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<document ID-DOI="10.26028/cybium/2020-443-003" ID-Zenodo-Dep="7806280" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1680797231632" checkinUser="diego" docAuthor="Denys *, Gaël P. J., Dettai, Agnès, Persat, Henri, Daszkiewicz, Piotr, Hautecoeur, Mélyne &amp; Keith, Philippe" docDate="2020" docId="038487D4C75DFFFAFEEEFD00FABAFB00" docLanguage="en" docName="Cybium.44.3.205-237.pdf" docOrigin="Cybium 44 (3)" docTitle="Phoxinus fayollarum Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith 2020, new species" docType="treatment" docVersion="4" lastPageNumber="225" masterDocId="FFBDFFACC74FFFEEFFE4FF99FFCAFFD3" masterDocTitle="Revision of Phoxinus in France with the description of two new species (Teleostei, Leuciscidae)" masterLastPageNumber="238" masterPageNumber="205" pageNumber="223" updateTime="1680800786029" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of Phoxinus in France with the description of two new species (Teleostei, Leuciscidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Denys *, Gaël P. J.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dettai, Agnès</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Persat, Henri</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Daszkiewicz, Piotr</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hautecoeur, Mélyne</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Keith, Philippe</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Cybium</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-09-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>44</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>205</mods:start>
<mods:end>238</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.26028/cybium/2020-443-003</mods:identifier>
<mods:identifier type="Zenodo-Dep">7806280</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7806158" ID-Zenodo-Dep="7806158" LSID="urn:lsid:plazi:treatment:038487D4C75DFFFAFEEEFD00FABAFB00" httpUri="http://treatment.plazi.org/id/038487D4C75DFFFAFEEEFD00FABAFB00" lastPageId="20" lastPageNumber="225" pageId="18" pageNumber="223">
<subSubSection box="[266,642,665,690]" pageId="18" pageNumber="223" type="nomenclature">
<paragraph blockId="18.[266,642,665,722]" box="[266,642,665,690]" pageId="18" pageNumber="223">
<heading bold="true" box="[266,642,665,690]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="223" reason="4">
<emphasis bold="true" box="[266,642,665,690]" pageId="18" pageNumber="223">
<taxonomicName authority="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith, 2020" authorityName="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith" authorityYear="2020" box="[266,498,665,689]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="223" phylum="Chordata" rank="species" species="fayollarum" status="new species">
<emphasis bold="true" box="[266,498,665,689]" italics="true" pageId="18" pageNumber="223">Phoxinus fayollarum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[511,642,665,690]" pageId="18" pageNumber="223" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[414,493,698,722]" pageId="18" pageNumber="223" type="description">
<paragraph blockId="18.[266,642,665,722]" box="[414,493,698,722]" pageId="18" pageNumber="223">
<heading box="[414,493,698,722]" centered="true" fontSize="10" level="4" pageId="18" pageNumber="223" reason="4">
(
<figureCitation box="[422,485,698,722]" captionStart="Figure 9" captionStartId="18.[813,878,1309,1331]" captionTargetBox="[813,1456,215,1292]" captionTargetId="figure-483@18.[813,1456,215,1292]" captionTargetPageId="18" captionText="Figure 9. Phoxinus fayollarum Holotype m (A), 58.13 mm SL from the Boron Stream, Pionsat 46°06.4N, 2°41.1E, MNHN 2019-0263; paratypes m (B) and f (C), same location as holotype, MNHN 2014-2729; nuptial colouration pattern for male (D) and female (E)." figureDoi="http://doi.org/10.5281/zenodo.7806298" httpUri="https://zenodo.org/record/7806298/files/figure.png" pageId="18" pageNumber="223">Fig. 9</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="19" lastPageNumber="224" pageId="18" pageNumber="223" type="materials_examined">
<paragraph blockId="18.[132,775,762,850]" box="[132,236,762,786]" pageId="18" pageNumber="223">
<heading bold="true" box="[132,236,762,786]" fontSize="10" level="3" pageId="18" pageNumber="223" reason="6">
<emphasis bold="true" box="[132,236,762,786]" pageId="18" pageNumber="223">
<typeStatus box="[132,236,762,786]" pageId="18" pageNumber="223">Holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="18.[132,775,762,850]" pageId="18" pageNumber="223">
<materialsCitation collectionCode="MNHN" country="France" elevation="522" latitude="46.106667" location="Pionsat" longLatPrecision="112" longitude="2.685" municipality="Pionsat" pageId="18" pageNumber="223" specimenCode="MNHN 2019-0263" specimenCount="1" stateProvince="Boron Stream">
<specimenCode box="[170,393,794,818]" pageId="18" pageNumber="223">MNHN 2019-0263</specimenCode>
,
<emphasis bold="true" box="[407,422,796,817]" pageId="18" pageNumber="223">
<specimenCount box="[407,422,796,817]" pageId="18" pageNumber="223" type="male">m</specimenCount>
</emphasis>
, 58.1 mm SL;
<collectingCountry box="[607,692,794,817]" name="France" pageId="18" pageNumber="223" value="France">France</collectingCountry>
:
<collectingRegion pageId="18" pageNumber="223">Boron Stream</collectingRegion>
,
<collectingMunicipality box="[220,301,826,850]" pageId="18" pageNumber="223">Pionsat</collectingMunicipality>
,
<geoCoordinate box="[312,424,826,850]" degrees="46" direction="north" minutes="06.4" orientation="latitude" pageId="18" pageNumber="223" precision="92" value="46.106667">46°06.4N</geoCoordinate>
<geoCoordinate box="[430,529,826,850]" degrees="2" direction="east" minutes="41.1" orientation="longitude" pageId="18" pageNumber="223" precision="92" value="2.685">2°41.1E</geoCoordinate>
;
<specimenCount box="[539,609,825,849]" pageId="18" pageNumber="223" type="generic">
<elevation box="[539,609,825,849]" metricMagnitude="2" metricUnit="m" metricValue="5.22" pageId="18" pageNumber="223" unit="m" value="522.0">522 m</elevation>
</specimenCount>
</materialsCitation>
.
</paragraph>
<paragraph blockId="18.[132,776,890,1074]" box="[132,246,890,914]" pageId="18" pageNumber="223">
<heading bold="true" box="[132,246,890,914]" fontSize="10" level="3" pageId="18" pageNumber="223" reason="6">
<emphasis bold="true" box="[132,246,890,914]" pageId="18" pageNumber="223">
<typeStatus box="[132,246,890,914]" pageId="18" pageNumber="223">Paratypes</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="18.[132,776,890,1074]" pageId="18" pageNumber="223">
<materialsCitation collectionCode="MNHN" country="France" elevation="522" latitude="46.106667" location="Pionsat" longLatPrecision="112" longitude="2.685" municipality="Pionsat" pageId="18" pageNumber="223" specimenCode="MNHN 2014-2729" specimenCount="8" specimenCount-female="3" specimenCount-male="5" stateProvince="Boron Stream">
<specimenCode box="[170,379,922,946]" pageId="18" pageNumber="223">MNHN 2014-2729</specimenCode>
,
<specimenCount box="[391,426,922,946]" pageId="18" pageNumber="223" type="male">
5
<emphasis bold="true" box="[411,426,924,945]" pageId="18" pageNumber="223">m</emphasis>
</specimenCount>
,
<specimenCount box="[440,473,922,946]" pageId="18" pageNumber="223" type="female">
3
<emphasis bold="true" box="[461,473,923,945]" pageId="18" pageNumber="223">f</emphasis>
</specimenCount>
, 57.5-68.1 mm SL;
<collectingCountry box="[693,771,922,945]" name="France" pageId="18" pageNumber="223" value="France">France</collectingCountry>
:
<collectingRegion box="[132,297,954,978]" pageId="18" pageNumber="223">Boron Stream</collectingRegion>
,
<collectingMunicipality box="[309,394,954,978]" pageId="18" pageNumber="223">Pionsat</collectingMunicipality>
<geoCoordinate box="[402,522,954,978]" degrees="46" direction="north" minutes="06.4" orientation="latitude" pageId="18" pageNumber="223" precision="92" value="46.106667">46°06.4N</geoCoordinate>
<geoCoordinate box="[530,637,954,978]" degrees="2" direction="east" minutes="41.1" orientation="longitude" pageId="18" pageNumber="223" precision="92" value="2.685">2°41.1E</geoCoordinate>
; alt.
<elevation box="[695,771,954,978]" metricMagnitude="2" metricUnit="m" metricValue="5.22" pageId="18" pageNumber="223" unit="m" value="522.0">522 m</elevation>
.
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="231" latitude="47.235" location="Dompierre-sur-Nievre" longLatPrecision="111" longitude="3.25" municipality="Dompierre-sur-Nievre" pageId="18" pageNumber="223" specimenCode="MNHN 2014-2732" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="stream Nievre">
<specimenCode box="[132,350,986,1010]" pageId="18" pageNumber="223">MNHN 2014-2732</specimenCode>
,
<specimenCount box="[363,400,986,1010]" pageId="18" pageNumber="223" type="male">
2
<emphasis bold="true" box="[385,400,988,1009]" pageId="18" pageNumber="223">m</emphasis>
</specimenCount>
,
<specimenCount box="[419,453,986,1010]" pageId="18" pageNumber="223" type="female">
2
<emphasis bold="true" box="[441,453,987,1009]" pageId="18" pageNumber="223">f</emphasis>
</specimenCount>
, 50.2-58.9 mm SL;
<collectingCountry box="[690,772,986,1009]" name="France" pageId="18" pageNumber="223" value="France">France</collectingCountry>
:
<collectingRegion box="[132,287,1018,1042]" pageId="18" pageNumber="223">stream Nièvre</collectingRegion>
,
<collectingMunicipality box="[297,541,1018,1042]" pageId="18" pageNumber="223">Dompierre-sur-Nièvre</collectingMunicipality>
,
<geoCoordinate box="[551,664,1018,1042]" degrees="47" direction="north" minutes="14.1" orientation="latitude" pageId="18" pageNumber="223" precision="92" value="47.235">47°14.1N</geoCoordinate>
<geoCoordinate box="[671,770,1018,1042]" degrees="3" direction="east" minutes="15.0" orientation="longitude" pageId="18" pageNumber="223" precision="92" value="3.25">3°15.0E</geoCoordinate>
; alt.
<specimenCount box="[172,242,1050,1074]" pageId="18" pageNumber="223" type="generic">
<elevation box="[172,242,1050,1074]" metricMagnitude="2" metricUnit="m" metricValue="2.31" pageId="18" pageNumber="223" unit="m" value="231.0">231 m</elevation>
</specimenCount>
</materialsCitation>
.
</paragraph>
<paragraph blockId="18.[132,776,1114,1650]" box="[132,356,1114,1138]" pageId="18" pageNumber="223">
<heading bold="true" box="[132,356,1114,1138]" fontSize="10" level="3" pageId="18" pageNumber="223" reason="6">
<emphasis bold="true" box="[132,356,1114,1138]" pageId="18" pageNumber="223">Additional material</emphasis>
</heading>
</paragraph>
<paragraph blockId="18.[132,776,1114,1650]" pageId="18" pageNumber="223">
<materialsCitation collectionCode="MNHN" country="France" elevation="800" latitude="45.385" location="Sauvessanges" longLatPrecision="113" longitude="3.8933334" municipality="Sauvessanges" pageId="18" pageNumber="223" specimenCode="MNHN 2010-0992" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Ance du Nord Stream">
<specimenCode box="[170,380,1146,1170]" pageId="18" pageNumber="223">MNHN 2010-0992</specimenCode>
,
<specimenCount box="[391,426,1146,1170]" pageId="18" pageNumber="223" type="male">
2
<emphasis bold="true" box="[411,426,1148,1169]" pageId="18" pageNumber="223">m</emphasis>
</specimenCount>
,
<specimenCount box="[440,473,1146,1170]" pageId="18" pageNumber="223" type="female">
1
<emphasis bold="true" box="[461,473,1147,1169]" pageId="18" pageNumber="223">f</emphasis>
</specimenCount>
, 50.2-56.1 mm SL;
<collectingCountry box="[693,771,1146,1169]" name="France" pageId="18" pageNumber="223" value="France">France</collectingCountry>
:
<collectingRegion box="[132,373,1178,1202]" pageId="18" pageNumber="223">Ance du Nord Stream</collectingRegion>
,
<collectingMunicipality box="[384,537,1178,1202]" pageId="18" pageNumber="223">Sauvessanges</collectingMunicipality>
,
<geoCoordinate box="[548,663,1178,1202]" degrees="45" direction="north" minutes="23.1" orientation="latitude" pageId="18" pageNumber="223" precision="92" value="45.385">45°23.1N</geoCoordinate>
<geoCoordinate box="[669,770,1178,1202]" degrees="3" direction="east" minutes="53.6" orientation="longitude" pageId="18" pageNumber="223" precision="92" value="3.8933334">3°53.6E</geoCoordinate>
; alt.
<elevation box="[178,254,1210,1234]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="18" pageNumber="223" unit="m" value="800.0">800 m</elevation>
.
