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<document id="45C0B18D424F9F8BB30EADCED010E70F" ID-DOI="10.59893/bjc.23(2).009" ID-ISSN="1407-8619" ID-Zenodo-Dep="13203106" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657248871" checkinUser="felipe" docAuthor="Sarker, Subir &amp; Talukder, Saranika" docDate="2023" docId="038C9D1E0461FFDDFFD9FDFA820073D8" docLanguage="en" docName="BalticJColeopterol.23.2.233-239.pdf" docOrigin="Baltic Journal of Coleopterology 23 (2)" docSource="http://dx.doi.org/10.1128/mra.00329-24" docStyle="DocumentStyle:514B3A4252435B0AFD496C900C6D7572.1:BalticJColeop.2023-.journal_article" docStyleId="514B3A4252435B0AFD496C900C6D7572" docStyleName="BalticJColeop.2023-.journal_article" docStyleVersion="1" docTitle="Cacomorphocerus incurvus Sarker &amp; Talukder, 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="239" masterDocId="FFB5E5660460FFDFFFABFFD081247205" masterDocTitle="Two new taxa of fossil Cantharidae from Baltic amber" masterLastPageNumber="239" masterPageNumber="233" pageNumber="233" updateTime="1722696919189" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="0932571BE9768F94E0BD68463FEC3B19">Two new taxa of fossil Cantharidae from Baltic amber</mods:title>
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<mods:namePart id="1F457A4494432CE48F36F8BC9EE28E8F">Sarker, Subir</mods:namePart>
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<mods:namePart id="D87E4CBAD5F528DCFB4C8284E7982406">Talukder, Saranika</mods:namePart>
</mods:name>
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<mods:title id="F523409C3FA0896BD81BF7B758E54EB9">Baltic Journal of Coleopterology</mods:title>
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<mods:date id="F9D8798B4A5F919F22D922BC1171B27A">2023</mods:date>
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<mods:identifier id="A4948E9DA82650A355A9B6BFEF0A398F" type="DOI">10.59893/bjc.23(2).009</mods:identifier>
<mods:identifier id="71A7C176BA8B2E6B31942A717CF64096" type="ISSN">1407-8619</mods:identifier>
<mods:identifier id="C21A24517027A86AA1A2EA8D1D04DD34" type="Zenodo-Dep">13203106</mods:identifier>
</mods:mods>
<treatment id="038C9D1E0461FFDDFFD9FDFA820073D8" LSID="urn:lsid:plazi:treatment:038C9D1E0461FFDDFFD9FDFA820073D8" httpUri="http://treatment.plazi.org/id/038C9D1E0461FFDDFFD9FDFA820073D8" lastPageId="2" pageId="1">
<subSubSection id="C33F7F830461FFDEFFD9FDFA83357047" box="[114,529,554,579]" pageId="1" type="nomenclature">
<paragraph id="8B9A2C080461FFDEFFD9FDFA83357047" blockId="1.[114,529,554,609]" box="[114,529,554,579]" pageId="1">
<heading id="D0D29B640461FFDEFFD9FDFA83357047" bold="true" box="[114,529,554,579]" fontSize="10" level="2" pageId="1" reason="4">
<emphasis id="B951F01A0461FFDEFFD9FDFA83357047" bold="true" box="[114,529,554,579]" pageId="1">
<taxonomicName id="4C25578B0461FFDEFFD4FDFB808B7046" authorityName="Sarker &amp; Talukder" authorityYear="2023" box="[127,431,555,579]" class="Insecta" family="Cantharidae" genus="Cacomorphocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="incurvus" status="sp. nov.">
<emphasis id="B951F01A0461FFDEFFD4FDFB808B7046" bold="true" box="[127,431,555,579]" italics="true" pageId="1">Cacomorphocerus incurvus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624D610461FFDEFE1DFDFA83357047" box="[438,529,554,578]" pageId="1" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33F7F830461FFDEFFD9FD9981E67064" box="[114,194,585,609]" pageId="1" type="description">
<paragraph id="8B9A2C080461FFDEFFD9FD9981E67064" blockId="1.[114,529,554,609]" box="[114,194,585,609]" pageId="1">
(
<figureCitation id="131E308D0461FFDEFFD1FD99819C7064" box="[122,184,585,609]" captionStart="Fig" captionStartId="1.[624,663,1112,1136]" captionTargetBox="[624,1070,263,1074]" captionTargetId="figure-205@1.[624,1070,263,1075]" captionTargetPageId="1" captionText="Fig. 1. Cacomorphocerus incurvussp. nov. in Baltic amber (code BaA08RU). A: Holotype, dorsal view, scale bar = 1.0 mm; B: Holotype, ventral view, scale bar = 1.0 mm; C: Holotype, detail of head (ventral view), scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13203108" httpUri="https://zenodo.org/record/13203108/files/figure.png" pageId="1">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33F7F830461FFDEFFD9FD57800971D4" pageId="1" type="materials_examined">
