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<document id="47A5659999081A86C4C79EA064F6640E" ID-CLB-Dataset="25600" ID-DOI="10.11646/zootaxa.4664.4.5" ID-GBIF-Dataset="844fafcf-8f84-4039-ade1-7a03eadaf1d4" ID-ISSN="1175-5326" ID-Zenodo-Dep="3386951" ID-ZooBank="B875497D-DF28-4639-880B-2B3F52E3631B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1567672571258" checkinUser="plazi" docAuthor="Boll, Piter Kehoma, Amaral, Silvana Vargas Do &amp; Leal-Zanchet, Ana Maria" docDate="2019" docId="03848782FF8DFFE3FF7EFC29FBEBFC1E" docLanguage="en" docName="zootaxa.4664.4.5.pdf.imf" docOrigin="Zootaxa 4664 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cratera obsidiana Boll &amp; Amaral &amp; Leal-Zanchet 2019, sp. nov." docType="treatment" docUuid="9E3C4246-0E1D-4838-81E9-6ADE0882CFF0" docUuidSource="ZooBank" docVersion="10" lastPageNumber="542" masterDocId="FFBDFFFAFF8CFFE4FFE9FF94FFC8FFD5" masterDocTitle="Two new land planarian species (Platyhelminthes: Tricladida: Geoplanidae) from protected areas in southern Brazil" masterLastPageNumber="550" masterPageNumber="535" pageNumber="536" updateTime="1698754694161" updateUser="ExternalLinkService">
<mods:mods id="57D26C57B51CDA1FDE90AC8F3BF7995F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="78EFC0C9C78F28F32814EA137CA0D0FB">
<mods:title id="8DD5B962F808F8DC94D2B94F045CD506">Two new land planarian species (Platyhelminthes: Tricladida: Geoplanidae) from protected areas in southern Brazil</mods:title>
</mods:titleInfo>
<mods:name id="575ED8DA8958621A87AC23274B0F62F0" type="personal">
<mods:role id="F7A5CE9D8A8502488F6EC26D286BEE06">
<mods:roleTerm id="0087FCCC47857B351D6F2A0FC6E93DBD">Author</mods:roleTerm>
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<mods:namePart id="D47FA5D0276BBEEA1522B78C62B4C949">Boll, Piter Kehoma</mods:namePart>
</mods:name>
<mods:name id="2A4053DFF0D86093707DF70D3217A02C" type="personal">
<mods:role id="749A4503E0EEB422FA723AE73C2925D3">
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</mods:role>
<mods:namePart id="A4A23221C3A5ACFCDD972829146B41F3">Amaral, Silvana Vargas Do</mods:namePart>
</mods:name>
<mods:name id="89E0527DE52E739EC3960C5FD474EEA8" type="personal">
<mods:role id="1FFE46D38E8566EE8286BBC61A527EA1">
<mods:roleTerm id="D1A7E54089067C9DA820BC2F99ECB903">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="66EB7DA6A4D823787C0E83D9A99FD200">Leal-Zanchet, Ana Maria</mods:namePart>
</mods:name>
<mods:typeOfResource id="5138121D06D2ABFB8066FD06D3F2F0DB">text</mods:typeOfResource>
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<mods:titleInfo id="590800A74A0DDD38CC22040E72ECD375">
<mods:title id="182488191FE2326B7C6D31D592B3A027">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="743E2A4DA33C155D08A95A749486BCCF">
<mods:date id="5BC0814CD1971309200C3D343CD05F7C">2019</mods:date>
<mods:detail id="049140C89412819BF6306E6F68FC18FA" type="pubDate">
<mods:number id="D49262EFB24708240F953C4C60D8DD9A">2019-09-05</mods:number>
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<mods:detail id="DC81D655340D974DEC9EBE56441FA921" type="volume">
<mods:number id="FABE3B76125400E22842B5800CE8A7E0">4664</mods:number>
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<mods:number id="184E73ACEB1B58DDB76425DC1393C5CB">4</mods:number>