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="205" latitude="45.798332" location="Aixe-sur-Vienne" longLatPrecision="112" longitude="1.1516666" municipality="Aixe-sur-Vienne" pageId="18" pageNumber="223" specimenCode="MNHN 2010-1055" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Aurence Stream">
<specimenCode box="[267,487,1210,1234]" pageId="18" pageNumber="223">MNHN 2010-1055</specimenCode>
,
<specimenCount box="[501,538,1210,1234]" pageId="18" pageNumber="223" type="male">
2
<emphasis bold="true" box="[523,538,1213,1234]" pageId="18" pageNumber="223">m</emphasis>
</specimenCount>
,
<specimenCount box="[558,593,1210,1234]" pageId="18" pageNumber="223" type="female">
1
<emphasis bold="true" box="[581,593,1211,1233]" pageId="18" pageNumber="223">f</emphasis>
</specimenCount>
, 48.9-54.8 mm SL;
<collectingCountry box="[179,259,1242,1265]" name="France" pageId="18" pageNumber="223" value="France">France</collectingCountry>
:
<collectingRegion box="[268,450,1242,1266]" pageId="18" pageNumber="223">Aurence Stream</collectingRegion>
,
<collectingMunicipality box="[460,647,1242,1266]" pageId="18" pageNumber="223">Aixe-sur-Vienne</collectingMunicipality>
,
<geoCoordinate box="[659,775,1242,1266]" degrees="45" direction="north" minutes="47.9" orientation="latitude" pageId="18" pageNumber="223" precision="92" value="45.798332">45°47.9N</geoCoordinate>
<geoCoordinate box="[132,237,1274,1298]" degrees="1" direction="east" minutes="09.1" orientation="longitude" pageId="18" pageNumber="223" precision="92" value="1.1516666">1°09.1E</geoCoordinate>
; alt.
<elevation box="[294,369,1274,1298]" metricMagnitude="2" metricUnit="m" metricValue="2.05" pageId="18" pageNumber="223" unit="m" value="205.0">205 m</elevation>
.
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="734" latitude="44.84" location="Saint-Haon" longLatPrecision="113" longitude="3.7366667" municipality="Saint-Haon" pageId="18" pageNumber="223" specimenCode="MNHN 2011-0819" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Allier Stream">
<specimenCode box="[380,596,1274,1298]" pageId="18" pageNumber="223">MNHN 2011-0819</specimenCode>
,
<specimenCount box="[609,646,1274,1298]" pageId="18" pageNumber="223" type="male">
2
<emphasis bold="true" box="[631,646,1277,1298]" pageId="18" pageNumber="223">m</emphasis>
</specimenCount>
,
<specimenCount box="[664,698,1274,1298]" pageId="18" pageNumber="223" type="female">
4
<emphasis bold="true" box="[686,698,1276,1298]" pageId="18" pageNumber="223">f</emphasis>
</specimenCount>
, 41.0- 60.5 mm SL;
<collectingCountry box="[280,358,1306,1329]" name="France" pageId="18" pageNumber="223" value="France">France</collectingCountry>
:
<collectingRegion box="[366,514,1306,1330]" pageId="18" pageNumber="223">Allier Stream</collectingRegion>
,
<collectingMunicipality box="[525,651,1306,1330]" pageId="18" pageNumber="223">Saint-Haon</collectingMunicipality>
,
<geoCoordinate box="[661,775,1306,1330]" degrees="44" direction="north" minutes="50.4" orientation="latitude" pageId="18" pageNumber="223" precision="92" value="44.84">44°50.4N</geoCoordinate>
<geoCoordinate box="[132,232,1338,1362]" degrees="3" direction="east" minutes="44.2" orientation="longitude" pageId="18" pageNumber="223" precision="92" value="3.7366667">3°44.2E</geoCoordinate>
; alt.
<elevation box="[282,353,1338,1362]" metricMagnitude="2" metricUnit="m" metricValue="7.34" pageId="18" pageNumber="223" unit="m" value="734.0">734 m</elevation>
.
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="114" latitude="48.468334" location="Souce" longLatPrecision="110" longitude="-0.65833336" municipality="Souce" pageId="18" pageNumber="223" specimenCode="MNHN 2011-0910" specimenCount="5" specimenCount-female="5" stateProvince="Varenne Stream">
<specimenCode box="[363,569,1338,1362]" pageId="18" pageNumber="223">MNHN 2011-0910</specimenCode>
,
<specimenCount box="[579,610,1338,1362]" pageId="18" pageNumber="223" type="female">
5
<emphasis bold="true" box="[598,610,1340,1362]" pageId="18" pageNumber="223">f</emphasis>
</specimenCount>
, 41.0-47.0 mm SL;
<collectingCountry box="[179,258,1371,1394]" name="France" pageId="18" pageNumber="223" value="France">France</collectingCountry>
:
<collectingRegion box="[268,446,1370,1394]" pageId="18" pageNumber="223">Varenne Stream</collectingRegion>
,
<collectingMunicipality box="[457,527,1370,1394]" pageId="18" pageNumber="223">Soucé</collectingMunicipality>
,
<geoCoordinate box="[538,654,1370,1394]" degrees="48" direction="north" minutes="28.1" orientation="latitude" pageId="18" pageNumber="223" precision="92" value="48.468334">48°28.1N</geoCoordinate>
<geoCoordinate box="[661,771,1370,1394]" degrees="0" direction="west" minutes="39.5" orientation="longitude" pageId="18" pageNumber="223" precision="92" value="-0.65833336">0°39.5W</geoCoordinate>
; alt.
<elevation box="[172,242,1402,1426]" metricMagnitude="2" metricUnit="m" metricValue="1.14" pageId="18" pageNumber="223" unit="m" value="114.0">114 m</elevation>
.
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="88" latitude="47.043335" location="Ciran" longLatPrecision="112" longitude="0.8933333" municipality="Ciran" pageId="18" pageNumber="223" specimenCode="MNHN 2014-2733" specimenCount="15" specimenCount-female="11" specimenCount-male="4" stateProvince="Esves Stream">
<specimenCode box="[252,461,1402,1426]" pageId="18" pageNumber="223">MNHN 2014-2733</specimenCode>
,
<specimenCount box="[471,506,1402,1426]" pageId="18" pageNumber="223" type="male">
4
<emphasis bold="true" box="[491,506,1405,1426]" pageId="18" pageNumber="223">m</emphasis>
</specimenCount>
,
<specimenCount box="[519,564,1402,1426]" pageId="18" pageNumber="223" type="female">
11
<emphasis bold="true" box="[552,564,1404,1426]" pageId="18" pageNumber="223">f</emphasis>
</specimenCount>
, 48.1-67.0 mm SL;
<collectingCountry box="[132,209,1435,1458]" name="France" pageId="18" pageNumber="223" value="France">France</collectingCountry>
:
<collectingRegion box="[218,365,1434,1458]" pageId="18" pageNumber="223">Esves Stream</collectingRegion>
,
<collectingMunicipality box="[375,437,1434,1458]" pageId="18" pageNumber="223">Ciran</collectingMunicipality>
,
<geoCoordinate box="[447,560,1434,1458]" degrees="47" direction="north" minutes="02.6" orientation="latitude" pageId="18" pageNumber="223" precision="92" value="47.043335">47°02.6N</geoCoordinate>
<geoCoordinate box="[566,664,1434,1458]" degrees="0" direction="east" minutes="53.6" orientation="longitude" pageId="18" pageNumber="223" precision="92" value="0.8933333">0°53.6E</geoCoordinate>
; alt.
<elevation box="[714,771,1434,1458]" metricMagnitude="1" metricUnit="m" metricValue="8.8" pageId="18" pageNumber="223" unit="m" value="88.0">88 m</elevation>
.
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="158" latitude="47.17" location="Rians" longLatPrecision="112" longitude="2.5933332" municipality="Rians" pageId="18" pageNumber="223" specimenCode="MNHN 2014-2740" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Oua- tier Stream">
<specimenCode box="[132,338,1466,1490]" pageId="18" pageNumber="223">MNHN 2014-2740</specimenCode>
,
<specimenCount box="[347,381,1466,1490]" pageId="18" pageNumber="223" type="male">
1
<emphasis bold="true" box="[366,381,1469,1490]" pageId="18" pageNumber="223">m</emphasis>
</specimenCount>
,
<specimenCount box="[393,424,1466,1490]" pageId="18" pageNumber="223" type="female">
4
<emphasis bold="true" box="[412,424,1468,1490]" pageId="18" pageNumber="223">f</emphasis>
</specimenCount>
, 59.1-73.6 mm SL;
<collectingCountry box="[636,713,1467,1490]" name="France" pageId="18" pageNumber="223" value="France">France</collectingCountry>
:
<collectingRegion pageId="18" pageNumber="223">Oua- tier Stream</collectingRegion>
,
<collectingMunicipality box="[265,329,1498,1522]" pageId="18" pageNumber="223">Rians</collectingMunicipality>
,
<geoCoordinate box="[340,453,1498,1522]" degrees="47" direction="north" minutes="10.2" orientation="latitude" pageId="18" pageNumber="223" precision="92" value="47.17">47°10.2N</geoCoordinate>
<geoCoordinate box="[460,560,1498,1522]" degrees="2" direction="east" minutes="35.6" orientation="longitude" pageId="18" pageNumber="223" precision="92" value="2.5933332">2°35.6E</geoCoordinate>
; alt.
<elevation box="[611,683,1498,1522]" metricMagnitude="2" metricUnit="m" metricValue="1.58" pageId="18" pageNumber="223" unit="m" value="158.0">158 m</elevation>
.
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="83" latitude="47.823334" location="Vance" longLatPrecision="111" longitude="0.6433333" municipality="Vance" pageId="18" pageNumber="223" specimenCode="MNHN 2014-2745" specimenCount="10" specimenCount-female="8" specimenCount-male="2" stateProvince="Tusson Stream">
<specimenCode pageId="18" pageNumber="223">MNHN 2014-2745</specimenCode>
,
<specimenCount box="[271,309,1530,1554]" pageId="18" pageNumber="223" type="male">
2
<emphasis bold="true" box="[294,309,1533,1554]" pageId="18" pageNumber="223">m</emphasis>
</specimenCount>
,
<specimenCount box="[328,362,1530,1554]" pageId="18" pageNumber="223" type="female">
8
<emphasis bold="true" box="[350,362,1532,1554]" pageId="18" pageNumber="223">f</emphasis>
</specimenCount>
, 43.4-70.1 mm SL;
<collectingCountry box="[600,683,1531,1554]" name="France" pageId="18" pageNumber="223" value="France">France</collectingCountry>
:
<collectingRegion pageId="18" pageNumber="223">Tusson Stream</collectingRegion>
,
<collectingMunicipality box="[219,287,1562,1586]" pageId="18" pageNumber="223">Vancé</collectingMunicipality>
,
<geoCoordinate box="[296,409,1562,1586]" degrees="47" direction="north" minutes="49.4" orientation="latitude" pageId="18" pageNumber="223" precision="92" value="47.823334">47°49.4N</geoCoordinate>
<geoCoordinate box="[414,512,1562,1586]" degrees="0" direction="east" minutes="38.6" orientation="longitude" pageId="18" pageNumber="223" precision="92" value="0.6433333">0°38.6E</geoCoordinate>
; alt.
<elevation box="[561,617,1562,1586]" metricMagnitude="1" metricUnit="m" metricValue="8.3" pageId="18" pageNumber="223" unit="m" value="83.0">83 m</elevation>
.
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="121" latitude="47.036667" location="Moulins-sur-Cephons" longLatPrecision="112" longitude="1.5516666" municipality="Moulins-sur-Cephons" pageId="18" pageNumber="223" specimenCode="MNHN 2014- 2767" specimenCount="8" specimenCount-female="4" specimenCount-male="4" stateProvince="Cephons Stream">
<specimenCode pageId="18" pageNumber="223">MNHN 2014- 2767</specimenCode>
,
<specimenCount box="[200,235,1594,1618]" pageId="18" pageNumber="223" type="male">
4
<emphasis bold="true" box="[220,235,1597,1618]" pageId="18" pageNumber="223">m</emphasis>
</specimenCount>
,
<specimenCount box="[249,282,1594,1618]" pageId="18" pageNumber="223" type="female">
4
<emphasis bold="true" box="[270,282,1596,1618]" pageId="18" pageNumber="223">f</emphasis>
</specimenCount>
, 41.2-51.9 mm SL;
<collectingCountry box="[502,580,1595,1618]" name="France" pageId="18" pageNumber="223" value="France">France</collectingCountry>
:
<collectingRegion box="[590,771,1594,1618]" pageId="18" pageNumber="223">Céphons Stream</collectingRegion>
,
<collectingMunicipality box="[132,367,1626,1650]" pageId="18" pageNumber="223">Moulins-sur-Céphons</collectingMunicipality>
,
<geoCoordinate box="[377,489,1626,1650]" degrees="47" direction="north" minutes="02.2" orientation="latitude" pageId="18" pageNumber="223" precision="92" value="47.036667">47°02.2N</geoCoordinate>
<geoCoordinate box="[495,594,1626,1650]" degrees="1" direction="east" minutes="33.1" orientation="longitude" pageId="18" pageNumber="223" precision="92" value="1.5516666">1°33.1E</geoCoordinate>
; alt.
