treatments-xml/data/03/84/87/038487D4C740FFFCFEF4FA9DFDE4FD82.xml

<document ID-DOI="10.26028/cybium/2020-443-003" ID-Zenodo-Dep="7806280" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1680797231632" checkinUser="diego" docAuthor="Denys *, Gaël P. J., Dettai, Agnès, Persat, Henri, Daszkiewicz, Piotr, Hautecoeur, Mélyne &amp; Keith, Philippe" docDate="2020" docId="038487D4C740FFFCFEF4FA9DFDE4FD82" docLanguage="en" docName="Cybium.44.3.205-237.pdf" docOrigin="Cybium 44 (3)" docTitle="Phoxinus dragarum Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith 2020, new species" docType="treatment" docVersion="4" lastPageNumber="223" masterDocId="FFBDFFACC74FFFEEFFE4FF99FFCAFFD3" masterDocTitle="Revision of Phoxinus in France with the description of two new species (Teleostei, Leuciscidae)" masterLastPageNumber="238" masterPageNumber="205" pageNumber="220" updateTime="1680800786029" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Revision of Phoxinus in France with the description of two new species (Teleostei, Leuciscidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Denys *, Gaël P. J.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dettai, Agnès</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Persat, Henri</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Daszkiewicz, Piotr</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hautecoeur, Mélyne</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Keith, Philippe</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Cybium</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-09-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>44</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>205</mods:start>
<mods:end>238</mods:end>
</mods:extent>
</mods:part>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.26028/cybium/2020-443-003</mods:identifier>
<mods:identifier type="Zenodo-Dep">7806280</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7806156" ID-Zenodo-Dep="7806156" LSID="urn:lsid:plazi:treatment:038487D4C740FFFCFEF4FA9DFDE4FD82" httpUri="http://treatment.plazi.org/id/038487D4C740FFFCFEF4FA9DFDE4FD82" lastPageId="18" lastPageNumber="223" pageId="15" pageNumber="220">
<subSubSection box="[272,636,1284,1309]" pageId="15" pageNumber="220" type="nomenclature">
<paragraph blockId="15.[272,636,1284,1342]" box="[272,636,1284,1309]" pageId="15" pageNumber="220">
<heading bold="true" box="[272,636,1284,1309]" centered="true" fontSize="10" level="2" pageId="15" pageNumber="220" reason="4">
<emphasis bold="true" box="[272,636,1284,1309]" pageId="15" pageNumber="220">
<taxonomicName authority="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith, 2020" authorityName="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith" authorityYear="2020" box="[272,493,1284,1308]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="220" phylum="Chordata" rank="species" species="dragarum" status="new species">
<emphasis bold="true" box="[272,493,1284,1308]" italics="true" pageId="15" pageNumber="220">Phoxinus dragarum</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[505,636,1284,1309]" pageId="15" pageNumber="220" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[414,493,1318,1342]" pageId="15" pageNumber="220" type="description">
<paragraph blockId="15.[272,636,1284,1342]" box="[414,493,1318,1342]" pageId="15" pageNumber="220">
<heading box="[414,493,1318,1342]" centered="true" fontSize="10" level="4" pageId="15" pageNumber="220" reason="4">
(
<figureCitation box="[422,484,1318,1342]" captionStart="Figure 8" captionStartId="15.[813,876,1315,1337]" captionTargetBox="[813,1456,215,1298]" captionTargetId="figure-514@15.[813,1456,215,1298]" captionTargetPageId="15" captionText="Figure 8. – Phoxinus dragarum Holotype m (A), 49.52 mm SL from the Arrat-Devant Stream, Manent-Montané 43°20.7’N, 0°37.5’E, MNHN 2019-0262; paratypes m (B) and f (C), same location as holotype, MNHN 2014-2725; nuptial colouration pattern for male (D) and female (E)." figureDoi="http://doi.org/10.5281/zenodo.7806296" httpUri="https://zenodo.org/record/7806296/files/figure.png" pageId="15" pageNumber="220">Fig. 8</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7806296" ID-Zenodo-Dep="7806296" httpUri="https://zenodo.org/record/7806296/files/figure.png" pageId="15" pageNumber="220" startId="15.[813,876,1315,1337]" targetBox="[813,1456,215,1298]" targetPageId="15" targetType="figure">
<paragraph blockId="15.[813,1455,1315,1433]" pageId="15" pageNumber="220">
Figure 8. 
<emphasis box="[904,1111,1315,1337]" italics="true" pageId="15" pageNumber="220">
– 
<taxonomicName authorityName="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith" authorityYear="2020" box="[921,1111,1315,1336]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="15" pageNumber="220" phylum="Chordata" rank="species" species="dragarum">Phoxinus dragarum</taxonomicName>
</emphasis>
Holotype 
<emphasis bold="true" box="[1211,1224,1317,1336]" pageId="15" pageNumber="220">m</emphasis>
(
<emphasis bold="true" box="[1238,1255,1315,1336]" pageId="15" pageNumber="220">A</emphasis>
), 49.52 mm SL from the Arrat-Devant Stream, Manent-Montané 43°20.7’N, 0°37.5’E, MNHN 2019-0262; paratypes 
<emphasis bold="true" box="[1119,1132,1365,1384]" pageId="15" pageNumber="220">m</emphasis>
(
<emphasis bold="true" box="[1149,1165,1363,1384]" pageId="15" pageNumber="220">B</emphasis>
) and 
<emphasis bold="true" box="[1223,1234,1364,1384]" pageId="15" pageNumber="220">f</emphasis>
(
<emphasis bold="true" box="[1251,1268,1363,1385]" pageId="15" pageNumber="220">C</emphasis>
), same location as holotype, MNHN 2014-2725; nuptial colouration pattern for male (
<emphasis bold="true" box="[821,838,1411,1432]" pageId="15" pageNumber="220">D</emphasis>
) and female (
<emphasis bold="true" box="[970,986,1411,1432]" pageId="15" pageNumber="220">E</emphasis>
).
</paragraph>
</caption>
<subSubSection lastPageId="16" lastPageNumber="221" pageId="15" pageNumber="220" type="materials_examined">
<paragraph blockId="15.[132,775,1383,1508]" box="[132,236,1383,1407]" pageId="15" pageNumber="220">
<heading bold="true" box="[132,236,1383,1407]" fontSize="10" level="3" pageId="15" pageNumber="220" reason="6">
<emphasis bold="true" box="[132,236,1383,1407]" pageId="15" pageNumber="220">
<typeStatus box="[132,236,1383,1407]" pageId="15" pageNumber="220">Holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="15.[132,775,1383,1508]" pageId="15" pageNumber="220">
<materialsCitation collectionCode="MNHN" country="France" elevation="275" latitude="43.345" location="Manent-Montane" longLatPrecision="114" longitude="0.625" pageId="15" pageNumber="220" specimenCode="MNHN 2019-0262" specimenCount="1" stateProvince="Arrat- Devant Stream">
<specimenCode box="[170,394,1417,1441]" pageId="15" pageNumber="220">MNHN 2019-0262</specimenCode>
, 
<emphasis bold="true" box="[408,423,1419,1440]" pageId="15" pageNumber="220">
<specimenCount box="[408,423,1419,1440]" pageId="15" pageNumber="220" type="male">m</specimenCount>
</emphasis>
, 49.5 mm SL; 
<collectingCountry box="[608,692,1417,1440]" name="France" pageId="15" pageNumber="220" value="France">France</collectingCountry>
: 
<collectingRegion pageId="15" pageNumber="220">Arrat- Devant Stream</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:038487D4C740FFFCFEF4FA9DFDE4FD82:8EF26019C740FFE1FED6FA33FE25FA11" box="[306,495,1450,1474]" country="France" latitude="43.345" longLatPrecision="114" longitude="0.625" name="Manent-Montane" pageId="15" pageNumber="220" stateProvince="Arrat- Devant Stream">Manent-Montané</location>
, 
<geoCoordinate box="[505,621,1450,1474]" degrees="43" direction="north" minutes="20.7" orientation="latitude" pageId="15" pageNumber="220" precision="92" value="43.345">43°20.7’N</geoCoordinate>
, 
<geoCoordinate box="[631,731,1450,1474]" degrees="0" direction="east" minutes="37.5" orientation="longitude" pageId="15" pageNumber="220" precision="92" value="0.625">0°37.5’E</geoCoordinate>
, alt. 
<specimenCount box="[132,202,1484,1508]" pageId="15" pageNumber="220" type="generic">
<elevation box="[132,202,1484,1508]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="15" pageNumber="220" unit="m" value="275.0">275 m</elevation>
</specimenCount>
</materialsCitation>
.
</paragraph>
<paragraph blockId="15.[132,776,1549,1908]" box="[132,246,1549,1573]" pageId="15" pageNumber="220">
<heading bold="true" box="[132,246,1549,1573]" fontSize="10" level="3" pageId="15" pageNumber="220" reason="6">
<emphasis bold="true" box="[132,246,1549,1573]" pageId="15" pageNumber="220">
<typeStatus box="[132,246,1549,1573]" pageId="15" pageNumber="220">Paratypes</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="15.[132,776,1549,1908]" pageId="15" pageNumber="220">
<materialsCitation collectionCode="MNHN" country="France" latitude="43.345" location="Manent-Montane" longLatPrecision="114" longitude="0.625" pageId="15" pageNumber="220" specimenCode="MNHN 2014-2725" specimenCount="23" specimenCount-female="18" specimenCount-male="5" stateProvince="Arrat-Devant Stream">
<specimenCode box="[170,376,1582,1606]" pageId="15" pageNumber="220">MNHN 2014-2725</specimenCode>
, 
<specimenCount box="[387,422,1582,1606]" pageId="15" pageNumber="220" type="male">
5 
<emphasis bold="true" box="[407,422,1585,1606]" pageId="15" pageNumber="220">m</emphasis>
</specimenCount>
, 
<specimenCount box="[435,480,1583,1607]" pageId="15" pageNumber="220" type="female">
18 
<emphasis bold="true" box="[468,480,1584,1606]" pageId="15" pageNumber="220">f</emphasis>
</specimenCount>
, 44.9-61.2 mm SL; 
<collectingCountry box="[695,772,1583,1606]" name="France" pageId="15" pageNumber="220" value="France">France</collectingCountry>
: 
<collectingRegion box="[132,396,1616,1640]" pageId="15" pageNumber="220">Arrat-Devant Stream</collectingRegion>
, 
<location LSID="urn:lsid:plazi:treatment:038487D4C740FFFCFEF4FA9DFDE4FD82:8EF26019C740FFE1FE7FF9C9FDBBF9BB" box="[411,625,1616,1640]" country="France" latitude="43.345" longLatPrecision="114" longitude="0.625" name="Manent-Montane" pageId="15" pageNumber="220" stateProvince="Arrat-Devant Stream">Manent-Montané</location>
, 
<geoCoordinate box="[640,766,1616,1640]" degrees="43" direction="north" minutes="20.7" orientation="latitude" pageId="15" pageNumber="220" precision="92" value="43.345">43°20.7’N</geoCoordinate>
, 
<geoCoordinate box="[132,235,1649,1673]" degrees="0" direction="east" minutes="37.5" orientation="longitude" pageId="15" pageNumber="220" precision="92" value="0.625">0°37.5’E</geoCoordinate>
.
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="76" latitude="44.32" location="Escaudes" longLatPrecision="113" longitude="-0.19" municipality="Escaudes" pageId="15" pageNumber="220" specimenCode="MNHN 2014-2813" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Ciron Stream">
<specimenCode box="[246,459,1650,1674]" pageId="15" pageNumber="220">MNHN 2014-2813</specimenCode>
, 
<specimenCount box="[471,507,1650,1674]" pageId="15" pageNumber="220" type="male">
2 
<emphasis bold="true" box="[492,507,1652,1673]" pageId="15" pageNumber="220">m</emphasis>
</specimenCount>
, 
<specimenCount box="[523,556,1650,1674]" pageId="15" pageNumber="220" type="female">
4 
<emphasis bold="true" box="[544,556,1651,1673]" pageId="15" pageNumber="220">f</emphasis>
</specimenCount>
, 49.3-57.5 mm SL; 
<collectingCountry box="[132,211,1683,1706]" name="France" pageId="15" pageNumber="220" value="France">France</collectingCountry>
: 
<collectingRegion box="[221,370,1683,1707]" pageId="15" pageNumber="220">Ciron Stream</collectingRegion>
, 
<collectingMunicipality box="[380,484,1683,1707]" pageId="15" pageNumber="220">Escaudes</collectingMunicipality>
, 
<geoCoordinate box="[495,612,1683,1707]" degrees="44" direction="north" minutes="19.2" orientation="latitude" pageId="15" pageNumber="220" precision="92" value="44.32">44°19.2’N</geoCoordinate>
, 
<geoCoordinate box="[623,730,1683,1707]" degrees="0" direction="west" minutes="11.4" orientation="longitude" pageId="15" pageNumber="220" precision="92" value="-0.19">0°11.4’W</geoCoordinate>
, alt. 
<elevation box="[132,190,1716,1740]" metricMagnitude="1" metricUnit="m" metricValue="7.6" pageId="15" pageNumber="220" unit="m" value="76.0">76 m</elevation>
.
