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<document id="21BB662D78C3A15BC5C9EB8301F7BC73" ID-CLB-Dataset="4095" ID-DOI="10.5852/ejt.2021.764.1477" ID-GBIF-Dataset="e9baa250-ec84-440f-96bf-06b55f6738e3" ID-ISSN="2118-9773" ID-Zenodo-Dep="5514464" ID-ZooBank="8B5799F9-487D-4A9F-BF9F-54ADA06056FF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1631926753962" checkinUser="felipe" docAuthor="Yang, Qi-cheng, Pan, Zhao-hui, Liu, Xiao-yan &amp; Yang, Ding" docDate="2021" docId="038487C8FFABFF88FDF8FB01FA95F8B5" docLanguage="en" docName="ejt-764_152-192.pdf" docOrigin="European Journal of Taxonomy 764 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.764.1477" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tipula (Vestiplex) uncinella Yang &amp; Pan &amp; Liu &amp; Yang 2021, sp. nov." docType="treatment" docUuid="635FCCE1-5615-49F6-B272-82142261A897" docUuidSource="ZooBank" docVersion="8" lastPageNumber="161" masterDocId="FFBDFFB0FFACFF81FFBAFFA1FFE6FF82" masterDocTitle="Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet" masterLastPageNumber="192" masterPageNumber="152" pageNumber="159" updateTime="1699449403584" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D8DA2A3FC2D8A952356E32CFA4BA69F5">Six new species of Tipula (Vestiplex) Bezzi (Diptera, Tipulidae) from Tibet</mods:title>
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<mods:namePart id="4622A87C9519A3ACA8090EFF082D1345">Yang, Qi-cheng</mods:namePart>
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<mods:affiliation id="D367095FA5F7FC05EED9658FCD75FD3E">Hubei Insect Resources Utilization and Sustainable Pest Management Key Laboratory, College of Plant Science &amp; Technology of Huazhong Agriculture University, Wuhan, Hubei, 430070, China. Plant Protection College, Shenyang Agricultural University, Shenyang, Liaoning, 110866, China. Institute of Plateau Ecology, Tibet Agriculture &amp; Animal Husbandry University, Linzhi, Xizang, P. R. 860000, China. China Agricultural University, Beijing, 100193, China.</mods:affiliation>
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<mods:namePart id="9113C5895152E42E4E4973939C5316C8">Pan, Zhao-hui</mods:namePart>
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<mods:affiliation id="5D4214EC17EA1413FCC76999793F33E7">Hubei Insect Resources Utilization and Sustainable Pest Management Key Laboratory, College of Plant Science &amp; Technology of Huazhong Agriculture University, Wuhan, Hubei, 430070, China. Plant Protection College, Shenyang Agricultural University, Shenyang, Liaoning, 110866, China. Institute of Plateau Ecology, Tibet Agriculture &amp; Animal Husbandry University, Linzhi, Xizang, P. R. 860000, China. China Agricultural University, Beijing, 100193, China.</mods:affiliation>
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<taxonomicName id="4C2D4D5DFFABFF86FDF8FB01FC76FB39" authority="Yang &amp; Pan &amp; Liu &amp; Yang, 2021" authorityName="Yang &amp; Pan &amp; Liu &amp; Yang" authorityYear="2021" box="[578,912,1184,1211]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="159" phylum="Arthropoda" rank="species" species="uncinella" status="sp. nov." subGenus="Vestiplex">
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<emphasis id="B959EACCFFABFF86FDF8FB01FD77FB39" bold="true" box="[578,657,1184,1211]" italics="true" pageId="7" pageNumber="159">Tipula</emphasis>
(
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)