<specimenCount box="[644,714,1626,1650]" pageId="18" pageNumber="223" type="generic">
<elevation box="[644,714,1626,1650]" metricMagnitude="2" metricUnit="m" metricValue="1.21" pageId="18" pageNumber="223" unit="m" value="121.0">121 m</elevation>
</specimenCount>
</materialsCitation>
.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7806298" ID-Zenodo-Dep="7806298" httpUri="https://zenodo.org/record/7806298/files/figure.png" pageId="18" pageNumber="223" startId="18.[813,878,1309,1331]" targetBox="[813,1456,215,1292]" targetPageId="18" targetType="figure">
<paragraph blockId="18.[813,1456,1309,1427]" pageId="18" pageNumber="223">
Figure 9.
<emphasis box="[911,1141,1309,1331]" italics="true" pageId="18" pageNumber="223">
<taxonomicName authorityName="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith" authorityYear="2020" box="[930,1141,1309,1330]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="223" phylum="Chordata" rank="species" species="fayollarum">Phoxinusfayollarum</taxonomicName>
</emphasis>
Holotype
<emphasis bold="true" box="[1248,1261,1312,1331]" pageId="18" pageNumber="223">m</emphasis>
(
<emphasis bold="true" box="[1279,1296,1309,1330]" pageId="18" pageNumber="223">A</emphasis>
), 58.13 mm SL from the Boron Stream, Pionsat 46°06.4N, 2°41.1E, MNHN 2019-0263; paratypes
<emphasis bold="true" box="[1031,1044,1360,1379]" pageId="18" pageNumber="223">m</emphasis>
(
<emphasis bold="true" box="[1060,1076,1357,1378]" pageId="18" pageNumber="223">B</emphasis>
) and
<emphasis bold="true" box="[1131,1142,1359,1379]" pageId="18" pageNumber="223">f</emphasis>
(
<emphasis bold="true" box="[1158,1175,1357,1379]" pageId="18" pageNumber="223">C</emphasis>
), same location as holotype, MNHN 2014-2729; nuptial colouration pattern for male (
<emphasis bold="true" box="[1387,1404,1381,1402]" pageId="18" pageNumber="223">D</emphasis>
) and female (
<emphasis bold="true" box="[891,907,1405,1426]" pageId="18" pageNumber="223">E</emphasis>
).
</paragraph>
</caption>
<paragraph blockId="18.[132,775,1690,1907]" box="[132,544,1690,1714]" pageId="18" pageNumber="223">
<heading bold="true" box="[132,544,1690,1714]" fontSize="10" level="3" pageId="18" pageNumber="223" reason="6">
<emphasis bold="true" box="[132,544,1690,1714]" pageId="18" pageNumber="223">Material used for molecular analysis</emphasis>
</heading>
</paragraph>
<paragraph blockId="18.[132,775,1690,1907]" lastBlockId="19.[132,776,217,882]" lastPageId="19" lastPageNumber="224" pageId="18" pageNumber="223">
<materialsCitation collectionCode="MNHN" country="France" elevation="522" latitude="46.106667" location="Pionsat" longLatPrecision="112" longitude="2.685" municipality="Pionsat" pageId="18" pageNumber="223" specimenCode="MNHN 2019-0263" specimenCount="1" stateProvince="Boron Stream" typeStatus="holotype">
<specimenCode box="[170,379,1722,1746]" pageId="18" pageNumber="223">MNHN 2019-0263</specimenCode>
,
<typeStatus box="[390,488,1722,1746]" pageId="18" pageNumber="223">holotype</typeStatus>
, tag 12474;
<collectingCountry box="[619,697,1723,1746]" name="France" pageId="18" pageNumber="223" value="France">France</collectingCountry>
:
<collectingRegion pageId="18" pageNumber="223">Boron Stream</collectingRegion>
,
<collectingMunicipality box="[220,301,1754,1778]" pageId="18" pageNumber="223">Pionsat</collectingMunicipality>
,
<geoCoordinate box="[311,423,1754,1778]" degrees="46" direction="north" minutes="06.4" orientation="latitude" pageId="18" pageNumber="223" precision="92" value="46.106667">46°06.4N</geoCoordinate>
<geoCoordinate box="[428,527,1754,1778]" degrees="2" direction="east" minutes="41.1" orientation="longitude" pageId="18" pageNumber="223" precision="92" value="2.685">2°41.1E</geoCoordinate>
; alt.
<elevation box="[576,645,1754,1778]" metricMagnitude="2" metricUnit="m" metricValue="5.22" pageId="18" pageNumber="223" unit="m" value="522.0">522 m</elevation>
. (COI: Gen- Bank accession number:
<accessionNumber box="[405,530,1786,1810]" pageId="18" pageNumber="223">MT975733</accessionNumber>
, 12S: GenBank acces- sion number:
<accessionNumber box="[290,421,1818,1842]" pageId="18" pageNumber="223">MT975362</accessionNumber>
).
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="522" latitude="46.106667" location="Pionsat" longLatPrecision="112" longitude="2.685" municipality="Pionsat" pageId="18" pageNumber="223" specimenCode="MNHN 2014-2729" specimenCount="1" stateProvince="Boron Stream" typeStatus="paratype">
<specimenCode box="[440,659,1819,1843]" pageId="18" pageNumber="223">MNHN 2014-2729</specimenCode>
,
<typeStatus box="[670,770,1819,1843]" pageId="18" pageNumber="223">paratype</typeStatus>
,
<specimenCount box="[132,146,1851,1875]" pageId="18" pageNumber="223">1</specimenCount>
, tag 12475;
<collectingCountry box="[291,373,1851,1874]" name="France" pageId="18" pageNumber="223" value="France">France</collectingCountry>
:
<collectingRegion box="[384,545,1851,1875]" pageId="18" pageNumber="223">Boron Stream</collectingRegion>
,
<collectingMunicipality box="[557,644,1850,1874]" pageId="18" pageNumber="223">Pionsat</collectingMunicipality>
,
<geoCoordinate box="[657,775,1850,1874]" degrees="46" direction="north" minutes="06.4" orientation="latitude" pageId="18" pageNumber="223" precision="92" value="46.106667">46°06.4N</geoCoordinate>
<geoCoordinate box="[132,237,1883,1907]" degrees="2" direction="east" minutes="41.1" orientation="longitude" pageId="18" pageNumber="223" precision="92" value="2.685">2°41.1E</geoCoordinate>
; alt.
<elevation box="[293,368,1882,1906]" metricMagnitude="2" metricUnit="m" metricValue="5.22" pageId="18" pageNumber="223" unit="m" value="522.0">522 m</elevation>
. (COI: GenBankaccessionnumber:
<accessionNumber box="[813,937,1461,1485]" pageId="18" pageNumber="223">MT975734</accessionNumber>
, 12S: GenBankaccessionnumber:
<accessionNumber box="[1318,1444,1461,1485]" pageId="18" pageNumber="223">MT975363</accessionNumber>
)
</materialsCitation>
.
<materialsCitation collectionCode="MNHN" country="France" elevation="800" latitude="45.385" location="Sauvessanges" longLatPrecision="113" longitude="3.8933334" municipality="Sauvessanges" pageId="18" pageNumber="223" specimenCode="MNHN 2010-0992" specimenCount="1" stateProvince="Ance du Nord Stream">
<specimenCode box="[813,1036,1493,1517]" pageId="18" pageNumber="223">MNHN 2010-0992</specimenCode>
,
<specimenCount box="[1048,1063,1493,1517]" pageId="18" pageNumber="223">1</specimenCount>
, tag 4209;
<collectingCountry box="[1197,1281,1494,1517]" name="France" pageId="18" pageNumber="223" value="France">France</collectingCountry>
:
<collectingRegion pageId="18" pageNumber="223">Ance du Nord Stream</collectingRegion>
,
<collectingMunicipality box="[909,1070,1526,1550]" pageId="18" pageNumber="223">Sauvessanges</collectingMunicipality>
,
<geoCoordinate box="[1083,1203,1526,1550]" degrees="45" direction="north" minutes="23.1" orientation="latitude" pageId="18" pageNumber="223" precision="92" value="45.385">45°23.1N</geoCoordinate>
<geoCoordinate box="[1211,1318,1526,1550]" degrees="3" direction="east" minutes="53.6" orientation="longitude" pageId="18" pageNumber="223" precision="92" value="3.8933334">3°53.6E</geoCoordinate>
; alt.
<elevation box="[1376,1452,1526,1550]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="18" pageNumber="223" unit="m" value="800.0">800 m</elevation>
. (COI: GenBankaccessionnumber:
<accessionNumber box="[1206,1332,1558,1582]" pageId="18" pageNumber="223">MT975768</accessionNumber>
, 12S: Gen- Bank accession number:
<accessionNumber box="[1091,1220,1590,1614]" pageId="18" pageNumber="223">MT975381</accessionNumber>
)
</materialsCitation>
.
<materialsCitation collectionCode="MNHN" country="France" elevation="205" latitude="45.798332" location="Aixe-sur-Vienne" longLatPrecision="112" longitude="1.1516666" municipality="Aixe-sur-Vienne" pageId="18" pageNumber="223" specimenCode="MNHN 2010-1055" specimenCount="1" stateProvince="Aurence Stream">
<specimenCode box="[1238,1451,1590,1614]" pageId="18" pageNumber="223">MNHN 2010-1055</specimenCode>
,
<specimenCount box="[813,828,1623,1647]" pageId="18" pageNumber="223">1</specimenCount>
, tag 4237;
<collectingCountry box="[961,1045,1623,1646]" name="France" pageId="18" pageNumber="223" value="France">France</collectingCountry>
:
<collectingRegion box="[1055,1245,1623,1647]" pageId="18" pageNumber="223">Aurence Stream</collectingRegion>
,
<collectingMunicipality box="[1256,1451,1623,1647]" pageId="18" pageNumber="223">Aixe-sur-Vienne</collectingMunicipality>
,
<geoCoordinate box="[813,932,1655,1679]" degrees="45" direction="north" minutes="47.9" orientation="latitude" pageId="18" pageNumber="223" precision="92" value="45.798332">45°47.9N</geoCoordinate>
<geoCoordinate box="[941,1046,1655,1679]" degrees="1" direction="east" minutes="09.1" orientation="longitude" pageId="18" pageNumber="223" precision="92" value="1.1516666">1°09.1E</geoCoordinate>
; alt.
<elevation box="[1105,1182,1655,1679]" metricMagnitude="2" metricUnit="m" metricValue="2.05" pageId="18" pageNumber="223" unit="m" value="205.0">205 m</elevation>
. (COI: GenBank acces- sion number:
<accessionNumber box="[968,1096,1688,1712]" pageId="18" pageNumber="223">MT975769</accessionNumber>
, 12S: GenBank accession num- ber:
<accessionNumber box="[862,989,1720,1744]" pageId="18" pageNumber="223">MT975382</accessionNumber>
)
</materialsCitation>
.
<materialsCitation collectionCode="1" country="France" elevation="734" latitude="44.84" location="Saint-Haon" longLatPrecision="113" longitude="3.7366667" municipality="Saint-Haon" pageId="18" pageNumber="223" specimenCode="MNHN 2011-0819" specimenCount="1" stateProvince="Allier Stream">
<specimenCode box="[1006,1214,1720,1744]" pageId="18" pageNumber="223">MNHN 2011-0819</specimenCode>
,
<collectionCode box="[1225,1239,1720,1744]" pageId="18" pageNumber="223">1</collectionCode>
, tag 10741;
<collectingCountry box="[1374,1452,1721,1744]" name="France" pageId="18" pageNumber="223" value="France">France</collectingCountry>
:
<collectingRegion box="[813,961,1753,1777]" pageId="18" pageNumber="223">Allier Stream</collectingRegion>
,
<collectingMunicipality box="[972,1098,1753,1777]" pageId="18" pageNumber="223">Saint-Haon</collectingMunicipality>
,
<geoCoordinate box="[1109,1222,1752,1776]" degrees="44" direction="north" minutes="50.4" orientation="latitude" pageId="18" pageNumber="223" precision="92" value="44.84">44°50.4N</geoCoordinate>
<geoCoordinate box="[1229,1329,1753,1777]" degrees="3" direction="east" minutes="44.2" orientation="longitude" pageId="18" pageNumber="223" precision="92" value="3.7366667">3°44.2E</geoCoordinate>
; alt.
<elevation box="[1380,1452,1753,1777]" metricMagnitude="2" metricUnit="m" metricValue="7.34" pageId="18" pageNumber="223" unit="m" value="734.0">734 m</elevation>
. (COI: GenBankaccessionnumber:
<accessionNumber box="[1206,1331,1785,1809]" pageId="18" pageNumber="223">MT975702</accessionNumber>
, 12S: Gen- Bank accession number:
<accessionNumber box="[1091,1221,1817,1841]" pageId="18" pageNumber="223">MT975347</accessionNumber>
)
</materialsCitation>
.
