[857,1444,1565,1590]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="15" reason="2">
<taxonomicName id="4C3D70F4FFC0A71BB7536C71EEDF1816" authority="Balakirev" authorityName="Balakirev" authorityYear="2017" box="[857,1444,1565,1590]" class="Mammalia" family="Muridae" genus="Maxomys" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="15" phylum="Chordata" rank="subSpecies" species="surifer" status="subsp." subSpecies="tonkinensis">
<emphasis id="B949D765FFC0A71BB7536C71EEDF1816" bold="true" box="[857,1444,1565,1590]" pageId="14" pageNumber="15">
<emphasis id="B949D765FFC0A71BB7536C71EFB21816" bold="true" box="[857,1225,1565,1590]" italics="true" pageId="14" pageNumber="15">Maxomys surifer tonkinensis</emphasis>
Balakirev subsp.
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B820B77FFC0A71BB06E6C51EFE21876" blockId="14.[857,1444,1565,1679]" box="[1124,1177,1597,1622]" pageId="14" pageNumber="15">
<emphasis id="B949D765FFC0A71BB06E6C51EFE21876" bold="true" box="[1124,1177,1597,1622]" pageId="14" pageNumber="15">nov.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C32758FCFFC0A705B76A6C30E94E1C76" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15" type="description">
<paragraph id="8B820B77FFC0A71BB76A6C30EFA118AF" blockId="14.[857,1444,1565,1679]" pageId="14" pageNumber="15">
<uri id="FFAC0775FFC0A71BB76A6C30EFA118AF" pageId="14" pageNumber="15">
urn:lsid:zoobank.org:act:
<uuid id="FF9B31A2FFC0A71BB0686C30EFA118AF" pageId="14" pageNumber="15">7E8A1428-2943-4A06-9FC5- 263DD4DCC7F5</uuid>
</uri>
</paragraph>
<paragraph id="8B820B77FFC0A71BB7856CD8EF1C190D" blockId="14.[850,1452,1716,1966]" pageId="14" pageNumber="15">
<materialsCitation id="3B55012AFFC0A71BB7856CD8EF1C190D" collectingDate="2014-03-08" collectionCode="ZMMU" collectorName="Alexander E. Balakirev." pageId="14" pageNumber="15" specimenCount="1" typeStatus="holotype">
<emphasis id="B949D765FFC0A71BB7856CD8EF7E18ED" box="[911,1029,1716,1741]" italics="true" pageId="14" pageNumber="15">
<typeStatus id="5486B5D5FFC0A71BB7856CD8EF7E18ED" box="[911,1029,1716,1741]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectionCode id="ED2C93B2FFC0A71BB0116CD8EF0B18ED" box="[1051,1136,1716,1741]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="14" pageNumber="15" type="Museum">ZMMU</collectionCode>
S-194718, adult male, skull, skin, body in ethanol, field number Na-52, genetic voucher Na-52, collected
<date id="FF832DB7FFC0A71BB0E86C98EEFC192D" box="[1250,1415,1780,1805]" pageId="14" pageNumber="15" value="2014-03-08">
<collectingDate id="EFC7D45FFFC0A71BB0E86C98EEFC192D" box="[1250,1415,1780,1805]" pageId="14" pageNumber="15" value="2014-03-08">8 March 2014</collectingDate>
</date>
by
<collectorName id="26C86EA1FFC0A71BB7586D78EF1C190D" box="[850,1127,1812,1837]" pageId="14" pageNumber="15">Alexander E. Balakirev.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8B820B77FFC0A71BB7856D58EE0119AD" blockId="14.[850,1452,1716,1966]" pageId="14" pageNumber="15">
<emphasis id="B949D765FFC0A71BB7856D58EF56196D" box="[911,1069,1844,1869]" italics="true" pageId="14" pageNumber="15">Type locality</emphasis>
:
<collectingCountry id="F32A4BE7FFC0A71BB0486D58EFD6196D" box="[1090,1197,1844,1869]" name="Vietnam" pageId="14" pageNumber="15">Vietnam</collectingCountry>
,
<collectingRegion id="49F9C595FFC0A71BB0B66D58EEDC196D" box="[1212,1447,1844,1869]" country="Vietnam" name="Nghe An" pageId="14" pageNumber="15">Nghe An Province</collectingRegion>
, Quy Chau District, vicinity of Ban Ke Can Village,
<geoCoordinate id="EE096DB0FFC0A71BB7586D18E89219AD" box="[850,1001,1908,1933]" degrees="19" direction="north" minutes="31.184" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="19.519733">19°31.184'N</geoCoordinate>
;
<geoCoordinate id="EE096DB0FFC0A71BB7F96D18EFEC19AD" box="[1011,1175,1908,1933]" degrees="105" direction="east" minutes="10.282" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="105.171364">105°10.282'E</geoCoordinate>
, altitude
<quantity id="4CC5A692FFC0A71BB0F56D18EE4119AD" box="[1279,1338,1908,1933]" metricMagnitude="1" metricUnit="m" metricValue="9.5" pageId="14" pageNumber="15" unit="m" value="95.0">95 m</quantity>
a.s.l.