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<mods:classification id="349C473410EB3F5877A65FF9F34DA729">journal article</mods:classification>
<mods:identifier id="ED929A127C26427ABD92D48F83AC6D4D" type="CLB-Dataset">25600</mods:identifier>
<mods:identifier id="9D5041161CF1B387A00B210B9DFE5D3E" type="DOI">10.11646/zootaxa.4664.4.5</mods:identifier>
<mods:identifier id="4EFC8DC4089FA6F84B19E14B28F6304D" type="GBIF-Dataset">844fafcf-8f84-4039-ade1-7a03eadaf1d4</mods:identifier>
<mods:identifier id="7A4A761C04A816810C19E26B78D974E7" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="9CDF9F1391F1CBEAE7B507B61D6DA85A" type="Zenodo-Dep">3386951</mods:identifier>
<mods:identifier id="E055A90CC6AD00D99BD3FFF243C6354E" type="ZooBank">B875497D-DF28-4639-880B-2B3F52E3631B</mods:identifier>
</mods:mods>
<treatment id="03848782FF8DFFE3FF7EFC29FBEBFC1E" ID-GBIF-Taxon="159611485" LSID="urn:lsid:zoobank.org:act:9E3C4246-0E1D-4838-81E9-6ADE0882CFF0" httpUri="http://treatment.plazi.org/id/03848782FF8DFFE3FF7EFC29FBEBFC1E" lastPageId="7" lastPageNumber="542" pageId="1" pageNumber="536">
<subSubSection id="C337651FFF8DFFE5FF7EFC29FC08FC2E" pageId="1" pageNumber="536" type="nomenclature">
<paragraph id="8B923694FF8DFFE5FF7EFC29FC9DFC02" blockId="1.[151,960,957,1019]" box="[151,853,957,984]" pageId="1" pageNumber="536">
<heading id="D0DA81F8FF8DFFE5FF7EFC29FC9DFC02" bold="true" box="[151,853,957,984]" fontSize="11" level="1" pageId="1" pageNumber="536" reason="1">
<emphasis id="B959EA86FF8DFFE5FF7EFC29FC9DFC02" bold="true" box="[151,853,957,984]" pageId="1" pageNumber="536">
<taxonomicName id="4C2D4D17FF8DFFE5FF7EFC29FD23FC02" authority="Amaral, Boll &amp; Leal-Zanchet" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[151,747,957,984]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="1" pageNumber="536" phylum="Platyhelminthes" rank="species" species="obsidiana" status="sp. nov.">
<emphasis id="B959EA86FF8DFFE5FF7EFC29FEB8FC0D" bold="true" box="[151,368,957,984]" italics="true" pageId="1" pageNumber="536">Cratera obsidiana</emphasis>
Amaral, Boll &amp; Leal-Zanchet
</taxonomicName>
,
<taxonomicNameLabel id="A26A57FDFF8DFFE5FD1EFC29FC9DFC02" box="[759,853,957,983]" pageId="1" pageNumber="536" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B923694FF8DFFE5FF7EFC75FC08FC2E" blockId="1.[151,960,957,1019]" box="[151,960,993,1019]" pageId="1" pageNumber="536">
<uri id="FFBC3A96FF8DFFE5FF7EFC75FC08FC2E" box="[151,960,993,1019]" pageId="1" pageNumber="536">
urn:lsid:zoobank.org:act:
<uuid id="FF8B0C41FF8DFFE5FE43FC75FC08FC2E" box="[426,960,993,1019]" pageId="1" pageNumber="536">9E3C4246-0E1D-4838-81E9-6ADE0882CFF0</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C337651FFF8DFFE5FF7EFBB5FD4DFB8D" pageId="1" pageNumber="536" type="description">
<paragraph id="8B923694FF8DFFE5FF7EFBB5FD2DFBEC" blockId="1.[151,741,1056,1112]" box="[151,741,1056,1081]" pageId="1" pageNumber="536">
<taxonomicName id="4C2D4D17FF8DFFE5FF7EFBB5FF33FBEC" box="[151,251,1057,1081]" class="Rhabditophora" family="Geoplanidae" genus="Geoplana" kingdom="Animalia" order="Seriata" pageId="1" pageNumber="536" phylum="Platyhelminthes" rank="genus">
<emphasis id="B959EA86FF8DFFE5FF7EFBB5FF33FBEC" box="[151,251,1057,1081]" italics="true" pageId="1" pageNumber="536">Geoplana</emphasis>
</taxonomicName>
sp.