<materialsCitation collectionCode="MNHN" country="France" elevation="114" lastPageId="19" lastPageNumber="224" latitude="48.468334" location="Souce" longLatPrecision="110" longitude="-0.65833336" municipality="Souce" pageId="18" pageNumber="223" specimenCode="MNHN 2011-0910" specimenCount="1" stateProvince="Varenne Stream">
<specimenCode box="[1239,1451,1818,1842]" pageId="18" pageNumber="223">MNHN 2011-0910</specimenCode>
,
<specimenCount box="[813,827,1850,1874]" pageId="18" pageNumber="223">1</specimenCount>
, tag 10752;
<collectingCountry box="[969,1050,1850,1873]" name="France" pageId="18" pageNumber="223" value="France">France</collectingCountry>
:
<collectingRegion box="[1061,1243,1850,1874]" pageId="18" pageNumber="223">Varenne Stream</collectingRegion>
,
<collectingMunicipality box="[1255,1327,1850,1874]" pageId="18" pageNumber="223">Soucé</collectingMunicipality>
,
<geoCoordinate box="[1338,1455,1850,1874]" degrees="48" direction="north" minutes="28.1" orientation="latitude" pageId="18" pageNumber="223" precision="92" value="48.468334">48°28.1N</geoCoordinate>
<geoCoordinate box="[813,924,1882,1906]" degrees="0" direction="west" minutes="39.5" orientation="longitude" pageId="18" pageNumber="223" precision="92" value="-0.65833336">0°39.5W</geoCoordinate>
; alt.
<elevation box="[979,1053,1882,1906]" metricMagnitude="2" metricUnit="m" metricValue="1.14" pageId="18" pageNumber="223" unit="m" value="114.0">114 m</elevation>
. (COI: GenBankaccessionnumber:
<accessionNumber box="[132,257,217,241]" pageId="19" pageNumber="224">MT975703</accessionNumber>
, 12S: GenBankaccessionnumber:
<accessionNumber box="[638,764,217,241]" pageId="19" pageNumber="224">MT975348</accessionNumber>
)
</materialsCitation>
.
<materialsCitation collectionCode="MNHN" country="France" elevation="158" latitude="47.17" location="Rians" longLatPrecision="112" longitude="2.5933332" municipality="Rians" pageId="19" pageNumber="224" specimenCode="MNHN 2014-2740" specimenCount="2" stateProvince="Oua- tier Stream">
<specimenCode box="[132,344,249,273]" pageId="19" pageNumber="224">MNHN 2014-2740</specimenCode>
,
<specimenCount box="[356,372,249,273]" pageId="19" pageNumber="224">2</specimenCount>
, tag 12347 and 12348;
<collectingCountry box="[631,710,250,273]" name="France" pageId="19" pageNumber="224" value="France">France</collectingCountry>
:
<collectingRegion pageId="19" pageNumber="224">Oua- tier Stream</collectingRegion>
,
<collectingMunicipality box="[269,335,281,305]" pageId="19" pageNumber="224">Rians</collectingMunicipality>
,
<geoCoordinate box="[347,463,281,305]" degrees="47" direction="north" minutes="10.2" orientation="latitude" pageId="19" pageNumber="224" precision="92" value="47.17">47°10.2N</geoCoordinate>
<geoCoordinate box="[470,573,281,305]" degrees="2" direction="east" minutes="35.6" orientation="longitude" pageId="19" pageNumber="224" precision="92" value="2.5933332">2°35.6E</geoCoordinate>
; alt.
<elevation box="[627,700,281,305]" metricMagnitude="2" metricUnit="m" metricValue="1.58" pageId="19" pageNumber="224" unit="m" value="158.0">158 m</elevation>
. (COI: GenBank accession numbers:
<accessionNumber box="[491,624,313,337]" pageId="19" pageNumber="224">MT975724</accessionNumber>
,
<accessionNumber box="[638,770,313,337]" pageId="19" pageNumber="224">MT975725</accessionNumber>
, 12S: GenBankaccessionnumber:
<accessionNumber box="[535,670,345,369]" pageId="19" pageNumber="224">MT975358</accessionNumber>
)
</materialsCitation>
.
<materialsCitation collectionCode="MNHN" country="France" elevation="578" latitude="45.841667" location="Felletin" longLatPrecision="112" longitude="2.2016666" municipality="Felletin" pageId="19" pageNumber="224" specimenCode="MNHN 2010-0474" specimenCount="1" stateProvince="Dejoune Stream">
<specimenCode pageId="19" pageNumber="224">MNHN 2010-0474</specimenCode>
,
<specimenCount box="[265,279,378,402]" pageId="19" pageNumber="224">1</specimenCount>
, tag 4029;
<collectingCountry box="[403,482,378,401]" name="France" pageId="19" pageNumber="224" value="France">France</collectingCountry>
:
<collectingRegion box="[492,673,377,402]" pageId="19" pageNumber="224">Dejoune Stream</collectingRegion>
,
<collectingMunicipality box="[684,771,377,401]" pageId="19" pageNumber="224">Felletin</collectingMunicipality>
,
<geoCoordinate box="[132,251,409,433]" degrees="45" direction="north" minutes="50.5" orientation="latitude" pageId="19" pageNumber="224" precision="92" value="45.841667">45°50.5N</geoCoordinate>
<geoCoordinate box="[261,367,410,434]" degrees="2" direction="east" minutes="12.1" orientation="longitude" pageId="19" pageNumber="224" precision="92" value="2.2016666">2°12.1E</geoCoordinate>
; alt.
<elevation box="[425,501,409,433]" metricMagnitude="2" metricUnit="m" metricValue="5.779999999999999" pageId="19" pageNumber="224" unit="m" value="578.0">578 m</elevation>
. (COI: GenBank acces- sion number:
<accessionNumber box="[281,409,441,465]" pageId="19" pageNumber="224">MT975752</accessionNumber>
)
</materialsCitation>
.
<materialsCitation collectionCode="MNHN" country="France" elevation="198" latitude="46.15167" location="Semme Stream" longLatPrecision="112" longitude="1.145" municipality="Semme Stream" pageId="19" pageNumber="224" specimenCode="MNHN 2010-1008" specimenCount="1">
<specimenCode box="[426,636,442,466]" pageId="19" pageNumber="224">MNHN 2010-1008</specimenCode>
,
<specimenCount box="[646,660,442,466]" pageId="19" pageNumber="224">1</specimenCount>
, tag 4189;
<collectingCountry box="[132,213,474,497]" name="France" pageId="19" pageNumber="224" value="France">France</collectingCountry>
:
<collectingMunicipality box="[225,399,474,498]" pageId="19" pageNumber="224">Semme Stream</collectingMunicipality>
,
<collectingMunicipality box="[410,485,474,497]" pageId="19" pageNumber="224">Droux</collectingMunicipality>
,
<geoCoordinate box="[497,615,473,497]" degrees="46" direction="north" minutes="09.1" orientation="latitude" pageId="19" pageNumber="224" precision="92" value="46.15167">46°09.1N</geoCoordinate>
<geoCoordinate box="[623,728,474,498]" degrees="1" direction="east" minutes="08.7" orientation="longitude" pageId="19" pageNumber="224" precision="92" value="1.145">1°08.7E</geoCoordinate>
; alt.
<elevation box="[132,208,506,530]" metricMagnitude="2" metricUnit="m" metricValue="1.98" pageId="19" pageNumber="224" unit="m" value="198.0">198 m</elevation>
. (COI: GenBankaccessionnumber:
<accessionNumber box="[631,764,505,529]" pageId="19" pageNumber="224">MT975766</accessionNumber>
)
</materialsCitation>
.
<materialsCitation collectionCode="MNHN" country="France" elevation="133" latitude="48.385" location="Sarthe Stream" longLatPrecision="110" longitude="-0.035" municipality="Moulin-le-Carbonel" pageId="19" pageNumber="224" specimenCode="MNHN 2011-0920" specimenCount="1">
<specimenCode box="[132,348,538,562]" pageId="19" pageNumber="224">MNHN 2011-0920</specimenCode>
,
<specimenCount box="[360,374,538,562]" pageId="19" pageNumber="224">1</specimenCount>
, tag 10755;
<collectingCountry box="[517,598,538,561]" name="France" pageId="19" pageNumber="224" value="France">France</collectingCountry>
:
<location LSID="urn:lsid:plazi:treatment:038487D4C75DFFFAFEEEFD00FABAFB00:8EF26019C75CFFFDFD85FD80FCC9FDE1" box="[609,771,537,562]" country="France" latitude="48.385" longLatPrecision="110" longitude="-0.035" municipality="Moulin-le-Carbonel" name="Sarthe Stream" pageId="19" pageNumber="224">Sarthe Stream</location>
,
<collectingMunicipality box="[132,348,570,594]" pageId="19" pageNumber="224">Moulin-le-Carbonel</collectingMunicipality>
,
<geoCoordinate box="[358,470,570,594]" degrees="48" direction="north" minutes="23.1" orientation="latitude" pageId="19" pageNumber="224" precision="92" value="48.385">48°23.1N</geoCoordinate>
<geoCoordinate box="[476,583,570,594]" degrees="0" direction="west" minutes="02.1" orientation="longitude" pageId="19" pageNumber="224" precision="92" value="-0.035">0°02.1W</geoCoordinate>
; alt.
<elevation box="[633,703,570,594]" metricMagnitude="2" metricUnit="m" metricValue="1.33" pageId="19" pageNumber="224" unit="m" value="133.0">133 m</elevation>
. (COI: GenBankaccessionnumber:
<accessionNumber box="[464,593,601,625]" pageId="19" pageNumber="224">MT975704</accessionNumber>
, COI: GenBank accession number:
<accessionNumber box="[344,472,633,657]" pageId="19" pageNumber="224">MT975349</accessionNumber>
)
</materialsCitation>
.
<materialsCitation collectionCode="MNHN" country="France" elevation="64" latitude="46.381668" location="Sevre niortaise River" longLatPrecision="112" longitude="-0.15666667" municipality="Sainte- Eanne" pageId="19" pageNumber="224" specimenCode="MNHN 2014-0015" specimenCount="2">
<specimenCode box="[490,702,633,657]" pageId="19" pageNumber="224">MNHN 2014-0015</specimenCode>
,
<specimenCount box="[713,729,634,658]" pageId="19" pageNumber="224">2</specimenCount>
, tag 18978 and 18979;
<collectingCountry box="[346,427,666,689]" name="France" pageId="19" pageNumber="224" value="France">France</collectingCountry>
:
<location LSID="urn:lsid:plazi:treatment:038487D4C75DFFFAFEEEFD00FABAFB00:8EF26019C75CFFFDFE53FD03FD61FD61" box="[439,683,666,690]" country="France" latitude="46.381668" longLatPrecision="112" longitude="-0.15666667" municipality="Sainte- Eanne" name="Sevre niortaise River" pageId="19" pageNumber="224">Sèvre niortaise River</location>
,
<collectingMunicipality pageId="19" pageNumber="224">Sainte- Eanne</collectingMunicipality>
,
<geoCoordinate box="[219,339,698,722]" degrees="46" direction="north" minutes="22.9" orientation="latitude" pageId="19" pageNumber="224" precision="92" value="46.381668">46°22.9N</geoCoordinate>
<geoCoordinate box="[347,462,698,722]" degrees="0" direction="west" minutes="09.4" orientation="longitude" pageId="19" pageNumber="224" precision="92" value="-0.15666667">0°09.4W</geoCoordinate>
; alt.
<elevation box="[520,583,698,722]" metricMagnitude="1" metricUnit="m" metricValue="6.4" pageId="19" pageNumber="224" unit="m" value="64.0">64 m</elevation>
. (COI: GenBank accession numbers:
<accessionNumber box="[349,473,730,754]" pageId="19" pageNumber="224">MT975735</accessionNumber>
,
<accessionNumber box="[483,607,730,754]" pageId="19" pageNumber="224">MT975736</accessionNumber>
, 12S: GenBank accession numbers:
<accessionNumber box="[348,471,762,786]" pageId="19" pageNumber="224">MT975365</accessionNumber>
,
<accessionNumber box="[482,608,762,786]" pageId="19" pageNumber="224">MT975366</accessionNumber>
)
</materialsCitation>
.
<materialsCitation collectionCode="MNHN" country="France" elevation="386" latitude="44.533333" location="de Flaujac Stream" longLatPrecision="927" longitude="2.7916667" municipality="Espalion" pageId="19" pageNumber="224" specimenCode="MNHN 2011- 0937" specimenCount="1">
<specimenCode pageId="19" pageNumber="224">MNHN 2011- 0937</specimenCode>
,
<specimenCount box="[205,220,794,818]" pageId="19" pageNumber="224">1</specimenCount>
, tag 10777;
<collectingCountry box="[366,449,794,817]" name="France" pageId="19" pageNumber="224" value="France">France</collectingCountry>
: Boralde
<location LSID="urn:lsid:plazi:treatment:038487D4C75DFFFAFEEEFD00FABAFB00:8EF26019C75CFFFDFDCBFC83FCC8FCE1" box="[559,770,794,818]" country="France" latitude="44.533333" longLatPrecision="927" longitude="2.7916667" municipality="Espalion" name="de Flaujac Stream" pageId="19" pageNumber="224">de Flaujac Stream</location>
,
<collectingMunicipality box="[132,234,826,850]" pageId="19" pageNumber="224">Espalion</collectingMunicipality>
,
<geoCoordinate box="[246,343,826,850]" degrees="44" direction="north" minutes="32" orientation="latitude" pageId="19" pageNumber="224" precision="925" value="44.533333">44°32N</geoCoordinate>
<geoCoordinate box="[351,454,826,850]" degrees="2" direction="east" minutes="47.5" orientation="longitude" pageId="19" pageNumber="224" precision="92" value="2.7916667">2°47.5E</geoCoordinate>
, alt.