</paragraph>
<paragraph id="8B820B77FFC0A71AB7856DF8E92E1D56" blockId="14.[850,1452,1716,1966]" lastBlockId="15.[136,738,253,1974]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="B949D765FFC0A71BB7856DF8EF71198D" box="[911,1034,1940,1965]" italics="true" pageId="14" pageNumber="15">
<typeStatus id="5486B5D5FFC0A71BB7856DF8EF71198D" box="[911,1034,1940,1965]" pageId="14" pageNumber="15" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectionCode id="ED2C93B2FFC0A71BB0166DF8EF16198D" box="[1052,1133,1940,1965]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="14" pageNumber="15" type="Museum">ZMMU</collectionCode>
S-194710 (male, body at ethanol, collected
<date id="FF832DB7FFC1A71AB5786A91EA8C1F36" box="[370,503,253,278]" pageId="15" pageNumber="16" value="2014-03-06">6.03.2014</date>
),
<collectionCode id="ED2C93B2FFC1A71AB6066A91E9251F36" box="[524,606,253,278]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="15" pageNumber="16" type="Museum">ZMMU</collectionCode>
S-194711 (female, skull and skin, collected
<date id="FF832DB7FFC1A71AB6466B71E9AE1F16" box="[588,725,285,310]" pageId="15" pageNumber="16" value="2014-03-07">7.03.2014</date>
),
<collectionCode id="ED2C93B2FFC1A71AB4826B51EBA21F76" box="[136,217,317,342]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="15" pageNumber="16" type="Museum">ZMMU</collectionCode>
S-194712 (male, skull and skin, collected
<date id="FF832DB7FFC1A71AB4826B31EA7E1F56" box="[136,261,349,374]" pageId="15" pageNumber="16" value="2014-03-07">7.03.2014</date>
),
<collectionCode id="ED2C93B2FFC1A71AB51C6B31EA1D1F56" box="[278,358,349,374]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="15" pageNumber="16" type="Museum">ZMMU</collectionCode>
S-194713 (male, skull and skin, collected
<date id="FF832DB7FFC1A71AB4FD6B11EA081FB6" box="[247,371,381,406]" pageId="15" pageNumber="16" value="2014-03-07">7.03.2014</date>
),
<collectionCode id="ED2C93B2FFC1A71AB58E6B11EAAF1FB6" box="[388,468,381,406]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="15" pageNumber="16" type="Museum">ZMMU</collectionCode>
S-194714 (male, skull, collected
<date id="FF832DB7FFC1A71AB4F26BF1EA0E1F96" box="[248,373,413,438]" pageId="15" pageNumber="16" value="2014-03-07">7.03.2014</date>
),
<collectionCode id="ED2C93B2FFC1A71AB58F6BF1EAAE1F96" box="[389,469,413,438]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="15" pageNumber="16" type="Museum">ZMMU</collectionCode>
S-194715 (male, body at ethanol, collected
<date id="FF832DB7FFC1A71AB5896BD1E97A1FF6" box="[387,513,445,470]" pageId="15" pageNumber="16" value="2014-03-08">8.03.2014</date>
),
<collectionCode id="ED2C93B2FFC1A71AB61E6BD1E91F1FF6" box="[532,612,445,470]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="15" pageNumber="16" type="Museum">ZMMU</collectionCode>
S-194716 (male, body at ethanol, collected
<date id="FF832DB7FFC1A71AB6466BB1E9AE1FD6" box="[588,725,477,502]" pageId="15" pageNumber="16" value="2014-03-08">8.03.2014</date>
),
<collectionCode id="ED2C93B2FFC1A71AB4826B91EBA31C36" box="[136,216,509,534]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="15" pageNumber="16" type="Museum">ZMMU</collectionCode>