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="64" latitude="44.805" location="Bayac" longLatPrecision="113" longitude="0.72999996" municipality="Bayac" pageId="15" pageNumber="220" specimenCode="MNHN 2014-2807" specimenCount="1" specimenCount-female="1" stateProvince="Couze Stream">
<specimenCode box="[201,410,1716,1740]" pageId="15" pageNumber="220">MNHN 2014-2807</specimenCode>
, 
<specimenCount box="[421,453,1716,1740]" pageId="15" pageNumber="220" type="female">
1 
<emphasis bold="true" box="[441,453,1718,1740]" pageId="15" pageNumber="220">f</emphasis>
</specimenCount>
, 62.7 mm SL; 
<collectingCountry box="[617,696,1717,1740]" name="France" pageId="15" pageNumber="220" value="France">France</collectingCountry>
: 
<collectingRegion pageId="15" pageNumber="220">Couze Stream</collectingRegion>
, 
<collectingMunicipality box="[229,304,1750,1774]" pageId="15" pageNumber="220">Bayac</collectingMunicipality>
, 
<geoCoordinate box="[316,436,1750,1774]" degrees="44" direction="north" minutes="48.3" orientation="latitude" pageId="15" pageNumber="220" precision="92" value="44.805">44°48.3’N</geoCoordinate>
, 
<geoCoordinate box="[452,554,1750,1774]" degrees="0" direction="east" minutes="43.8" orientation="longitude" pageId="15" pageNumber="220" precision="92" value="0.72999996">0°43.8’E</geoCoordinate>
, alt. 
<elevation box="[617,679,1750,1774]" metricMagnitude="1" metricUnit="m" metricValue="6.4" pageId="15" pageNumber="220" unit="m" value="64.0">64 m</elevation>
.
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="123" latitude="44.54" location="Montcabrier" longLatPrecision="113" longitude="1.065" municipality="Montcabrier" pageId="15" pageNumber="220" specimenCode="MNHN 2014-2735" specimenCount="7" specimenCount-female="6" specimenCount-male="1" stateProvince="Theze Stream">
<specimenCode pageId="15" pageNumber="220">MNHN 2014-2735</specimenCode>
, 
<specimenCount box="[274,312,1783,1807]" pageId="15" pageNumber="220" type="male">
1 
<emphasis bold="true" box="[297,312,1786,1807]" pageId="15" pageNumber="220">m</emphasis>
</specimenCount>
, 
<specimenCount box="[332,367,1783,1807]" pageId="15" pageNumber="220" type="female">
6 
<emphasis bold="true" box="[355,367,1785,1807]" pageId="15" pageNumber="220">f</emphasis>
</specimenCount>
, 43.6-60.8 mm SL; 
<collectingCountry box="[610,694,1784,1807]" name="France" pageId="15" pageNumber="220" value="France">France</collectingCountry>
: 
<collectingRegion pageId="15" pageNumber="220">Thèze Stream</collectingRegion>
, 
<collectingMunicipality box="[231,377,1817,1841]" pageId="15" pageNumber="220">Montcabrier</collectingMunicipality>
, 
<geoCoordinate box="[390,515,1817,1841]" degrees="44" direction="north" minutes="32.4" orientation="latitude" pageId="15" pageNumber="220" precision="92" value="44.54">44°32.4’N</geoCoordinate>
, 
<geoCoordinate box="[528,635,1817,1841]" degrees="1" direction="east" minutes="03.9" orientation="longitude" pageId="15" pageNumber="220" precision="92" value="1.065">1°03.9’E</geoCoordinate>
, alt. 
<elevation box="[695,771,1817,1841]" metricMagnitude="2" metricUnit="m" metricValue="1.23" pageId="15" pageNumber="220" unit="m" value="123.0">123 m</elevation>
.
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="189" latitude="43.986668" location="Cahuzac-sur-Vere" longLatPrecision="113" longitude="1.895" municipality="Cahuzac-sur-Vere" pageId="15" pageNumber="220" specimenCode="MNHN 2014-2738" specimenCount="3" specimenCount-male="3" stateProvince="Vere Stream">
<specimenCode box="[132,348,1850,1874]" pageId="15" pageNumber="220">MNHN 2014-2738</specimenCode>
, 
<specimenCount box="[361,397,1850,1874]" pageId="15" pageNumber="220" type="male">
3 
<emphasis bold="true" box="[382,397,1853,1874]" pageId="15" pageNumber="220">m</emphasis>
</specimenCount>
, 45.5-51.9 mm SL; 
<collectingCountry box="[629,710,1851,1874]" name="France" pageId="15" pageNumber="220" value="France">France</collectingCountry>
: 
<collectingRegion pageId="15" pageNumber="220">Vère Stream</collectingRegion>
, 
<collectingMunicipality box="[221,417,1883,1907]" pageId="15" pageNumber="220">Cahuzac-sur-Vère</collectingMunicipality>
, 
<geoCoordinate box="[427,541,1883,1907]" degrees="43" direction="north" minutes="59.2" orientation="latitude" pageId="15" pageNumber="220" precision="92" value="43.986668">43°59.2’N</geoCoordinate>
, 
<geoCoordinate box="[551,649,1883,1907]" degrees="1" direction="east" minutes="53.7" orientation="longitude" pageId="15" pageNumber="220" precision="92" value="1.895">1°53.7’E</geoCoordinate>
, alt. 
<specimenCount box="[699,769,1884,1908]" pageId="15" pageNumber="220" type="generic">
<elevation box="[699,769,1884,1908]" metricMagnitude="2" metricUnit="m" metricValue="1.89" pageId="15" pageNumber="220" unit="m" value="189.0">189 m</elevation>
</specimenCount>
</materialsCitation>
.
</paragraph>
<paragraph blockId="15.[813,1456,1467,1701]" box="[813,1036,1467,1491]" pageId="15" pageNumber="220">
<heading bold="true" box="[813,1036,1467,1491]" fontSize="10" level="3" pageId="15" pageNumber="220" reason="6">
<emphasis bold="true" box="[813,1036,1467,1491]" pageId="15" pageNumber="220">Additional material</emphasis>
</heading>
</paragraph>
<paragraph blockId="15.[813,1456,1467,1701]" pageId="15" pageNumber="220">
<materialsCitation collectionCode="MNHN" country="France" elevation="94" latitude="45.323334" location="Bourdeilles" longLatPrecision="113" longitude="0.585" municipality="Bourdeilles" pageId="15" pageNumber="220" specimenCode="MNHN 2014-2761" specimenCount="3" specimenCount-female="3" stateProvince="Dronne Stream">
<specimenCode box="[851,1073,1502,1526]" pageId="15" pageNumber="220">MNHN 2014-2761</specimenCode>
, 
<specimenCount box="[1087,1123,1503,1527]" pageId="15" pageNumber="220" type="female">
3 
<emphasis bold="true" box="[1111,1123,1504,1526]" pageId="15" pageNumber="220">f</emphasis>
</specimenCount>
, 44.7-53.7 mm SL; 
<collectingCountry box="[1368,1451,1503,1526]" name="France" pageId="15" pageNumber="220" value="France">France</collectingCountry>
: 
<collectingRegion box="[813,977,1538,1562]" pageId="15" pageNumber="220">Dronne Stream</collectingRegion>
, 
<collectingMunicipality box="[987,1113,1537,1561]" pageId="15" pageNumber="220">Bourdeilles</collectingMunicipality>
, 
<geoCoordinate box="[1122,1237,1537,1561]" degrees="45" direction="north" minutes="19.4" orientation="latitude" pageId="15" pageNumber="220" precision="92" value="45.323334">45°19.4’N</geoCoordinate>
, 
<geoCoordinate box="[1247,1346,1537,1561]" degrees="0" direction="east" minutes="35.1" orientation="longitude" pageId="15" pageNumber="220" precision="92" value="0.585">0°35.1’E</geoCoordinate>
, alt. 
<elevation box="[1395,1452,1538,1562]" metricMagnitude="1" metricUnit="m" metricValue="9.4" pageId="15" pageNumber="220" unit="m" value="94.0">94 m</elevation>
.
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="134" latitude="44.661667" location="Gaugeac" longLatPrecision="113" longitude="0.875" municipality="Gaugeac" pageId="15" pageNumber="220" specimenCode="MNHN 2014-2759" specimenCount="4" specimenCount-female="4" stateProvince="Dropt Stream">
<specimenCode box="[813,1027,1572,1596]" pageId="15" pageNumber="220">MNHN 2014-2759</specimenCode>
, 
<specimenCount box="[1038,1071,1572,1596]" pageId="15" pageNumber="220" type="female">
4 
<emphasis bold="true" box="[1059,1071,1574,1596]" pageId="15" pageNumber="220">f</emphasis>
</specimenCount>
, 57.6-66.5 mm SL; 
<collectingCountry box="[1300,1380,1573,1596]" name="France" pageId="15" pageNumber="220" value="France">France</collectingCountry>
: 
<collectingRegion pageId="15" pageNumber="220">Dropt Stream</collectingRegion>
, 
<collectingMunicipality box="[903,1001,1607,1631]" pageId="15" pageNumber="220">Gaugeac</collectingMunicipality>
, 
<geoCoordinate box="[1011,1129,1607,1631]" degrees="44" direction="north" minutes="39.7" orientation="latitude" pageId="15" pageNumber="220" precision="92" value="44.661667">44°39.7’N</geoCoordinate>
, 
<geoCoordinate box="[1139,1240,1607,1631]" degrees="0" direction="east" minutes="52.5" orientation="longitude" pageId="15" pageNumber="220" precision="92" value="0.875">0°52.5’E</geoCoordinate>
, alt. 
<elevation box="[1291,1362,1607,1631]" metricMagnitude="2" metricUnit="m" metricValue="1.34" pageId="15" pageNumber="220" unit="m" value="134.0">134 m</elevation>
.
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="451" latitude="43.045" location="Montesquieu-Avantes" longLatPrecision="114" longitude="1.1866667" municipality="Montesquieu-Avantes" pageId="15" pageNumber="220" specimenCode="MNHN 2014-2731" specimenCount="4" specimenCount-female="4" stateProvince="Volp Stream">
<specimenCode pageId="15" pageNumber="220">MNHN 2014-2731</specimenCode>
, 
<specimenCount box="[950,983,1642,1666]" pageId="15" pageNumber="220" type="female">
4 
<emphasis bold="true" box="[971,983,1644,1666]" pageId="15" pageNumber="220">f</emphasis>
</specimenCount>
, 51.0-57.6 mm SL; 
<collectingCountry box="[1216,1298,1643,1666]" name="France" pageId="15" pageNumber="220" value="France">France</collectingCountry>
: 
<collectingRegion box="[1308,1451,1642,1666]" pageId="15" pageNumber="220">Volp Stream</collectingRegion>
, 
<collectingMunicipality box="[813,1049,1677,1701]" pageId="15" pageNumber="220">Montesquieu-Avantès</collectingMunicipality>
, 
<geoCoordinate box="[1059,1173,1677,1701]" degrees="43" direction="north" minutes="02.7" orientation="latitude" pageId="15" pageNumber="220" precision="92" value="43.045">43°02.7’N</geoCoordinate>
, 
<geoCoordinate box="[1184,1280,1677,1701]" degrees="1" direction="east" minutes="11.2" orientation="longitude" pageId="15" pageNumber="220" precision="92" value="1.1866667">1°11.2’E</geoCoordinate>
, alt. 
<specimenCount box="[1330,1400,1677,1701]" pageId="15" pageNumber="220" type="generic">
<elevation box="[1330,1400,1677,1701]" metricMagnitude="2" metricUnit="m" metricValue="4.51" pageId="15" pageNumber="220" unit="m" value="451.0">451 m</elevation>
</specimenCount>
</materialsCitation>
.
</paragraph>
<paragraph blockId="15.[813,1455,1744,1908]" box="[813,1225,1744,1768]" pageId="15" pageNumber="220">
<heading bold="true" box="[813,1225,1744,1768]" fontSize="10" level="3" pageId="15" pageNumber="220" reason="6">
<emphasis bold="true" box="[813,1225,1744,1768]" pageId="15" pageNumber="220">Material used for molecular analysis</emphasis>
</heading>
</paragraph>
<paragraph blockId="15.[813,1455,1744,1908]" lastBlockId="16.[132,776,217,1426]" lastPageId="16" lastPageNumber="221" pageId="15" pageNumber="220">
<materialsCitation collectionCode="MNHN" country="France" elevation="275" latitude="43.345" location="Manent-Montane" longLatPrecision="114" longitude="0.625" municipality="Manent-Montane" pageId="15" pageNumber="220" specimenCode="MNHN 2019-0262" specimenCount="1" stateProvince="Arrat- Devant Stream" typeStatus="holotype">
<specimenCode box="[850,1060,1779,1803]" pageId="15" pageNumber="220">MNHN 2019-0262</specimenCode>
, 
<typeStatus box="[1071,1169,1779,1803]" pageId="15" pageNumber="220">holotype</typeStatus>
, tag 12470; 
<collectingCountry box="[1302,1380,1779,1802]" name="France" pageId="15" pageNumber="220" value="France">France</collectingCountry>
: 
<collectingRegion pageId="15" pageNumber="220">Arrat- Devant Stream</collectingRegion>
, 
<collectingMunicipality box="[986,1175,1814,1838]" pageId="15" pageNumber="220">Manent-Montané</collectingMunicipality>
, 
<geoCoordinate box="[1185,1301,1814,1838]" degrees="43" direction="north" minutes="20.7" orientation="latitude" pageId="15" pageNumber="220" precision="92" value="43.345">43°20.7’N</geoCoordinate>
, 
<geoCoordinate box="[1312,1411,1814,1838]" degrees="0" direction="east" minutes="37.5" orientation="longitude" pageId="15" pageNumber="220" precision="92" value="0.625">0°37.5’E</geoCoordinate>
, alt. 