<quantity id="4CD59B71FF8DFFE5FECCFBB4FE84FBEC" box="[293,332,1056,1081]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="1" pageNumber="536" unit="in" value="6.0">6 in</quantity>
<bibRefCitation id="EFBC4B65FF8DFFE5FEBBFBB4FD2DFBEC" author="Baptista, V. dos &amp; Oliveira, S. M. de &amp; Leal-Zanchet, A. M." box="[338,741,1056,1081]" pageId="1" pageNumber="536" pagination="247 - 252" refId="ref10192" refString="Baptista, V. dos A., Oliveira, S. M. de &amp; Leal-Zanchet, A. M. (2010) Inventario de planarias terrestres (Platyhelminthes, Tricladida) em remanescente de Floresta Estacional Decidual do Sul do Brasil. Biota Neotropica, 10, 247 - 252. https: // doi. org / 10.1590 / S 1676 - 0603201000020002 7" type="journal article" year="2010">Baptista, Oliveira &amp; Leal-Zanchet 2010</bibRefCitation>
</paragraph>
<paragraph id="8B923694FF8DFFE5FF7EFBD4FD4DFB8D" blockId="1.[151,741,1056,1112]" box="[151,645,1087,1112]" pageId="1" pageNumber="536">
<taxonomicName id="4C2D4D17FF8DFFE5FF7EFBD4FF33FB8D" box="[151,251,1088,1112]" class="Rhabditophora" family="Geoplanidae" genus="Geoplana" kingdom="Animalia" order="Seriata" pageId="1" pageNumber="536" phylum="Platyhelminthes" rank="genus">
<emphasis id="B959EA86FF8DFFE5FF7EFBD4FF33FB8D" box="[151,251,1088,1112]" italics="true" pageId="1" pageNumber="536">Geoplana</emphasis>
</taxonomicName>
sp.
<quantity id="4CD59B71FF8DFFE5FECCFBABFE84FB8D" box="[293,332,1087,1112]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="1" pageNumber="536" unit="in" value="6.0">6 in</quantity>
<bibRefCitation id="EFBC4B65FF8DFFE5FEBBFBABFD4DFB8D" author="Baptista, V. A. &amp; Leal-Zanchet, A. M." box="[338,645,1087,1112]" pageId="1" pageNumber="536" pagination="83 - 90" refId="ref10260" refString="Baptista, V. A. &amp; Leal-Zanchet, A. M. (2010) Land flatworm community structure in a subtropical deciduous forest in Southern Brazil. Belgian Journal of Zoology, 140, 83 - 90." type="journal article" year="2010">Baptista &amp; Leal-Zanchet 2010</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C337651FFF8DFFE5FF7EFB16FD81FB15" pageId="1" pageNumber="536" type="etymology">
<paragraph id="8B923694FF8DFFE5FF7EFB16FD81FB15" blockId="1.[151,1437,1154,2045]" pageId="1" pageNumber="536">
<emphasis id="B959EA86FF8DFFE5FF7EFB16FEEAFB49" bold="true" box="[151,290,1154,1180]" pageId="1" pageNumber="536">Etymology:</emphasis>
The specific name is a noun in apposition and refers to the bright glossy black color of the dorsum, resembling the volcanic rock obsidian.