<paragraph id="8B9A2C080461FFDEFFD9FD5781ED70FE" blockId="1.[114,593,647,763]" pageId="1">
<materialsCitation id="3B4D26550461FFDEFFD9FD5781ED70FE" country="Russia" county="Baltic" location="Baltic" municipality="Fanti" pageId="1" specimenCount="1" typeStatus="holotype">
<emphasis id="B951F01A0461FFDEFFD9FD5781C3709A" bold="true" box="[114,231,647,671]" pageId="1">
<typeStatus id="549E92AA0461FFDEFFD9FD5781C6709A" box="[114,226,647,671]" pageId="1" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="8EFA7AD30461FFDEFF44FD57800A709A" LSID="urn:lsid:plazi:treatment:038C9D1E0461FFDDFFD9FDFA820073D8:8EFA7AD30461FFDEFF44FD57800A709A" box="[239,302,647,671]" country="Russia" county="Baltic" municipality="Fanti" name="Baltic" pageId="1">Baltic</location>
amber, collection
<collectingMunicipality id="6BFEB6720461FFDEFDA6FD578368709A" box="[525,588,647,671]" pageId="1">Fanti</collectingMunicipality>
, access code BaA08RU (BaA =
<collectingCounty id="62FB54840461FFDEFDA0FD75837570B8" box="[523,593,677,701]" pageId="1">Baltic</collectingCounty>
amber; 08 = sequential number; RU =
<collectingCountry id="F3326C980461FFDEFFD9FD33819A70FE" box="[114,190,739,763]" name="Russia" pageId="1">Russia</collectingCountry>
).
</materialsCitation>
</paragraph>
<paragraph id="8B9A2C080461FFDEFFD9FCF080367152" blockId="1.[114,593,800,855]" pageId="1">
<emphasis id="B951F01A0461FFDEFFD9FCF08030713D" bold="true" box="[114,276,800,824]" pageId="1">Type locality.</emphasis>
Yantarny mine,
<collectingRegion id="49E1E2EA0461FFDEFE60FCF0819D7152" country="Russia" name="Kaliningrad" pageId="1">Kaliningrad region</collectingRegion>
,
<collectingCountry id="F3326C980461FFDEFF6FFCEF802A7152" box="[196,270,831,855]" name="Russia" pageId="1">Russia</collectingCountry>
.
</paragraph>
<paragraph id="8B9A2C080461FFDEFFD9FCAC800971D4" blockId="1.[114,592,892,977]" pageId="1">
<emphasis id="B951F01A0461FFDEFFD9FCAC803D7191" bold="true" box="[114,281,892,916]" pageId="1">Type horizon.</emphasis>
Middle Eocene: Lutetian (47.841.2 Mya) to late Eocene: Priabonian (37.833.9 Mya).
</paragraph>
</subSubSection>
<subSubSection id="C33F7F830461FFDDFFD9FC2783A077D4" lastPageId="2" pageId="1" type="diagnosis">
<paragraph id="8B9A2C080461FFDEFFD9FC2783757450" blockId="1.[114,593,1015,1621]" pageId="1">
<emphasis id="B951F01A0461FFDEFFD9FC278051760A" bold="true" box="[114,373,1015,1039]" pageId="1">Differential diagnosis.</emphasis>
Compared to the new species described here,
<taxonomicName id="4C25578B0461FFDEFFD9FBE4818C766E" authority="Schaufuss, 1892" authorityName="Schaufuss" authorityYear="1892" class="Insecta" family="Cantharidae" genus="Cacomorphocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="cerambyx">
<emphasis id="B951F01A0461FFDEFFD9FBE480E57649" box="[114,449,1076,1100]" italics="true" pageId="1">Cacomorphocerus cerambyx</emphasis>
<bibRefCitation id="EFB451F90461FFDEFE7CFBE4818C766E" author="Schaufuss C. F. C." pageId="1" refId="ref3297" refString="Schaufuss C. F. C. 1892. Preussens Bernstein-Kafer. Neue Formen aus der Helm'schen Sammlung im Danziger Provinzialmuseum. Berliner Entomologische Zeitschrift 36 (I): 53 - 64 [1891]." type="book" year="1892">Schaufuss, 1892</bibRefCitation>
</taxonomicName>
shows a different pronotum, which is square, with narrower apical margin and a longitudinal impression on the disc (
<bibRefCitation id="EFB451F90461FFDEFFD2FB7F800976C2" author="Schaufuss C. F. C." box="[121,301,1199,1223]" pageId="1" refId="ref3297" refString="Schaufuss C. F. C. 1892. Preussens Bernstein-Kafer. Neue Formen aus der Helm'schen Sammlung im Danziger Provinzialmuseum. Berliner Entomologische Zeitschrift 36 (I): 53 - 64 [1891]." type="book" year="1892">Schaufuss 1892</bibRefCitation>
;
<bibRefCitation id="EFB451F90461FFDEFE94FB7F819676E0" author="Fanti F. &amp; Kupryjanowicz J." pageId="1" pagination="149 - 153" refId="ref2527" refString="Fanti F., Kupryjanowicz J. 2018. Discovery of a new fossil soldier beetle in Eocene Baltic amber, with the establishment of the new tribe Cacomorphocerini. Annales de Paleontologie 104: 149 - 153." type="journal article" year="2018">Fanti &amp; Kupryjanowicz 2018</bibRefCitation>
).
