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<mods:title id="A601D0FEE021255DA640A59493477794">New or rare Madagascar tiger beetles- 21. Physodeutera (Microlepidia) propripenis sp. nov., Ph. (M.) marginemaculata (W. Horn) and Ph. (M.) peyrierasi Rivalier (Coleoptera: Cicindelidae)</mods:title>
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<mods:affiliation id="370C02656B13812B19E8C9228DD12E5D">Mendel University in Brno, Department of Forest Ecology, Zemědělská 3, 613 00 Brno, Czech Republic</mods:affiliation>
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<mods:namePart id="B5DC104424E0D16F7800DA486E38F2EE">Brzoska, David</mods:namePart>
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<mods:namePart id="CC31AE695F2DC7E7BB7970F0EF9D7FFE">Vybíral, Jan</mods:namePart>
<mods:affiliation id="DA78AFF393C82D619D1C1912C98E3A0C">Biosférická rezervace Dolní Morava, o. p. s., Národních hrdinů 23, 690 02 Břeclav, Czech Republic</mods:affiliation>
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<paragraph id="8B923695FFD1FF88FF0EFE49FCECFD9B" blockId="12.[151,803,510,573]" box="[151,803,510,537]" pageId="12" pageNumber="45">
<heading id="D0DA81F9FFD1FF88FF0EFE49FCECFD9B" bold="true" box="[151,803,510,537]" fontSize="11" level="1" pageId="12" pageNumber="45" reason="1">
<taxonomicName id="4C2D4D16FFD1FF88FF0EFE49FCECFD9B" ID-CoL="77HMY" authority="Rivalier, 1967" authorityName="Rivalier" authorityYear="1967" box="[151,803,510,537]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="peyrierasi" subGenus="Microlepidia">
<emphasis id="B959EA87FFD1FF88FF0EFE49FCECFD9B" bold="true" box="[151,803,510,537]" pageId="12" pageNumber="45">
<emphasis id="B959EA87FFD1FF88FF0EFE49FEF3FD9A" bold="true" box="[151,316,510,537]" italics="true" pageId="12" pageNumber="45">Physodeutera</emphasis>
(
<emphasis id="B959EA87FFD1FF88FED4FE49FE27FD9A" bold="true" box="[333,488,510,537]" italics="true" pageId="12" pageNumber="45">Microlepidia</emphasis>
)
<emphasis id="B959EA87FFD1FF88FE60FE48FDBFFD9A" bold="true" box="[505,624,511,537]" italics="true" pageId="12" pageNumber="45">peyrierasi</emphasis>
<bibRefCitation id="EFBC4B64FFD1FF88FDEEFE49FCECFD9B" author="Rivalier, E." box="[631,803,510,536]" pageId="12" pageNumber="45" pagination="261 - 296" refId="ref12291" refString="Rivalier, E. (1967) Le genre Physodeutera Lacordaire (Col. Cicindelidae) revision et description d'especes nouvelles. Annales de la Societe entomologique de France, New Series, 3, 261 - 296." type="journal article" year="1967">Rivalier, 1967</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C337651EFFD1FF88FF0EFD94FE9EFDBE" box="[151,337,546,573]" pageId="12" pageNumber="45" type="description">
<paragraph id="8B923695FFD1FF88FF0EFD94FE9EFDBE" blockId="12.[151,803,510,573]" box="[151,337,546,573]" pageId="12" pageNumber="45">
(
<figureCitation id="13162A10FFD1FF88FF06FD94FEEEFDBE" box="[159,289,547,573]" captionStart="FIGURES 4049" captionStartId="13.[151,264,1867,1891]" captionTargetBox="[157,1429,210,1832]" captionTargetId="figure-17@13.[151,1435,200,1844]" captionTargetPageId="13" captionText="FIGURES 4049. Physodeutera (Microlepidia) peyrierasi Rivalier. 40—habitus, ♁, 10.3 mm, Mahamasina (CCJM); 41— part of head, type locality Ankarana (CCJM ex APCA); 4244—labrum: 42—♁, type locality (CCJM ex APCA); 43—♁, Mahamasina (CCJM); 44—♀, ibid.; 45—pronotum, ♁, ibid.; 46—aedeagus showing internal sac, type locality (CCJM ex APCA); 4749—elytron: 47—♁, type locality (CCJM ex APCA); 48—♁, Mahamasina (CCJM); 49—♁, ibid. (CCJM). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4594665" httpUri="https://zenodo.org/record/4594665/files/figure.png" pageId="12" pageNumber="45">Figs 4049</figureCitation>
,
<figureCitation id="13162A10FFD1FF88FEB5FD95FE86FDBE" box="[300,329,546,573]" captionStart="FIGURES 5052" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[201,1384,194,1906]" captionTargetId="figure-27@14.[188,1400,181,1919]" captionTargetPageId="14" captionText="FIGURES 5052. Distribution of three Physodeutera species in North Madagascar. 50— Physodeutera (Microlepidia) propripenis sp. nov.; 51— Physodeutera (Microlepidia) marginemaculata (W. Horn); 52— Physodeutera (Microlepidia) peyrierasi Rivalier." figureDoi="http://doi.org/10.5281/zenodo.4594667" httpUri="https://zenodo.org/record/4594667/files/figure.png" pageId="12" pageNumber="45">52</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C337651EFFD1FF88FF0EFDD0FC00FD1D" pageId="12" pageNumber="45" type="reference_group">