<emphasis id="B959EACCFFABFF86FCA4FB01FC76FB39" bold="true" box="[798,912,1184,1211]" italics="true" pageId="7" pageNumber="159">uncinella</emphasis>
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<figureCitation id="13162A5BFFABFF86FD78FB46FCAEFA80" box="[706,840,1255,1282]" captionStart-0="Figs 1012" captionStart-1="Figs 1320" captionStartId-0="8.[189,242,1959,1985]" captionStartId-1="10.[189,242,1855,1881]" captionTargetBox-0="[249,1322,283,1845]" captionTargetBox-1="[268,1297,279,1797]" captionTargetId-0="figure-11@8.[231,1357,265,1921]" captionTargetId-1="figure-12@10.[266,1322,265,1818]" captionTargetPageId-0="8" captionTargetPageId-1="10" captionText-0="Figs 1012. Tipula (Vestiplex) uncinella sp. nov., paratypes, ♂♂ (CAU). 10. Habitus, lateral view. 11. Wing. 12. Habitus, dorsal view. Abbreviations: see Material and methods. Scale bars = 1.0 mm." captionText-1="Figs 1320. Tipula (Vestiplex) uncinella sp. nov., holotype, ♂ (CAU). 13. Hypopygium, lateral view. 14. Inner gonostylus, lateral external view. 15. Adminiculum, ventral view and oblique view. 16. Outer gonostylus, lateral external view. 17. Tergite nine, dorsal view. 1819. Semen pump, lateral view and dorsal view. 20. Hypopygium, ventral view. Abbreviations: see Material and methods. Scale bars: 13, 17, 20 = 0.5 mm; 1416, 1819 = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5514470" figureDoi-1="http://doi.org/10.5281/zenodo.5514472" httpUri-0="https://zenodo.org/record/5514470/files/figure.png" httpUri-1="https://zenodo.org/record/5514472/files/figure.png" pageId="7" pageNumber="159">Figs 1020</figureCitation>
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<figureCitation id="13162A5BFFABFF86FCEEFB46FC97FA83" box="[852,881,1255,1281]" captionStart="Figs 8486. 84" captionStartId="37.[189,242,1955,1981]" captionTargetBox="[241,1309,264,1888]" captionTargetId="figure-15@37.[233,1355,264,1914]" captionTargetPageId="37" captionText="Figs 8486. 84. Tipula (Vestiplex) bucera sp. nov., mating. 85. Habitat of 80K, Muotuo, Tibet, China. 86. Habitat of Yigong, Bomi, Tibet, China." figureDoi="http://doi.org/10.5281/zenodo.5514523" httpUri="https://zenodo.org/record/5514523/files/figure.png" pageId="7" pageNumber="159">86</figureCitation>
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<emphasis id="B959EACCFFABFF86FF07FA88FED0FAC1" bold="true" box="[189,310,1321,1347]" pageId="7" pageNumber="159">Diagnosis</emphasis>
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<paragraph id="8B9236DEFFABFF86FF07FAFBFA91F98B" blockId="7.[189,1401,1369,1545]" pageId="7" pageNumber="159">Scape and pedicel yellow, flagellum grayish yellow. Prescutum with four grayish yellow stripes, all stripes with brown margins. Scutellum yellow with dark brown mid-longitudinal stripe. Mediotergite yellow with brown mid-longitudinal stripe. Tergite 9 separated medially, pair of angular arms arising from ventral surface of tergite 9, arms with granulated margins. Base of posterior margin of inner gonostylus without a small protrusion; upper beak slightly twisted, lower beak curved, middle concave.</paragraph>
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<emphasis id="B959EACCFFABFF86FF07F98CFEA2F9C5" bold="true" box="[189,324,1581,1607]" pageId="7" pageNumber="159">Etymology</emphasis>
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The species is named after the uncinatus of the adminiculum, the Latin adjective
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means bearing hooks, barbed.