S-194717 (male, skull, collected
<date id="FF832DB7FFC1A71AB6566B91E9A31C36" box="[604,728,509,534]" pageId="15" pageNumber="16" value="2014-03-08">8.03.2014</date>
),
<collectionCode id="ED2C93B2FFC1A71AB4826871EBA31C16" box="[136,216,541,566]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="15" pageNumber="16" type="Museum">ZMMU</collectionCode>
S-194719 (male, skull, collected
<date id="FF832DB7FFC1A71AB6696871E9A51C16" box="[611,734,541,566]" pageId="15" pageNumber="16" value="2014-03-09">9.03.2014</date>
, trapped near Ban Dom1 Village),
<collectionCode id="ED2C93B2FFC1A71AB61C6851E91D1C76" box="[534,614,573,598]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="15" pageNumber="16" type="Museum">ZMMU</collectionCode>
S-194720 (male, skull, collected
<date id="FF832DB7FFC1A71AB5846831E9721C56" box="[398,521,605,630]" pageId="15" pageNumber="16" value="2014-03-09">9.03.2014</date>
, trapped near Ban Dom1 Village),
<collectionCode id="ED2C93B2FFC1A71AB54E6811EAEF1CB6" box="[324,404,637,662]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="15" pageNumber="16" type="Museum">ZMMU</collectionCode>
S-194721 (female, body at ethanol, collected
<date id="FF832DB7FFC1A71AB57868F1EA8D1C96" box="[370,502,669,694]" pageId="15" pageNumber="16" value="2014-03-09">9.03.2014</date>
),
<collectionCode id="ED2C93B2FFC1A71AB60168F1E9261C96" box="[523,605,669,694]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="15" pageNumber="16" type="Museum">ZMMU</collectionCode>
S-194722 (male, skull, collected
<date id="FF832DB7FFC1A71AB5C768D1E91C1CF6" box="[461,615,701,726]" pageId="15" pageNumber="16" value="2014-03-10">10.03.2014</date>
),
<collectionCode id="ED2C93B2FFC1A71AB68068D1E9A41CF6" box="[650,735,701,726]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="15" pageNumber="16" type="Museum">ZMMU</collectionCode>
S-194723 (male, skull, collected
<date id="FF832DB7FFC1A71AB63768B1E9AE1CD6" box="[573,725,733,758]" pageId="15" pageNumber="16" value="2014-10-03">10.03.2014</date>
),
<collectionCode id="ED2C93B2FFC1A71AB4826891EB9B1D36" box="[136,224,765,790]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="15" pageNumber="16" type="Museum">ZMMU</collectionCode>
S-194724 (female, skull, collected
<date id="FF832DB7FFC1A71AB4826971EA511D16" box="[136,298,797,822]" pageId="15" pageNumber="16" value="2014-10-03">10.03.2014</date>
),
<collectionCode id="ED2C93B2FFC1A71AB5496971EAE21D16" box="[323,409,797,822]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="15" pageNumber="16" type="Museum">ZMMU</collectionCode>
S-194725 (male, skull, collected
<date id="FF832DB7FFC1A71AB4F56951EAF61D76" box="[255,397,829,854]" pageId="15" pageNumber="16" value="2014-11-03">11.03.2014</date>
),
<collectionCode id="ED2C93B2FFC1A71AB5AB6951EA891D76" box="[417,498,829,854]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="15" pageNumber="16" type="Museum">ZMMU</collectionCode>
S-194726 (female, body at ethanol, collected
<date id="FF832DB7FFC1A71AB5B46931E9311D56" box="[446,586,861,886]" pageId="15" pageNumber="16" value="2014-12-03">12.03.2014</date>
).