<paragraph id="8B923695FFD1FF88FF0EFDD0FD34FDFC" blockId="12.[151,975,615,670]" box="[151,763,615,639]" pageId="12" pageNumber="45">
<treatmentCitationGroup id="AB3D11BBFFD1FF88FF0EFDD0FD34FDFC" box="[151,763,615,639]" pageId="12" pageNumber="45">
<treatmentCitation id="0A8C1084FFD1FF88FF0EFDD0FD38FDFC" author="Rivalier, E." box="[151,759,615,639]" page="268" pageId="12" pageNumber="45" year="1967">
<taxonomicName id="4C2D4D16FFD1FF88FF0EFDD0FD38FDFC" ID-CoL="77HMY" authority="Rivalier, 1967: 268" authorityName="Rivalier" authorityPageNumber="268" authorityYear="1967" box="[151,759,615,639]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="peyrierasi" subGenus="Microlepidia">
<emphasis id="B959EA87FFD1FF88FF0EFDD0FEEDFDFC" box="[151,290,615,639]" italics="true" pageId="12" pageNumber="45">Physodeutera</emphasis>
(
<emphasis id="B959EA87FFD1FF88FEA8FDD0FE7BFDFC" box="[305,436,615,639]" italics="true" pageId="12" pageNumber="45">Microlepidia</emphasis>
)
<emphasis id="B959EA87FFD1FF88FE5AFDDFFDE5FDFC" box="[451,554,616,639]" italics="true" pageId="12" pageNumber="45">peyrierasi</emphasis>
<bibRefCitation id="EFBC4B64FFD1FF88FDA9FDD0FD38FDFC" author="Rivalier, E." box="[560,759,615,639]" pageId="12" pageNumber="45" pagination="261 - 296" refId="ref12291" refString="Rivalier, E. (1967) Le genre Physodeutera Lacordaire (Col. Cicindelidae) revision et description d'especes nouvelles. Annales de la Societe entomologique de France, New Series, 3, 261 - 296." type="journal article" year="1967">Rivalier, 1967: 268</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8B923695FFD1FF88FF0EFD31FC00FD1D" blockId="12.[151,975,615,670]" box="[151,975,646,670]" pageId="12" pageNumber="45">
<treatmentCitationGroup id="AB3D11BBFFD1FF88FF0EFD31FC00FD1D" box="[151,975,646,670]" pageId="12" pageNumber="45">
<treatmentCitation id="0A8C1084FFD1FF88FF0EFD31FD31FD1D" author="Moravec, J." box="[151,766,646,670]" page="91" pageId="12" pageNumber="45" year="2002">
<taxonomicName id="4C2D4D16FFD1FF88FF0EFD31FD31FD1D" ID-CoL="77HMY" authority=": Moravec 2002 a: 91" authorityName="Moravec" authorityPageNumber="91" authorityYear="2002" box="[151,766,646,670]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="peyrierasi" subGenus="Microlepidia">
<emphasis id="B959EA87FFD1FF88FF0EFD31FEEDFD1D" box="[151,290,646,670]" italics="true" pageId="12" pageNumber="45">Physodeutera</emphasis>
(
<emphasis id="B959EA87FFD1FF88FEA8FD31FE7BFD1D" box="[305,436,646,670]" italics="true" pageId="12" pageNumber="45">Microlepidia</emphasis>
)
<emphasis id="B959EA87FFD1FF88FE5AFD31FDE5FD1E" box="[451,554,646,669]" italics="true" pageId="12" pageNumber="45">peyrierasi</emphasis>
:
<bibRefCitation id="EFBC4B64FFD1FF88FDAEFD31FD31FD1D" author="Moravec, J." box="[567,766,646,670]" pageId="12" pageNumber="45" refId="ref11194" refString="Moravec, J. (2002 a) Tiger beetles of Madagascar 2. A monograph of the genus Physodeutera (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, Czech Republic, 290 pp." type="book" year="2002">Moravec 2002a: 91</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs: 234243, 712.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C337651EFFD1FF88FF0EFD7BFF2CFB15" pageId="12" pageNumber="45" type="materials_examined">
<paragraph id="8B923695FFD1FF88FF0EFD7BFE04FD65" blockId="12.[151,1437,716,2039]" box="[151,459,716,743]" pageId="12" pageNumber="45">
<emphasis id="B959EA87FFD1FF88FF0EFD7BFEFFFD65" bold="true" box="[151,304,716,742]" pageId="12" pageNumber="45">
<typeStatus id="54968837FFD1FF88FF0EFD7BFF1DFD65" box="[151,210,716,742]" pageId="12" pageNumber="45">Type</typeStatus>
locality
</emphasis>
. “Ankarana”.
</paragraph>
<paragraph id="8B923695FFD1FF88FF5EFD47FE2EFC19" blockId="12.[151,1437,716,2039]" pageId="12" pageNumber="45">
<emphasis id="B959EA87FFD1FF88FF5EFD47FEBDFC89" bold="true" box="[199,370,752,778]" pageId="12" pageNumber="45">Type material</emphasis>
.