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<emphasis id="B959EACCFFABFF86FF07F961FE4CF958" bold="true" box="[189,426,1728,1754]" pageId="7" pageNumber="159">Material examined</emphasis>
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</paragraph>
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<materialsCitation id="3B453C83FFABFF86FF50F8D9FC60F835" ID-GBIF-Occurrence="3382544313" collectingDate="2017-05-05" collectingDateMax="2017-05-07" collectingDateMin="2017-05-05" collectingMethod="sweeping net" collectionCode="CAU" collectorName="Qicheng Yang" country="China" elevation="2270" latitude="30.271944" location="CHINA" longLatPrecision="20" longitude="94.80778" municipality="Bomi" pageId="7" pageNumber="159" specimenCount="12" specimenCount-male="12" stateProvince="Xizang" typeStatus="paratype">
<emphasis id="B959EACCFFABFF86FF50F8D9FE81F810" bold="true" box="[234,359,1912,1938]" pageId="7" pageNumber="159">
<typeStatus id="5496887CFFABFF86FF50F8D9FE81F810" box="[234,359,1912,1938]" pageId="7" pageNumber="159" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F33A764EFFABFF86FF07F83CFEFEF835" box="[189,280,1949,1975]" name="China" pageId="7" pageNumber="159">CHINA</collectingCountry>
<specimenCount id="9D2BFD57FFABFF86FE8BF83CFE63F83A" box="[305,389,1949,1976]" pageId="7" pageNumber="159" type="male">12 ♂♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED3CAE1BFFABFF86FCF2F83CFC60F835" box="[840,902,1949,1975]" collectionName="China, Beijing, China Agricultural University" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="7" pageNumber="159">CAU</collectionCode>
</materialsCitation>
<materialsCitation id="3B453C83FFABFF86FC25F83CFBB5F85E" ID-GBIF-Occurrence="3382544314" collectingDate="2017-05-17" collectingMethod="light trap" collectionCode="CAU" collectorName="Qicheng Yang" country="China" elevation="2290" latitude="29.834446" location="Bomi" longLatPrecision="20" longitude="94.74333" pageId="7" pageNumber="159" specimenCount="9" specimenCount-male="9" stateProvince="Tibet" typeStatus="paratype">
<specimenCount id="9D2BFD57FFABFF86FC25F83CFC02F83A" box="[927,996,1949,1976]" pageId="7" pageNumber="159" type="male">9 ♂♂</specimenCount>
;
<collectingRegion id="49E9F83CFFABFF86FC4AF83CFBD6F835" box="[1008,1072,1949,1975]" country="China" name="Xizang" pageId="7" pageNumber="159">Tibet</collectingRegion>
,
<location id="8EF26005FFABFF86FB86F83CFB66F835" LSID="urn:lsid:plazi:treatment:038487C8FFABFF88FDF8FB01FA95F8B5:8EF26005FFABFF86FB86F83CFB66F835" box="[1084,1152,1949,1975]" country="China" latitude="29.834446" longLatPrecision="20" longitude="94.74333" name="Bomi" pageId="7" pageNumber="159" stateProvince="Tibet">Bomi</location>
,
<locationDeviation id="9A202FFCFFABFF86FB31F83CFB36F835" box="[1163,1232,1949,1975]" pageId="7" pageNumber="159">106K</locationDeviation>
;
<geoCoordinate id="EE195019FFABFF86FB66F83AFA94F83A" box="[1244,1394,1947,1976]" degrees="29" direction="north" minutes="50" orientation="latitude" pageId="7" pageNumber="159" precision="15" seconds="04" value="29.834446">29°5004″ N</geoCoordinate>
,
<geoCoordinate id="EE195019FFABFF86FF07F860FEB6F85F" box="[189,336,1985,2013]" degrees="94" direction="east" minutes="44" orientation="longitude" pageId="7" pageNumber="159" precision="15" seconds="36" value="94.74333">94°4436″ E</geoCoordinate>
;
<elevation id="0000D1EDFFABFF86FEE6F863FE13F85E" box="[348,501,1986,2013]" metricMagnitude="3" metricUnit="m" metricValue="2.29" pageId="7" pageNumber="159" unit="m" value="2290.0">
<quantity id="4CD59B3BFFABFF86FEE6F863FE53F85F" box="[348,437,1986,2013]" metricMagnitude="3" metricUnit="m" metricValue="2.29" pageId="7" pageNumber="159" unit="m" value="2290.0">2290 m</quantity>
a.s.l.