<taxonomicName id="4C25578B0461FFDEFF70FB1E80987701" authority="Fanti &amp; Kupryjanowicz, 2018" authorityName="Fanti &amp; Kupryjanowicz" authorityYear="2018" class="Insecta" family="Cantharidae" genus="Cacomorphocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="wiszniewskii">
<emphasis id="B951F01A0461FFDEFF70FB1E837476E3" box="[219,592,1230,1254]" italics="true" pageId="1">Cacomorphocerus wiszniewskii</emphasis>
<bibRefCitation id="EFB451F90461FFDEFFD9FB3C80987701" author="Fanti F. &amp; Kupryjanowicz J." box="[114,444,1260,1284]" pageId="1" pagination="149 - 153" refId="ref2527" refString="Fanti F., Kupryjanowicz J. 2018. Discovery of a new fossil soldier beetle in Eocene Baltic amber, with the establishment of the new tribe Cacomorphocerini. Annales de Paleontologie 104: 149 - 153." type="journal article" year="2018">Fanti &amp; Kupryjanowicz, 2018</bibRefCitation>
</taxonomicName>
differs from the new species in the subrectangular pronotum, with sides slightly concave from base to the apical third, where a very small denticle is present, and in the less saucer-shaped antennomeres IV-VI (
<bibRefCitation id="EFB451F90461FFDEFE48FA55804F77B9" author="Fanti F. &amp; Kupryjanowicz J." pageId="1" pagination="149 - 153" refId="ref2527" refString="Fanti F., Kupryjanowicz J. 2018. Discovery of a new fossil soldier beetle in Eocene Baltic amber, with the establishment of the new tribe Cacomorphocerini. Annales de Paleontologie 104: 149 - 153." type="journal article" year="2018">Fanti &amp; Kupryjanowicz 2018</bibRefCitation>
).
<taxonomicName id="4C25578B0461FFDEFE2EFA7480DE77DE" authority="(Kuska &amp; Kania, 2010)" baseAuthorityName="Kuska &amp; Kania" baseAuthorityYear="2010" class="Insecta" family="Cantharidae" genus="Cacomorphocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="jantaricus">
<emphasis id="B951F01A0461FFDEFE2EFA7481C177DF" italics="true" pageId="1">Cacomorphocerus jantaricus</emphasis>
(
<bibRefCitation id="EFB451F90461FFDEFF58FA1380D777DE" author="Kuska A. &amp; Kania I." box="[243,499,1475,1499]" pageId="1" pagination="49 - 56" refId="ref2935" refString="Kuska A., Kania I. 2010. New soldier beetles (Coleoptera, Cantharidae) from the Eocene Baltic amber. Zootaxa 2400: 49 - 56." type="journal article" year="2010">Kuśka &amp; Kania, 2010</bibRefCitation>
)
</taxonomicName>
shows antennomeres differently shaped and pronotum more rectangular, with sides slightly protruding laterally near the apical margin (
<bibRefCitation id="EFB451F90461FFDEFF7BF9ED80EC7450" author="Kuska A. &amp; Kania I." box="[208,456,1597,1621]" pageId="1" pagination="49 - 56" refId="ref2935" refString="Kuska A., Kania I. 2010. New soldier beetles (Coleoptera, Cantharidae) from the Eocene Baltic amber. Zootaxa 2400: 49 - 56." type="journal article" year="2010">Kuśka &amp; Kania 2010</bibRefCitation>
). The other
</paragraph>
<caption id="DF5A7C800461FFDEFDDBFB8882A9770F" ID-DOI="http://doi.org/10.5281/zenodo.13203108" ID-Zenodo-Dep="13203108" httpUri="https://zenodo.org/record/13203108/files/figure.png" pageId="1" startId="1.[624,663,1112,1136]" targetBox="[624,1070,263,1074]" targetPageId="1" targetType="figure">
<paragraph id="8B9A2C080461FFDEFDDBFB8882A9770F" blockId="1.[624,1103,1112,1290]" pageId="1">
Fig. 1.