<typeStatus id="54968837FFD1FF88FE1CFD47FE22FC89" box="[389,493,752,778]" pageId="12" pageNumber="45" type="holotype">Holotype</typeStatus>
♁ (by monotypy) in
<collectionCode id="ED3CAE50FFD1FF88FD68FD46FC86FC88" box="[753,841,753,779]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="45">MNHN</collectionCode>
, labelled: “Ankarana” [handwritten // “Type” [red, printed] // “Peyrierasi / Rivalier / type” [handwritten // “aedéage 1814” [handwritten, referring to the aedeagus mounted separately by Rivalier] // “
<typeStatus id="54968837FFD1FF88FDA9FC8FFD54FCD1" box="[560,667,824,850]" pageId="12" pageNumber="45" type="holotype">Holotype</typeStatus>
(by monotypy) /
<taxonomicName id="4C2D4D16FFD1FF88FCF1FC8FFB65FCD0" authority="" authorityName="Rivalier" authorityYear="1967" box="[872,1194,824,851]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="subGenus" subGenus="Microlepidia">Physodeutera (Microlepidia)</taxonomicName>
/
<taxonomicName id="4C2D4D16FFD1FF88FB5CFC8FFF00FCF5" authority="Rivalier, 1967" authorityName="Rivalier" authorityYear="1967" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="peyrierasi">
peyrierasi
<bibRefCitation id="EFBC4B64FFD1FF88FAA5FC8FFF00FCF5" author="Rivalier, E." pageId="12" pageNumber="45" pagination="261 - 296" refId="ref12291" refString="Rivalier, E. (1967) Le genre Physodeutera Lacordaire (Col. Cicindelidae) revision et description d'especes nouvelles. Annales de la Societe entomologique de France, New Series, 3, 261 - 296." type="journal article" year="1967">Rivalier, 1967</bibRefCitation>
</taxonomicName>
/ design. by J. Moravec, 2000” [red, printed] // “
<taxonomicName id="4C2D4D16FFD1FF88FD76FCEBFBFCFCF4" authority="" authorityName="Rivalier" authorityYear="1967" box="[751,1075,860,887]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="subGenus" subGenus="Microlepidia">Physodeutera (Microlepidia)</taxonomicName>
/
<taxonomicName id="4C2D4D16FFD1FF88FBD0FCEBFA92FCF5" authority="Rivalier, 1967" authorityName="Rivalier" authorityYear="1967" box="[1097,1373,860,886]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="peyrierasi">
peyrierasi
<bibRefCitation id="EFBC4B64FFD1FF88FB27FCEBFA92FCF5" author="Rivalier, E." box="[1214,1373,860,886]" pageId="12" pageNumber="45" pagination="261 - 296" refId="ref12291" refString="Rivalier, E. (1967) Le genre Physodeutera Lacordaire (Col. Cicindelidae) revision et description d'especes nouvelles. Annales de la Societe entomologique de France, New Series, 3, 261 - 296." type="journal article" year="1967">Rivalier, 1967</bibRefCitation>
</taxonomicName>
/ det. Jiří Moravec 2000” [printed].
</paragraph>
<paragraph id="8B923695FFD1FF88FF5EFC13FBF9FB85" blockId="12.[151,1437,716,2039]" pageId="12" pageNumber="45">
<emphasis id="B959EA87FFD1FF88FF5EFC13FE33FC3D" bold="true" box="[199,508,932,958]" pageId="12" pageNumber="45">Other material examined</emphasis>
. 2 ♁♁,
<materialsCitation id="3B453CC8FFD1FF88FDFAFC13FBFDFB85" collectionCode="MNHN, CCJM, APCA" determinerName="Rivalier &amp; Jiri Moravec" pageId="12" pageNumber="45" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D2BFD1CFFD1FF88FDFAFC13FD5EFC3C" box="[611,657,932,959]" pageId="12" pageNumber="45" type="female">1 ♀</specimenCount>
in
<collectionCode id="ED3CAE50FFD1FF88FD24FC12FCD8FC3C" box="[701,791,933,959]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="45">MNHN</collectionCode>
: “Mus. Paris / Ankarana / XII.67, Vadon et Peyrieras”. 1 ♁ in
<collectionCode id="ED3CAE50FFD1FF88FF75FC7FFEFAFC61" box="[236,309,968,994]" pageId="12" pageNumber="45">CCJM</collectionCode>
(ex
<collectionCode id="ED3CAE50FFD1FF88FEF3FC7FFE74FC61" box="[362,443,968,994]" pageId="12" pageNumber="45">APCA</collectionCode>
): “Ankarana / XII.67 / Vadon et Peyrieras” // “
<taxonomicName id="4C2D4D16FFD1FF88FC4FFC7FFBBBFC61" box="[982,1140,968,994]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="peyrierasi">Ph. peyrierasi</taxonomicName>
/ Rivalier /
<determinerName id="7E4EDA51FFD1FF88FA9AFC7FFA92FC61" box="[1283,1373,968,994]" pageId="12" pageNumber="45">Rivalier</determinerName>
det.” “
<taxonomicName id="4C2D4D16FFD1FF88FF39FC5BFE2BFB84" authority="" authorityName="Rivalier" authorityYear="1967" box="[160,484,1004,1031]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="subGenus" subGenus="Microlepidia">Physodeutera (Microlepidia)</taxonomicName>
/
<taxonomicName id="4C2D4D16FFD1FF88FE62FC5BFCC0FB85" authority="Rivalier, 1967" authorityName="Rivalier" authorityYear="1967" box="[507,783,1004,1030]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="peyrierasi">
peyrierasi
<bibRefCitation id="EFBC4B64FFD1FF88FDE9FC5BFCC0FB85" author="Rivalier, E." box="[624,783,1004,1030]" pageId="12" pageNumber="45" pagination="261 - 296" refId="ref12291" refString="Rivalier, E. (1967) Le genre Physodeutera Lacordaire (Col. Cicindelidae) revision et description d'especes nouvelles. Annales de la Societe entomologique de France, New Series, 3, 261 - 296." type="journal article" year="1967">Rivalier, 1967</bibRefCitation>
</taxonomicName>
/ det.
<determinerName id="7E4EDA51FFD1FF88FCCCFC5BFC2BFB84" box="[853,996,1004,1031]" pageId="12" pageNumber="45">Jiří Moravec</determinerName>
1999”
</materialsCitation>
.