<elevation box="[511,585,826,850]" metricMagnitude="2" metricUnit="m" metricValue="3.86" pageId="19" pageNumber="224" unit="m" value="386.0">386 m</elevation>
. (COI: GenBank accession number:
<accessionNumber box="[335,460,858,882]" pageId="19" pageNumber="224">MT975707</accessionNumber>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="224" type="diagnosis">
<paragraph blockId="19.[132,775,922,1907]" box="[132,242,922,946]" pageId="19" pageNumber="224">
<heading bold="true" box="[132,242,922,946]" fontSize="10" level="3" pageId="19" pageNumber="224" reason="6">
<emphasis bold="true" box="[132,242,922,946]" pageId="19" pageNumber="224">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="19.[132,775,922,1907]" pageId="19" pageNumber="224">
<taxonomicName authorityName="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith" authorityYear="2020" box="[170,410,954,978]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="224" phylum="Chordata" rank="species" species="fayollarum">
<emphasis box="[170,410,954,978]" italics="true" pageId="19" pageNumber="224">Phoxinus fayollarum</emphasis>
</taxonomicName>
is distinguished from the other French
<taxonomicName box="[217,408,986,1010]" pageId="19" pageNumber="224">
<emphasis box="[217,320,986,1010]" italics="true" pageId="19" pageNumber="224">Phoxinus</emphasis>
species
</taxonomicName>
by its unique nuptial colour pattern. The male has black bars in Z1 to Z2 and green in Z3 to Z4 (vs. black in Z2 to Z3). Z4 is bright yellow (vs. greyish of green) and the belly is red (vs. white, pinkish to light orange or greyish to black) (
<figureCitation box="[446,567,1114,1138]" captionStart="Figure 9" captionStartId="18.[813,878,1309,1331]" captionTargetBox="[813,1456,215,1292]" captionTargetId="figure-483@18.[813,1456,215,1292]" captionTargetPageId="18" captionText="Figure 9. Phoxinus fayollarum Holotype m (A), 58.13 mm SL from the Boron Stream, Pionsat 46°06.4N, 2°41.1E, MNHN 2019-0263; paratypes m (B) and f (C), same location as holotype, MNHN 2014-2729; nuptial colouration pattern for male (D) and female (E)." figureDoi="http://doi.org/10.5281/zenodo.7806298" httpUri="https://zenodo.org/record/7806298/files/figure.png" pageId="19" pageNumber="224">Fig. 9B, D</figureCitation>
). The female has a green stripe along Z3 (vs. black if present), the Z4 is yellowpinkish (vs. greyish or green), the belly red (vs. white), and the female has red spots at the pectoral, pelvic and anal fins bases (vs. pinkish) (
<figureCitation box="[342,449,1242,1266]" captionStart="Figure 9" captionStartId="18.[813,878,1309,1331]" captionTargetBox="[813,1456,215,1292]" captionTargetId="figure-483@18.[813,1456,215,1292]" captionTargetPageId="18" captionText="Figure 9. Phoxinus fayollarum Holotype m (A), 58.13 mm SL from the Boron Stream, Pionsat 46°06.4N, 2°41.1E, MNHN 2019-0263; paratypes m (B) and f (C), same location as holotype, MNHN 2014-2729; nuptial colouration pattern for male (D) and female (E)." figureDoi="http://doi.org/10.5281/zenodo.7806298" httpUri="https://zenodo.org/record/7806298/files/figure.png" pageId="19" pageNumber="224">Fig 9C, E</figureCitation>
).
</paragraph>
<caption ID-Table-UUID="DF52664AC75CFFFDFCC9FC21FB23FB95" httpUri="http://table.plazi.org/id/DF52664AC75CFFFDFCC9FC21FB23FB95" pageId="19" pageNumber="224" startId="19.[813,867,952,974]" targetBox="[819,1450,1110,1898]" targetIsTable="true" targetPageId="19" targetType="table">
<paragraph blockId="19.[813,1456,952,1094]" pageId="19" pageNumber="224">
Table V.
<emphasis box="[900,912,953,974]" italics="true" pageId="19" pageNumber="224"></emphasis>
Morphometric data of
<taxonomicName authorityName="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith" authorityYear="2020" box="[1141,1347,952,973]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="224" phylum="Chordata" rank="species" species="fayollarum">
<emphasis box="[1141,1347,952,973]" italics="true" pageId="19" pageNumber="224">Phoxinus fayollarum</emphasis>
</taxonomicName>
(holotype: MNHN 2019-0263; paratypes: MNHN 2014-2729, MNHN 2014- 2732; and additional materials (MNHN 2010-0992, MNHN 2010- 1055, MNHN 2011-0819, MNHN 2011-0910, MNHN 2014-2733, MNHN 2014-02740, MNHN 2014-2745, MNHN 2014-2767; n = 69). The calculations include the holotype.
</paragraph>
</caption>
<paragraph pageId="19" pageNumber="224">
<table box="[819,1450,1110,1898]" gridcols="6" gridrows="23" pageId="19" pageNumber="224">
<tr box="[819,1450,1110,1167]" gridrow="0" pageId="19" pageNumber="224" rowspan-0="1">
<th box="[1064,1153,1110,1167]" gridcol="1" gridrow="0" pageId="19" pageNumber="224">Holotype</th>
<th box="[1168,1450,1110,1167]" colspan="4" colspanRight="3" gridcol="2" gridrow="0" pageId="19" pageNumber="224">Holotype &amp; other individuals Mean Min Max SD</th>
</tr>
<tr box="[819,1450,1179,1201]" gridrow="1" pageId="19" pageNumber="224">
<th box="[819,1050,1179,1201]" gridcol="0" gridrow="1" pageId="19" pageNumber="224">SL (mm)</th>
<td box="[1064,1153,1179,1201]" gridcol="1" gridrow="1" pageId="19" pageNumber="224">58.1</td>
<td box="[1168,1227,1179,1201]" gridcol="2" gridrow="1" pageId="19" pageNumber="224">53.5</td>
<td box="[1265,1307,1179,1201]" gridcol="3" gridrow="1" pageId="19" pageNumber="224">41.0</td>
<td box="[1339,1381,1179,1201]" gridcol="4" gridrow="1" pageId="19" pageNumber="224">73.6</td>
<td box="[1412,1450,1179,1201]" gridcol="5" gridrow="1" pageId="19" pageNumber="224">8.0</td>
</tr>
<tr box="[819,1450,1212,1234]" gridrow="2" pageId="19" pageNumber="224">
<th box="[819,1450,1212,1234]" colspan="6" colspanRight="5" gridcol="0" gridrow="2" pageId="19" pageNumber="224">
<emphasis box="[819,1097,1212,1234]" italics="true" pageId="19" pageNumber="224">In percent of standard length</emphasis>
</th>
</tr>
<tr box="[819,1450,1246,1268]" gridrow="3" pageId="19" pageNumber="224">
<th box="[819,1050,1246,1268]" gridcol="0" gridrow="3" pageId="19" pageNumber="224">Head length</th>
<td box="[1064,1153,1246,1268]" gridcol="1" gridrow="3" pageId="19" pageNumber="224">27.6</td>
<td box="[1168,1227,1246,1268]" gridcol="2" gridrow="3" pageId="19" pageNumber="224">26.0</td>
<td box="[1265,1307,1246,1268]" gridcol="3" gridrow="3" pageId="19" pageNumber="224">23.6</td>
<td box="[1339,1381,1246,1268]" gridcol="4" gridrow="3" pageId="19" pageNumber="224">28.2</td>
<td box="[1412,1450,1246,1268]" gridcol="5" gridrow="3" pageId="19" pageNumber="224">1.0</td>
</tr>
<tr box="[819,1450,1279,1301]" gridrow="4" pageId="19" pageNumber="224">
<th box="[819,1050,1279,1301]" gridcol="0" gridrow="4" pageId="19" pageNumber="224">Predorsal length</th>
<td box="[1064,1153,1279,1301]" gridcol="1" gridrow="4" pageId="19" pageNumber="224">51.9</td>
<td box="[1168,1227,1279,1301]" gridcol="2" gridrow="4" pageId="19" pageNumber="224">55.5</td>
<td box="[1265,1307,1279,1301]" gridcol="3" gridrow="4" pageId="19" pageNumber="224">51.9</td>
<td box="[1339,1381,1279,1301]" gridcol="4" gridrow="4" pageId="19" pageNumber="224">59.1</td>
<td box="[1412,1450,1279,1301]" gridcol="5" gridrow="4" pageId="19" pageNumber="224">1.6</td>
</tr>
<tr box="[819,1450,1312,1334]" gridrow="5" pageId="19" pageNumber="224">
<th box="[819,1050,1312,1334]" gridcol="0" gridrow="5" pageId="19" pageNumber="224">Post dorsal length</th>
<td box="[1064,1153,1312,1334]" gridcol="1" gridrow="5" pageId="19" pageNumber="224">36.2</td>
<td box="[1168,1227,1312,1334]" gridcol="2" gridrow="5" pageId="19" pageNumber="224">35.9</td>
<td box="[1265,1307,1312,1334]" gridcol="3" gridrow="5" pageId="19" pageNumber="224">28.0</td>
<td box="[1339,1381,1312,1334]" gridcol="4" gridrow="5" pageId="19" pageNumber="224">40.0</td>
<td box="[1412,1450,1312,1334]" gridcol="5" gridrow="5" pageId="19" pageNumber="224">2.0</td>
</tr>
<tr box="[819,1450,1345,1367]" gridrow="6" pageId="19" pageNumber="224">
<th box="[819,1050,1345,1367]" gridcol="0" gridrow="6" pageId="19" pageNumber="224">Prepectoral length</th>
<td box="[1064,1153,1345,1367]" gridcol="1" gridrow="6" pageId="19" pageNumber="224">24.6</td>
<td box="[1168,1227,1345,1367]" gridcol="2" gridrow="6" pageId="19" pageNumber="224">26.0</td>
<td box="[1265,1307,1345,1367]" gridcol="3" gridrow="6" pageId="19" pageNumber="224">22.8</td>
<td box="[1339,1381,1345,1367]" gridcol="4" gridrow="6" pageId="19" pageNumber="224">30.3</td>
<td box="[1412,1450,1345,1367]" gridcol="5" gridrow="6" pageId="19" pageNumber="224">1.3</td>
</tr>
<tr box="[819,1450,1378,1400]" gridrow="7" pageId="19" pageNumber="224">
<th box="[819,1050,1378,1400]" gridcol="0" gridrow="7" pageId="19" pageNumber="224">Prepelvic length</th>
<td box="[1064,1153,1378,1400]" gridcol="1" gridrow="7" pageId="19" pageNumber="224">45.1</td>
<td box="[1168,1227,1378,1400]" gridcol="2" gridrow="7" pageId="19" pageNumber="224">48.3</td>
<td box="[1265,1307,1378,1400]" gridcol="3" gridrow="7" pageId="19" pageNumber="224">42.7</td>
<td box="[1339,1381,1378,1400]" gridcol="4" gridrow="7" pageId="19" pageNumber="224">53.4</td>
<td box="[1412,1450,1378,1400]" gridcol="5" gridrow="7" pageId="19" pageNumber="224">1.9</td>
</tr>
<tr box="[819,1450,1411,1434]" gridrow="8" pageId="19" pageNumber="224">
<th box="[819,1050,1411,1434]" gridcol="0" gridrow="8" pageId="19" pageNumber="224">Preanal length</th>
<td box="[1064,1153,1411,1434]" gridcol="1" gridrow="8" pageId="19" pageNumber="224">63.5</td>
<td box="[1168,1227,1411,1434]" gridcol="2" gridrow="8" pageId="19" pageNumber="224">65.3</td>
<td box="[1265,1307,1411,1434]" gridcol="3" gridrow="8" pageId="19" pageNumber="224">61.2</td>
<td box="[1339,1381,1411,1434]" gridcol="4" gridrow="8" pageId="19" pageNumber="224">69.5</td>
<td box="[1412,1450,1411,1434]" gridcol="5" gridrow="8" pageId="19" pageNumber="224">1.9</td>
</tr>
<tr box="[819,1450,1445,1467]" gridrow="9" pageId="19" pageNumber="224">
<th box="[819,1050,1445,1467]" gridcol="0" gridrow="9" pageId="19" pageNumber="224">Body depth</th>
<td box="[1064,1153,1445,1467]" gridcol="1" gridrow="9" pageId="19" pageNumber="224">21.5</td>
<td box="[1168,1227,1445,1467]" gridcol="2" gridrow="9" pageId="19" pageNumber="224">21.3</td>
<td box="[1265,1307,1445,1467]" gridcol="3" gridrow="9" pageId="19" pageNumber="224">15.2</td>
<td box="[1339,1381,1445,1467]" gridcol="4" gridrow="9" pageId="19" pageNumber="224">28.3</td>
<td box="[1412,1450,1445,1467]" gridcol="5" gridrow="9" pageId="19" pageNumber="224">2.3</td>
</tr>
<tr box="[819,1450,1478,1500]" gridrow="10" pageId="19" pageNumber="224">
<th box="[819,1050,1478,1500]" gridcol="0" gridrow="10" pageId="19" pageNumber="224">Distance dorsal anal</th>
<td box="[1064,1153,1478,1500]" gridcol="1" gridrow="10" pageId="19" pageNumber="224">23.7</td>
<td box="[1168,1227,1478,1500]" gridcol="2" gridrow="10" pageId="19" pageNumber="224">22.2</td>
<td box="[1265,1307,1478,1500]" gridcol="3" gridrow="10" pageId="19" pageNumber="224">16.8</td>