</elevation>
;
<date id="FF93101EFFABFF86FDB8F863FD44F85E" box="[514,674,1986,2013]" pageId="7" pageNumber="159" value="2017-05-17">
<collectingDate id="EFD7E9F6FFABFF86FDB8F863FD44F85E" box="[514,674,1986,2013]" pageId="7" pageNumber="159" value="2017-05-17">17 May 2017</collectingDate>
</date>
;
<collectorName id="26D85308FFABFF86FD14F863FCB3F85F" box="[686,853,1986,2013]" pageId="7" pageNumber="159">Qicheng Yang</collectorName>
leg.;
<collectingMethod id="526C4EC9FFABFF86FC2CF863FBE3F85F" box="[918,1029,1986,2013]" pageId="7" pageNumber="159">light trap</collectingMethod>
;
<collectionCode id="ED3CAE1BFFABFF86FBABF863FBB5F85E" box="[1041,1107,1986,2012]" collectionName="China, Beijing, China Agricultural University" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="7" pageNumber="159">CAU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF526656FFA4FF89FF07F806FAAEF867" ID-DOI="http://doi.org/10.5281/zenodo.5514470" ID-Zenodo-Dep="5514470" httpUri="https://zenodo.org/record/5514470/files/figure.png" pageId="8" pageNumber="160" startId="8.[189,242,1959,1985]" targetBox="[249,1322,283,1845]" targetPageId="8">
<paragraph id="8B9236DEFFA4FF89FF07F806FAAEF867" blockId="8.[189,1399,1959,2021]" pageId="8" pageNumber="160">
<emphasis id="B959EACCFFA4FF89FF07F806FEA9F843" bold="true" box="[189,335,1959,1985]" pageId="8" pageNumber="160">Figs 1012.</emphasis>
<taxonomicName id="4C2D4D5DFFA4FF89FEE0F806FD4EF843" authority="Yang &amp; Pan &amp; Liu &amp; Yang, 2021" authorityName="Yang &amp; Pan &amp; Liu &amp; Yang" authorityYear="2021" box="[346,680,1959,1986]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="160" phylum="Arthropoda" rank="species" species="uncinella" status="sp. nov." subGenus="Vestiplex">
<emphasis id="B959EACCFFA4FF89FEE0F806FE40F843" box="[346,422,1959,1985]" italics="true" pageId="8" pageNumber="160">Tipula</emphasis>
(
<emphasis id="B959EACCFFA4FF89FE01F806FDC2F843" box="[443,548,1959,1985]" italics="true" pageId="8" pageNumber="160">Vestiplex</emphasis>
)
<emphasis id="B959EACCFFA4FF89FD80F806FD4EF843" box="[570,680,1959,1985]" italics="true" pageId="8" pageNumber="160">uncinella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A57B7FFA4FF89FD0EF809FCF2F843" box="[692,788,1960,1986]" pageId="8" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
, paratypes, ♂♂ (CAU).
<emphasis id="B959EACCFFA4FF89FBFCF806FB85F843" bold="true" box="[1094,1123,1959,1985]" pageId="8" pageNumber="160">10</emphasis>
. Habitus, lateral view.
<emphasis id="B959EACCFFA4FF89FF07F86BFF3FF866" bold="true" box="[189,217,1994,2020]" pageId="8" pageNumber="160">11</emphasis>
. Wing.
<emphasis id="B959EACCFFA4FF89FE8FF86BFEB4F866" bold="true" box="[309,338,1994,2020]" pageId="8" pageNumber="160">12</emphasis>
. Habitus, dorsal view. Abbreviations: see Material and methods. Scale bars = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C3376555FFA5FF88FF07FEA9FE18FAFF" pageId="9" pageNumber="161" type="description">
<paragraph id="8B9236DEFFA5FF88FF07FEA9FEA8FEA0" blockId="9.[189,334,264,290]" box="[189,334,264,290]" pageId="9" pageNumber="161">
<heading id="D0DA81B2FFA5FF88FF07FEA9FEA8FEA0" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="9" pageNumber="161" reason="3">
<emphasis id="B959EACCFFA5FF88FF07FEA9FEA8FEA0" bold="true" box="[189,334,264,290]" pageId="9" pageNumber="161">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B9236DEFFA5FF88FF50FE96FECCFED3" blockId="9.[189,1398,311,373]" box="[234,298,311,337]" pageId="9" pageNumber="161">