</paragraph>
<paragraph id="8B923695FFD1FF88FF5EFBA6FF2CFB15" blockId="12.[151,1437,716,2039]" pageId="12" pageNumber="45">
<materialsCitation id="3B453CC8FFD1FF88FF5EFBA6FF10FB15" collectingDate="2007-01-05" collectingDateMax="2007-01-06" collectingDateMin="2007-01-05" collectionCode="MHCW, CJVB, CCJM" collectorName="Jan Vybiral &amp; F. Pavel" country="Madagascar" determinerName="Jiri Moravec" latitude="-12.969083" location="Recent" longLatPrecision="1" longitude="49.136196" pageId="12" pageNumber="45" specimenCount="3" specimenCount-female="3">
<location id="8EF2604EFFD1FF88FF5EFBA6FEDAFBA8" LSID="urn:lsid:plazi:treatment:03848783FFD1FF88FF0EFE49FA56F875:8EF2604EFFD1FF88FF5EFBA6FEDAFBA8" box="[199,277,1041,1067]" country="Madagascar" latitude="-12.969083" longLatPrecision="1" longitude="49.136196" name="Recent" pageId="12" pageNumber="45">Recent</location>
data: 1 ♁,
<specimenCount id="9D2BFD1CFFD1FF88FE0BFBA7FE73FBA8" box="[402,444,1040,1067]" pageId="12" pageNumber="45" type="female">1 ♀</specimenCount>
in
<collectionCode id="ED3CAE50FFD1FF88FE46FBA7FDF5FBA9" box="[479,570,1040,1066]" pageId="12" pageNumber="45">MHCW</collectionCode>
,
<specimenCount id="9D2BFD1CFFD1FF88FDDCFBA7FDA0FBA8" box="[581,623,1040,1067]" pageId="12" pageNumber="45" type="female">1 ♀</specimenCount>
in
<collectionCode id="ED3CAE50FFD1FF88FD0AFBA7FD15FBA9" box="[659,730,1040,1066]" pageId="12" pageNumber="45">CJVB</collectionCode>
,
<specimenCount id="9D2BFD1CFFD1FF88FD7FFBA7FCDFFBA8" box="[742,784,1040,1067]" pageId="12" pageNumber="45" type="female">1 ♀</specimenCount>
in
<collectionCode id="ED3CAE50FFD1FF88FCADFBA7FC4DFBA9" box="[820,898,1040,1066]" pageId="12" pageNumber="45">CCJM</collectionCode>
: “N.
<collectingCountry id="F33A7605FFD1FF88FC23FBA7FB8CFBA9" box="[954,1091,1040,1066]" name="Madagascar" pageId="12" pageNumber="45">Madagascar</collectingCountry>
, Ankarana N.P. / Mahamatsina [Mahamasina] env.,
<date id="FF931055FFD1FF88FEE2FB83FE3FFBCC" box="[379,496,1076,1103]" pageId="12" pageNumber="45" value="2007-01-05" valueMax="2007-01-06" valueMin="2007-01-05">
<collectingDate id="EFD7E9BDFFD1FF88FEE2FB83FE3FFBCC" box="[379,496,1076,1103]" pageId="12" pageNumber="45" value="2007-01-05" valueMax="2007-01-06" valueMin="2007-01-05">5-6.I.2007</collectingDate>
</date>
/
<geoCoordinate id="EE195052FFD1FF88FD9CFB83FD67FBCC" box="[517,680,1076,1103]" degrees="12" direction="south" minutes="58" orientation="latitude" pageId="12" pageNumber="45" precision="1" seconds="08.7" value="-12.969083">S12°58´08.7´´</geoCoordinate>
;
<geoCoordinate id="EE195052FFD1FF88FD2AFB83FC9BFBCD" box="[691,852,1076,1102]" degrees="49" direction="east" minutes="08" orientation="longitude" pageId="12" pageNumber="45" precision="1" seconds="10.3" value="49.136196">E49°08´10.3´´</geoCoordinate>
/
<collectorName id="26D85343FFD1FF88FCF3FB82FC27FBCD" box="[874,1000,1076,1103]" pageId="12" pageNumber="45">Jan Vybíral</collectorName>
leg.”. 1 ♁ in
<collectionCode id="ED3CAE50FFD1FF88FBE6FB83FB07FBCD" box="[1151,1224,1076,1102]" pageId="12" pageNumber="45">CCJM</collectionCode>
with same locality label except for: “
<collectorName id="26D85343FFD1FF88FEF3FBEFFE06FBF1" box="[362,457,1112,1138]" pageId="12" pageNumber="45">F. Pavel</collectorName>
leg.” // “
<taxonomicName id="4C2D4D16FFD1FF88FDA1FBEFFCB0FBF0" authority="" authorityName="Rivalier" authorityYear="1967" box="[568,895,1112,1139]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="subGenus" subGenus="Microlepidia">Physodeutera (Microlepidia)</taxonomicName>
/
<taxonomicName id="4C2D4D16FFD1FF88FC04FBEFFB76FBF1" authority="Rivalier, 1967" authorityName="Rivalier" authorityYear="1967" box="[925,1209,1112,1138]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="peyrierasi">
peyrierasi
<bibRefCitation id="EFBC4B64FFD1FF88FB8FFBEFFB76FBF1" author="Rivalier, E." box="[1046,1209,1112,1138]" pageId="12" pageNumber="45" pagination="261 - 296" refId="ref12291" refString="Rivalier, E. (1967) Le genre Physodeutera Lacordaire (Col. Cicindelidae) revision et description d'especes nouvelles. Annales de la Societe entomologique de France, New Series, 3, 261 - 296." type="journal article" year="1967">Rivalier, 1967</bibRefCitation>
</taxonomicName>
/ det.