</paragraph>
<paragraph id="8B820B77FFC1A71AB4CF6911EA9B1DF6" blockId="15.[136,738,253,1974]" pageId="15" pageNumber="16">
All collected by Alexander E. Balakirev and Tran Quang Tien from the same locality as the
<typeStatus id="5486B5D5FFC1A71AB48269D1EB961DF6" box="[136,237,957,982]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
or in closest vicinity.
</paragraph>
<paragraph id="8B820B77FFC1A71AB4CF69B1EA481A76" blockId="15.[136,738,253,1974]" pageId="15" pageNumber="16">
<emphasis id="B949D765FFC1A71AB4CF69B1EA331DD6" box="[197,328,989,1014]" italics="true" pageId="15" pageNumber="16">Etymology</emphasis>
: The new subspecies is named after
<collectingCountry id="F32A4BE7FFC1A71AB4C76991EA5E1A36" box="[205,293,1021,1046]" name="Vietnam" pageId="15" pageNumber="16">Tonkin</collectingCountry>
, the former name of northern part of Eastern Indochina, with the Latin suffix -
<emphasis id="B949D765FFC1A71AB6AB6E71E99A1A16" box="[673,737,1053,1078]" italics="true" pageId="15" pageNumber="16">ensis</emphasis>
(belonging to).
</paragraph>
<paragraph id="8B820B77FFC1A71AB4CF6E31E9CF1AF6" blockId="15.[136,738,253,1974]" pageId="15" pageNumber="16">
<emphasis id="B949D765FFC1A71AB4CF6E31EA311A56" box="[197,330,1117,1142]" italics="true" pageId="15" pageNumber="16">Diagnosis</emphasis>
: Medium-sized rat, larger on average in its external and cranial measurements than the nominotypical
<taxonomicName id="4C3D70F4FFC1A71AB5AA6EF1E96A1A96" baseAuthorityName="Miller" baseAuthorityYear="1900" box="[416,529,1181,1206]" class="Mammalia" family="Muridae" genus="Maxomys" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="surifer">
<emphasis id="B949D765FFC1A71AB5AA6EF1E96A1A96" box="[416,529,1181,1206]" italics="true" pageId="15" pageNumber="16">M. surifer</emphasis>
</taxonomicName>
with a longer tail (106-115% of body length for most individuals).
</paragraph>
<paragraph id="8B820B77FFC1A71AB4CF6EB1E9081B16" blockId="15.[136,738,253,1974]" pageId="15" pageNumber="16">
<emphasis id="B949D765FFC1A71AB4CF6EB1E9311AD6" box="[197,586,1245,1270]" italics="true" pageId="15" pageNumber="16">Description and comparisons</emphasis>
: Head and body length
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, tail
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, ear
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, weight
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. figure 10 B.