</paragraph>
</subSubSection>
<subSubSection id="C337651FFF8DFFE5FF2EFB5EFC1EF9FD" pageId="1" pageNumber="536" type="materials_examined">
<paragraph id="8B923694FF8DFFE5FF2EFB5EFA9EFA85" blockId="1.[151,1437,1154,2045]" pageId="1" pageNumber="536">
<emphasis id="B959EA86FF8DFFE5FF2EFB5EFEB8FB31" bold="true" box="[199,368,1225,1252]" pageId="1" pageNumber="536">Type material.</emphasis>
<materialsCitation id="3B453CC9FF8DFFE5FE9FFB5EFA9EFA85" ID-GBIF-Occurrence="2382692858" collectingDate="2006-09-10" collectionCode="MZUSP, PL" collectorName="S. M. Oliveira" county="State Park of Turvo" latitude="-27.333334" location="Brazil-anterior" longLatPrecision="1237" longitude="-54.166668" municipality="Derrubadas" pageId="1" pageNumber="536" specimenCount="1" stateProvince="Rio Grande do Sul" typeStatus="holotype">
<emphasis id="B959EA86FF8DFFE5FE9FFB5EFE22FB31" bold="true" box="[374,490,1226,1252]" pageId="1" pageNumber="536">
<typeStatus id="54968836FF8DFFE5FE9FFB5EFE2DFB31" box="[374,485,1226,1252]" pageId="1" pageNumber="536" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED3CAE51FF8DFFE5FE19FB5EFD83FB31" box="[496,587,1226,1252]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="536">MZUSP</collectionCode>
<collectionCode id="ED3CAE51FF8DFFE5FDB8FB5DFDBDFB31" box="[593,629,1225,1252]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="1" pageNumber="536" type="Museum">PL</collectionCode>
.2158:
<emphasis id="B959EA86FF8DFFE5FD55FB5EFD2CFB31" box="[700,740,1226,1252]" italics="true" pageId="1" pageNumber="536">leg.</emphasis>
<collectorName id="26D85342FF8DFFE5FD03FB5EFC4FFB31" box="[746,903,1226,1253]" pageId="1" pageNumber="536">S. M. Oliveira</collectorName>
,
<date id="FF931054FF8DFFE5FC78FB5EFBAFFB31" box="[913,1127,1226,1252]" pageId="1" pageNumber="536" value="2006-09-10">
<collectingDate id="EFD7E9BCFF8DFFE5FC78FB5EFBAFFB31" box="[913,1127,1226,1252]" pageId="1" pageNumber="536" value="2006-09-10">10 September 2006</collectingDate>
</date>
,
<collectingCounty id="62F34E18FF8DFFE5FB98FB5EFA8CFB30" box="[1137,1348,1226,1253]" pageId="1" pageNumber="536">State Park of Turvo</collectingCounty>
(27°00’ to
<geoCoordinate id="EE195053FF8DFFE5FF5DFB7AFED8FADD" box="[180,272,1262,1288]" degrees="27" direction="south" minutes="20" orientation="latitude" pageId="1" pageNumber="536" precision="925" value="-27.333334">27°20’S</geoCoordinate>
, 53°40’ to
<geoCoordinate id="EE195053FF8DFFE5FE60FB79FE39FADD" box="[393,497,1261,1288]" degrees="54" direction="west" minutes="10" orientation="longitude" pageId="1" pageNumber="536" precision="925" value="-54.166668">54°10’W</geoCoordinate>
),
<collectingMunicipality id="6BF6ACEEFF8DFFE5FDECFB7AFD4EFADD" box="[517,646,1262,1288]" pageId="1" pageNumber="536">Derrubadas</collectingMunicipality>
, state of
<collectingRegion id="49E9F876FF8DFFE5FD00FB7AFC71FADD" box="[745,953,1262,1288]" country="Brazil" name="Rio Grande do Sul" pageId="1" pageNumber="536">Rio Grande do Sul</collectingRegion>
,
<location id="8EF2604FFF8DFFE5FC2CFB7AFBB0FADD" LSID="urn:lsid:plazi:treatment:03848782FF8DFFE3FF7EFC29FBEBFC1E:8EF2604FFF8DFFE5FC2CFB7AFBB0FADD" box="[965,1144,1262,1288]" county="State Park of Turvo" latitude="-27.333334" longLatPrecision="1237" longitude="-54.166668" municipality="Derrubadas" name="Brazil-anterior" pageId="1" pageNumber="536" stateProvince="Rio Grande do Sul">Brazil—anterior</location>
tip: transverse sections on 18 slides; anterior region at the level of the ovaries: sagittal sections on 30 slides; pre-pharyngeal region: transverse sections on 6 slides; pharynx: sagittal sections on 14 slides; copulatory apparatus: sagittal sections on 26 slides.