<determinerName id="7E4EDA51FFD1FF88FA93FBEFFA52FBF0" box="[1290,1437,1112,1139]" pageId="12" pageNumber="45">Jiří Moravec</determinerName>
2020”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C337651EFFD1FF88FF5EFB17FB76F9E9" pageId="12" pageNumber="45" type="diagnosis">
<paragraph id="8B923695FFD1FF88FF5EFB17FE70FA58" blockId="12.[151,1437,716,2039]" pageId="12" pageNumber="45">
<emphasis id="B959EA87FFD1FF88FF5EFB17FE02FB39" bold="true" box="[199,461,1184,1210]" pageId="12" pageNumber="45">Differential diagnosis.</emphasis>
<taxonomicName id="4C2D4D16FFD1FF88FE4DFB17FC5FFB39" box="[468,912,1184,1211]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="peyrierasi" subGenus="Microlepidia">
<emphasis id="B959EA87FFD1FF88FE4DFB17FDA1FB39" box="[468,622,1184,1210]" italics="true" pageId="12" pageNumber="45">Physodeutera</emphasis>
(
<emphasis id="B959EA87FFD1FF88FDE4FB17FCC1FB39" box="[637,782,1184,1210]" italics="true" pageId="12" pageNumber="45">Microlepidia</emphasis>
)
<emphasis id="B959EA87FFD1FF88FC87FB16FC5FFB39" box="[798,912,1185,1210]" italics="true" pageId="12" pageNumber="45">peyrierasi</emphasis>
</taxonomicName>
shares the sexually dimorphic shape of labrum (
<figureCitation id="13162A10FFD1FF88FF06FB73FEEEFB5D" box="[159,289,1220,1246]" captionStart="FIGURES 4049" captionStartId="13.[151,264,1867,1891]" captionTargetBox="[157,1429,210,1832]" captionTargetId="figure-17@13.[151,1435,200,1844]" captionTargetPageId="13" captionText="FIGURES 4049. Physodeutera (Microlepidia) peyrierasi Rivalier. 40—habitus, ♁, 10.3 mm, Mahamasina (CCJM); 41— part of head, type locality Ankarana (CCJM ex APCA); 4244—labrum: 42—♁, type locality (CCJM ex APCA); 43—♁, Mahamasina (CCJM); 44—♀, ibid.; 45—pronotum, ♁, ibid.; 46—aedeagus showing internal sac, type locality (CCJM ex APCA); 4749—elytron: 47—♁, type locality (CCJM ex APCA); 48—♁, Mahamasina (CCJM); 49—♁, ibid. (CCJM). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4594665" httpUri="https://zenodo.org/record/4594665/files/figure.png" pageId="12" pageNumber="45">Figs 4244</figureCitation>
) and truncate aedeagus apex (
<figureCitation id="13162A10FFD1FF88FDE5FB73FD00FB5C" box="[636,719,1220,1247]" captionStart="FIGURES 4049" captionStartId="13.[151,264,1867,1891]" captionTargetBox="[157,1429,210,1832]" captionTargetId="figure-17@13.[151,1435,200,1844]" captionTargetPageId="13" captionText="FIGURES 4049. Physodeutera (Microlepidia) peyrierasi Rivalier. 40—habitus, ♁, 10.3 mm, Mahamasina (CCJM); 41— part of head, type locality Ankarana (CCJM ex APCA); 4244—labrum: 42—♁, type locality (CCJM ex APCA); 43—♁, Mahamasina (CCJM); 44—♀, ibid.; 45—pronotum, ♁, ibid.; 46—aedeagus showing internal sac, type locality (CCJM ex APCA); 4749—elytron: 47—♁, type locality (CCJM ex APCA); 48—♁, Mahamasina (CCJM); 49—♁, ibid. (CCJM). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4594665" httpUri="https://zenodo.org/record/4594665/files/figure.png" pageId="12" pageNumber="45">Fig. 46</figureCitation>
) with
<taxonomicName id="4C2D4D16FFD1FF88FC82FB73FB87FB5D" baseAuthorityName="W. Horn" baseAuthorityYear="1934" box="[795,1096,1220,1247]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="33" phylum="Arthropoda" rank="species" species="marginemaculata" subGenus="Microlepidia">
<emphasis id="B959EA87FFD1FF88FC82FB73FCF5FB5D" box="[795,826,1220,1246]" italics="true" pageId="12" pageNumber="45">Ph</emphasis>
. (
<emphasis id="B959EA87FFD1FF88FCCDFB72FCA4FB5D" box="[852,875,1221,1246]" italics="true" pageId="12" pageNumber="45">M</emphasis>
.)