</paragraph>
<paragraph id="8B820B77FFC1A71AB4CF6F51E8D41F96" blockId="15.[136,738,253,1974]" lastBlockId="15.[813,1414,253,694]" pageId="15" pageNumber="16">
New subspecies differs from the
<emphasis id="B949D765FFC1A71AB66C6F51E99B1B76" box="[614,736,1341,1366]" italics="true" pageId="15" pageNumber="16">M. s. finis</emphasis>
which distributed over southern Indochina in the larger skull sizes and in the relatively short upper and lower toothrows. The most evident morphological features at skull construction are also be noticed the size and shape of incisive foramina. In contrast with
<emphasis id="B949D765FFC1A71AB5FC6F91E9051836" box="[502,638,1533,1558]" italics="true" pageId="15" pageNumber="16">M. s. finis</emphasis>
, which usually has shorter, reniform openings with clearly pointed cranial edge of notch,
<emphasis id="B949D765FFC1A71AB6066C51E99A1876" box="[524,737,1597,1622]" italics="true" pageId="15" pageNumber="16">M. s. tonkinensis</emphasis>
has more elongated foramina with more or less rounded cranial and caudal edges of notch figures 7-9, its somewhat reassemble in shape to the foramina characteristic to representatives of genus
<taxonomicName id="4C3D70F4FFC1A71AB4826CB1EA5318D6" authorityName="Ellerman" authorityYear="1947" box="[136,296,1757,1782]" class="Mammalia" family="Muridae" genus="Leopoldamys" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="B949D765FFC1A71AB4826CB1EA5318D6" box="[136,296,1757,1782]" italics="true" pageId="15" pageNumber="16">Leopoldamys</emphasis>
</taxonomicName>
but much wider at caudal side. The most remarkable external feature is the pattern of tail coloration, namely the character of tail tip discoloration. This progressive discoloration, characteristic for all
<taxonomicName id="4C3D70F4FFC1A71AB58C6D31EA871956" baseAuthorityName="Miller" baseAuthorityYear="1900" box="[390,508,1885,1910]" class="Mammalia" family="Muridae" genus="Maxomys" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="surifer">
<emphasis id="B949D765FFC1A71AB58C6D31EA871956" box="[390,508,1885,1910]" italics="true" pageId="15" pageNumber="16">M. surifer</emphasis>
</taxonomicName>
starts appreciably earlier than it usually happens with representatives of southern populations, discolorations starts approximately from the middle of tail and terminal one third usually became completely white, whereas for
<emphasis id="B949D765FFC1A71AB7C56B51EF361F76" box="[975,1101,317,342]" italics="true" pageId="15" pageNumber="16">M. s. finis</emphasis>
discolorations launched from approximately terminal third with only very tip about 1/5 of tail length being completely discolored.
</paragraph>
<paragraph id="8B820B77FFC1A71AB7636BD1EF651C96" blockId="15.[813,1414,253,694]" pageId="15" pageNumber="16">
<emphasis id="B949D765FFC1A71AB7636BD1E88A1FF6" box="[873,1009,445,470]" italics="true" pageId="15" pageNumber="16">Distribution</emphasis>
: Northern
<collectingCountry id="F32A4BE7FFC1A71AB0726BD1EFA01FF6" box="[1144,1243,445,470]" name="Vietnam" pageId="15" pageNumber="16">Vietnam</collectingCountry>
from
<collectingRegion id="49F9C595FFC1A71AB12C6BD1E8E61FD6" country="Vietnam" name="Ha Tinh" pageId="15" pageNumber="16">Ha Tinh Province</collectingRegion>
in its extreme south to Ba Vi (
<collectingRegion id="49F9C595FFC1A71AB1326BB1EEFF1FD6" box="[1336,1412,477,502]" country="Vietnam" name="Ha Noi" pageId="15" pageNumber="16">Hanoi</collectingRegion>
area) in the north; central
<collectingCountry id="F32A4BE7FFC1A71AB06B6B91EFE01C36" box="[1121,1179,509,534]" name="Laos" pageId="15" pageNumber="16">Laos</collectingCountry>
(
<collectingRegion id="49F9C595FFC1A71AB0A26B91EE2A1C36" box="[1192,1361,509,534]" country="Laos" name="Khammouan" pageId="15" pageNumber="16">Khammouane</collectingRegion>
and
<collectingRegion id="49F9C595FFC1A71AB7276871E8C51C16" box="[813,958,541,566]" country="Laos" name="Bolikhamxai" pageId="15" pageNumber="16">Bolikhamxai</collectingRegion>
provinces). It may be probably found in southernmost
<collectingRegion id="49F9C595FFC1A71AB7FD6851EF2D1C76" box="[1015,1110,573,598]" country="China" name="Yunnan" pageId="15" pageNumber="16">Yunnan</collectingRegion>
,
<collectingCountry id="F32A4BE7FFC1A71AB0696851EFD21C77" box="[1123,1193,573,599]" name="China" pageId="15" pageNumber="16">China</collectingCountry>
(
<bibRefCitation id="EFAC7686FFC1A71AB0B36851EE321C76" author="Wang YX" box="[1209,1353,573,598]" pageId="15" pageNumber="16" refId="ref11175" refString="Wang YX. 2003. A complete checklist of mammals species and subspecies of China: a taxonomic and geographic references. China Forestry Publishing House, Beijing." type="book" year="2003">Wang 2003</bibRefCitation>
) and northern
<collectingCountry id="F32A4BE7FFC1A71AB7956831E8A41C56" box="[927,991,605,630]" name="Laos" pageId="15" pageNumber="16">Laos</collectingCountry>
, but its presence there has to be approved by investigation of museum specimens or new field records.