<emphasis id="B959EACCFFA5FF88FF50FE96FECCFED3" bold="true" box="[234,298,311,337]" pageId="9" pageNumber="161">Male</emphasis>
</paragraph>
<paragraph id="8B9236DEFFA5FF88FF07FEFAFA90FEF6" blockId="9.[189,1398,311,373]" box="[189,1398,346,373]" pageId="9" pageNumber="161">MEASUREMENTS. Body length 14.0 mm, wing length 18.020.0 mm, antenna length 3.03.5 mm (n = 22).</paragraph>
<paragraph id="8B9236DEFFA5FF88FF07FE3DFAA1FE5B" blockId="9.[188,1398,412,473]" pageId="9" pageNumber="161">
HEAD (
<figureCitation id="13162A5BFFA5FF88FEB4FE3DFE72FE34" box="[270,404,412,438]" captionStart="Figs 1012" captionStartId="8.[189,242,1959,1985]" captionTargetBox="[249,1322,283,1845]" captionTargetId="figure-11@8.[231,1357,265,1921]" captionTargetPageId="8" captionText="Figs 1012. Tipula (Vestiplex) uncinella sp. nov., paratypes, ♂♂ (CAU). 10. Habitus, lateral view. 11. Wing. 12. Habitus, dorsal view. Abbreviations: see Material and methods. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5514470" httpUri="https://zenodo.org/record/5514470/files/figure.png" pageId="9" pageNumber="161">Figs 10, 12</figureCitation>
). Mainly yellow. Occipital stripe dark gray. Rostrum and nasus yellow. Setae on head brown. Scape and pedicel yellow, flagellum grayish yellow. Proboscis and palpus brownish yellow.
</paragraph>
<paragraph id="8B9236DEFFA5FF88FF07FDA0FCC4FCB6" blockId="9.[188,1401,512,820]" pageId="9" pageNumber="161">
THORAX (
<figureCitation id="13162A5BFFA5FF88FE91FDA1FE53FD98" box="[299,437,512,538]" captionStart="Figs 1012" captionStartId="8.[189,242,1959,1985]" captionTargetBox="[249,1322,283,1845]" captionTargetId="figure-11@8.[231,1357,265,1921]" captionTargetPageId="8" captionText="Figs 1012. Tipula (Vestiplex) uncinella sp. nov., paratypes, ♂♂ (CAU). 10. Habitus, lateral view. 11. Wing. 12. Habitus, dorsal view. Abbreviations: see Material and methods. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5514470" httpUri="https://zenodo.org/record/5514470/files/figure.png" pageId="9" pageNumber="161">Figs 10, 12</figureCitation>
). Mainly yellow, heavily pruinose. Pronotum brownish yellow with yellow setae. Prescutum with four grayish yellow stripes, all stripes with brown margins, median stripe broadened apically and gradually narrowing to base, lateral stripes long-oval. Scutum yellow with four grayish yellow spots, anterior margins of anterior two spots brown. Scutellum yellow with dark brown midlongitudinal stripe. Mediotergite yellow with brown mid-longitudinal stripe. Thoracic pleuron yellow. Parascutellum yellow. Meron yellow. Setae on thorax yellow. Legs with coxae and femora yellow, femora with dark brown apices, tibiae and tarsi dark brown. Setae on legs dark brown. Wing variegated light brown; pterostigma brown; petiole of cell m 1 distinctly shorter than discal cell (
<figureCitation id="13162A5BFFA5FF88FB30FD56FB39FC93" box="[1162,1247,759,785]" captionStart="Figs 1012" captionStartId="8.[189,242,1959,1985]" captionTargetBox="[249,1322,283,1845]" captionTargetId="figure-11@8.[231,1357,265,1921]" captionTargetPageId="8" captionText="Figs 1012. Tipula (Vestiplex) uncinella sp. nov., paratypes, ♂♂ (CAU). 10. Habitus, lateral view. 11. Wing. 12. Habitus, dorsal view. Abbreviations: see Material and methods. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5514470" httpUri="https://zenodo.org/record/5514470/files/figure.png" pageId="9" pageNumber="161">Fig. 11</figureCitation>
). Halter with stem yellow, knob pale black with pale yellow apex.