<emphasis id="B959EA87FFD1FF88FC1DFB73FB87FB5D" box="[900,1096,1220,1246]" italics="true" pageId="12" pageNumber="45">marginemaculata</emphasis>
</taxonomicName>
, but may be distinguished by its generally somewhat smaller body (particularly in males), longer male labrum with better indicated (
<figureCitation id="13162A10FFD1FF88FAD9FB5FFA5BFA81" box="[1344,1428,1256,1282]" captionStart="FIGURES 4049" captionStartId="13.[151,264,1867,1891]" captionTargetBox="[157,1429,210,1832]" captionTargetId="figure-17@13.[151,1435,200,1844]" captionTargetPageId="13" captionText="FIGURES 4049. Physodeutera (Microlepidia) peyrierasi Rivalier. 40—habitus, ♁, 10.3 mm, Mahamasina (CCJM); 41— part of head, type locality Ankarana (CCJM ex APCA); 4244—labrum: 42—♁, type locality (CCJM ex APCA); 43—♁, Mahamasina (CCJM); 44—♀, ibid.; 45—pronotum, ♁, ibid.; 46—aedeagus showing internal sac, type locality (CCJM ex APCA); 4749—elytron: 47—♁, type locality (CCJM ex APCA); 48—♁, Mahamasina (CCJM); 49—♁, ibid. (CCJM). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4594665" httpUri="https://zenodo.org/record/4594665/files/figure.png" pageId="12" pageNumber="45">Fig. 43</figureCitation>
) or even wide, blunt median tooth (
<figureCitation id="13162A10FFD1FF88FD86FABBFDBDFAA5" box="[543,626,1292,1318]" captionStart="FIGURES 4049" captionStartId="13.[151,264,1867,1891]" captionTargetBox="[157,1429,210,1832]" captionTargetId="figure-17@13.[151,1435,200,1844]" captionTargetPageId="13" captionText="FIGURES 4049. Physodeutera (Microlepidia) peyrierasi Rivalier. 40—habitus, ♁, 10.3 mm, Mahamasina (CCJM); 41— part of head, type locality Ankarana (CCJM ex APCA); 4244—labrum: 42—♁, type locality (CCJM ex APCA); 43—♁, Mahamasina (CCJM); 44—♀, ibid.; 45—pronotum, ♁, ibid.; 46—aedeagus showing internal sac, type locality (CCJM ex APCA); 4749—elytron: 47—♁, type locality (CCJM ex APCA); 48—♁, Mahamasina (CCJM); 49—♁, ibid. (CCJM). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4594665" httpUri="https://zenodo.org/record/4594665/files/figure.png" pageId="12" pageNumber="45">Fig. 42</figureCitation>
) and more elongate, parallel-sided elytra with wider humeri and elytral surface evenly convex with indistinct juxtahumeral and discal impressions and markedly reduced elytral maculation (
<figureCitation id="13162A10FFD1FF88FF06FAE3FEEBFAED" box="[159,292,1364,1390]" captionStart="FIGURES 4049" captionStartId="13.[151,264,1867,1891]" captionTargetBox="[157,1429,210,1832]" captionTargetId="figure-17@13.[151,1435,200,1844]" captionTargetPageId="13" captionText="FIGURES 4049. Physodeutera (Microlepidia) peyrierasi Rivalier. 40—habitus, ♁, 10.3 mm, Mahamasina (CCJM); 41— part of head, type locality Ankarana (CCJM ex APCA); 4244—labrum: 42—♁, type locality (CCJM ex APCA); 43—♁, Mahamasina (CCJM); 44—♀, ibid.; 45—pronotum, ♁, ibid.; 46—aedeagus showing internal sac, type locality (CCJM ex APCA); 4749—elytron: 47—♁, type locality (CCJM ex APCA); 48—♁, Mahamasina (CCJM); 49—♁, ibid. (CCJM). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4594665" httpUri="https://zenodo.org/record/4594665/files/figure.png" pageId="12" pageNumber="45">Figs 4749</figureCitation>
). Nevertheless, similarly to
<taxonomicName id="4C2D4D16FFD1FF88FDFAFAE3FC5FFAED" baseAuthorityName="W. Horn" baseAuthorityYear="1934" box="[611,912,1364,1391]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="33" phylum="Arthropoda" rank="species" species="marginemaculata" subGenus="Microlepidia">
<emphasis id="B959EA87FFD1FF88FDFAFAE3FD4DFAED" box="[611,642,1364,1390]" italics="true" pageId="12" pageNumber="45">Ph</emphasis>
. (
<emphasis id="B959EA87FFD1FF88FD02FAE2FD7DFAED" box="[667,690,1365,1390]" italics="true" pageId="12" pageNumber="45">M</emphasis>
.)
<emphasis id="B959EA87FFD1FF88FD52FAE3FC5FFAED" box="[715,912,1364,1390]" italics="true" pageId="12" pageNumber="45">marginemaculata</emphasis>
</taxonomicName>
treated above, the elytral maculation is rather variable also in
<taxonomicName id="4C2D4D16FFD1FF88FED3FACFFDEEFA11" box="[330,545,1400,1427]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="33" phylum="Arthropoda" rank="species" species="peyrierasi" subGenus="Microlepidia">
<emphasis id="B959EA87FFD1FF88FED3FACFFEA6FA11" box="[330,361,1400,1426]" italics="true" pageId="12" pageNumber="45">Ph</emphasis>
. (
<emphasis id="B959EA87FFD1FF88FE19FACEFE58FA11" box="[384,407,1401,1426]" italics="true" pageId="12" pageNumber="45">M</emphasis>
.)
<emphasis id="B959EA87FFD1FF88FE36FACEFDEEFA11" box="[431,545,1401,1426]" italics="true" pageId="12" pageNumber="45">peyrierasi</emphasis>
</taxonomicName>
. It may consist of only small humeral macula in males or possessing also small sublateral-median macula in both sexes, or the elytra may be immaculate in some females (the humeral macula is always absent in females).