<td box="[1339,1381,1478,1500]" gridcol="4" gridrow="10" pageId="19" pageNumber="224">27.0</td>
<td box="[1412,1450,1478,1500]" gridcol="5" gridrow="10" pageId="19" pageNumber="224">2.0</td>
</tr>
<tr box="[819,1450,1511,1533]" gridrow="11" pageId="19" pageNumber="224">
<th box="[819,1050,1511,1533]" gridcol="0" gridrow="11" pageId="19" pageNumber="224">Distance pectoral pelvic</th>
<td box="[1064,1153,1511,1533]" gridcol="1" gridrow="11" pageId="19" pageNumber="224">17.1</td>
<td box="[1168,1227,1511,1533]" gridcol="2" gridrow="11" pageId="19" pageNumber="224">20.8</td>
<td box="[1265,1307,1511,1533]" gridcol="3" gridrow="11" pageId="19" pageNumber="224">16.2</td>
<td box="[1339,1381,1511,1533]" gridcol="4" gridrow="11" pageId="19" pageNumber="224">25.0</td>
<td box="[1412,1450,1511,1533]" gridcol="5" gridrow="11" pageId="19" pageNumber="224">2.2</td>
</tr>
<tr box="[819,1450,1544,1566]" gridrow="12" pageId="19" pageNumber="224">
<th box="[819,1050,1544,1566]" gridcol="0" gridrow="12" pageId="19" pageNumber="224">Pectoral fin length</th>
<td box="[1064,1153,1544,1566]" gridcol="1" gridrow="12" pageId="19" pageNumber="224">18.5</td>
<td box="[1168,1227,1544,1566]" gridcol="2" gridrow="12" pageId="19" pageNumber="224">18.1</td>
<td box="[1265,1307,1544,1566]" gridcol="3" gridrow="12" pageId="19" pageNumber="224">14.2</td>
<td box="[1339,1381,1544,1566]" gridcol="4" gridrow="12" pageId="19" pageNumber="224">21.4</td>
<td box="[1412,1450,1544,1566]" gridcol="5" gridrow="12" pageId="19" pageNumber="224">1.6</td>
</tr>
<tr box="[819,1450,1577,1600]" gridrow="13" pageId="19" pageNumber="224">
<th box="[819,1050,1577,1600]" gridcol="0" gridrow="13" pageId="19" pageNumber="224">Pelvic fin length</th>
<td box="[1064,1153,1577,1600]" gridcol="1" gridrow="13" pageId="19" pageNumber="224">15.7</td>
<td box="[1168,1227,1577,1600]" gridcol="2" gridrow="13" pageId="19" pageNumber="224">14.8</td>
<td box="[1265,1307,1577,1600]" gridcol="3" gridrow="13" pageId="19" pageNumber="224">12.1</td>
<td box="[1339,1381,1577,1600]" gridcol="4" gridrow="13" pageId="19" pageNumber="224">18.3</td>
<td box="[1412,1450,1577,1600]" gridcol="5" gridrow="13" pageId="19" pageNumber="224">1.4</td>
</tr>
<tr box="[819,1450,1611,1633]" gridrow="14" pageId="19" pageNumber="224">
<th box="[819,1050,1611,1633]" gridcol="0" gridrow="14" pageId="19" pageNumber="224">Dorsal fin length</th>
<td box="[1064,1153,1611,1633]" gridcol="1" gridrow="14" pageId="19" pageNumber="224">21.8</td>
<td box="[1168,1227,1611,1633]" gridcol="2" gridrow="14" pageId="19" pageNumber="224">20.3</td>
<td box="[1265,1307,1611,1633]" gridcol="3" gridrow="14" pageId="19" pageNumber="224">16.4</td>
<td box="[1339,1381,1611,1633]" gridcol="4" gridrow="14" pageId="19" pageNumber="224">22.9</td>
<td box="[1412,1450,1611,1633]" gridcol="5" gridrow="14" pageId="19" pageNumber="224">1.3</td>
</tr>
<tr box="[819,1450,1644,1666]" gridrow="15" pageId="19" pageNumber="224">
<th box="[819,1050,1644,1666]" gridcol="0" gridrow="15" pageId="19" pageNumber="224">Anal fin length</th>
<td box="[1064,1153,1644,1666]" gridcol="1" gridrow="15" pageId="19" pageNumber="224">19.3</td>
<td box="[1168,1227,1644,1666]" gridcol="2" gridrow="15" pageId="19" pageNumber="224">19.9</td>
<td box="[1265,1307,1644,1666]" gridcol="3" gridrow="15" pageId="19" pageNumber="224">16.6</td>
<td box="[1339,1381,1644,1666]" gridcol="4" gridrow="15" pageId="19" pageNumber="224">23.5</td>
<td box="[1412,1450,1644,1666]" gridcol="5" gridrow="15" pageId="19" pageNumber="224">1.4</td>
</tr>
<tr box="[819,1450,1677,1699]" gridrow="16" pageId="19" pageNumber="224">
<th box="[819,1050,1677,1699]" gridcol="0" gridrow="16" pageId="19" pageNumber="224">Caudal peduncle length</th>
<td box="[1064,1153,1677,1699]" gridcol="1" gridrow="16" pageId="19" pageNumber="224">25.9</td>
<td box="[1168,1227,1677,1699]" gridcol="2" gridrow="16" pageId="19" pageNumber="224">25.5</td>
<td box="[1265,1307,1677,1699]" gridcol="3" gridrow="16" pageId="19" pageNumber="224">20.9</td>
<td box="[1339,1381,1677,1699]" gridcol="4" gridrow="16" pageId="19" pageNumber="224">28.9</td>
<td box="[1412,1450,1677,1699]" gridcol="5" gridrow="16" pageId="19" pageNumber="224">1.6</td>
</tr>
<tr box="[819,1450,1710,1732]" gridrow="17" pageId="19" pageNumber="224">
<th box="[819,1050,1710,1732]" gridcol="0" gridrow="17" pageId="19" pageNumber="224">Caudal peduncle depth</th>
<td box="[1064,1153,1710,1732]" gridcol="1" gridrow="17" pageId="19" pageNumber="224">9.4</td>
<td box="[1168,1227,1710,1732]" gridcol="2" gridrow="17" pageId="19" pageNumber="224">9.1</td>
<td box="[1265,1307,1710,1732]" gridcol="3" gridrow="17" pageId="19" pageNumber="224">6.9</td>
<td box="[1339,1381,1710,1732]" gridcol="4" gridrow="17" pageId="19" pageNumber="224">11.2</td>
<td box="[1412,1450,1710,1732]" gridcol="5" gridrow="17" pageId="19" pageNumber="224">0.8</td>
</tr>
<tr box="[819,1450,1744,1765]" gridrow="18" pageId="19" pageNumber="224">
<th box="[819,1450,1744,1765]" colspan="6" colspanRight="5" gridcol="0" gridrow="18" pageId="19" pageNumber="224">
<emphasis box="[819,1059,1744,1765]" italics="true" pageId="19" pageNumber="224">In percent of head length</emphasis>
</th>
</tr>
<tr box="[819,1450,1777,1799]" gridrow="19" pageId="19" pageNumber="224">
<th box="[819,1050,1777,1799]" gridcol="0" gridrow="19" pageId="19" pageNumber="224">Head depth</th>
<td box="[1064,1153,1777,1799]" gridcol="1" gridrow="19" pageId="19" pageNumber="224">50.0</td>
<td box="[1168,1227,1777,1799]" gridcol="2" gridrow="19" pageId="19" pageNumber="224">52.0</td>
<td box="[1265,1307,1777,1799]" gridcol="3" gridrow="19" pageId="19" pageNumber="224">45.6</td>
<td box="[1339,1381,1777,1799]" gridcol="4" gridrow="19" pageId="19" pageNumber="224">57.7</td>
<td box="[1412,1450,1777,1799]" gridcol="5" gridrow="19" pageId="19" pageNumber="224">2.9</td>
</tr>
<tr box="[819,1450,1810,1832]" gridrow="20" pageId="19" pageNumber="224">
<th box="[819,1050,1810,1832]" gridcol="0" gridrow="20" pageId="19" pageNumber="224">Eye diameter</th>
<td box="[1064,1153,1810,1832]" gridcol="1" gridrow="20" pageId="19" pageNumber="224">22.3</td>
<td box="[1168,1227,1810,1832]" gridcol="2" gridrow="20" pageId="19" pageNumber="224">26.4</td>
<td box="[1265,1307,1810,1832]" gridcol="3" gridrow="20" pageId="19" pageNumber="224">21.5</td>
<td box="[1339,1381,1810,1832]" gridcol="4" gridrow="20" pageId="19" pageNumber="224">32.3</td>
<td box="[1412,1450,1810,1832]" gridcol="5" gridrow="20" pageId="19" pageNumber="224">2.6</td>
</tr>
<tr box="[819,1450,1843,1865]" gridrow="21" pageId="19" pageNumber="224">
<th box="[819,1050,1843,1865]" gridcol="0" gridrow="21" pageId="19" pageNumber="224">Snout length</th>
<td box="[1064,1153,1843,1865]" gridcol="1" gridrow="21" pageId="19" pageNumber="224">30.9</td>
<td box="[1168,1227,1843,1865]" gridcol="2" gridrow="21" pageId="19" pageNumber="224">29.0</td>
<td box="[1265,1307,1843,1865]" gridcol="3" gridrow="21" pageId="19" pageNumber="224">18.8</td>
<td box="[1339,1381,1843,1865]" gridcol="4" gridrow="21" pageId="19" pageNumber="224">36.8</td>
<td box="[1412,1450,1843,1865]" gridcol="5" gridrow="21" pageId="19" pageNumber="224">2.9</td>
</tr>
<tr box="[819,1450,1876,1898]" gridrow="22" pageId="19" pageNumber="224">
<th box="[819,1050,1876,1898]" gridcol="0" gridrow="22" pageId="19" pageNumber="224">Inter-orbital width</th>
<td box="[1064,1153,1876,1898]" gridcol="1" gridrow="22" pageId="19" pageNumber="224">32.1</td>
<td box="[1168,1227,1876,1898]" gridcol="2" gridrow="22" pageId="19" pageNumber="224">32.5</td>
<td box="[1265,1307,1876,1898]" gridcol="3" gridrow="22" pageId="19" pageNumber="224">26.1</td>
<td box="[1339,1381,1876,1898]" gridcol="4" gridrow="22" pageId="19" pageNumber="224">38.3</td>
<td box="[1412,1450,1876,1898]" gridcol="5" gridrow="22" pageId="19" pageNumber="224">2.7</td>
</tr>
</table>
</paragraph>
<paragraph blockId="19.[132,775,922,1907]" pageId="19" pageNumber="224">
<taxonomicName authorityName="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith" authorityYear="2020" box="[170,418,1274,1298]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="224" phylum="Chordata" rank="species" species="fayollarum">
<emphasis box="[170,418,1274,1298]" italics="true" pageId="19" pageNumber="224">Phoxinus fayollarum</emphasis>
</taxonomicName>
is further distinguished from
<taxonomicName authorityName="Kottelat" authorityYear="2007" box="[132,234,1306,1330]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="224" phylum="Chordata" rank="species" species="bigerri">
<emphasis box="[132,234,1306,1330]" italics="true" pageId="19" pageNumber="224">P. bigerri</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Hanko" authorityYear="1922" box="[287,370,1306,1330]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="224" phylum="Chordata" rank="species" species="csikii">
<emphasis box="[287,370,1306,1330]" italics="true" pageId="19" pageNumber="224">P. csikii</emphasis>
</taxonomicName>
by a straight to slightly concave anal fin margin (vs. straight to convex), from
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[597,727,1338,1362]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="224" phylum="Chordata" rank="species" species="phoxinus">
<emphasis box="[597,727,1338,1362]" italics="true" pageId="19" pageNumber="224">P. phoxinus</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Kottelat" authorityYear="2007" box="[132,288,1371,1394]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="224" phylum="Chordata" rank="species" species="septimaniae">
<emphasis box="[132,288,1371,1394]" italics="true" pageId="19" pageNumber="224">P. septimaniae</emphasis>
</taxonomicName>
by a stout and blunt snout (vs. slightly point- ed, and two scaled patches at the pectoral fin bases separat- ed by a nonscaled area (vs. scaled breath between the pectoral fin insertions in some populations of
<taxonomicName authorityName="Kottelat" authorityYear="2007" box="[599,759,1467,1490]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="224" phylum="Chordata" rank="species" species="septimaniae">
<emphasis box="[599,759,1467,1490]" italics="true" pageId="19" pageNumber="224">P. septimaniae</emphasis>
</taxonomicName>
), and from
<taxonomicName authorityName="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith" authorityYear="2020" box="[238,373,1498,1522]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="224" phylum="Chordata" rank="species" species="dragarum">
<emphasis box="[238,373,1498,1522]" italics="true" pageId="19" pageNumber="224">P. dragarum</emphasis>
</taxonomicName>
by an incomplete lateral line usually reaching to the caudal peduncle (vs. usually reaching to the anal fin base or just slightly behind the last anal fin ray), and 56-84 pored scales (vs. 50-74).