<elevation box="[813,883,1849,1873]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="15" pageNumber="220" unit="m" value="275.0">275 m</elevation>
. (COI: GenBankaccessionnumber: 
<accessionNumber box="[1273,1396,1849,1873]" pageId="15" pageNumber="220">MT975386</accessionNumber>
, 12S: GenBankaccessionnumber: 
<accessionNumber box="[1146,1278,1883,1907]" pageId="15" pageNumber="220">MT975361</accessionNumber>
).
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="76" lastPageId="16" lastPageNumber="221" latitude="44.32" location="Escaudes" longLatPrecision="113" longitude="-0.19" municipality="Escaudes" pageId="15" pageNumber="220" specimenCode="MNHN 2014" specimenCount="1" stateProvince="Ciron Stream" typeStatus="paratype">
<specimenCode box="[1298,1448,1884,1908]" pageId="15" pageNumber="220">MNHN 2014</specimenCode>
- 2813, 
<typeStatus box="[197,302,218,242]" pageId="16" pageNumber="221">paratypes</typeStatus>
, 2, tag 1989 and 1990; 
<collectingCountry box="[542,619,218,241]" name="France" pageId="16" pageNumber="221" value="France">France</collectingCountry>
: 
<collectingRegion box="[627,771,217,241]" pageId="16" pageNumber="221">Ciron Stream</collectingRegion>
, 
<collectingMunicipality box="[132,242,249,273]" pageId="16" pageNumber="221">Escaudes</collectingMunicipality>
, 
<geoCoordinate box="[256,380,249,273]" degrees="44" direction="north" minutes="19.2" orientation="latitude" pageId="16" pageNumber="221" precision="92" value="44.32">44°19.2’N</geoCoordinate>
, 
<geoCoordinate box="[395,506,249,273]" degrees="0" direction="west" minutes="11.4" orientation="longitude" pageId="16" pageNumber="221" precision="92" value="-0.19">0°11.4’W</geoCoordinate>
, alt. 
<elevation box="[567,629,249,273]" metricMagnitude="1" metricUnit="m" metricValue="7.6" pageId="16" pageNumber="221" unit="m" value="76.0">76 m</elevation>
. (COI: Gen- Bank accession numbers: 
<accessionNumber box="[414,537,281,305]" pageId="16" pageNumber="221">MT975361</accessionNumber>
, 
<accessionNumber box="[548,675,281,305]" pageId="16" pageNumber="221">MT975746</accessionNumber>
).
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="189" latitude="43.986668" location="Cahuzac-sur-Vere" longLatPrecision="113" longitude="1.895" municipality="Cahuzac-sur-Vere" pageId="16" pageNumber="221" specimenCode="MNHN 2014-2738" specimenCount="2" stateProvince="Vere Stream" typeStatus="paratype">
<specimenCode pageId="16" pageNumber="221">MNHN 2014-2738</specimenCode>
, 
<typeStatus box="[275,379,314,338]" pageId="16" pageNumber="221">paratype</typeStatus>
, 
<specimenCount box="[392,409,313,337]" pageId="16" pageNumber="221">2</specimenCount>
, tag 12342 and 12343; 
<collectingCountry box="[688,771,314,337]" name="France" pageId="16" pageNumber="221" value="France">France</collectingCountry>
: 
<collectingRegion box="[132,274,345,369]" pageId="16" pageNumber="221">Vère Stream</collectingRegion>
, 
<collectingMunicipality box="[284,487,345,369]" pageId="16" pageNumber="221">Cahuzac-sur-Vère</collectingMunicipality>
, 
<geoCoordinate box="[498,616,345,369]" degrees="43" direction="north" minutes="59.2" orientation="latitude" pageId="16" pageNumber="221" precision="92" value="43.986668">43°59.2’N</geoCoordinate>
, 
<geoCoordinate box="[627,729,345,369]" degrees="1" direction="east" minutes="53.7" orientation="longitude" pageId="16" pageNumber="221" precision="92" value="1.895">1°53.7’E</geoCoordinate>
, alt. 
<elevation box="[132,203,378,402]" metricMagnitude="2" metricUnit="m" metricValue="1.89" pageId="16" pageNumber="221" unit="m" value="189.0">189 m</elevation>
. (COI: GenBankaccessionnumber: 
<accessionNumber box="[593,715,377,401]" pageId="16" pageNumber="221">MT975720</accessionNumber>
, 12S: GenBank accession numbers: 
<accessionNumber box="[485,617,409,433]" pageId="16" pageNumber="221">MT975353</accessionNumber>
, 
<accessionNumber box="[630,764,409,433]" pageId="16" pageNumber="221">MT975354</accessionNumber>
).
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="25" latitude="44.468334" location="Montpouillan" longLatPrecision="113" longitude="0.12666667" municipality="Montpouillan" pageId="16" pageNumber="221" specimenCode="MNHN 2011-0933" specimenCount="1" stateProvince="Avance Stream">
<specimenCode box="[132,345,442,466]" pageId="16" pageNumber="221">MNHN 2011-0933</specimenCode>
, 
<specimenCount box="[357,371,442,466]" pageId="16" pageNumber="221">1</specimenCount>
, tag 10781; 
<collectingCountry box="[510,590,442,465]" name="France" pageId="16" pageNumber="221" value="France">France</collectingCountry>
: 
<collectingRegion box="[600,770,442,466]" pageId="16" pageNumber="221">Avance Stream</collectingRegion>
, 
<collectingMunicipality box="[132,284,473,497]" pageId="16" pageNumber="221">Montpouillan</collectingMunicipality>
, 
<geoCoordinate box="[296,413,474,498]" degrees="44" direction="north" minutes="28.1" orientation="latitude" pageId="16" pageNumber="221" precision="92" value="44.468334">44°28.1’N</geoCoordinate>
, 
<geoCoordinate box="[425,527,473,497]" degrees="0" direction="east" minutes="07.6" orientation="longitude" pageId="16" pageNumber="221" precision="92" value="0.12666667">0°07.6’E</geoCoordinate>
, alt. 
<elevation box="[580,638,473,497]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="16" pageNumber="221" unit="m" value="25.0">25 m</elevation>
. (COI: Gen- Bank accession number: 
<accessionNumber box="[405,529,505,529]" pageId="16" pageNumber="221">MT975709</accessionNumber>
, 12S: GenBank acces- sion number: 
<accessionNumber box="[278,404,537,561]" pageId="16" pageNumber="221">MT975350</accessionNumber>
).
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="386" latitude="44.533333" location="Espalion" longLatPrecision="927" longitude="2.7916667" municipality="Espalion" pageId="16" pageNumber="221" specimenCode="MNHN 2011-0937" specimenCount="1" stateProvince="Boralde de Flaujac Stream">
<specimenCode box="[420,625,538,562]" pageId="16" pageNumber="221">MNHN 2011-0937</specimenCode>
, 
<specimenCount box="[635,648,538,562]" pageId="16" pageNumber="221">1</specimenCount>
, tag 10778; 
<collectingCountry box="[132,215,570,593]" name="France" pageId="16" pageNumber="221" value="France">France</collectingCountry>
: 
<collectingRegion box="[227,540,570,594]" pageId="16" pageNumber="221">Boralde de Flaujac Stream</collectingRegion>
, 
<collectingMunicipality box="[552,656,570,594]" pageId="16" pageNumber="221">Espalion</collectingMunicipality>
, 
<geoCoordinate box="[669,770,570,594]" degrees="44" direction="north" minutes="32" orientation="latitude" pageId="16" pageNumber="221" precision="925" value="44.533333">44°32’N</geoCoordinate>
, 
<geoCoordinate box="[132,236,601,625]" degrees="2" direction="east" minutes="47.5" orientation="longitude" pageId="16" pageNumber="221" precision="92" value="2.7916667">2°47.5’E</geoCoordinate>
, alt. 
<elevation box="[293,367,602,626]" metricMagnitude="2" metricUnit="m" metricValue="3.86" pageId="16" pageNumber="221" unit="m" value="386.0">386 m</elevation>
. (COI: GenBankaccessionnumber: 
<accessionNumber box="[132,257,633,657]" pageId="16" pageNumber="221">MT975708</accessionNumber>
).
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="321" latitude="43.038334" location="Dun" longLatPrecision="114" longitude="1.7933333" municipality="Dun" pageId="16" pageNumber="221" specimenCode="MNHN 2010-0491" specimenCount="1" stateProvince="Douc- touyre Stream">
<specimenCode box="[274,479,634,658]" pageId="16" pageNumber="221">MNHN 2010-0491</specimenCode>
, 
<specimenCount box="[490,503,634,658]" pageId="16" pageNumber="221">1</specimenCount>
, tag 4103; 
<collectingCountry box="[622,699,634,657]" name="France" pageId="16" pageNumber="221" value="France">France</collectingCountry>
: 
<collectingRegion pageId="16" pageNumber="221">Douc- touyre Stream</collectingRegion>
, 
<collectingMunicipality box="[296,345,666,689]" pageId="16" pageNumber="221">Dun</collectingMunicipality>
, 
<geoCoordinate box="[356,468,666,690]" degrees="43" direction="north" minutes="02.3" orientation="latitude" pageId="16" pageNumber="221" precision="92" value="43.038334">43°02.3’N</geoCoordinate>
, 
<geoCoordinate box="[482,578,666,690]" degrees="1" direction="east" minutes="47.6" orientation="longitude" pageId="16" pageNumber="221" precision="92" value="1.7933333">1°47.6’E</geoCoordinate>
, alt. 
<elevation box="[632,702,666,690]" metricMagnitude="2" metricUnit="m" metricValue="3.21" pageId="16" pageNumber="221" unit="m" value="321.0">321 m</elevation>
. (COI: GenBankaccessionnumber: 
<accessionNumber box="[466,598,698,722]" pageId="16" pageNumber="221">MT975757</accessionNumber>
).
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="525" latitude="45.141666" location="Saint-Bazilede-la-Roche" longLatPrecision="113" longitude="1.96" municipality="Saint-Bazilede-la-Roche" pageId="16" pageNumber="221" specimenCode="MNHN 2011- 0416" specimenCount="1" stateProvince="Doustre Stream">
<specimenCode pageId="16" pageNumber="221">MNHN 2011- 0416</specimenCode>
, 
<specimenCount box="[203,217,730,754]" pageId="16" pageNumber="221">1</specimenCount>
, tag 4427; 
<collectingCountry box="[344,424,730,753]" name="France" pageId="16" pageNumber="221" value="France">France</collectingCountry>
: 
<collectingRegion box="[436,614,730,754]" pageId="16" pageNumber="221">Doustre Stream</collectingRegion>
, 
<collectingMunicipality pageId="16" pageNumber="221">Saint-Bazilede-la-Roche</collectingMunicipality>
, 
<geoCoordinate box="[281,399,762,786]" degrees="45" direction="north" minutes="08.5" orientation="latitude" pageId="16" pageNumber="221" precision="92" value="45.141666">45°08.5’N</geoCoordinate>
, 
<geoCoordinate box="[411,512,762,786]" degrees="1" direction="east" minutes="57.6" orientation="longitude" pageId="16" pageNumber="221" precision="92" value="1.96">1°57.6’E</geoCoordinate>
, alt. 
<elevation box="[565,638,762,786]" metricMagnitude="2" metricUnit="m" metricValue="5.25" pageId="16" pageNumber="221" unit="m" value="525.0">525 m</elevation>
. (COI: Gen- Bank accession number: 
<accessionNumber box="[410,540,794,818]" pageId="16" pageNumber="221">MT975793</accessionNumber>
).
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="367" latitude="43.068333" location="Lespiteau" longLatPrecision="114" longitude="0.7583333" municipality="Lespiteau" pageId="16" pageNumber="221" specimenCode="MNHN 2010-0513" specimenCount="1" stateProvince="Job Stream">
<specimenCode box="[558,771,794,818]" pageId="16" pageNumber="221">MNHN 2010-0513</specimenCode>
, 
<specimenCount box="[132,147,826,850]" pageId="16" pageNumber="221">1</specimenCount>
, tag 4124; 
<collectingCountry box="[280,363,826,849]" name="France" pageId="16" pageNumber="221" value="France">France</collectingCountry>
: 
<collectingRegion box="[374,507,826,850]" pageId="16" pageNumber="221">Job Stream</collectingRegion>
, 
<collectingMunicipality box="[519,635,826,850]" pageId="16" pageNumber="221">Lespiteau</collectingMunicipality>
, 
<geoCoordinate box="[648,770,826,850]" degrees="43" direction="north" minutes="04.1" orientation="latitude" pageId="16" pageNumber="221" precision="92" value="43.068333">43°04.1’N</geoCoordinate>
, 
<geoCoordinate box="[132,236,858,882]" degrees="0" direction="east" minutes="45.5" orientation="longitude" pageId="16" pageNumber="221" precision="92" value="0.7583333">0°45.5’E</geoCoordinate>
, alt. 