</materialsCitation>
</paragraph>
<paragraph id="8B923694FF8DFFE5FF2EFACEFC1EF9FD" blockId="1.[151,1437,1154,2045]" pageId="1" pageNumber="536">
<materialsCitation id="3B453CC9FF8DFFE5FF2EFACEFC1AF9FD" ID-GBIF-Occurrence="2382692861" collectingDate="2006-09-10" collectingDateMax="2006-11-17" collectingDateMin="2006-09-10" collectionCode="PL" collectorName="Three &amp; C. Palacios &amp; S. M. Oliveira &amp; M. B. Antunes" county="State Park of Turvo" latitude="-27.333334" location="Brazil-anterior" longLatPrecision="1237" longitude="-54.166668" municipality="Derrubadas" pageId="1" pageNumber="536" specimenCount="1" stateProvince="Rio Grande do Sul" typeStatus="paratype">
<emphasis id="B959EA86FF8DFFE5FF2EFACEFE80FAA1" bold="true" box="[199,328,1370,1396]" pageId="1" pageNumber="536">
<typeStatus id="54968836FF8DFFE5FF2EFACEFE8BFAA1" box="[199,323,1370,1396]" pageId="1" pageNumber="536" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectorName id="26D85342FF8DFFE5FEA6FACEFE58FAA1" box="[335,400,1370,1396]" pageId="1" pageNumber="536">Three</collectorName>
specimens collected in the same locality as the holotype. MZU
<collectionCode id="ED3CAE51FF8DFFE5FBB4FACFFB49FAA0" box="[1117,1153,1371,1397]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="1" pageNumber="536" type="Museum">PL</collectionCode>
. 270:
<emphasis id="B959EA86FF8DFFE5FB2DFACEFB2EFAA1" box="[1220,1254,1370,1396]" italics="true" pageId="1" pageNumber="536">leg</emphasis>
.
<collectorName id="26D85342FF8DFFE5FB1DFACEFABDFAA1" box="[1268,1397,1370,1396]" pageId="1" pageNumber="536">C. Palacios</collectorName>
,
<date id="FF931054FF8DFFE5FA69FACEFE82FA4D" pageId="1" pageNumber="536" value="2006-11-17">
<collectingDate id="EFD7E9BCFF8DFFE5FA69FACEFE82FA4D" pageId="1" pageNumber="536" value="2006-11-17">17 November 2006</collectingDate>
</date>
—copulatory apparatus: sagittal sections on 9 slides; MZU
<collectionCode id="ED3CAE51FF8DFFE5FC34FAEBFBC9FA4C" box="[989,1025,1407,1433]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="1" pageNumber="536" type="Museum">PL</collectionCode>
. 271:
<emphasis id="B959EA86FF8DFFE5FBABFAEAFBACFA4D" box="[1090,1124,1406,1432]" italics="true" pageId="1" pageNumber="536">leg</emphasis>
.
<collectorName id="26D85342FF8DFFE5FB9BFAEAFADBFA4D" box="[1138,1299,1406,1433]" pageId="1" pageNumber="536">S. M. Oliveira</collectorName>
,
<date id="FF931054FF8DFFE5FAF4FAEAFF30FA69" pageId="1" pageNumber="536" value="2006-09-10">
<collectingDate id="EFD7E9BCFF8DFFE5FAF4FAEAFF30FA69" pageId="1" pageNumber="536" value="2006-09-10">10 September 2006</collectingDate>
</date>
—anterior tip: sagittal sections on 8 slides; anterior region at the level of the ovaries: sagittal sections on 49 slides; pre-pharyngeal region: transverse sections on 15 slides; pharynx: sagittal sections on 15 slides; copulatory apparatus: horizontal sections on 9 slides. MZU
<collectionCode id="ED3CAE51FF8DFFE5FD2FFA7FFD22F9D0" box="[710,746,1515,1541]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="1" pageNumber="536" type="Museum">PL</collectionCode>
. 272:
<emphasis id="B959EA86FF8DFFE5FCDAFA7EFC9DF9D1" box="[819,853,1514,1540]" italics="true" pageId="1" pageNumber="536">leg</emphasis>
.