</paragraph>
<paragraph id="8B9236DEFFA5FF88FF07FCFDFBEFFC1A" blockId="9.[189,1398,859,921]" pageId="9" pageNumber="161">
ABDOMEN (
<figureCitation id="13162A5BFFA5FF88FE85FCFAFE2CFCF7" box="[319,458,859,885]" captionStart="Figs 1012" captionStartId="8.[189,242,1959,1985]" captionTargetBox="[249,1322,283,1845]" captionTargetId="figure-11@8.[231,1357,265,1921]" captionTargetPageId="8" captionText="Figs 1012. Tipula (Vestiplex) uncinella sp. nov., paratypes, ♂♂ (CAU). 10. Habitus, lateral view. 11. Wing. 12. Habitus, dorsal view. Abbreviations: see Material and methods. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5514470" httpUri="https://zenodo.org/record/5514470/files/figure.png" pageId="9" pageNumber="161">Figs 10, 12</figureCitation>
). Mainly yellow. Abdominal tergites each with three brown stripes. Abdominal segments 59 dark brown to black. Setae on abdomen brownish yellow.
</paragraph>
<paragraph id="8B9236DEFFA5FF88FF07FC60FB5EFA95" blockId="9.[187,1401,960,1304]" pageId="9" pageNumber="161">
HYPOPYGIUM (
<figureCitation id="13162A5BFFA5FF88FEE7FC61FDF6FC58" box="[349,528,960,986]" captionStart="Figs 1320" captionStartId="10.[189,242,1855,1881]" captionTargetBox="[268,1297,279,1797]" captionTargetId="figure-12@10.[266,1322,265,1818]" captionTargetPageId="10" captionText="Figs 1320. Tipula (Vestiplex) uncinella sp. nov., holotype, ♂ (CAU). 13. Hypopygium, lateral view. 14. Inner gonostylus, lateral external view. 15. Adminiculum, ventral view and oblique view. 16. Outer gonostylus, lateral external view. 17. Tergite nine, dorsal view. 1819. Semen pump, lateral view and dorsal view. 20. Hypopygium, ventral view. Abbreviations: see Material and methods. Scale bars: 13, 17, 20 = 0.5 mm; 1416, 1819 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514472" httpUri="https://zenodo.org/record/5514472/files/figure.png" pageId="9" pageNumber="161">Figs 1317, 20</figureCitation>
). Gonocoxite elongated, separated from sternite 9, posterior margin with an unusually strong and slender black spine, directed chiefly dorsally, slightly twisted. Tergite 9 separated medially, middle of posterior margin of lobe slightly protrude; a pair of angular black arms arising from ventral surface of tergite 9, surface of arms not smooth, with granulated margin. Sternite 9 with a median prominence. Adminiculum narrow spade-like, apex slightly furcal, ventral caudal part with a small uncinate projection. Outer gonostylus anteriorly bent, posterior margin of basal half truncated (
<figureCitation id="13162A5BFFA5FF88FF7DFB32FEF9FB2C" box="[199,287,1171,1198]" captionStart="Figs 1320" captionStartId="10.[189,242,1855,1881]" captionTargetBox="[268,1297,279,1797]" captionTargetId="figure-12@10.[266,1322,265,1818]" captionTargetPageId="10" captionText="Figs 1320. Tipula (Vestiplex) uncinella sp. nov., holotype, ♂ (CAU). 13. Hypopygium, lateral view. 14. Inner gonostylus, lateral external view. 15. Adminiculum, ventral view and oblique view. 16. Outer gonostylus, lateral external view. 17. Tergite nine, dorsal view. 1819. Semen pump, lateral view and dorsal view. 20. Hypopygium, ventral view. Abbreviations: see Material and methods. Scale bars: 13, 17, 20 = 0.5 mm; 1416, 1819 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514472" httpUri="https://zenodo.org/record/5514472/files/figure.png" pageId="9" pageNumber="161">Fig. 16</figureCitation>
). Inner gonostylus with concavity at middle, posterior margin with big semicircular bulge; base of posterior margin of inner gonostylus without a small protrusion; upper beak slightly twisted, lower beak curved, middle concave (