</paragraph>
<paragraph id="8B923695FFD1FF88FF5EFA53FB76F9E9" blockId="12.[151,1437,716,2039]" pageId="12" pageNumber="45">
<emphasis id="B959EA87FFD1FF88FF5EFA53FE88FA7D" bold="true" box="[199,327,1508,1534]" pageId="12" pageNumber="45">Variability</emphasis>
. Besides the above-mentioned variability in the number of whitish elytral maculae, the male labrum may also possess somewhat prolonged anteromedian lobe (but never forming acute tooth) as in
<figureCitation id="13162A10FFD1FF88FB4CF9BFFAE7F9A1" box="[1237,1320,1544,1570]" captionStart="FIGURES 4049" captionStartId="13.[151,264,1867,1891]" captionTargetBox="[157,1429,210,1832]" captionTargetId="figure-17@13.[151,1435,200,1844]" captionTargetPageId="13" captionText="FIGURES 4049. Physodeutera (Microlepidia) peyrierasi Rivalier. 40—habitus, ♁, 10.3 mm, Mahamasina (CCJM); 41— part of head, type locality Ankarana (CCJM ex APCA); 4244—labrum: 42—♁, type locality (CCJM ex APCA); 43—♁, Mahamasina (CCJM); 44—♀, ibid.; 45—pronotum, ♁, ibid.; 46—aedeagus showing internal sac, type locality (CCJM ex APCA); 4749—elytron: 47—♁, type locality (CCJM ex APCA); 48—♁, Mahamasina (CCJM); 49—♁, ibid. (CCJM). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4594665" httpUri="https://zenodo.org/record/4594665/files/figure.png" pageId="12" pageNumber="45">Fig. 42</figureCitation>
, while the anteromedian lobe in other males may be truncate in middle and with only indicated minute tooth (as in
<figureCitation id="13162A10FFD1FF88FADBF99BFA40F9C5" box="[1346,1423,1580,1606]" captionStart="FIGURES 4049" captionStartId="13.[151,264,1867,1891]" captionTargetBox="[157,1429,210,1832]" captionTargetId="figure-17@13.[151,1435,200,1844]" captionTargetPageId="13" captionText="FIGURES 4049. Physodeutera (Microlepidia) peyrierasi Rivalier. 40—habitus, ♁, 10.3 mm, Mahamasina (CCJM); 41— part of head, type locality Ankarana (CCJM ex APCA); 4244—labrum: 42—♁, type locality (CCJM ex APCA); 43—♁, Mahamasina (CCJM); 44—♀, ibid.; 45—pronotum, ♁, ibid.; 46—aedeagus showing internal sac, type locality (CCJM ex APCA); 4749—elytron: 47—♁, type locality (CCJM ex APCA); 48—♁, Mahamasina (CCJM); 49—♁, ibid. (CCJM). Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4594665" httpUri="https://zenodo.org/record/4594665/files/figure.png" pageId="12" pageNumber="45">Fig 43</figureCitation>
), corresponding also with the male labrum illustrated by
<bibRefCitation id="EFBC4B64FFD1FF88FC9BF9E7FC67F9E9" author="Rivalier, E." box="[770,936,1616,1642]" pageId="12" pageNumber="45" pagination="261 - 296" refId="ref12291" refString="Rivalier, E. (1967) Le genre Physodeutera Lacordaire (Col. Cicindelidae) revision et description d'especes nouvelles. Annales de la Societe entomologique de France, New Series, 3, 261 - 296." type="journal article" year="1967">Rivalier (1967</bibRefCitation>
. fig. 7) for the
<typeStatus id="54968837FFD1FF88FBD6F9E7FB7AF9E9" box="[1103,1205,1616,1642]" pageId="12" pageNumber="45" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C337651EFFD1FF88FF5EF9C3FD72F8C0" pageId="12" pageNumber="45" type="distribution">
<paragraph id="8B923695FFD1FF88FF5EF9C3FD72F8C0" blockId="12.[151,1437,716,2039]" pageId="12" pageNumber="45">
<emphasis id="B959EA87FFD1FF88FF5EF9C3FE3BF90D" bold="true" box="[199,500,1652,1678]" pageId="12" pageNumber="45">Distribution and biology</emphasis>
(
<figureCitation id="13162A10FFD1FF88FD91F9C3FDAEF90C" box="[520,609,1652,1679]" captionStart="FIGURES 5052" captionStartId="14.[151,264,1943,1967]" captionTargetBox="[201,1384,194,1906]" captionTargetId="figure-27@14.[188,1400,181,1919]" captionTargetPageId="14" captionText="FIGURES 5052. Distribution of three Physodeutera species in North Madagascar. 50— Physodeutera (Microlepidia) propripenis sp. nov.; 51— Physodeutera (Microlepidia) marginemaculata (W. Horn); 52— Physodeutera (Microlepidia) peyrierasi Rivalier." figureDoi="http://doi.org/10.5281/zenodo.4594667" httpUri="https://zenodo.org/record/4594667/files/figure.png" pageId="12" pageNumber="45">Fig. 52</figureCitation>
).