</paragraph>
<paragraph id="8B820B77FFC1A705B72768B1E9551F56" blockId="15.[813,1415,733,1975]" lastBlockId="16.[174,775,253,374]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<emphasis id="B949D765FFC1A71AB72768B1EF661CD6" bold="true" box="[813,1053,733,758]" pageId="15" pageNumber="16">Acknowledgment:</emphasis>
This work and the one new subspecies name have been registered with Zoo Bank under
<uri id="FFAC0775FFC1A71AB7CF6971EF8B1D77" pageId="15" pageNumber="16">
urn:lsid:zoobank.org:pub:
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</uri>
. This study was realized with the support of the Joint Russian-Vietnamese Tropical Research and Technological Centre,
<collectingRegion id="49F9C595FFC1A71AB79E69F1E89A1D96" box="[916,993,925,950]" country="Vietnam" name="Ha Noi" pageId="15" pageNumber="16">Hanoi</collectingRegion>
,
<collectingCountry id="F32A4BE7FFC1A71AB7FA69F1EF201D96" box="[1008,1115,925,950]" name="Vietnam" pageId="15" pageNumber="16">Vietnam</collectingCountry>
. We thank Dr. Victor
<collectionCode id="ED2C93B2FFC1A71AB16469F1EEFA1D96" box="[1390,1409,925,950]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="15" pageNumber="16" type="Museum">V</collectionCode>
. Suntsov (A.N. Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences,
<collectingRegion id="49F9C595FFC1A71AB11469B1EEF81DD6" box="[1310,1411,989,1014]" country="Russia" name="Moscow" pageId="15" pageNumber="16">Moscow</collectingRegion>
,
<collectingCountry id="F32A4BE7FFC1A71AB7276991E8E91A36" box="[813,914,1021,1046]" name="Russia" pageId="15" pageNumber="16">Russia</collectingCountry>
), Dr. Sergey
<collectionCode id="ED2C93B2FFC1A71AB0466991EF1A1A36" box="[1100,1121,1021,1046]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="15" pageNumber="16" type="Museum">V</collectionCode>
. Kruskop (Zoological Museum of
<collectingRegion id="49F9C595FFC1A71AB7CD6E71EF001A17" box="[967,1147,1053,1079]" country="Russia" name="Moscow" pageId="15" pageNumber="16">Moscow State</collectingRegion>
University,
<collectingRegion id="49F9C595FFC1A71AB1126E71EEF91A16" box="[1304,1410,1053,1078]" country="Russia" name="Moscow" pageId="15" pageNumber="16">Moscow</collectingRegion>
,
<collectingCountry id="F32A4BE7FFC1A71AB7276E51E8E81A76" box="[813,915,1085,1110]" name="Russia" pageId="15" pageNumber="16">Russia</collectingCountry>
), Dr. Charles M. Francis (Canadian Wildlife Service, Environment
<collectingCountry id="F32A4BE7FFC1A71AB0A56E31EE6D1A57" box="[1199,1302,1117,1143]" name="Canada" pageId="15" pageNumber="16">Canada</collectingCountry>
, Ottawa,
<collectingCountry id="F32A4BE7FFC1A71AB7276E11E8E91AB7" box="[813,914,1149,1175]" name="Canada" pageId="15" pageNumber="16">Canada</collectingCountry>
), Dr. Vu Dinh Thong (Zoological Museum of Institute of Ecology and Biological Resources, Vietnamese Academy of Science and Technology) who allow us to access for samples and museum collections and datasets, Dr. Nguyen Dang Hoi, Dr. Bui Xuan Phuong, Mr. Tran Quang Tien and Ms. Pham Mai Phuong (all from the Joint Russian-Vietnamese Tropical Research and Technological Centre,