<elevation box="[293,367,858,882]" metricMagnitude="2" metricUnit="m" metricValue="3.67" pageId="16" pageNumber="221" unit="m" value="367.0">367 m</elevation>
. (COI: GenBankaccessionnumber: 
<accessionNumber box="[132,267,890,914]" pageId="16" pageNumber="221">MT975762</accessionNumber>
).
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="493" latitude="42.911667" location="Cierp-Gaud" longLatPrecision="114" longitude="0.06333333" municipality="Cierp-Gaud" pageId="16" pageNumber="221" specimenCode="MNHN 2011-0278" specimenCount="2" stateProvince="Pique Stream">
<specimenCode box="[286,508,890,914]" pageId="16" pageNumber="221">MNHN 2011-0278</specimenCode>
, 
<specimenCount box="[520,537,890,914]" pageId="16" pageNumber="221">2</specimenCount>
, tag 4390 and 4391; 
<collectingCountry box="[132,213,922,945]" name="France" pageId="16" pageNumber="221" value="France">France</collectingCountry>
: 
<collectingRegion box="[224,377,922,946]" pageId="16" pageNumber="221">Pique Stream</collectingRegion>
, 
<collectingMunicipality box="[389,524,922,946]" pageId="16" pageNumber="221">Cierp-Gaud</collectingMunicipality>
, 
<geoCoordinate box="[535,655,922,946]" degrees="42" direction="north" minutes="54.7" orientation="latitude" pageId="16" pageNumber="221" precision="92" value="42.911667">42°54.7’N</geoCoordinate>
, 
<geoCoordinate box="[667,770,922,946]" degrees="0" direction="east" minutes="03.8" orientation="longitude" pageId="16" pageNumber="221" precision="92" value="0.06333333">0°03.8’E</geoCoordinate>
, alt. 
<elevation box="[172,243,954,978]" metricMagnitude="2" metricUnit="m" metricValue="4.93" pageId="16" pageNumber="221" unit="m" value="493.0">493 m</elevation>
. (COI: GenBank accession numbers: 
<accessionNumber box="[647,771,954,978]" pageId="16" pageNumber="221">MT975787</accessionNumber>
, 
<accessionNumber box="[132,262,986,1010]" pageId="16" pageNumber="221">MT975788</accessionNumber>
).
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="109" latitude="44.018333" location="Saint-Pe-Saint-Simon" longLatPrecision="113" longitude="0.061666667" municipality="Saint-Pe-Saint-Simon" pageId="16" pageNumber="221" specimenCode="MNHN 2011-0928" specimenCount="1" stateProvince="Rimbez Stream">
<specimenCode box="[280,493,986,1010]" pageId="16" pageNumber="221">MNHN 2011-0928</specimenCode>
, 
<specimenCount box="[505,519,986,1010]" pageId="16" pageNumber="221">1</specimenCount>
, tag 10786 and 10787; 
<collectingCountry box="[132,211,1018,1041]" name="France" pageId="16" pageNumber="221" value="France">France</collectingCountry>
: 
<collectingRegion box="[221,392,1018,1042]" pageId="16" pageNumber="221">Rimbez Stream</collectingRegion>
, 
<collectingMunicipality box="[402,643,1018,1042]" pageId="16" pageNumber="221">Saint-Pe-Saint-Simon</collectingMunicipality>
, 
<geoCoordinate box="[654,771,1018,1042]" degrees="44" direction="north" minutes="01.1" orientation="latitude" pageId="16" pageNumber="221" precision="92" value="44.018333">44°01.1’N</geoCoordinate>
, 
<geoCoordinate box="[132,236,1050,1074]" degrees="0" direction="east" minutes="03.7" orientation="longitude" pageId="16" pageNumber="221" precision="92" value="0.061666667">0°03.7’E</geoCoordinate>
, alt. 
<elevation box="[293,367,1050,1074]" metricMagnitude="2" metricUnit="m" metricValue="1.09" pageId="16" pageNumber="221" unit="m" value="109.0">109 m</elevation>
. (COI: GenBankaccessionnumber: 
<accessionNumber box="[132,256,1082,1106]" pageId="16" pageNumber="221">MT975710</accessionNumber>
, 12S: GenBank accession numbers: 
<accessionNumber box="[647,771,1082,1106]" pageId="16" pageNumber="221">MT975351</accessionNumber>
, 
<accessionNumber box="[132,259,1114,1138]" pageId="16" pageNumber="221">MT975352</accessionNumber>
).
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="412" latitude="42.956665" location="Riverenert" longLatPrecision="114" longitude="1.1766667" municipality="Riverenert" pageId="16" pageNumber="221" specimenCode="MNHN 2010-0504" specimenCount="1" stateProvince="Salat Stream">
<specimenCode box="[277,487,1114,1138]" pageId="16" pageNumber="221">MNHN 2010-0504</specimenCode>
, 
<specimenCount box="[497,511,1114,1138]" pageId="16" pageNumber="221">1</specimenCount>
, tag 4116; 
<collectingCountry box="[632,710,1115,1138]" name="France" pageId="16" pageNumber="221" value="France">France</collectingCountry>
: 
<collectingRegion pageId="16" pageNumber="221">Salat Stream</collectingRegion>
, 
<collectingMunicipality box="[222,340,1146,1170]" pageId="16" pageNumber="221">Rivèrenert</collectingMunicipality>
, 
<geoCoordinate box="[351,467,1146,1170]" degrees="42" direction="north" minutes="57.4" orientation="latitude" pageId="16" pageNumber="221" precision="92" value="42.956665">42°57.4’N</geoCoordinate>
, 
<geoCoordinate box="[478,578,1146,1170]" degrees="1" direction="east" minutes="10.6" orientation="longitude" pageId="16" pageNumber="221" precision="92" value="1.1766667">1°10.6’E</geoCoordinate>
, alt. 
<elevation box="[630,701,1146,1170]" metricMagnitude="2" metricUnit="m" metricValue="4.12" pageId="16" pageNumber="221" unit="m" value="412.0">412 m</elevation>
. (COI: GenBankaccessionnumber: 
<accessionNumber box="[467,597,1178,1202]" pageId="16" pageNumber="221">MT975760</accessionNumber>
, 12S: GenBank accession number: 
<accessionNumber box="[344,472,1210,1234]" pageId="16" pageNumber="221">MT975378</accessionNumber>
).
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="637" latitude="45.503334" location="Saint-Angel" longLatPrecision="112" longitude="2.235" municipality="Saint-Angel" pageId="16" pageNumber="221" specimenCode="MNHN 2010-0476" specimenCount="1" stateProvince="Triouzoune Stream">
<specimenCode box="[490,702,1210,1234]" pageId="16" pageNumber="221">MNHN 2010-0476</specimenCode>
, 
<specimenCount box="[713,727,1210,1234]" pageId="16" pageNumber="221">1</specimenCount>
, tag 4031; 
<collectingCountry box="[201,279,1243,1266]" name="France" pageId="16" pageNumber="221" value="France">France</collectingCountry>
: 
<collectingRegion box="[288,499,1242,1266]" pageId="16" pageNumber="221">Triouzoune Stream</collectingRegion>
, 
<collectingMunicipality box="[509,643,1242,1266]" pageId="16" pageNumber="221">Saint-Angel</collectingMunicipality>
, 
<geoCoordinate box="[654,770,1242,1266]" degrees="45" direction="north" minutes="30.2" orientation="latitude" pageId="16" pageNumber="221" precision="92" value="45.503334">45°30.2’N</geoCoordinate>
, 
<geoCoordinate box="[132,233,1274,1298]" degrees="2" direction="east" minutes="14.1" orientation="longitude" pageId="16" pageNumber="221" precision="92" value="2.235">2°14.1’E</geoCoordinate>
, alt. 
<elevation box="[292,367,1274,1298]" metricMagnitude="2" metricUnit="m" metricValue="6.37" pageId="16" pageNumber="221" unit="m" value="637.0">637 m</elevation>
. (COI: GenBankaccessionnumber: 
<accessionNumber box="[132,263,1306,1330]" pageId="16" pageNumber="221">MT975753</accessionNumber>
).
</materialsCitation>
<materialsCitation collectionCode="MNHN" country="France" elevation="153" latitude="42.213333" location="Tavignano River" longLatPrecision="114" longitude="9.3133335" municipality="Piedicorte-di-Gaccio" pageId="16" pageNumber="221" specimenCode="MNHN 2011-1570" specimenCount="2">
<specimenCode box="[280,493,1306,1330]" pageId="16" pageNumber="221">MNHN 2011-1570</specimenCode>
, 
<specimenCount box="[505,521,1306,1330]" pageId="16" pageNumber="221">2</specimenCount>
, tag 10931 and 10932; 
<collectingCountry box="[132,210,1339,1362]" name="France" pageId="16" pageNumber="221" value="France">France</collectingCountry>
: 
<location LSID="urn:lsid:plazi:treatment:038487D4C740FFFCFEF4FA9DFDE4FD82:8EF26019C75FFFFEFF38FAA3FE59FA81" box="[220,403,1338,1362]" country="France" latitude="42.213333" longLatPrecision="114" longitude="9.3133335" municipality="Piedicorte-di-Gaccio" name="Tavignano River" pageId="16" pageNumber="221">Tavignano River</location>
, 
<collectingMunicipality box="[414,644,1338,1362]" pageId="16" pageNumber="221">Piedicorte-di-Gaccio</collectingMunicipality>
, 
<geoCoordinate box="[654,768,1338,1362]" degrees="42" direction="north" minutes="12.8" orientation="latitude" pageId="16" pageNumber="221" precision="92" value="42.213333">42°12.8’N</geoCoordinate>
, 
<geoCoordinate box="[132,235,1370,1394]" degrees="9" direction="east" minutes="18.8" orientation="longitude" pageId="16" pageNumber="221" precision="92" value="9.3133335">9°18.8’E</geoCoordinate>
, alt. 
<elevation box="[289,363,1370,1394]" metricMagnitude="2" metricUnit="m" metricValue="1.53" pageId="16" pageNumber="221" unit="m" value="153.0">153 m</elevation>
. (COI: GenBank accession numbers: 
<accessionNumber box="[132,255,1402,1426]" pageId="16" pageNumber="221">MT975715</accessionNumber>
, 
<accessionNumber box="[265,390,1402,1426]" pageId="16" pageNumber="221">MT975716</accessionNumber>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="221" type="diagnosis">
<paragraph blockId="16.[132,776,1466,1907]" box="[132,242,1466,1490]" pageId="16" pageNumber="221">
<heading bold="true" box="[132,242,1466,1490]" fontSize="10" level="3" pageId="16" pageNumber="221" reason="6">
<emphasis bold="true" box="[132,242,1466,1490]" pageId="16" pageNumber="221">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="16.[132,776,1466,1907]" pageId="16" pageNumber="221">
<taxonomicName authorityName="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith" authorityYear="2020" box="[170,402,1498,1522]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="dragarum">
<emphasis box="[170,402,1498,1522]" italics="true" pageId="16" pageNumber="221">Phoxinus dragarum</emphasis>
</taxonomicName>
is distinguished from the other French 
<taxonomicName box="[212,394,1530,1554]" pageId="16" pageNumber="221">
<emphasis box="[212,311,1530,1554]" italics="true" pageId="16" pageNumber="221">Phoxinus</emphasis>
species
</taxonomicName>
by its unique nuptial colour pattern. The male has a green stripe in Z3 reaching anal fin base (vs. stripe black or absent), and greenish yellow Z4 (
<figureCitation box="[652,766,1594,1619]" captionStart="Figure 8" captionStartId="15.[813,876,1315,1337]" captionTargetBox="[813,1456,215,1298]" captionTargetId="figure-514@15.[813,1456,215,1298]" captionTargetPageId="15" captionText="Figure 8. – Phoxinus dragarum Holotype m (A), 49.52 mm SL from the Arrat-Devant Stream, Manent-Montané 43°20.7’N, 0°37.5’E, MNHN 2019-0262; paratypes m (B) and f (C), same location as holotype, MNHN 2014-2725; nuptial colouration pattern for male (D) and female (E)." figureDoi="http://doi.org/10.5281/zenodo.7806296" httpUri="https://zenodo.org/record/7806296/files/figure.png" pageId="16" pageNumber="221">Fig. 8B, D</figureCitation>
) (vs. greenor golden yellow). The female has a green-ochre colouration in Z4 (vs. yellow, green, greyish or white), and a black line in Z5 from pectoral fin bases to anal fin base (
<figureCitation box="[140,249,1722,1747]" captionStart="Figure 8" captionStartId="15.[813,876,1315,1337]" captionTargetBox="[813,1456,215,1298]" captionTargetId="figure-514@15.[813,1456,215,1298]" captionTargetPageId="15" captionText="Figure 8. – Phoxinus dragarum Holotype m (A), 49.52 mm SL from the Arrat-Devant Stream, Manent-Montané 43°20.7’N, 0°37.5’E, MNHN 2019-0262; paratypes m (B) and f (C), same location as holotype, MNHN 2014-2725; nuptial colouration pattern for male (D) and female (E)." figureDoi="http://doi.org/10.5281/zenodo.7806296" httpUri="https://zenodo.org/record/7806296/files/figure.png" pageId="16" pageNumber="221">Fig. 8C, E</figureCitation>
) (vs. absence of black line).