<collectorName id="26D85342FF8DFFE5FC8CFA7FFBD8F9D1" box="[869,1040,1514,1541]" pageId="1" pageNumber="536">M. B. Antunes</collectorName>
,
<date id="FF931054FF8DFFE5FBF7FA7EFB31F9D1" box="[1054,1273,1514,1540]" pageId="1" pageNumber="536" value="2006-11-17">
<collectingDate id="EFD7E9BCFF8DFFE5FBF7FA7EFB31F9D1" box="[1054,1273,1514,1540]" pageId="1" pageNumber="536" value="2006-11-17">17 November 2006</collectingDate>
</date>
—anterior tip: preserved in clove oil; copulatory apparatus: sagittal sections on 11 slides
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C337651FFF8DFFE5FF2EF9A5FBA6F941" pageId="1" pageNumber="536" type="diagnosis">
<paragraph id="8B923694FF8DFFE5FF2EF9A5FAA5F9A5" blockId="1.[151,1437,1154,2045]" pageId="1" pageNumber="536">
<emphasis id="B959EA86FF8DFFE5FF2EF9A5FE8CF999" bold="true" box="[199,324,1585,1612]" pageId="1" pageNumber="536">Diagnosis:</emphasis>
species of
<taxonomicName id="4C2D4D17FF8DFFE5FE55F9A7FDDDF998" authorityName="Carbayo, Alvarez-Presas, Olivares, Marques, Froehlich &amp; Riutort" authorityYear="2013" box="[444,533,1587,1613]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="1" pageNumber="536" phylum="Platyhelminthes" rank="genus">
<emphasis id="B959EA86FF8DFFE5FE55F9A7FDDDF998" box="[444,533,1587,1613]" italics="true" pageId="1" pageNumber="536">Cratera</emphasis>
</taxonomicName>
with dorsal color homogeneously black; pharynx cylindrical with dorsal insertion shifted posteriorly; prostatic vesicle extrabulbar with proximal portion T-shaped and slightly dorsally oriented.
</paragraph>
<paragraph id="8B923694FF8DFFE5FF2EF9EDFBA6F941" blockId="1.[151,1437,1154,2045]" box="[199,1134,1657,1685]" pageId="1" pageNumber="536">
<emphasis id="B959EA86FF8DFFE5FF2EF9EDFEA4F941" bold="true" box="[199,364,1657,1684]" pageId="1" pageNumber="536">Type-locality:</emphasis>
State Park of Turvo, Derrubadas, state of
<collectingRegion id="49E9F876FF8DFFE5FCAAF9EEFBDFF941" box="[835,1047,1658,1684]" country="Brazil" name="Rio Grande do Sul" pageId="1" pageNumber="536">Rio Grande do Sul</collectingRegion>
,
<collectingCountry id="F33A7604FF8DFFE5FBCBF9EEFBA2F941" box="[1058,1130,1658,1684]" name="Brazil" pageId="1" pageNumber="536">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C337651FFF8DFFE5FF2EF909FD38F96D" box="[199,752,1693,1720]" pageId="1" pageNumber="536" type="distribution">
<paragraph id="8B923694FF8DFFE5FF2EF909FD38F96D" blockId="1.[151,1437,1154,2045]" box="[199,752,1693,1720]" pageId="1" pageNumber="536">
<emphasis id="B959EA86FF8DFFE5FF2EF909FEABF96D" bold="true" box="[199,355,1693,1720]" pageId="1" pageNumber="536">Distribution:</emphasis>
known only from the type-locality.