<taxonomicName id="4C25578B0461FFDEFD6AFB8882D57675" authorityName="Sarker &amp; Talukder" authorityYear="2023" box="[705,1009,1112,1136]" class="Insecta" family="Cantharidae" genus="Cacomorphocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" phylum="Arthropoda" rank="species" species="incurvus" status="sp. nov.">
<emphasis id="B951F01A0461FFDEFD6AFB8882D57675" box="[705,1009,1112,1136]" italics="true" pageId="1">Cacomorphocerus incurvus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624D610461FFDEFC5FFB88856B7675" box="[1012,1103,1112,1136]" pageId="1" rank="species">sp. nov.</taxonomicNameLabel>
in Baltic amber (code BaA08RU). A: Holotype, dorsal view, scale bar = 1.0 mm; B: Holotype, ventral view, scale bar = 1.0 mm; C: Holotype, detail of head (ventral view), scale bar = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8B9A2C080461FFDEFFD9F976829477A6" blockId="1.[114,154,1702,1726]" lastBlockId="1.[624,1103,1327,1627]" pageId="1">
234
<emphasis id="B951F01A0461FFDEFDDBFAFF82257742" bold="true" box="[624,769,1327,1351]" pageId="1">Description.</emphasis>
Sex undefined, adult, robust, winged. Body length: about
<quantity id="4CDD81ED0461FFDEFC6FFA9E856B7763" box="[964,1103,1358,1382]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" metricValueMax="7.5" metricValueMin="7.3" pageId="1" unit="mm" value="7.4" valueMax="7.5" valueMin="7.3">7.3-7.5 mm</quantity>
(the specimen is folded), elytra
<quantity id="4CDD81ED0461FFDEFC7DFABC856F7781" box="[982,1099,1388,1412]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" metricValueMax="5.1" metricValueMin="5.0" pageId="1" unit="mm" value="5.05" valueMax="5.1" valueMin="5.0">5-5.1 mm</quantity>
. Entirely blackish-dark brown.
</paragraph>
<paragraph id="8B9A2C080461FFDDFDDBFA7A801372ED" blockId="1.[624,1103,1327,1627]" lastBlockId="2.[189,669,178,1489]" lastPageId="2" pageId="1">Head elongated, rounded behind the eyes, covered with long setae and a shallow punctation. Eyes roundish, convex and prominent, inserted in the upper-lateral part of the head. Mandibles elongated, falciform. Maxillary palps 4-segmented; last palpomere securiform. Labial palps 3- segmented.</paragraph>
<paragraph id="8B9A2C080462FFDDFF16FF3F80CA7090" blockId="2.[189,669,178,1489]" pageId="2">Antennae 12-segmented, short, reaching about the elytral half, strongly pubescent; scape short, stout, club-shaped; antennomere II very short, globular, about 2.7 times shorter than scape; antennomeres IIIVII saucer-shaped and asymmetrically inflated; antennomere VIII strongly saucer-shaped and asymmetrically dilated with apical lobes, sturdier than previous ones; antennomere IX squared, thinner than previous one; antennomeres XXII elongated, filiform, rather robust, X-XI slightly enlarged apically, last antennomere thinner and rounded at apex.</paragraph>
<paragraph id="8B9A2C080462FFDDFF16FD4C804B7169" blockId="2.[189,669,178,1489]" pageId="2">Pronotum transverse, about as wide as head; apical and basal margins straight; sides straight, without denticles; surface without impressions, covered with shallow granules and several setae. Scutellum triangular, with pointed apex which is rounded at apex.</paragraph>
<paragraph id="8B9A2C080462FFDDFF16FCA3837F7600" blockId="2.[189,669,178,1489]" pageId="2">Elytra long, covering and surpassing the last abdominal segment, parallel-sided and with rounded apex, wider than pronotum; surface smooth and pubescent. Hind wings transparent, slightly longer than elytra.</paragraph>
<paragraph id="8B9A2C080462FFDDFF16FBDC831E771C" blockId="2.[189,669,178,1489]" pageId="2">Metasternum wide, short, robust, rounded apically and pubescent. Sternites transverse and pubescent. Legs slender and short; coxae very large; trochanters elongated, rounded apically; femora cylindrical, straight, more robust than tibiae; tibiae cylindrical, pro- and mesotibiae very slightly longer than pro- and mesofemora, metatibiae longer than metafemora.</paragraph>
<caption id="DF5A7C800462FFDDFD10FBEA823076AB" ID-DOI="http://doi.org/10.5281/zenodo.13203112" ID-Zenodo-Dep="13203112" httpUri="https://zenodo.org/record/13203112/files/figure.png" pageId="2" startId="2.[699,738,1082,1106]" targetBox="[699,1184,754,1045]" targetPageId="2" targetType="figure">
<paragraph id="8B9A2C080462FFDDFD10FBEA823076AB" blockId="2.[699,1179,1082,1198]" pageId="2">
Fig. 2.