</paragraph>
<paragraph blockId="16.[132,776,1466,1907]" lastBlockId="16.[813,1456,217,1202]" pageId="16" pageNumber="221">
<taxonomicName authorityName="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith" authorityYear="2020" box="[170,410,1754,1778]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="dragarum">
<emphasis box="[170,410,1754,1778]" italics="true" pageId="16" pageNumber="221">Phoxinus dragarum</emphasis>
</taxonomicName>
is further distinguished from 
<taxonomicName authorityName="Kottelat" authorityYear="2007" box="[132,232,1787,1811]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="bigerri">
<emphasis box="[132,232,1787,1811]" italics="true" pageId="16" pageNumber="221">P. bigerri</emphasis>
</taxonomicName>
by having 50-74 pored scales (vs. 68-87), a slight- ly stout snout (vs. very stout), and a straight to slightly concave anal fin margin (vs. straight to convex). It is further distinguished from 
<taxonomicName authorityName="Hanko" authorityYear="1922" box="[344,427,1883,1907]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="csikii">
<emphasis box="[344,427,1883,1907]" italics="true" pageId="16" pageNumber="221">P. csikii</emphasis>
</taxonomicName>
by having a snout slightly stout (vs. very stout and blunt), and a straight to slightly concave anal fin margin (vs. straight to slightly convex). The new species is further distinguished from 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1216,1340,281,305]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="phoxinus">
<emphasis box="[1216,1340,281,305]" italics="true" pageId="16" pageNumber="221">P. phoxinus</emphasis>
</taxonomicName>
by having an incomplete lateral line usually reaching anal fin base or just after the last ray (vs. reaching to caudal peduncle), and a slightly stout snout (vs. slightly pointed).
</paragraph>
<paragraph blockId="16.[813,1456,217,1202]" pageId="16" pageNumber="221">
<taxonomicName authorityName="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith" authorityYear="2020" box="[850,1064,409,433]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="dragarum">
<emphasis box="[850,1064,409,433]" italics="true" pageId="16" pageNumber="221">Phoxinus dragarum</emphasis>
</taxonomicName>
is further distinguished from 
<taxonomicName authorityName="Kottelat" authorityYear="2007" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="septimaniae">
<emphasis italics="true" pageId="16" pageNumber="221">P. septimaniae</emphasis>
</taxonomicName>
by having an incomplete lateral line usually reaching anal fin base or just after the last ray (vs. reaching cau- dal peduncle or almost complete), 50-74 pored scales (vs. 73-86), a slightly stout snout (vs. slightly pointed), and 2 scaled patches at the pectoral fin bases separated by an unscaled area (vs. scaled breath between the pectoral fin bases in some populations).
</paragraph>
<paragraph blockId="16.[813,1456,217,1202]" pageId="16" pageNumber="221">
The nuptial male 
<taxonomicName authorityName="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith" authorityYear="2020" box="[1037,1170,665,689]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="dragarum">
<emphasis box="[1037,1170,665,689]" italics="true" pageId="16" pageNumber="221">P. dragarum</emphasis>
</taxonomicName>
is further distinguished by having a red belly, the red colour ending before the caudal peduncle (vs. belly greyish to black in 
<taxonomicName authorityName="Kottelat" authorityYear="2007" box="[1240,1343,729,753]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="bigerri">
<emphasis box="[1240,1343,729,753]" italics="true" pageId="16" pageNumber="221">P. bigerri</emphasis>
</taxonomicName>
; red belly extending to the caudal peduncle in 
<taxonomicName authorityName="Hanko" authorityYear="1922" box="[1195,1276,761,785]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="csikii">
<emphasis box="[1195,1276,761,785]" italics="true" pageId="16" pageNumber="221">P. csikii</emphasis>
</taxonomicName>
; belly pinkish to light orange in 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[973,1095,793,817]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="phoxinus">
<emphasis box="[973,1095,793,817]" italics="true" pageId="16" pageNumber="221">P. phoxinus</emphasis>
</taxonomicName>
; white in 
<taxonomicName authorityName="Kottelat" authorityYear="2007" box="[1201,1355,794,817]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="septimaniae">
<emphasis box="[1201,1355,794,817]" italics="true" pageId="16" pageNumber="221">P. septimaniae</emphasis>
</taxonomicName>
), lacking a yellow dot between the mouth and the eye (vs. presence in 
<taxonomicName authorityName="Kottelat" authorityYear="2007" box="[841,999,858,881]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="septimaniae">
<emphasis box="[841,999,858,881]" italics="true" pageId="16" pageNumber="221">P. septimaniae</emphasis>
</taxonomicName>
), having red lips (vs. pinkish in 
<taxonomicName authorityName="Kottelat" authorityYear="2007" box="[1353,1455,857,881]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="bigerri">
<emphasis box="[1353,1455,857,881]" italics="true" pageId="16" pageNumber="221">P. bigerri</emphasis>
</taxonomicName>
and 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[859,985,889,913]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="phoxinus">
<emphasis box="[859,985,889,913]" italics="true" pageId="16" pageNumber="221">P. phoxinus</emphasis>
</taxonomicName>
), yellow pectoral, pelvic and anal fins (vs. orange in 
<taxonomicName authorityName="Kottelat" authorityYear="2007" box="[924,1029,921,945]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="bigerri">
<emphasis box="[924,1029,921,945]" italics="true" pageId="16" pageNumber="221">P. bigerri</emphasis>
</taxonomicName>
; yellow-orange pectoral fin with a red disc at the interior side in 
<taxonomicName authorityName="Kottelat" authorityYear="2007" box="[1107,1270,954,977]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="septimaniae">
<emphasis box="[1107,1270,954,977]" italics="true" pageId="16" pageNumber="221">P. septimaniae</emphasis>
</taxonomicName>
), and red pectoral, pelvic and anal fins bases (
<emphasis box="[1152,1175,986,1009]" italics="true" pageId="16" pageNumber="221">vs</emphasis>
. pinkish in 
<taxonomicName authorityName="Kottelat" authorityYear="2007" box="[1305,1408,986,1010]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="bigerri">
<emphasis box="[1305,1408,986,1010]" italics="true" pageId="16" pageNumber="221">P. bigerri</emphasis>
</taxonomicName>
and 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[813,937,1018,1042]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="phoxinus">
<emphasis box="[813,937,1018,1042]" italics="true" pageId="16" pageNumber="221">P. phoxinus</emphasis>
</taxonomicName>
). The nuptial female is further distinguished by having a black stripe along Z3 (vs. no stripe in 
<taxonomicName authorityName="Kottelat" authorityYear="2007" box="[1320,1421,1050,1074]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="bigerri">
<emphasis box="[1320,1421,1050,1074]" italics="true" pageId="16" pageNumber="221">P. bigerri</emphasis>
</taxonomicName>
), a black Z5 (vs. white in 
<taxonomicName authorityName="Kottelat" authorityYear="2007" box="[1054,1155,1082,1106]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="bigerri">
<emphasis box="[1054,1155,1082,1106]" italics="true" pageId="16" pageNumber="221">P. bigerri</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Hanko" authorityYear="1922" box="[1207,1290,1082,1106]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="csikii">
<emphasis box="[1207,1290,1082,1106]" italics="true" pageId="16" pageNumber="221">P. csikii</emphasis>
</taxonomicName>
), a pale yellow spot on the operculum (vs. absence in 
<taxonomicName authorityName="Kottelat" authorityYear="2007" box="[1215,1315,1114,1138]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="bigerri">
<emphasis box="[1215,1315,1114,1138]" italics="true" pageId="16" pageNumber="221">P. bigerri</emphasis>
</taxonomicName>
; marked yel- low in 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[886,1007,1146,1170]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="phoxinus">
<emphasis box="[886,1007,1146,1170]" italics="true" pageId="16" pageNumber="221">P. phoxinus</emphasis>
</taxonomicName>
), and pale yellow pectoral fin (vs. marked yellow in 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[918,1039,1178,1202]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="221" phylum="Chordata" rank="species" species="phoxinus">
<emphasis box="[918,1039,1178,1202]" italics="true" pageId="16" pageNumber="221">P. phoxinus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="17" lastPageNumber="222" pageId="16" pageNumber="221" type="description">
<paragraph blockId="16.[813,1456,1242,1907]" box="[813,945,1242,1266]" pageId="16" pageNumber="221">
<heading bold="true" box="[813,945,1242,1266]" fontSize="10" level="3" pageId="16" pageNumber="221" reason="6">
<emphasis bold="true" box="[813,945,1242,1266]" pageId="16" pageNumber="221">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="16.[813,1456,1242,1907]" pageId="16" pageNumber="221">Morphometric data is shown in table IV. Largest examined specimen 66.5 mm SL. Body moderately elongate, dorsal and ventral profile about symmetric or ventral pro- file slightly more convex. Dorsal profile of head markedly more convex than ventral one. Caudal peduncle length 2.3- 3.5 times in depth of caudal peduncle. Snout slightly stout. Mouth terminal or slightly subterminal, upper lip and snout not projecting beyond tip of lower jaw. Tip of upper lip at level or above level of lowest point of eye. In lateral view, eye not or almost flushed with dorsal profile. No supraorbital ridge.</paragraph>
<paragraph blockId="16.[813,1456,1242,1907]" lastBlockId="17.[132,775,1178,1458]" lastPageId="17" lastPageNumber="222" pageId="16" pageNumber="221">
Dorsal fin with 3 simple and 6½-8½ branched rays. Dis- tal dorsal fin margin markedly convex, second branched ray longest, dorsal fin origin behind vertical of pelvic fin origin. Pectoral fin with 1 simple and 13-18 branched rays (10 for 
<specimenCount box="[813,966,1754,1778]" pageId="16" pageNumber="221" type="generic">one specimen</specimenCount>
; usually 14-15), reaching to pelvic fin origin in male, not reaching it in female. Pelvic fin with 1 simple and 5-8 branched rays (usually 7-8), reaching slightly beyond anal fin origin in male, not reaching in female. Anal fin with 3 simple and 6½-8½ branched rays. Anal fin origin slightly in front of vertical through insertion of dorsal fin or behind. Anal fin margin straight to slightly concave, second branched ray longest.
</paragraph>
<caption ID-Table-UUID="DF52664AC75EFFFFFF60FF40FCC9FE9C" httpUri="http://table.plazi.org/id/DF52664AC75EFFFFFF60FF40FCC9FE9C" pageId="17" pageNumber="222" startId="17.[132,186,217,239]" targetBox="[139,769,351,1138]" targetIsTable="true" targetPageId="17" targetType="table">
<paragraph blockId="17.[132,776,217,335]" pageId="17" pageNumber="222">
Table IV. 
<emphasis box="[228,240,218,239]" italics="true" pageId="17" pageNumber="222">–</emphasis>
Morphometric data of 
<taxonomicName authorityName="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith" authorityYear="2020" box="[470,666,217,238]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="222" phylum="Chordata" rank="species" species="dragarum">
<emphasis box="[470,666,217,238]" italics="true" pageId="17" pageNumber="222">Phoxinus dragarum</emphasis>
</taxonomicName>
(holotype: MNHN 2019-0262; paratypes: MNHN 2014-2725, MNHN 2014- 2813, MNHN 2014-2807, MNHN 2014-2735, MNHN 2014-2738; and additional materials (MNHN 2014-2761, MNHN 2014-2759, MNHN 2014-2731; n = 52). The calculations include the holotype.