</paragraph>
<paragraph blockId="19.[132,775,922,1907]" lastBlockId="19.[812,1455,217,306]" pageId="19" pageNumber="224">
The nuptial male
<taxonomicName authorityName="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith" authorityYear="2020" box="[364,512,1626,1650]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="224" phylum="Chordata" rank="species" species="fayollarum">
<emphasis box="[364,512,1626,1650]" italics="true" pageId="19" pageNumber="224">P. fayollarum</emphasis>
</taxonomicName>
is further distinguished by its clearly distinct green bars crossing Z3 and Z4 (vs. indistinct in
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[268,393,1690,1714]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="224" phylum="Chordata" rank="species" species="phoxinus">
<emphasis box="[268,393,1690,1714]" italics="true" pageId="19" pageNumber="224">P. phoxinus</emphasis>
</taxonomicName>
), the red belly not extending to the caudal peduncle (vs. extending in
<taxonomicName authorityName="Hanko" authorityYear="1922" box="[501,584,1722,1746]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="224" phylum="Chordata" rank="species" species="csikii">
<emphasis box="[501,584,1722,1746]" italics="true" pageId="19" pageNumber="224">P. csikii</emphasis>
</taxonomicName>
), having red pectoral, pelvic and anal fins bases (
<emphasis box="[481,503,1755,1778]" italics="true" pageId="19" pageNumber="224">vs</emphasis>
. pinkish in
<taxonomicName authorityName="Kottelat" authorityYear="2007" box="[629,729,1754,1778]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="224" phylum="Chordata" rank="species" species="bigerri">
<emphasis box="[629,729,1754,1778]" italics="true" pageId="19" pageNumber="224">P. bigerri</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[132,258,1787,1811]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="224" phylum="Chordata" rank="species" species="phoxinus">
<emphasis box="[132,258,1787,1811]" italics="true" pageId="19" pageNumber="224">P. phoxinus</emphasis>
</taxonomicName>
), orange pectoral, pelvic and anal fins (vs. yel- low in
<taxonomicName authorityName="Hanko" authorityYear="1922" box="[210,296,1819,1843]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="224" phylum="Chordata" rank="species" species="csikii">
<emphasis box="[210,296,1819,1843]" italics="true" pageId="19" pageNumber="224">P. csikii</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith" authorityYear="2020" box="[351,488,1819,1843]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="224" phylum="Chordata" rank="species" species="dragarum">
<emphasis box="[351,488,1819,1843]" italics="true" pageId="19" pageNumber="224">P. dragarum</emphasis>
</taxonomicName>
), no red disc on the inner side of the pectoral fin (presence in
<taxonomicName authorityName="Kottelat" authorityYear="2007" box="[517,673,1851,1874]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="224" phylum="Chordata" rank="species" species="septimaniae">
<emphasis box="[517,673,1851,1874]" italics="true" pageId="19" pageNumber="224">P. septimaniae</emphasis>
</taxonomicName>
), and the absence of a yellow dot between the mouth and the eye (vs. the presence in
<taxonomicName authorityName="Kottelat" authorityYear="2007" box="[978,1133,218,241]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="224" phylum="Chordata" rank="species" species="septimaniae">
<emphasis box="[978,1133,218,241]" italics="true" pageId="19" pageNumber="224">P. septimaniae</emphasis>
</taxonomicName>
). The nuptial female is distinguished by a white Z5 (vs. black in
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1198,1322,249,273]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="224" phylum="Chordata" rank="species" species="phoxinus">
<emphasis box="[1198,1322,249,273]" italics="true" pageId="19" pageNumber="224">P. phoxinus</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Kottelat" authorityYear="2007" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="224" phylum="Chordata" rank="species" species="septimaniae">
<emphasis italics="true" pageId="19" pageNumber="224">P. septimaniae</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith" authorityYear="2020" box="[909,1041,281,305]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="19" pageNumber="224" phylum="Chordata" rank="species" species="dragarum">
<emphasis box="[909,1041,281,305]" italics="true" pageId="19" pageNumber="224">P. dragarum</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="20" lastPageNumber="225" pageId="19" pageNumber="224" type="description">
<paragraph blockId="19.[813,1456,345,918]" box="[813,945,345,369]" pageId="19" pageNumber="224">
<heading bold="true" box="[813,945,345,369]" fontSize="10" level="3" pageId="19" pageNumber="224" reason="6">
<emphasis bold="true" box="[813,945,345,369]" pageId="19" pageNumber="224">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="19.[813,1456,345,918]" pageId="19" pageNumber="224">Morphometric data is shown in table V. Largest examined specimen 73.6 mm SL. Body moderately elongated, dorsal and ventral profile about symmetric or ventral profile slightly more convex. Dorsal head profile markedly more convex than ventral one. Caudal peduncle length 2.2-3.5 times in its depth. Snout stout and blunt. Mouth slightly subterminal, upper lip and snout projecting beyond tip of lower jaw. Tip of upper lip at level or above level of lowest point of eye. In lateral view, eye not or almost flushed with dorsal profile. No supraorbital ridge.</paragraph>
<paragraph blockId="19.[813,1456,345,918]" lastBlockId="20.[132,775,216,509]" lastPageId="20" lastPageNumber="225" pageId="19" pageNumber="224">Dorsal fin with 3 simple and 6½-8½ branched rays. Dis- tal dorsal-fin margin markedly convex, second branched ray longest, dorsal-fin origin behind pelvicfin insertion. Pectoral fin with 1 simple and 11-16 branched rays (usually 14-15), reaching to pelvicfin origin in male, not reaching in female. Pelvicfin with 1 simple and 6-9 branched rays (mainly 7-8), reaching slightly beyond anal fin origin in male, not reaching in female. Anal fin with 3 simple and 6½-8½ branched rays. Anal fin origin slightly in front of vertical through dorsal-fin origin. Anal fin margin straight to slightly concave, second branched ray longest.</paragraph>
<paragraph blockId="20.[132,775,216,509]" pageId="20" pageNumber="225">Lateral line incomplete reaching caudal peduncle. 72-91 lateral scales, including 56-84 pored scales. Two scaled patches at pectoral fin origin separated by an unscaled area. No scale between pectoral fin bases. Nuptial tubercles present in both sexes, smaller in female.</paragraph>
<paragraph blockId="20.[132,776,550,1143]" box="[132,354,550,574]" pageId="20" pageNumber="225">
<heading bold="true" box="[132,354,550,574]" fontSize="10" level="3" pageId="20" pageNumber="225" reason="6">
<emphasis bold="true" box="[132,354,550,574]" pageId="20" pageNumber="225">Nuptial colouration</emphasis>
</heading>
</paragraph>
<paragraph blockId="20.[132,776,550,1143]" pageId="20" pageNumber="225">
<emphasis box="[170,226,584,608]" italics="true" pageId="20" pageNumber="225">Male</emphasis>
. Snout and top of head dark brown. Ablack bar on operculum reaching from top of head to branchiostegal rays. Cheeks white. Lips dark pink or red. Large yellow spot on operculum almost covering whole opercular and a white spot above at upper corner. Z1 brown to black. Z2 pale brown. Flank bars crossing Z1 to Z4, flank bars black from Z1 to Z2 and green from Z3 to Z4. Z4 bright yellow. Ablack line along Z5 between pectoral fin base and caudal peduncle, from pelvic fin base to caudal peduncle in some individuals. Pectoral, pelvic and anal fins yellow-orange. Pectoral, pel- vic and anal fin bases red. (
<figureCitation box="[420,536,918,942]" captionStart="Figure 9" captionStartId="18.[813,878,1309,1331]" captionTargetBox="[813,1456,215,1292]" captionTargetId="figure-483@18.[813,1456,215,1292]" captionTargetPageId="18" captionText="Figure 9. Phoxinus fayollarum Holotype m (A), 58.13 mm SL from the Boron Stream, Pionsat 46°06.4N, 2°41.1E, MNHN 2019-0263; paratypes m (B) and f (C), same location as holotype, MNHN 2014-2729; nuptial colouration pattern for male (D) and female (E)." figureDoi="http://doi.org/10.5281/zenodo.7806298" httpUri="https://zenodo.org/record/7806298/files/figure.png" pageId="20" pageNumber="225">Fig. 9B, D</figureCitation>
). Female: Nuptial col- ouration pattern similar to male but paler with black bars in Z1 to Z2, a green stripe along Z3, Z4 yellow-pinkish, and Z5 white. Lips pinkish and belly red. Presence of “male” characters in the female: white and yellow spots on operculum and red spots at pectoral, pelvic and anal fin bases (
<figureCitation captionStart="Figure 9" captionStartId="18.[813,878,1309,1331]" captionTargetBox="[813,1456,215,1292]" captionTargetId="figure-483@18.[813,1456,215,1292]" captionTargetPageId="18" captionText="Figure 9. Phoxinus fayollarum Holotype m (A), 58.13 mm SL from the Boron Stream, Pionsat 46°06.4N, 2°41.1E, MNHN 2019-0263; paratypes m (B) and f (C), same location as holotype, MNHN 2014-2729; nuptial colouration pattern for male (D) and female (E)." figureDoi="http://doi.org/10.5281/zenodo.7806298" httpUri="https://zenodo.org/record/7806298/files/figure.png" pageId="20" pageNumber="225">Fig 9C, E</figureCitation>
).
</paragraph>
<paragraph blockId="20.[132,775,1184,1342]" box="[132,447,1184,1208]" pageId="20" pageNumber="225">
<heading bold="true" box="[132,447,1184,1208]" fontSize="10" level="3" pageId="20" pageNumber="225" reason="6">
<emphasis bold="true" box="[132,447,1184,1208]" pageId="20" pageNumber="225">Colouration in preservation</emphasis>
</heading>
</paragraph>
<paragraph blockId="20.[132,775,1184,1342]" pageId="20" pageNumber="225">
Dark green, brown and black bars and dots remain after preservation in ethanol. Green colouration turns to dark brown, whereas yellow and red colourations are faded (
<figureCitation box="[140,225,1318,1342]" captionStart="Figure 9" captionStartId="18.[813,878,1309,1331]" captionTargetBox="[813,1456,215,1292]" captionTargetId="figure-483@18.[813,1456,215,1292]" captionTargetPageId="18" captionText="Figure 9. Phoxinus fayollarum Holotype m (A), 58.13 mm SL from the Boron Stream, Pionsat 46°06.4N, 2°41.1E, MNHN 2019-0263; paratypes m (B) and f (C), same location as holotype, MNHN 2014-2729; nuptial colouration pattern for male (D) and female (E)." figureDoi="http://doi.org/10.5281/zenodo.7806298" httpUri="https://zenodo.org/record/7806298/files/figure.png" pageId="20" pageNumber="225">Fig. 9A</figureCitation>
).