<figureCitation id="13162A5BFFA5FF88FD9EFB78FD9CFB76" box="[548,634,1241,1268]" captionStart="Figs 1320" captionStartId="10.[189,242,1855,1881]" captionTargetBox="[268,1297,279,1797]" captionTargetId="figure-12@10.[266,1322,265,1818]" captionTargetPageId="10" captionText="Figs 1320. Tipula (Vestiplex) uncinella sp. nov., holotype, ♂ (CAU). 13. Hypopygium, lateral view. 14. Inner gonostylus, lateral external view. 15. Adminiculum, ventral view and oblique view. 16. Outer gonostylus, lateral external view. 17. Tergite nine, dorsal view. 1819. Semen pump, lateral view and dorsal view. 20. Hypopygium, ventral view. Abbreviations: see Material and methods. Scale bars: 13, 17, 20 = 0.5 mm; 1416, 1819 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514472" httpUri="https://zenodo.org/record/5514472/files/figure.png" pageId="9" pageNumber="161">Fig. 14</figureCitation>
). Semen pump (
<figureCitation id="13162A5BFFA5FF88FC82FB78FC5BFB76" box="[824,957,1241,1268]" captionStart="Figs 1320" captionStartId="10.[189,242,1855,1881]" captionTargetBox="[268,1297,279,1797]" captionTargetId="figure-12@10.[266,1322,265,1818]" captionTargetPageId="10" captionText="Figs 1320. Tipula (Vestiplex) uncinella sp. nov., holotype, ♂ (CAU). 13. Hypopygium, lateral view. 14. Inner gonostylus, lateral external view. 15. Adminiculum, ventral view and oblique view. 16. Outer gonostylus, lateral external view. 17. Tergite nine, dorsal view. 1819. Semen pump, lateral view and dorsal view. 20. Hypopygium, ventral view. Abbreviations: see Material and methods. Scale bars: 13, 17, 20 = 0.5 mm; 1416, 1819 = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514472" httpUri="https://zenodo.org/record/5514472/files/figure.png" pageId="9" pageNumber="161">Figs 1819</figureCitation>
) as shown in figures. Aedeagus wide, basal width about 0.5 mm, gradually narrowing towards apex, and apex not bifurcated.
</paragraph>
<paragraph id="8B9236DEFFA5FF88FF50FA9FFEA3FADA" blockId="9.[189,510,1342,1406]" box="[234,325,1342,1368]" pageId="9" pageNumber="161">
<emphasis id="B959EACCFFA5FF88FF50FA9FFEA3FADA" bold="true" box="[234,325,1342,1368]" pageId="9" pageNumber="161">Female</emphasis>
</paragraph>
<paragraph id="8B9236DEFFA5FF88FF07FAC2FE18FAFF" blockId="9.[189,510,1342,1406]" box="[189,510,1379,1406]" pageId="9" pageNumber="161">Unknown or unrecognized.</paragraph>
</subSubSection>
<subSubSection id="C3376555FFA5FF88FF07FA3CFE85FA6B" box="[189,355,1437,1513]" pageId="9" pageNumber="161" type="distribution">
<paragraph id="8B9236DEFFA5FF88FF07FA3CFEB0FA35" blockId="9.[189,342,1437,1463]" box="[189,342,1437,1463]" pageId="9" pageNumber="161">
<heading id="D0DA81B2FFA5FF88FF07FA3CFEB0FA35" bold="true" box="[189,342,1437,1463]" fontSize="11" level="3" pageId="9" pageNumber="161" reason="3">
<emphasis id="B959EACCFFA5FF88FF07FA3CFEB0FA35" bold="true" box="[189,342,1437,1463]" pageId="9" pageNumber="161">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B9236DEFFA5FF88FF07FA6FFE85FA6B" blockId="9.[189,355,1486,1513]" box="[189,355,1486,1513]" pageId="9" pageNumber="161">
<collectingCountry id="F33A764EFFA5FF88FF07FA6FFEE5FA6A" box="[189,259,1486,1512]" name="China" pageId="9" pageNumber="161">China</collectingCountry>
(
<collectingRegion id="49E9F83CFFA5FF88FEA9FA6FFEB1FA6B" box="[275,343,1486,1513]" country="China" name="Xizang" pageId="9" pageNumber="161">Tibet</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3376555FFA5FF88FF07F9ADFA95F8B5" pageId="9" pageNumber="161" type="discussion">
<paragraph id="8B9236DEFFA5FF88FF07F9ADFEC9F9A4" blockId="9.[189,303,1548,1574]" box="[189,303,1548,1574]" pageId="9" pageNumber="161">
<heading id="D0DA81B2FFA5FF88FF07F9ADFEC9F9A4" bold="true" box="[189,303,1548,1574]" fontSize="11" level="3" pageId="9" pageNumber="161" reason="3">
<emphasis id="B959EACCFFA5FF88FF07F9ADFEC9F9A4" bold="true" box="[189,303,1548,1574]" pageId="9" pageNumber="161">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B9236DEFFA5FF88FF07F99CFA95F8B5" blockId="9.[188,1401,1597,1847]" pageId="9" pageNumber="161">
The new species is similar to
<emphasis id="B959EACCFFA5FF88FD8CF99CFDA0F9D4" box="[566,582,1597,1622]" italics="true" pageId="9" pageNumber="161">T</emphasis>
. (
<emphasis id="B959EACCFFA5FF88FDD9F99CFD93F9D4" box="[611,629,1597,1622]" italics="true" pageId="9" pageNumber="161">V</emphasis>
.)