</paragraph>
</caption>
<paragraph pageId="17" pageNumber="222">
<table box="[139,769,351,1138]" gridcols="6" gridrows="23" pageId="17" pageNumber="222">
<tr box="[139,769,351,408]" gridrow="0" pageId="17" pageNumber="222" rowspan-0="1">
<th box="[384,473,351,408]" gridcol="1" gridrow="0" pageId="17" pageNumber="222">Holotype</th>
<th box="[487,769,351,408]" colspan="4" colspanRight="3" gridcol="2" gridrow="0" pageId="17" pageNumber="222">Holotype &amp; other individuals Mean Min Max SD</th>
</tr>
<tr box="[139,769,420,442]" gridrow="1" pageId="17" pageNumber="222">
<th box="[139,370,420,442]" gridcol="0" gridrow="1" pageId="17" pageNumber="222">SL (mm)</th>
<td box="[384,473,420,442]" gridcol="1" gridrow="1" pageId="17" pageNumber="222">49.5</td>
<td box="[487,546,420,442]" gridcol="2" gridrow="1" pageId="17" pageNumber="222">52.8</td>
<td box="[584,626,420,442]" gridcol="3" gridrow="1" pageId="17" pageNumber="222">43.6</td>
<td box="[658,700,420,442]" gridcol="4" gridrow="1" pageId="17" pageNumber="222">66.5</td>
<td box="[731,769,420,442]" gridcol="5" gridrow="1" pageId="17" pageNumber="222">5.6</td>
</tr>
<tr box="[139,769,453,475]" gridrow="2" pageId="17" pageNumber="222">
<th box="[139,769,453,475]" colspan="6" colspanRight="5" gridcol="0" gridrow="2" pageId="17" pageNumber="222">
<emphasis box="[139,417,453,475]" italics="true" pageId="17" pageNumber="222">In percent of standard length</emphasis>
</th>
</tr>
<tr box="[139,769,486,508]" gridrow="3" pageId="17" pageNumber="222">
<th box="[139,370,486,508]" gridcol="0" gridrow="3" pageId="17" pageNumber="222">Head length</th>
<td box="[384,473,486,508]" gridcol="1" gridrow="3" pageId="17" pageNumber="222">26.5</td>
<td box="[487,546,486,508]" gridcol="2" gridrow="3" pageId="17" pageNumber="222">26.7</td>
<td box="[584,626,486,508]" gridcol="3" gridrow="3" pageId="17" pageNumber="222">24.0</td>
<td box="[658,700,486,508]" gridcol="4" gridrow="3" pageId="17" pageNumber="222">29.7</td>
<td box="[731,769,486,508]" gridcol="5" gridrow="3" pageId="17" pageNumber="222">1.1</td>
</tr>
<tr box="[139,769,519,542]" gridrow="4" pageId="17" pageNumber="222">
<th box="[139,370,519,542]" gridcol="0" gridrow="4" pageId="17" pageNumber="222">Predorsal length</th>
<td box="[384,473,519,542]" gridcol="1" gridrow="4" pageId="17" pageNumber="222">52.7</td>
<td box="[487,546,519,542]" gridcol="2" gridrow="4" pageId="17" pageNumber="222">56.3</td>
<td box="[584,626,519,542]" gridcol="3" gridrow="4" pageId="17" pageNumber="222">52.7</td>
<td box="[658,700,519,542]" gridcol="4" gridrow="4" pageId="17" pageNumber="222">59.6</td>
<td box="[731,769,519,542]" gridcol="5" gridrow="4" pageId="17" pageNumber="222">1.5</td>
</tr>
<tr box="[139,769,553,575]" gridrow="5" pageId="17" pageNumber="222">
<th box="[139,370,553,575]" gridcol="0" gridrow="5" pageId="17" pageNumber="222">Post dorsal length</th>
<td box="[384,473,553,575]" gridcol="1" gridrow="5" pageId="17" pageNumber="222">36.9</td>
<td box="[487,546,553,575]" gridcol="2" gridrow="5" pageId="17" pageNumber="222">36.2</td>
<td box="[584,626,553,575]" gridcol="3" gridrow="5" pageId="17" pageNumber="222">33.7</td>
<td box="[658,700,553,575]" gridcol="4" gridrow="5" pageId="17" pageNumber="222">39.9</td>
<td box="[731,769,553,575]" gridcol="5" gridrow="5" pageId="17" pageNumber="222">1.4</td>
</tr>
<tr box="[139,769,586,608]" gridrow="6" pageId="17" pageNumber="222">
<th box="[139,370,586,608]" gridcol="0" gridrow="6" pageId="17" pageNumber="222">Prepectoral length</th>
<td box="[384,473,586,608]" gridcol="1" gridrow="6" pageId="17" pageNumber="222">27.5</td>
<td box="[487,546,586,608]" gridcol="2" gridrow="6" pageId="17" pageNumber="222">26.1</td>
<td box="[584,626,586,608]" gridcol="3" gridrow="6" pageId="17" pageNumber="222">23.7</td>
<td box="[658,700,586,608]" gridcol="4" gridrow="6" pageId="17" pageNumber="222">28.2</td>
<td box="[731,769,586,608]" gridcol="5" gridrow="6" pageId="17" pageNumber="222">1.1</td>
</tr>
<tr box="[139,769,619,641]" gridrow="7" pageId="17" pageNumber="222">
<th box="[139,370,619,641]" gridcol="0" gridrow="7" pageId="17" pageNumber="222">Prepelvic length</th>
<td box="[384,473,619,641]" gridcol="1" gridrow="7" pageId="17" pageNumber="222">46.8</td>
<td box="[487,546,619,641]" gridcol="2" gridrow="7" pageId="17" pageNumber="222">48.6</td>
<td box="[584,626,619,641]" gridcol="3" gridrow="7" pageId="17" pageNumber="222">44.3</td>
<td box="[658,700,619,641]" gridcol="4" gridrow="7" pageId="17" pageNumber="222">52.2</td>
<td box="[731,769,619,641]" gridcol="5" gridrow="7" pageId="17" pageNumber="222">1.8</td>
</tr>
<tr box="[139,769,652,674]" gridrow="8" pageId="17" pageNumber="222">
<th box="[139,370,652,674]" gridcol="0" gridrow="8" pageId="17" pageNumber="222">Preanal length</th>
<td box="[384,473,652,674]" gridcol="1" gridrow="8" pageId="17" pageNumber="222">65.2</td>
<td box="[487,546,652,674]" gridcol="2" gridrow="8" pageId="17" pageNumber="222">64.9</td>
<td box="[584,626,652,674]" gridcol="3" gridrow="8" pageId="17" pageNumber="222">57.2</td>
<td box="[658,700,652,674]" gridcol="4" gridrow="8" pageId="17" pageNumber="222">68.4</td>
<td box="[731,769,652,674]" gridcol="5" gridrow="8" pageId="17" pageNumber="222">2.0</td>
</tr>
<tr box="[139,769,685,707]" gridrow="9" pageId="17" pageNumber="222">
<th box="[139,370,685,707]" gridcol="0" gridrow="9" pageId="17" pageNumber="222">Body depth</th>
<td box="[384,473,685,707]" gridcol="1" gridrow="9" pageId="17" pageNumber="222">21.0</td>
<td box="[487,546,685,707]" gridcol="2" gridrow="9" pageId="17" pageNumber="222">23.2</td>
<td box="[584,626,685,707]" gridcol="3" gridrow="9" pageId="17" pageNumber="222">19.5</td>
<td box="[658,700,685,707]" gridcol="4" gridrow="9" pageId="17" pageNumber="222">26.4</td>
<td box="[731,769,685,707]" gridcol="5" gridrow="9" pageId="17" pageNumber="222">1.7</td>
</tr>
<tr box="[139,769,718,740]" gridrow="10" pageId="17" pageNumber="222">
<th box="[139,370,718,740]" gridcol="0" gridrow="10" pageId="17" pageNumber="222">Distance dorsal anal</th>
<td box="[384,473,718,740]" gridcol="1" gridrow="10" pageId="17" pageNumber="222">20.6</td>
<td box="[487,546,718,740]" gridcol="2" gridrow="10" pageId="17" pageNumber="222">23.3</td>
<td box="[584,626,718,740]" gridcol="3" gridrow="10" pageId="17" pageNumber="222">19.9</td>
<td box="[658,700,718,740]" gridcol="4" gridrow="10" pageId="17" pageNumber="222">26.4</td>
<td box="[731,769,718,740]" gridcol="5" gridrow="10" pageId="17" pageNumber="222">1.4</td>
</tr>
<tr box="[139,769,751,773]" gridrow="11" pageId="17" pageNumber="222">
<th box="[139,370,751,773]" gridcol="0" gridrow="11" pageId="17" pageNumber="222">Distance pectoral pelvic</th>
<td box="[384,473,751,773]" gridcol="1" gridrow="11" pageId="17" pageNumber="222">18.9</td>
<td box="[487,546,751,773]" gridcol="2" gridrow="11" pageId="17" pageNumber="222">20.8</td>
<td box="[584,626,751,773]" gridcol="3" gridrow="11" pageId="17" pageNumber="222">16.4</td>
<td box="[658,700,751,773]" gridcol="4" gridrow="11" pageId="17" pageNumber="222">25.7</td>
<td box="[731,769,751,773]" gridcol="5" gridrow="11" pageId="17" pageNumber="222">2.4</td>
</tr>
<tr box="[139,769,784,807]" gridrow="12" pageId="17" pageNumber="222">
<th box="[139,370,784,807]" gridcol="0" gridrow="12" pageId="17" pageNumber="222">Pectoral fin length</th>
<td box="[384,473,784,807]" gridcol="1" gridrow="12" pageId="17" pageNumber="222">20.2</td>
<td box="[487,546,784,807]" gridcol="2" gridrow="12" pageId="17" pageNumber="222">18.2</td>
<td box="[584,626,784,807]" gridcol="3" gridrow="12" pageId="17" pageNumber="222">16.0</td>
<td box="[658,700,784,807]" gridcol="4" gridrow="12" pageId="17" pageNumber="222">21.1</td>
<td box="[731,769,784,807]" gridcol="5" gridrow="12" pageId="17" pageNumber="222">1.3</td>
</tr>
<tr box="[139,769,817,840]" gridrow="13" pageId="17" pageNumber="222">
<th box="[139,370,817,840]" gridcol="0" gridrow="13" pageId="17" pageNumber="222">Pelvic fin length</th>
<td box="[384,473,817,840]" gridcol="1" gridrow="13" pageId="17" pageNumber="222">16.4</td>
<td box="[487,546,817,840]" gridcol="2" gridrow="13" pageId="17" pageNumber="222">14.7</td>
<td box="[584,626,817,840]" gridcol="3" gridrow="13" pageId="17" pageNumber="222">12.5</td>
<td box="[658,700,817,840]" gridcol="4" gridrow="13" pageId="17" pageNumber="222">18.9</td>
<td box="[731,769,817,840]" gridcol="5" gridrow="13" pageId="17" pageNumber="222">1.5</td>
</tr>
<tr box="[139,769,851,873]" gridrow="14" pageId="17" pageNumber="222">
<th box="[139,370,851,873]" gridcol="0" gridrow="14" pageId="17" pageNumber="222">Dorsal fin length</th>
<td box="[384,473,851,873]" gridcol="1" gridrow="14" pageId="17" pageNumber="222">22.3</td>
<td box="[487,546,851,873]" gridcol="2" gridrow="14" pageId="17" pageNumber="222">20.3</td>
<td box="[584,626,851,873]" gridcol="3" gridrow="14" pageId="17" pageNumber="222">18.1</td>
<td box="[658,700,851,873]" gridcol="4" gridrow="14" pageId="17" pageNumber="222">24.2</td>
<td box="[731,769,851,873]" gridcol="5" gridrow="14" pageId="17" pageNumber="222">1.4</td>
</tr>
<tr box="[139,769,884,906]" gridrow="15" pageId="17" pageNumber="222">
<th box="[139,370,884,906]" gridcol="0" gridrow="15" pageId="17" pageNumber="222">Anal fin length</th>
<td box="[384,473,884,906]" gridcol="1" gridrow="15" pageId="17" pageNumber="222">22.3</td>
<td box="[487,546,884,906]" gridcol="2" gridrow="15" pageId="17" pageNumber="222">20.0</td>
<td box="[584,626,884,906]" gridcol="3" gridrow="15" pageId="17" pageNumber="222">18.1</td>
<td box="[658,700,884,906]" gridcol="4" gridrow="15" pageId="17" pageNumber="222">22.8</td>
<td box="[731,769,884,906]" gridcol="5" gridrow="15" pageId="17" pageNumber="222">1.1</td>
</tr>
<tr box="[139,769,917,939]" gridrow="16" pageId="17" pageNumber="222">
<th box="[139,370,917,939]" gridcol="0" gridrow="16" pageId="17" pageNumber="222">Caudal peduncle length</th>
<td box="[384,473,917,939]" gridcol="1" gridrow="16" pageId="17" pageNumber="222">28.0</td>
<td box="[487,546,917,939]" gridcol="2" gridrow="16" pageId="17" pageNumber="222">25.9</td>
<td box="[584,626,917,939]" gridcol="3" gridrow="16" pageId="17" pageNumber="222">23.5</td>
<td box="[658,700,917,939]" gridcol="4" gridrow="16" pageId="17" pageNumber="222">28.1</td>
<td box="[731,769,917,939]" gridcol="5" gridrow="16" pageId="17" pageNumber="222">1.2</td>
</tr>
<tr box="[139,769,950,972]" gridrow="17" pageId="17" pageNumber="222">
<th box="[139,370,950,972]" gridcol="0" gridrow="17" pageId="17" pageNumber="222">Caudal peduncle depth</th>
<td box="[384,473,950,972]" gridcol="1" gridrow="17" pageId="17" pageNumber="222">9.1</td>
<td box="[487,546,950,972]" gridcol="2" gridrow="17" pageId="17" pageNumber="222">9.3</td>
<td box="[584,626,950,972]" gridcol="3" gridrow="17" pageId="17" pageNumber="222">7.7</td>
<td box="[658,700,950,972]" gridcol="4" gridrow="17" pageId="17" pageNumber="222">14.1</td>
<td box="[731,769,950,972]" gridcol="5" gridrow="17" pageId="17" pageNumber="222">0.9</td>
</tr>
<tr box="[139,769,983,1005]" gridrow="18" pageId="17" pageNumber="222">
<th box="[139,769,983,1005]" colspan="6" colspanRight="5" gridcol="0" gridrow="18" pageId="17" pageNumber="222">
<emphasis box="[139,379,983,1005]" italics="true" pageId="17" pageNumber="222">In percent of head length</emphasis>