<taxonomicName id="4C25578B0462FFDDFCA7FBEA85577654" authorityName="Sarker &amp; Talukder" authorityYear="2023" box="[780,1139,1082,1106]" class="Insecta" family="Cantharidae" genus="Malthinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="amicitiae" status="sp. nov." subGenus="Malthinus">
<emphasis id="B951F01A0462FFDDFCA7FBEA825B7657" box="[780,895,1082,1106]" italics="true" pageId="2">Malthinus</emphasis>
(
<emphasis id="B951F01A0462FFDDFC20FBEA82D37657" box="[907,1015,1082,1106]" italics="true" pageId="2">Malthinus</emphasis>
)
<emphasis id="B951F01A0462FFDDFBA2FBEA85577654" box="[1033,1139,1082,1105]" italics="true" pageId="2">amicitiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2624D610462FFDDFBDEFBEA83CD7675" pageId="2" rank="species">sp. nov.</taxonomicNameLabel>
in Baltic amber (code BaA09RU). Holotype, dorso-lateral view, scale bar = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8B9A2C080462FFDDFF16FAF083A077D4" blockId="2.[189,669,178,1489]" pageId="2">Tarsi 5-segmented, short; tarsomere I elongated, tarsomeres II and III triangular and shorter than I, tarsomere IV bilobed, tarsomere V elongated but not extremely thin and enlarged apically; claws simple with a very small and obtuse tooth at base.</paragraph>
</subSubSection>
<subSubSection id="C33F7F830462FFDDFF16FA27850D72ED" pageId="2" type="etymology">
<paragraph id="8B9A2C080462FFDDFF16FA27800F746E" blockId="2.[189,668,1527,1643]" pageId="2">
<emphasis id="B951F01A0462FFDDFF16FA278060740A" bold="true" box="[189,324,1527,1551]" pageId="2">Etymology.</emphasis>
The specific epithet derives from the Latin adjective
<taxonomicName id="4C25578B0462FFDDFE55F9C583427428" authorityName="Sarker &amp; Talukder" authorityYear="2023" box="[510,614,1557,1581]" class="Insecta" family="Cantharidae" genus="Cacomorphocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" phylum="Arthropoda" rank="species" species="incurvus">
<emphasis id="B951F01A0462FFDDFDA1F9C683427428" box="[522,614,1558,1581]" italics="true" pageId="2">incurvus</emphasis>
</taxonomicName>
” = curved, in reference to the pronotum and head bent.
</paragraph>
<paragraph id="8B9A2C080462FFDDFD10FF61850D72ED" blockId="2.[699,1178,177,232]" pageId="2">
<emphasis id="B951F01A0462FFDDFD10FF61824072CC" bold="true" box="[699,868,177,201]" pageId="2">Syninclusions.</emphasis>
Enhydro and air bubbles, botanical fragments, stellate hairs.
</paragraph>
</subSubSection>
<subSubSection id="C33F7F830462FFDDFD10FEDD820073D8" pageId="2" type="discussion">
<paragraph id="8B9A2C080462FFDDFD10FEDD820073D8" blockId="2.[699,1179,269,477]" pageId="2">
<emphasis id="B951F01A0462FFDDFD10FEDD82147320" bold="true" box="[699,816,269,293]" pageId="2">Remarks.</emphasis>
The oblong reddish yellow amber piece measures approximately 41x20x
<quantity id="4CDD81ED0462FFDDFCA6FE9B827B7366" box="[781,863,331,355]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" unit="mm" value="10.0">10 mm</quantity>
. The inclusion is complete, with some folded parts and an open elytron showing the hind wing. Some parts of the body are slightly covered with white emulsion.
</paragraph>
</subSubSection>
</treatment>
</document>