<taxonomicName id="4C2D4D16FFD1FF88FDE0F9C3FBF0F90D" box="[633,1087,1652,1679]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="peyrierasi" subGenus="Microlepidia">
<emphasis id="B959EA87FFD1FF88FDE0F9C3FCDCF90D" box="[633,787,1652,1678]" italics="true" pageId="12" pageNumber="45">Physodeutera</emphasis>
(
<emphasis id="B959EA87FFD1FF88FCBEF9C3FC77F90D" box="[807,952,1652,1678]" italics="true" pageId="12" pageNumber="45">Microlepidia</emphasis>
)
<emphasis id="B959EA87FFD1FF88FC54F9C2FBF0F90D" box="[973,1087,1653,1678]" italics="true" pageId="12" pageNumber="45">peyrierasi</emphasis>
</taxonomicName>
is a very rare species known hitherto only from the
<typeStatus id="54968837FFD1FF88FE04F92FFE30F931" box="[413,511,1688,1714]" pageId="12" pageNumber="45" type="holotype">holotype</typeStatus>
and three above-listed historical specimens, all with “Ankarana” on their labels. Four other examined specimens were caught quite recently in a dry deciduous forest inside the Special Reserve of Ankarana near the eastern entry to the reserve (
<geoCoordinate id="EE195052FFD1FF88FD57F957FCBDF978" box="[718,882,1760,1787]" degrees="12" direction="south" minutes="58" orientation="latitude" pageId="12" pageNumber="45" precision="1" seconds="08.7" value="-12.969083">S12°58´08.7´´</geoCoordinate>
;
<geoCoordinate id="EE195052FFD1FF88FCE7F957FBFDF979" box="[894,1074,1760,1786]" degrees="049" direction="east" minutes="08" orientation="longitude" pageId="12" pageNumber="45" precision="1" seconds="10.3" value="49.136196">E049°08´10.3´´</geoCoordinate>
). The locality is near a camping site with bungalows called Mahamasina (in error written as “Mahamatsina” on the labels). Adults had the same behaviour as those of
<taxonomicName id="4C2D4D16FFD1FF88FE14F89FFD79F8C1" baseAuthorityName="W. Horn" baseAuthorityYear="1934" box="[397,694,1832,1859]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="33" phylum="Arthropoda" rank="species" species="marginemaculata" subGenus="Microlepidia">
<emphasis id="B959EA87FFD1FF88FE14F89FFE63F8C1" box="[397,428,1832,1858]" italics="true" pageId="12" pageNumber="45">Ph</emphasis>
. (
<emphasis id="B959EA87FFD1FF88FE5AF89EFE15F8C1" box="[451,474,1833,1858]" italics="true" pageId="12" pageNumber="45">M</emphasis>
.)
<emphasis id="B959EA87FFD1FF88FE6BF89FFD79F8C1" box="[498,694,1832,1858]" italics="true" pageId="12" pageNumber="45">marginemaculata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C337651EFFD1FF88FF5EF8FBFA56F875" pageId="12" pageNumber="45" type="discussion">
<paragraph id="8B923695FFD1FF88FF5EF8FBFBC3F8E5" blockId="12.[151,1437,716,2039]" box="[199,1036,1868,1895]" pageId="12" pageNumber="45">
<emphasis id="B959EA87FFD1FF88FF5EF8FBFEFBF8E5" bold="true" box="[199,308,1868,1894]" pageId="12" pageNumber="45">Remarks</emphasis>
. For detailed redescription see the monograph (
<bibRefCitation id="EFBC4B64FFD1FF88FCD2F8FBFBCEF8E5" author="Moravec, J." box="[843,1025,1868,1894]" pageId="12" pageNumber="45" refId="ref11194" refString="Moravec, J. (2002 a) Tiger beetles of Madagascar 2. A monograph of the genus Physodeutera (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, Czech Republic, 290 pp." type="book" year="2002">Moravec 2002a</bibRefCitation>
).
</paragraph>
<paragraph id="8B923695FFD1FF88FF5EF8C7FA56F875" blockId="12.[151,1437,716,2039]" pageId="12" pageNumber="45">
As discussed previously (
<bibRefCitation id="EFBC4B64FFD1FF88FE77F8C7FD68F809" author="Moravec, J." box="[494,679,1904,1930]" pageId="12" pageNumber="45" refId="ref11194" refString="Moravec, J. (2002 a) Tiger beetles of Madagascar 2. A monograph of the genus Physodeutera (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, Czech Republic, 290 pp." type="book" year="2002">Moravec 2002a</bibRefCitation>
), the sympatric occurrence in the area of the massif of Ankarana apparently excludes the possibility to consider
<taxonomicName id="4C2D4D16FFD1FF88FD59F823FC6EF82D" box="[704,929,1940,1967]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="33" phylum="Arthropoda" rank="species" species="peyrierasi" subGenus="Microlepidia">
<emphasis id="B959EA87FFD1FF88FD59F823FD10F82D" box="[704,735,1940,1966]" italics="true" pageId="12" pageNumber="45">Ph</emphasis>
. (
<emphasis id="B959EA87FFD1FF88FD62F822FCDDF82D" box="[763,786,1941,1966]" italics="true" pageId="12" pageNumber="45">M</emphasis>
.)
<emphasis id="B959EA87FFD1FF88FCB6F822FC6EF82D" box="[815,929,1941,1966]" italics="true" pageId="12" pageNumber="45">peyrierasi</emphasis>
</taxonomicName>
to be a geographic subspecies of
<taxonomicName id="4C2D4D16FFD1FF88FAA3F823FE94F851" baseAuthorityName="W. Horn" baseAuthorityYear="1934" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="33" phylum="Arthropoda" rank="species" species="marginemaculata" subGenus="Microlepidia">
<emphasis id="B959EA87FFD1FF88FAA3F823FA96F82D" box="[1338,1369,1940,1966]" italics="true" pageId="12" pageNumber="45">Ph</emphasis>
. (
<emphasis id="B959EA87FFD1FF88FAECF822FA43F82D" box="[1397,1420,1941,1966]" italics="true" pageId="12" pageNumber="45">M</emphasis>
.)
<emphasis id="B959EA87FFD1FF88FF0EF80FFE94F851" box="[151,347,1976,2002]" italics="true" pageId="12" pageNumber="45">marginemaculata</emphasis>
</taxonomicName>
. Consequently, and taking into consideration the tremendous biodiversity in
<collectingCountry id="F33A7605FFD1FF88FB7AF80FFAA3F851" box="[1251,1388,1976,2002]" name="Madagascar" pageId="12" pageNumber="45">Madagascar</collectingCountry>
, we prefer also here to consider the differences between these two taxa sufficient to keep them as two separate species.
</paragraph>
</subSubSection>
</treatment>
</document>