</paragraph>
<paragraph blockId="20.[132,775,1383,1608]" box="[132,440,1383,1407]" pageId="20" pageNumber="225">
<heading bold="true" box="[132,440,1383,1407]" fontSize="10" level="3" pageId="20" pageNumber="225" reason="6">
<emphasis bold="true" box="[132,440,1383,1407]" pageId="20" pageNumber="225">Molecular characterisation</emphasis>
</heading>
</paragraph>
<paragraph blockId="20.[132,775,1383,1608]" pageId="20" pageNumber="225">
On the COI marker,
<taxonomicName authorityName="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith" authorityYear="2020" box="[386,528,1417,1441]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="225" phylum="Chordata" rank="species" species="fayollarum">
<emphasis box="[386,528,1417,1441]" italics="true" pageId="20" pageNumber="225">P. fayollarum</emphasis>
</taxonomicName>
has six diagnostic sites including two synapomorphies (Tab. II). For the 12S rDNA marker, 6 diagnostic sites characterise this species: C (vs. A or G) in position 122, C (
<emphasis box="[407,429,1518,1541]" italics="true" pageId="20" pageNumber="225">vs</emphasis>
. T) in position 357, T (vs. C) in position 454, C (vs. T) in position 649, G (vs. A) in position 675 and A (vs. G) in position 874.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="225" type="distribution">
<paragraph blockId="20.[132,775,1649,1908]" box="[132,271,1649,1673]" pageId="20" pageNumber="225">
<heading bold="true" box="[132,271,1649,1673]" fontSize="10" level="3" pageId="20" pageNumber="225" reason="6">
<emphasis bold="true" box="[132,271,1649,1673]" pageId="20" pageNumber="225">Distribution</emphasis>
</heading>
</paragraph>
<paragraph blockId="20.[132,775,1649,1908]" pageId="20" pageNumber="225">
This species is endemic to the
<collectingRegion box="[503,562,1683,1707]" country="France" name="Pays de la Loire" pageId="20" pageNumber="225">Loire</collectingRegion>
drainage as well as the Sèvre Niortaise catchment (
<figureCitation box="[489,556,1716,1740]" captionStart="Figure 1" captionStartId="3.[132,197,1741,1763]" captionTargetBox="[548,1456,1050,1902]" captionTargetId="figure-410@3.[548,1455,1051,1902]" captionTargetPageId="3" captionText="Figure 1. Records of Phoxinus used in this study. Coloured squares represent the six species delineated in this study: P. phoxinus (blue), P. bigerri (red), P. csikii (green), P. septimaniae (yellow), P. dragarum (purple), and P. fayollarum (orange)." figureDoi="http://doi.org/10.5281/zenodo.7806282" httpUri="https://zenodo.org/record/7806282/files/figure.png" pageId="20" pageNumber="225">Fig. 1</figureCitation>
). It has been introduced in Garonne and Rhône drainages adjacent to the
<collectingRegion box="[717,775,1750,1774]" country="France" name="Pays de la Loire" pageId="20" pageNumber="225">Loire</collectingRegion>
catchment, for example in the stream Boralde de Flaujac and in the Chassezac River, Grevieres (
<bibRefCitation author="GEIGER M. F. &amp; HERDER F. &amp; MONAGHAN M. T. &amp; ALMADA V. &amp; BARBIERI R. &amp; BARICHE M. &amp; BERREBI P. &amp; BOHLEN J. &amp; CASAL-LOPEZ M. &amp; DELMASTRO G. B. &amp; DENYS G. P. J. &amp; DETTAI A. &amp; DOADRIO I., KALOGIANNI E. &amp; KARST H. &amp; KOTTELAT M. &amp; KOVACIC M. &amp; LAPORTE M. &amp; LORENZONI M. &amp; MARCIC Z. &amp; OZULUG M. &amp; PERDICES A. &amp; PEREA S. &amp; PERSAT H. &amp; PORCELOTTI S. &amp; PUZZI C. &amp; ROBALO J. &amp; SANDA R. &amp; SCHNEIDER M. &amp; SLECHTOVA V. &amp; STOUM- BOUDI M. &amp; WALTER S. &amp; FREYHOF J." box="[554,771,1817,1841]" journalOrPublisher="Mol. Ecol. Res." pageId="20" pageNumber="225" pagination="1210 - 1221" part="14" refId="ref25514" refString="GEIGER M. F., HERDER F., MONAGHAN M. T., ALMADA V., BARBIERI R., BARICHE M., BERREBI P., BOHLEN J., CASAL-LOPEZ M., DELMASTRO G. B., DENYS G. P. J., DETTAI A., DOADRIO I., KALOGIANNI E., KARST H., KOTTELAT M., KOVACIC M., LAPORTE M., LORENZONI M., MARCIC Z., OZULUG M., PERDICES A., PEREA S., PERSAT H., PORCELOTTI S., PUZZI C., ROBALO J., SANDA R., SCHNEIDER M., SLECHTOVA V., STOUM- BOUDI M., WALTER S. &amp; FREYHOF J., 2014. - Spatial heterogeneity in the Mediterranean Biodiversity Hotspot affects barcoding accuracy of its freshwater fishes. Mol. Ecol. Res., 14 (6): 1210 - 1221. DOI: 10.1111 / 1755 - 0998.12257" title="Spatial heterogeneity in the Mediterranean Biodiversity Hotspot affects barcoding accuracy of its freshwater fishes" type="journal article" year="2014">
Geiger
<emphasis box="[636,690,1817,1841]" italics="true" pageId="20" pageNumber="225">et al.</emphasis>
(2014)
</bibRefCitation>
. This lineage was named
<taxonomicName box="[393,529,1850,1874]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="225" phylum="Chordata" rank="species" species="undetermined">
<emphasis box="[393,529,1850,1874]" italics="true" pageId="20" pageNumber="225">Phoxinus sp.</emphasis>
</taxonomicName>
16 by
<bibRefCitation author="PALANDACIC A. &amp; NASEKA A. &amp; RAMLER D. &amp; AHNELT H." journalOrPublisher="BMC Evol. Biol." pageId="20" pageNumber="225" pagination="184" part="17" refId="ref27292" refString="PALANDACIC A., NASEKA A., RAMLER D. &amp; AHNELT H., 2017. - Contrasting morphology with molecular data: an approach to revision of species complexes based on the example of European Phoxinus (Cyprinidae). BMC Evol. Biol., 17: 184. DOI: 10.1186 / s 12862 - 017 - 1032 - x" title="Contrasting morphology with molecular data: an approach to revision of species complexes based on the example of European Phoxinus (Cyprinidae)" type="journal article" year="2017">
Palandačić
<emphasis box="[722,775,1850,1874]" italics="true" pageId="20" pageNumber="225">et al.</emphasis>
(2017)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="225" type="biology_ecology">
<paragraph blockId="20.[813,1456,218,434]" box="[813,903,218,242]" pageId="20" pageNumber="225">
<heading bold="true" box="[813,903,218,242]" fontSize="10" level="3" pageId="20" pageNumber="225" reason="6">
<emphasis bold="true" box="[813,903,218,242]" pageId="20" pageNumber="225">Ecology</emphasis>
</heading>
</paragraph>
<paragraph blockId="20.[813,1456,218,434]" pageId="20" pageNumber="225">
This species is common in the whole
<collectingRegion box="[1281,1343,250,274]" country="France" name="Pays de la Loire" pageId="20" pageNumber="225">Loire</collectingRegion>
drainage, and is especially abundant in the greyling zone (
<bibRefCitation author="LASNE E. &amp; BERGEROT B. &amp; LEK S. &amp; LAFFAILLE P." journalOrPublisher="River Res. Appl." pageId="20" pageNumber="225" pagination="877 - 890" part="23" refId="ref26417" refString="LASNE E., BERGEROT B., LEK S. &amp; LAFFAILLE P., 2007. - Fish zonation and indicator species for the evaluation of the ecological status of rivers: example of the Loire basin (France). River Res. Appl., 23: 877 - 890. DOI: 10.1002 / rra. 1030" title="Fish zonation and indicator species for the evaluation of the ecological status of rivers: example of the Loire basin (France)" type="journal article" year="2007">
Lasne
<emphasis box="[1395,1455,282,306]" italics="true" pageId="20" pageNumber="225">et al.,</emphasis>
2007
</bibRefCitation>
). It inhabits mostly clear and shallow streams with a slow discharge and a sandy or gravely ground. The spawing period is from April to May (
<bibRefCitation author="MAUDUYT F." box="[1134,1305,378,402]" journalOrPublisher="Bull. Soc. Acad. Agric., Belles-Lett., Sci. Arts Poitiers" pageId="20" pageNumber="225" pagination="8 - 49" part="3" refId="ref26906" refString="MAUDUYT F., 1848. - Ichtyologie de la Vienne, ou tableau methodique et descriptif des poissons qui vivent habituellement dans les eaux de ce departement ou qui y remontent periodiquement ou accidentellement. Bull. Soc. Acad. Agric., Belles-Lett., Sci. Arts Poitiers, 3: 8 - 49." title="Ichtyologie de la Vienne, ou tableau methodique et descriptif des poissons qui vivent habituellement dans les eaux de ce departement ou qui y remontent periodiquement ou accidentellement" type="journal article" year="1848">Mauduyt, 1848</bibRefCitation>
) and in June- July in higher altitudes (Denys, pers. obs., 2014).
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="225" type="etymology">
<paragraph blockId="20.[813,1455,474,691]" box="[813,936,474,498]" pageId="20" pageNumber="225">
<heading bold="true" box="[813,936,474,498]" fontSize="10" level="3" pageId="20" pageNumber="225" reason="6">
<emphasis bold="true" box="[813,936,474,498]" pageId="20" pageNumber="225">Etymology</emphasis>
</heading>
</paragraph>
<paragraph blockId="20.[813,1455,474,691]" pageId="20" pageNumber="225">
The species name
<taxonomicName authorityName="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith" authorityYear="2020" box="[1043,1160,506,530]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="225" phylum="Chordata" rank="species" species="fayollarum">
<emphasis box="[1043,1160,506,530]" italics="true" pageId="20" pageNumber="225">fayollarum</emphasis>
</taxonomicName>
is the genitive of a latinized name “fayolles”, fairies from of the
<collectingRegion box="[1210,1394,538,562]" country="France" name="Auvergne" pageId="20" pageNumber="225">Auvergne region</collectingRegion>
folklore, in the headwater of
<collectingRegion box="[1090,1149,570,594]" country="France" name="Pays de la Loire" pageId="20" pageNumber="225">Loire</collectingRegion>
drainage. Fayolles are fairies who create spring waters. And we owe to one of them, called “Grande Fayolle”, the thermal waters of Vichy (
<bibRefCitation author="ELY R. &amp; TSAAG VALREN A." bookContentInfo="313 p." journalOrPublisher="Bibliotheque du Merveilleux" pageId="20" pageNumber="225" refId="ref25117" refString="ELY R. &amp; TSAAG VALREN A., 2013. - Bestiaire fantastique et creatures feeriques de France. Bibliotheque du Merveilleux. 313 p. Dinan: Terre de Brume." title="Bestiaire fantastique et creatures feeriques de France" type="book" year="2013">Ely and Tsaag Valren, 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="225" type="vernacular_names">
<paragraph blockId="20.[813,1383,730,786]" box="[813,1008,730,754]" pageId="20" pageNumber="225">
<heading bold="true" box="[813,1008,730,754]" fontSize="10" level="3" pageId="20" pageNumber="225" reason="6">
<emphasis bold="true" box="[813,1008,730,754]" pageId="20" pageNumber="225">Vernacular name</emphasis>
</heading>
</paragraph>
<paragraph blockId="20.[813,1383,730,786]" box="[850,1383,762,786]" pageId="20" pageNumber="225">
<collectingRegion box="[850,908,762,786]" country="France" name="Pays de la Loire" pageId="20" pageNumber="225">Loire</collectingRegion>
minnow (English), Vairon ligérien (French).
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="225" type="discussion">
<paragraph blockId="20.[813,1455,826,1235]" box="[813,1038,826,850]" pageId="20" pageNumber="225">
<heading bold="true" box="[813,1038,826,850]" fontSize="10" level="3" pageId="20" pageNumber="225" reason="6">
<emphasis bold="true" box="[813,1038,826,850]" pageId="20" pageNumber="225">Nomenclatural note</emphasis>
</heading>
</paragraph>
<paragraph blockId="20.[813,1455,826,1235]" pageId="20" pageNumber="225">
<bibRefCitation author="DE LA PYLAIE J. M." box="[850,1115,858,882]" editor="Saurin F. A." pageId="20" pageNumber="225" pagination="524 - 534" part="1834" refId="ref24811" refString="DE LA PYLAIE J. M., 1835. - Recherches, en France, sur les pois- sons de l'Ocean, pendant les annees 1832 et 1833. In: Congres scientifique de France, seconde session tenue a Poitiers, en sep- tembre 1834 (Saurin F. A., ed.), pp. 524 - 534. Poitiers." title="Recherches, en France, sur les pois- sons de l'Ocean, pendant les annees 1832 et 1833" type="book" volumeTitle="Congres scientifique de France, seconde session tenue a Poitiers" year="1835">De la Pylaie (1835: 533)</bibRefCitation>
described
<taxonomicName authorityName="De la Pylaie" authorityYear="1835" box="[1231,1426,858,882]" class="Actinopterygii" family="Cyprinidae" genus="Leuciscus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="225" phylum="Chordata" rank="species" species="obtusus">
<emphasis box="[1231,1426,858,882]" italics="true" pageId="20" pageNumber="225">Leuciscus obtusus</emphasis>
</taxonomicName>
as a little leuciscid or small minnow without spots (“Able ou petit Verdon sans taches” in French) in the Vendée department (so possibly in the Loire drainage). This name is a
<emphasis box="[813,963,986,1010]" italics="true" pageId="20" pageNumber="225">nomen nudum</emphasis>
and potentially synonym of
<taxonomicName authority="(Kottelat, 1997)" baseAuthorityName="Kottelat" baseAuthorityYear="1997" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="225" phylum="Chordata" rank="species" species="phoxinus">
<emphasis box="[1268,1391,986,1010]" italics="true" pageId="20" pageNumber="225">P. phoxinus</emphasis>
(
<bibRefCitation author="KOTTELAT M." journalOrPublisher="Biol. Bratislava" pageId="20" pageNumber="225" pagination="1 - 271" part="52" refId="ref26148" refString="KOTTELAT M., 1997. - European freshwater fishes. An heuristic checklist of the freshwater fishes in Europe (exclusive of former USSR), with an introduction for non-systematists and comments on nomenclature and conservation. Biol. Bratislava, 52 (5): 1 - 271." title="European freshwater fishes. An heuristic checklist of the freshwater fishes in Europe (exclusive of former USSR), with an introduction for non-systematists and comments on nomenclature and conservation" type="journal article" year="1997">Kottelat, 1997</bibRefCitation>
)
</taxonomicName>
. However, a minnow is characterised by the pres- ence of spots on the snout, the flanks and the tail. According to the original description (the absence of spots), this taxon could correspond to another leuciscid species but not to a
<taxonomicName authorityName="Rafinesque" authorityYear="1820" box="[813,914,1146,1170]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="225" phylum="Chordata" rank="genus">
<emphasis box="[813,914,1146,1170]" italics="true" pageId="20" pageNumber="225">Phoxinus</emphasis>
</taxonomicName>
, even if the ancient French word “verdon” means minnow. Moreover, no
<typeStatus box="[1072,1120,1179,1203]" pageId="20" pageNumber="225">type</typeStatus>
specimen is known. Thus, we consider this taxon as an
<emphasis box="[1078,1224,1211,1235]" italics="true" pageId="20" pageNumber="225">incertae sedis</emphasis>
in
<taxonomicName box="[1257,1388,1211,1235]" family="Leuciscidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="20" pageNumber="225" phylum="Chordata" rank="family">Leuciscidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>