<collectingRegion id="49F9C595FFC1A71AB7856F11E8A11BB6" box="[911,986,1405,1430]" country="Vietnam" name="Ha Noi" pageId="15" pageNumber="16">Hanoi</collectingRegion>
,
<collectingCountry id="F32A4BE7FFC1A71AB7ED6F11EF2B1BB6" box="[999,1104,1405,1430]" name="Vietnam" pageId="15" pageNumber="16">Vietnam</collectingCountry>
), who made considerable efforts in preparing for a number of expeditions and who supplied us with a significant number of specimens. We also want to express our warmest gratitude to Dr. Kristofer M. Helgen, Mr. Craig Ludwig and Mr. Renee Regan (all from the Smithsonian Institution; National Museum of Natural History, Division of Mammals,
<collectingRegion id="49F9C595FFC1A71AB0F86C31EEF91856" box="[1266,1410,1629,1654]" country="United States of America" name="Washington" pageId="15" pageNumber="16">Washington</collectingRegion>
,
<collectingCountry id="F32A4BE7FFC1A71AB7276C11E81C18B7" box="[813,871,1661,1687]" name="United States of America" pageId="15" pageNumber="16">USA</collectingCountry>
) for their kind assistance in the investigation and photography of
<typeStatus id="5486B5D5FFC1A71AB0116CF1EFFB1896" box="[1051,1152,1693,1718]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
specimen. Finally, we thank the administrations of Ba Vi, Nam Cat Tien, Bi Dup-Nui Ba, Bu Gia Map, Lo Go Xa Mat, Dong Nai, Phu Quoc and Binh Chau National Parks and Nature Reserves for their aid in the management of our studies. This study was supported in part by the Research Program “Living nature: modern state and problems of development” of the Presidium of the Russian Academy of Sciences, by the Zoological Institute’ Program No.0125-2016- 004, the Russian Foundation for Basic Research (grant 16-04-00085) and the Russian Science Foundation (grant 17-14-01160).
</paragraph>
<paragraph id="8B820B77FFDEA705B4A46BF1E9C41F96" blockId="16.[174,703,413,438]" box="[174,703,413,438]" pageId="16" pageNumber="17">
<emphasis id="B949D765FFDEA705B4A46BF1E9C41F96" bold="true" box="[174,703,413,438]" pageId="16" pageNumber="17">List of abbreviation (not explained in text)</emphasis>
</paragraph>
<paragraph id="8B820B77FFDEA705B4A46BB1EA801FD6" blockId="16.[174,774,477,598]" box="[174,507,477,502]" pageId="16" pageNumber="17">DNA - deoxyribonucleic acid</paragraph>
<paragraph id="8B820B77FFDEA705B4A46B90EB851C16" blockId="16.[174,774,477,598]" pageId="16" pageNumber="17">
<collectionCode id="ED2C93B2FFDEA705B4A46B90EB951C36" box="[174,238,508,534]" pageId="16" pageNumber="17">ICZN</collectionCode>
- International Code of Zoological Nomenclature
</paragraph>
<paragraph id="8B820B77FFDEA705B4A46850E94E1C76" blockId="16.[174,774,477,598]" box="[174,565,572,598]" pageId="16" pageNumber="17">PCR - polymerase chain reaction</paragraph>
</subSubSection>
</treatment>
</document>
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