<emphasis id="B959EACCFFA5FF88FD29F99CFCD5F9D5" box="[659,819,1597,1623]" italics="true" pageId="9" pageNumber="161">schizophallus</emphasis>
(Alexander, 1973), but the latter differs in the following characters: arms of tergite 9 with only inner margin roughened; gonocoxite with an additional small spine (the one near posterior margin of basistylar suture); outer gonostylus with very abundant black setae from conspicuous punctures; apex of adminiculum profoundly divided into two long parallel spines, ventral caudal part without uncinatus. It is also similar
<emphasis id="B959EACCFFA5FF88FC0AF972FC58F96E" box="[944,958,1747,1772]" italics="true" pageId="9" pageNumber="161">T</emphasis>
.
<taxonomicName id="4C2D4D5DFFA5FF88FC75F973FAA5F96E" authority="Alexander, 1934" authorityName="Alexander" authorityYear="1934" box="[975,1347,1746,1772]" class="Insecta" family="Tipulidae" genus="Tipula" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="161" phylum="Arthropoda" rank="species" species="deserrata" subGenus="Vestiplex">
(
<emphasis id="B959EACCFFA5FF88FC62F972FC0CF96E" box="[984,1002,1747,1772]" italics="true" pageId="9" pageNumber="161">V</emphasis>
.)
<emphasis id="B959EACCFFA5FF88FBBFF973FB93F96E" box="[1029,1141,1746,1772]" italics="true" pageId="9" pageNumber="161">deserrata</emphasis>
Alexander, 1934
</taxonomicName>
and
<emphasis id="B959EACCFFA5FF88FF07F959FF2BF893" box="[189,205,1784,1809]" italics="true" pageId="9" pageNumber="161">T</emphasis>
. (
<emphasis id="B959EACCFFA5FF88FF51F959FF1BF893" box="[235,253,1784,1809]" italics="true" pageId="9" pageNumber="161">V</emphasis>
.)
<emphasis id="B959EACCFFA5FF88FEA0F956FE91F893" box="[282,375,1783,1809]" italics="true" pageId="9" pageNumber="161">factiosa</emphasis>
Alexander, 1940, but the posterior margin of inner gonostylus in the latter has no protrusion. The posterior margin of tergite 9 of
<emphasis id="B959EACCFFA5FF88FD53F8BCFD1FF8B4" box="[745,761,1821,1846]" italics="true" pageId="9" pageNumber="161">T</emphasis>
. (
<emphasis id="B959EACCFFA5FF88FCA8F8BCFCC2F8B4" box="[786,804,1821,1846]" italics="true" pageId="9" pageNumber="161">V</emphasis>
.)
<emphasis id="B959EACCFFA5FF88FC86F8BCFC7FF8B5" box="[828,921,1821,1847]" italics="true" pageId="9" pageNumber="161">factiosa</emphasis>
is smooth, its arms are large and strong.
</paragraph>
</subSubSection>
</treatment>
</document>