</th>
</tr>
<tr box="[139,769,1016,1038]" gridrow="19" pageId="17" pageNumber="222">
<th box="[139,370,1016,1038]" gridcol="0" gridrow="19" pageId="17" pageNumber="222">Head depth</th>
<td box="[384,473,1016,1038]" gridcol="1" gridrow="19" pageId="17" pageNumber="222">51.3</td>
<td box="[487,546,1016,1038]" gridcol="2" gridrow="19" pageId="17" pageNumber="222">51.1</td>
<td box="[584,626,1016,1038]" gridcol="3" gridrow="19" pageId="17" pageNumber="222">46.9</td>
<td box="[658,700,1016,1038]" gridcol="4" gridrow="19" pageId="17" pageNumber="222">54.3</td>
<td box="[731,769,1016,1038]" gridcol="5" gridrow="19" pageId="17" pageNumber="222">1.9</td>
</tr>
<tr box="[139,769,1049,1071]" gridrow="20" pageId="17" pageNumber="222">
<th box="[139,370,1049,1071]" gridcol="0" gridrow="20" pageId="17" pageNumber="222">Eye diameter</th>
<td box="[384,473,1049,1071]" gridcol="1" gridrow="20" pageId="17" pageNumber="222">25.5</td>
<td box="[487,546,1049,1071]" gridcol="2" gridrow="20" pageId="17" pageNumber="222">25.8</td>
<td box="[584,626,1049,1071]" gridcol="3" gridrow="20" pageId="17" pageNumber="222">21.5</td>
<td box="[658,700,1049,1071]" gridcol="4" gridrow="20" pageId="17" pageNumber="222">30.6</td>
<td box="[731,769,1049,1071]" gridcol="5" gridrow="20" pageId="17" pageNumber="222">2.0</td>
</tr>
<tr box="[139,769,1082,1105]" gridrow="21" pageId="17" pageNumber="222">
<th box="[139,370,1082,1105]" gridcol="0" gridrow="21" pageId="17" pageNumber="222">Snout length</th>
<td box="[384,473,1082,1105]" gridcol="1" gridrow="21" pageId="17" pageNumber="222">29.8</td>
<td box="[487,546,1082,1105]" gridcol="2" gridrow="21" pageId="17" pageNumber="222">29.2</td>
<td box="[584,626,1082,1105]" gridcol="3" gridrow="21" pageId="17" pageNumber="222">23.7</td>
<td box="[658,700,1082,1105]" gridcol="4" gridrow="21" pageId="17" pageNumber="222">35.6</td>
<td box="[731,769,1082,1105]" gridcol="5" gridrow="21" pageId="17" pageNumber="222">2.5</td>
</tr>
<tr box="[139,769,1116,1138]" gridrow="22" pageId="17" pageNumber="222">
<th box="[139,370,1116,1138]" gridcol="0" gridrow="22" pageId="17" pageNumber="222">Inter-orbital width</th>
<td box="[384,473,1116,1138]" gridcol="1" gridrow="22" pageId="17" pageNumber="222">27.1</td>
<td box="[487,546,1116,1138]" gridcol="2" gridrow="22" pageId="17" pageNumber="222">33.3</td>
<td box="[584,626,1116,1138]" gridcol="3" gridrow="22" pageId="17" pageNumber="222">25.8</td>
<td box="[658,700,1116,1138]" gridcol="4" gridrow="22" pageId="17" pageNumber="222">40.4</td>
<td box="[731,769,1116,1138]" gridcol="5" gridrow="22" pageId="17" pageNumber="222">3.0</td>
</tr>
</table>
</paragraph>
<paragraph blockId="17.[132,775,1178,1458]" pageId="17" pageNumber="222">Lateral line incomplete usually reaching to vertical of dorsal-fin base or to vertical of base of last anal fin ray. 70-97 lateral scales, including 50-74 pored scales. Two scaled patches at pectoral fin insertions separated by an unscaled area. No scales between pectoral fin bases. Nuptial tubercles observed in both sexes.</paragraph>
<paragraph blockId="17.[132,775,1498,1906]" box="[132,354,1498,1522]" pageId="17" pageNumber="222">
<heading bold="true" box="[132,354,1498,1522]" fontSize="10" level="3" pageId="17" pageNumber="222" reason="6">
<emphasis bold="true" box="[132,354,1498,1522]" pageId="17" pageNumber="222">Nuptial colouration</emphasis>
</heading>
</paragraph>
<paragraph blockId="17.[132,775,1498,1906]" pageId="17" pageNumber="222">
<emphasis box="[170,226,1530,1554]" italics="true" pageId="17" pageNumber="222">Male</emphasis>
. – Snout and top of head dark-brown. Ablack bar on operculum reaching from top of head to branchiostegal rays. Cheeks white. Operculum with a white spot at uppermost, posterior corner. Ayellow spot on suboperculum. Lips red. Z1 brown to black. Z2 pale brown. Black vermiculations in Z1 to Z2. Z3 green. Z4 yellow or yellowish-green. Ablack line along Z5 from pectoral fin origin to caudal peduncle. Belly red, red couloration not extending to caudal peduncle. Fins yellow-orange. (
<figureCitation box="[359,474,1786,1811]" captionStart="Figure 8" captionStartId="15.[813,876,1315,1337]" captionTargetBox="[813,1456,215,1298]" captionTargetId="figure-514@15.[813,1456,215,1298]" captionTargetPageId="15" captionText="Figure 8. – Phoxinus dragarum Holotype m (A), 49.52 mm SL from the Arrat-Devant Stream, Manent-Montané 43°20.7’N, 0°37.5’E, MNHN 2019-0262; paratypes m (B) and f (C), same location as holotype, MNHN 2014-2725; nuptial colouration pattern for male (D) and female (E)." figureDoi="http://doi.org/10.5281/zenodo.7806296" httpUri="https://zenodo.org/record/7806296/files/figure.png" pageId="17" pageNumber="222">Fig. 8B, D</figureCitation>
).
</paragraph>
<paragraph blockId="17.[132,775,1498,1906]" lastBlockId="17.[813,1455,217,305]" pageId="17" pageNumber="222">
<emphasis box="[170,253,1818,1842]" italics="true" pageId="17" pageNumber="222">Female</emphasis>
. – Z3 with a black stripe. Z4 green-ochre until anal fin. Ablack line along Z5 from pectoral fin origin to caudal peduncle. Pinkish tinge visible at base of pectoral, pelvic and anal fins and on mouth. Belly white. White and yellow spots on operculum and white spots at origin of pectoral, pelvic and anal fins (
<figureCitation box="[1093,1206,281,305]" captionStart="Figure 8" captionStartId="15.[813,876,1315,1337]" captionTargetBox="[813,1456,215,1298]" captionTargetId="figure-514@15.[813,1456,215,1298]" captionTargetPageId="15" captionText="Figure 8. – Phoxinus dragarum Holotype m (A), 49.52 mm SL from the Arrat-Devant Stream, Manent-Montané 43°20.7’N, 0°37.5’E, MNHN 2019-0262; paratypes m (B) and f (C), same location as holotype, MNHN 2014-2725; nuptial colouration pattern for male (D) and female (E)." figureDoi="http://doi.org/10.5281/zenodo.7806296" httpUri="https://zenodo.org/record/7806296/files/figure.png" pageId="17" pageNumber="222">Fig. 8C, E</figureCitation>
).
</paragraph>
<paragraph blockId="17.[813,1455,345,466]" box="[813,1128,345,369]" pageId="17" pageNumber="222">
<heading bold="true" box="[813,1128,345,369]" fontSize="10" level="3" pageId="17" pageNumber="222" reason="6">
<emphasis bold="true" box="[813,1128,345,369]" pageId="17" pageNumber="222">Colouration in preservation</emphasis>
</heading>
</paragraph>
<paragraph blockId="17.[813,1455,345,466]" pageId="17" pageNumber="222">
Dark bars remain after preservation in ethanol. Green colouration turns to dark grey, yellow and red colourations faded and usually absent (
<figureCitation box="[1087,1173,441,466]" captionStart="Figure 8" captionStartId="15.[813,876,1315,1337]" captionTargetBox="[813,1456,215,1298]" captionTargetId="figure-514@15.[813,1456,215,1298]" captionTargetPageId="15" captionText="Figure 8. – Phoxinus dragarum Holotype m (A), 49.52 mm SL from the Arrat-Devant Stream, Manent-Montané 43°20.7’N, 0°37.5’E, MNHN 2019-0262; paratypes m (B) and f (C), same location as holotype, MNHN 2014-2725; nuptial colouration pattern for male (D) and female (E)." figureDoi="http://doi.org/10.5281/zenodo.7806296" httpUri="https://zenodo.org/record/7806296/files/figure.png" pageId="17" pageNumber="222">Fig. 8A</figureCitation>
).
</paragraph>
<paragraph blockId="17.[813,1456,505,754]" box="[813,1122,505,530]" pageId="17" pageNumber="222">
<heading bold="true" box="[813,1122,505,530]" fontSize="10" level="3" pageId="17" pageNumber="222" reason="6">
<emphasis bold="true" box="[813,1122,505,530]" pageId="17" pageNumber="222">Molecular characterization</emphasis>
</heading>
</paragraph>
<paragraph blockId="17.[813,1456,505,754]" pageId="17" pageNumber="222">
On the COI marker, 
<taxonomicName authorityName="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith" authorityYear="2020" box="[1068,1200,537,561]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="222" phylum="Chordata" rank="species" species="dragarum">
<emphasis box="[1068,1200,537,561]" italics="true" pageId="17" pageNumber="222">P. dragarum</emphasis>
</taxonomicName>
has six diagnostic sites, including two synapomorphies. However, the Garonne and Dordogne populations are characterised both by three diagnostic sites (Tab. II). For the 12S rDNA marker, five diag- nostic sites characterise this species: G (
<emphasis box="[1264,1287,666,689]" italics="true" pageId="17" pageNumber="222">vs</emphasis>
. A) in position 179, A (vs. G) in position 307, T (vs. Aor G) in position 310, C (vs. Aor G) in position 412 and A (vs. G) in position 478.
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="222" type="distribution">
<paragraph blockId="17.[813,1456,793,978]" box="[813,952,793,817]" pageId="17" pageNumber="222">
<heading bold="true" box="[813,952,793,817]" fontSize="10" level="3" pageId="17" pageNumber="222" reason="6">
<emphasis bold="true" box="[813,952,793,817]" pageId="17" pageNumber="222">Distribution</emphasis>
</heading>
</paragraph>
<paragraph blockId="17.[813,1456,793,978]" pageId="17" pageNumber="222">
This species is endemic to the Garonne drainage (
<figureCitation box="[1377,1439,826,850]" captionStart="Figure 1" captionStartId="3.[132,197,1741,1763]" captionTargetBox="[548,1456,1050,1902]" captionTargetId="figure-410@3.[548,1455,1051,1902]" captionTargetPageId="3" captionText="Figure 1. – Records of Phoxinus used in this study. Coloured squares represent the six species delineated in this study: P. phoxinus (blue), P. bigerri (red), P. csikii (green), P. septimaniae (yellow), P. dragarum (purple), and P. fayollarum (orange)." figureDoi="http://doi.org/10.5281/zenodo.7806282" httpUri="https://zenodo.org/record/7806282/files/figure.png" pageId="17" pageNumber="222">Fig. 1</figureCitation>
). It has been introduced to Spain, in the Ebro drainage along the Pyrenean side (
<bibRefCitation author="CORRAL-LOU A. &amp; PEREA S. &amp; APARICIO E. &amp; DOADRIO I." box="[1023,1281,890,914]" journalOrPublisher="J. Zool. Syst. Evol. Res." pageId="17" pageNumber="222" pagination="926 - 994" part="57" refId="ref24596" refString="CORRAL-LOU A., PEREA S., APARICIO E. &amp; DOADRIO I., 2019. - Phylogeography and species delineation of the genus Phoxinus Rafinesque, 1820 (Actinopterygii: Leuciscidae) in the Iberian Peninsula. J. Zool. Syst. Evol. Res., 57 (4): 926 - 994. DOI: 10.1111 / jzs. 12320" title="Phylogeography and species delineation of the genus Phoxinus Rafinesque, 1820 (Actinopterygii: Leuciscidae) in the Iberian Peninsula" type="journal article" year="2019">
Corral-Lou 
<emphasis box="[1154,1217,890,914]" italics="true" pageId="17" pageNumber="222">et al.,</emphasis>
2019
</bibRefCitation>
) and also in the Guadalquivir drainage (
<bibRefCitation author="SAEZ-GOMEZ P. &amp; PRENDA J." box="[1088,1451,922,951]" journalOrPublisher="BioInvasions Rec." pageId="17" pageNumber="222" pagination="924 - 932" part="8" refId="ref28010" refString="SAEZ-GOMEZ P. &amp; PRENDA J., 2019. - Updating the distribu- tion data of recently introduced freshwater fish in the Guadalquivir River Basin (Spain). BioInvasions Rec., 8 (4): 924 - 932. DOI: 10.3391 / bir. 2019.8.4.21" title="Updating the distribu- tion data of recently introduced freshwater fish in the Guadalquivir River Basin (Spain)" type="journal article" year="2019">SÁez-Gómez and Prenda, 2019</bibRefCitation>
: fig. 2).
</paragraph>
</subSubSection>
<subSubSection pageId="17" pageNumber="222" type="biology_ecology">
<paragraph blockId="17.[813,1456,1018,1586]" box="[813,903,1018,1042]" pageId="17" pageNumber="222">
<heading bold="true" box="[813,903,1018,1042]" fontSize="10" level="3" pageId="17" pageNumber="222" reason="6">
<emphasis bold="true" box="[813,903,1018,1042]" pageId="17" pageNumber="222">Ecology</emphasis>
</heading>
</paragraph>
<paragraph blockId="17.[813,1456,1018,1586]" pageId="17" pageNumber="222">
<taxonomicName authorityName="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith" authorityYear="2020" box="[850,1062,1050,1074]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="222" phylum="Chordata" rank="species" species="dragarum">
<emphasis box="[850,1062,1050,1074]" italics="true" pageId="17" pageNumber="222">Phoxinus dragarum</emphasis>
</taxonomicName>
isabundant in the barbelzone (
<bibRefCitation author="IBARRA A. A. &amp; PARK Y. S. &amp; BROSSE S &amp; REYJOL Y. &amp; LIM P. &amp; LEK S." journalOrPublisher="Ecol. Freshw. Fish" pageId="17" pageNumber="222" pagination="233 - 242" part="14" refId="ref25840" refString="IBARRA A. A., PARK Y. S., BROSSE S, REYJOL Y., LIM P. &amp; LEK S., 2005. - Nested patterns of spatial diversity revealed for fish assemblages in a west European river. Ecol. Freshw. Fish, 14: 233 - 242. DOI: 10.1111 / j. 1600 - 0633.2005.00096. x" title="Nested patterns of spatial diversity revealed for fish assemblages in a west European river" type="journal article" year="2005">
Ibarra 
<emphasis box="[813,872,1082,1106]" italics="true" pageId="17" pageNumber="222">et al.,</emphasis>
2005
</bibRefCitation>
). It is usually found in headwater streams (
<bibRefCitation author="IBARRA A. A. &amp; PARK Y. S. &amp; BROSSE S &amp; REYJOL Y. &amp; LIM P. &amp; LEK S." journalOrPublisher="Ecol. Freshw. Fish" pageId="17" pageNumber="222" pagination="233 - 242" part="14" refId="ref25840" refString="IBARRA A. A., PARK Y. S., BROSSE S, REYJOL Y., LIM P. &amp; LEK S., 2005. - Nested patterns of spatial diversity revealed for fish assemblages in a west European river. Ecol. Freshw. Fish, 14: 233 - 242. DOI: 10.1111 / j. 1600 - 0633.2005.00096. x" title="Nested patterns of spatial diversity revealed for fish assemblages in a west European river" type="journal article" year="2005">
Ibarra 
<emphasis box="[813,872,1114,1138]" italics="true" pageId="17" pageNumber="222">et al.,</emphasis>
2005
</bibRefCitation>
; 
<bibRefCitation author="SANTOUL F. &amp; CAYROU J. &amp; MASTRORILLO S. &amp; CEREGHINO R." box="[946,1154,1114,1138]" journalOrPublisher="J. Fish Biol." pageId="17" pageNumber="222" pagination="301 - 314" part="66" refId="ref28060" refString="SANTOUL F., CAYROU J., MASTRORILLO S. &amp; CEREGHINO R., 2005. - Spatial patterns of the biological traits of freshwater fish communities in south-west France. J. Fish Biol., 66: 301 - 314. DOI: 10.1111 / j. 1095 - 8649.2004.00579. x" title="Spatial patterns of the biological traits of freshwater fish communities in south-west France" type="journal article" year="2005">
Santoul 
<emphasis box="[1033,1093,1114,1138]" italics="true" pageId="17" pageNumber="222">et al.,</emphasis>
2005
</bibRefCitation>
) with a low slope in altitude and a high fish species richness (
<bibRefCitation author="CEREGHINO R. &amp; SANTOUL F. &amp; COMPIN A. &amp; FIGUEROLA J. &amp; MASTRORILLO S." box="[1169,1420,1146,1170]" journalOrPublisher="AMBIO" pageId="17" pageNumber="222" pagination="440 - 444" part="34" refId="ref24371" refString="CEREGHINO R., SANTOUL F., COMPIN A., FIGUEROLA J. &amp; MASTRORILLO S., 2005. - Co-occurrence patterns of some small-bodied freshwater fishes in Southwestern France: impli- cations for fish conservation and environmental management. AMBIO, 34 (6): 440 - 444. DOI: 10.1579 / 0044 - 7447 - 34.6.440" title="Co-occurrence patterns of some small-bodied freshwater fishes in Southwestern France: impli- cations for fish conservation and environmental management" type="journal article" year="2005">
Céréghino 
<emphasis box="[1292,1354,1146,1170]" italics="true" pageId="17" pageNumber="222">et al.,</emphasis>
2005
</bibRefCitation>
). It is often found in sympatry with 
<taxonomicName authority="Kottelat &amp; Persat, 2005" authorityName="Kottelat &amp; Persat" authorityYear="2005" class="Actinopterygii" family="Cyprinidae" genus="Gobio" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="222" phylum="Chordata" rank="species" species="occitaniae">
<emphasis box="[1168,1359,1178,1202]" italics="true" pageId="17" pageNumber="222">Gobio occitaniae</emphasis>
Kottelat &amp; Persat, 2005
</taxonomicName>
(
<bibRefCitation author="CEREGHINO R. &amp; SANTOUL F. &amp; COMPIN A. &amp; FIGUEROLA J. &amp; MASTRORILLO S." box="[994,1252,1210,1234]" journalOrPublisher="AMBIO" pageId="17" pageNumber="222" pagination="440 - 444" part="34" refId="ref24371" refString="CEREGHINO R., SANTOUL F., COMPIN A., FIGUEROLA J. &amp; MASTRORILLO S., 2005. - Co-occurrence patterns of some small-bodied freshwater fishes in Southwestern France: impli- cations for fish conservation and environmental management. AMBIO, 34 (6): 440 - 444. DOI: 10.1579 / 0044 - 7447 - 34.6.440" title="Co-occurrence patterns of some small-bodied freshwater fishes in Southwestern France: impli- cations for fish conservation and environmental management" type="journal article" year="2005">
Céréghino 
<emphasis box="[1120,1183,1210,1234]" italics="true" pageId="17" pageNumber="222">et al.,</emphasis>
2005
</bibRefCitation>
). Studies made on populations from the stream Ariège and the Garonne River demonstrated that 
<taxonomicName authorityName="Denys* &amp; Dettai &amp; Persat &amp; Daszkiewicz &amp; Hautecoeur &amp; Keith" authorityYear="2020" box="[1008,1140,1274,1298]" class="Actinopterygii" family="Cyprinidae" genus="Phoxinus" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="222" phylum="Chordata" rank="species" species="dragarum">
<emphasis box="[1008,1140,1274,1298]" italics="true" pageId="17" pageNumber="222">P. dragarum</emphasis>
</taxonomicName>
prefers shallow water located along the bank, with no or low velocity and a mixed substrate (pebbles, gravel, sand and sludge) (
<bibRefCitation author="MASTRORILLO S. &amp; DAUBA F. &amp; BELAUD A." box="[1195,1451,1338,1362]" journalOrPublisher="Int. J. Lim." pageId="17" pageNumber="222" pagination="185 - 195" part="32" refId="ref26789" refString="MASTRORILLO S., DAUBA F. &amp; BELAUD A., 1996. - Utilisa- tion des microhabitats par le vairon, le goujon et la loche fran- che dans trois rivieres du sud-ouest de la France. Int. J. Lim., 32 (3): 185 - 195. DOI: 10.1051 / limn / 1996017" title="Utilisa- tion des microhabitats par le vairon, le goujon et la loche fran- che dans trois rivieres du sud-ouest de la France" type="journal article" year="1996">
Mastrorillo 
<emphasis box="[1326,1388,1338,1362]" italics="true" pageId="17" pageNumber="222">et al.,</emphasis>
1996
</bibRefCitation>
, 
<bibRefCitation author="MASTRORILLO S. &amp; LEK S. &amp; DAUBA F." box="[813,871,1370,1394]" journalOrPublisher="Aquat. Living Resour." pageId="17" pageNumber="222" pagination="169 - 176" part="10" refId="ref26851" refString="MASTRORILLO S., LEK S. &amp; DAUBA F., 1997. - Predicting the abundance of minnow Phoxinus phoxinus (Cyprinidae) in the Ariege (France) using artificial neural networks. Aquat. Living Resour., 10: 169 - 176. DOI: 10.1051 / alr: 1997018" title="Predicting the abundance of minnow Phoxinus phoxinus (Cyprinidae) in the Ariege (France) using artificial neural networks" type="journal article" year="1997">1997</bibRefCitation>
; 
<bibRefCitation author="REYJOL Y. &amp; LIM P. &amp; BELAUD A. &amp; LEK S." box="[882,1090,1370,1394]" journalOrPublisher="Ecol. Model." pageId="17" pageNumber="222" pagination="131 - 142" part="146" refId="ref27832" refString="REYJOL Y., LIM P., BELAUD A. &amp; LEK S., 2001. - Modelling of microhabitat used by fish in natural and regulated flows in the river Garonne (France). Ecol. Model., 146: 131 - 142. DOI: 10.1016 / S 0304 - 3800 (01) 00301 - 5" title="Modelling of microhabitat used by fish in natural and regulated flows in the river Garonne (France)" type="journal article" year="2001">
Reyjol 
<emphasis box="[962,1024,1370,1394]" italics="true" pageId="17" pageNumber="222">et al.,</emphasis>
2001
</bibRefCitation>
). Intraspecific variations of func- tional traits (such as body mass, metabolic rate, excretion rate and boldness) were observed within several populations due to local adaptations to water temperature or predation pressure, or co-variations driven by genetic drift (
<bibRefCitation author="RAFFARD A. &amp; CUCHEROUSSET J. &amp; PRUNIER J. G. &amp; LOOT G. &amp; SANTOUL F. &amp; BLANCHET S." journalOrPublisher="Ecol. Evol." pageId="17" pageNumber="222" pagination="2833 - 2846" part="9" refId="ref27665" refString="RAFFARD A., CUCHEROUSSET J., PRUNIER J. G., LOOT G., SANTOUL F. &amp; BLANCHET S., 2019. - Variability of func- tional traits and their syndromes in a freshwater fish species (Phoxinus phoxinus): The role of adaptive and nonadaptive processes. Ecol. Evol., 9 (5): 2833 - 2846. DOI: 10.1002 / ece 3.4961" title="Variability of func- tional traits and their syndromes in a freshwater fish species (Phoxinus phoxinus): The role of adaptive and nonadaptive processes" type="journal article" year="2019">
Raffard 
<emphasis italics="true" pageId="17" pageNumber="222">et al.,</emphasis>
2019
</bibRefCitation>
). The spawning season occurs from April to May, June in higher altitude (Denys, pers. obs., 2014).
</paragraph>
</subSubSection>
<subSubSection lastPageId="18" lastPageNumber="223" pageId="17" pageNumber="222" type="etymology">
<paragraph blockId="17.[813,1456,1626,1906]" box="[813,936,1626,1650]" pageId="17" pageNumber="222">
<heading bold="true" box="[813,936,1626,1650]" fontSize="10" level="3" pageId="17" pageNumber="222" reason="6">
<emphasis bold="true" box="[813,936,1626,1650]" pageId="17" pageNumber="222">Etymology</emphasis>
</heading>
</paragraph>
<paragraph blockId="17.[813,1456,1626,1906]" lastBlockId="18.[132,775,217,274]" lastPageId="18" lastPageNumber="223" pageId="17" pageNumber="222">
The species name dragarum is the genitive of a latinized name “dragas” which are fairies from of the Occitan folklore. Dragas are the companions of the Drac, a kind of dragon living in the streams of the Ariège department (Garonne drainage). In the land of Foix, these melusian serpentiform fairies attack intruders crossing their territory. The sound of bells alone can stop them. They come out only to wash their clothes with a gold beater, not without having closely watched the area with their only eye in the middle of the forehead (
<bibRefCitation author="ELY R. &amp; TSAAG VALREN A." bookContentInfo="313 p." box="[240,535,249,274]" journalOrPublisher="Bibliotheque du Merveilleux" pageId="18" pageNumber="223" refId="ref25117" refString="ELY R. &amp; TSAAG VALREN A., 2013. - Bestiaire fantastique et creatures feeriques de France. Bibliotheque du Merveilleux. 313 p. Dinan: Terre de Brume." title="Bestiaire fantastique et creatures feeriques de France" type="book" year="2013">Ely and Tsaag Valren, 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="223" type="vernacular_names">
<paragraph blockId="18.[132,775,313,401]" box="[132,328,313,337]" pageId="18" pageNumber="223">
<heading bold="true" box="[132,328,313,337]" fontSize="10" level="3" pageId="18" pageNumber="223" reason="6">
<emphasis bold="true" box="[132,328,313,337]" pageId="18" pageNumber="223">Vernacular name</emphasis>
</heading>
</paragraph>
<paragraph blockId="18.[132,775,313,401]" pageId="18" pageNumber="223">Garonne minnow (English), Vairon de la Garonne (French).</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="223" type="discussion">
<paragraph blockId="18.[132,775,442,593]" box="[132,236,442,466]" pageId="18" pageNumber="223">
<heading bold="true" box="[132,236,442,466]" fontSize="10" level="3" pageId="18" pageNumber="223" reason="6">
<emphasis bold="true" box="[132,236,442,466]" pageId="18" pageNumber="223">Remarks</emphasis>
</heading>
</paragraph>
<paragraph blockId="18.[132,775,442,593]" pageId="18" pageNumber="223">The Garonne and Dordogne clades are separated by a divergence of 1.6%. As we find no other character states dis- tinguishing both population groups, we consider them as two evolutionary lineages of a same species.</paragraph>
</subSubSection>
</treatment>
</document>