</paragraph>
</subSubSection>
<subSubSection id="C337651FFF8DFFE3FF2EF955FBEBFC1E" lastPageId="7" lastPageNumber="542" pageId="1" pageNumber="536" type="description">
<paragraph id="8B923694FF8DFFE5FF2EF955FE09F8B8" blockId="1.[151,1437,1154,2045]" pageId="1" pageNumber="536">
<emphasis id="B959EA86FF8DFFE5FF2EF955FDF2F909" bold="true" box="[199,570,1729,1756]" pageId="1" pageNumber="536">Description. External features.</emphasis>
Body elongate with parallel margins; anterior tip rounded and posterior tip pointed. While creeping, the largest specimen reached a length of
<quantity id="4CD59B71FF8DFFE5FC68F972FC1CF8D4" box="[897,980,1766,1793]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="536" unit="mm" value="40.0">40 mm</quantity>
. After fixation, maximum length was
<quantity id="4CD59B71FF8DFFE5FA69F972FF0EF8F0" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="536" unit="mm" value="38.0">38 mm</quantity>
. Mouth and gonopore located at the posterior third of the body (
<tableCitation id="C6AF032FFF8DFFE5FC4AF89EFC30F8F1" box="[931,1016,1802,1828]" captionStart="TABLE 1" captionStartId="2.[151,239,931,955]" captionTargetPageId="2" captionText="TABLE 1. Measurements, in mm, of specimens of Cratera obsidiana sp. nov. –: not measured; *: after fixation; DG: distance of gonopore from anterior end; DM: distance of mouth from anterior end; DMG: distance between mouth and gonopore; DPVP: distance between prostatic vesicle and pharyngeal pouch. The numbers given in parentheses represent the position relative to body length." httpUri="http://table.plazi.org/id/DF52661CFF8EFFE6FF7EFC37FDDBFBF2" pageId="1" pageNumber="536" tableUuid="DF52661CFF8EFFE6FF7EFC37FDDBFBF2">Table 1</tableCitation>
). In living specimens, dorsal surface homogeneously black (
<figureCitation id="13162A11FF8DFFE5FE77F8BAFE29F89D" box="[414,481,1838,1864]" captionStart="FIGURE 1" captionStartId="2.[151,250,858,883]" captionTargetBox="[162,1423,526,831]" captionTargetId="figure@2.[151,1436,517,839]" captionTargetPageId="2" captionText="FIGURE 1. Cratera obsidiana sp. nov.: dorsal aspect and eye pattern of fixed specimen (paratype MZU PL. 272)." httpUri="https://zenodo.org/record/3386953/files/figure.png" pageId="1" pageNumber="536">Fig. 1</figureCitation>
); venter light gray. After fixation, dorsal color becomes light brown and the ventral surface remains light gray.
</paragraph>
<paragraph id="8B923694FF8DFFE5FF2EF8E3FEC5F828" blockId="1.[151,1437,1154,2045]" pageId="1" pageNumber="536">
Eyes monolobate (with pigment cups of about
<quantity id="4CD59B71FF8DFFE5FD3EF8E1FC83F844" box="[727,843,1909,1937]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="1" pageNumber="536" unit="mm" value="22.5" valueMax="30.0" valueMin="15.0">15–30 µm</quantity>
) (
<figureCitation id="13162A11FF8DFFE5FC8DF8E2FC61F845" box="[868,937,1910,1936]" captionStart="FIGURES 2–8" captionStartId="4.[151,264,1934,1959]" captionTargetBox="[217,1387,166,1915]" captionTargetId="figure@4.[199,1387,166,1915]" captionTargetPageId="4" captionText="FIGURES 2–8. Cratera obsidiana sp. nov., holotype in transverse (2–6) and sagittal sections with the anterior end to the left (7, 8): (2) cephalic region; (3) detail of body-margin in pre-pharyngeal region; (4) pre-pharyngeal region; (5) detail of the dorsal surface in the pre-pharyngeal region; (6) detail of the ventral surface in the pre-pharyngeal region; (7) pharynx; (8) ovary." httpUri="https://zenodo.org/record/3386955/files/figure.png" pageId="1" pageNumber="536">Fig. 2</figureCitation>
), initially uniserial, surrounding the anterior tip (
<figureCitation id="13162A11FF8DFFE5FF2FF80EFEC2F861" box="[198,266,1946,1972]" captionStart="FIGURE 1" captionStartId="2.[151,250,858,883]" captionTargetBox="[162,1423,526,831]" captionTargetId="figure@2.[151,1436,517,839]" captionTargetPageId="2" captionText="FIGURE 1. Cratera obsidiana sp. nov.: dorsal aspect and eye pattern of fixed specimen (paratype MZU PL. 272)." httpUri="https://zenodo.org/record/3386953/files/figure.png" pageId="1" pageNumber="536">Fig. 1</figureCitation>
). After that, the eyes become larger and surrounded by clear halos, spreading onto the dorsal surface, and may occupy the whole body width (
<figureCitation id="13162A11FF8DFFE5FDB7F82AFD68F80D" box="[606,672,1982,2008]" captionStart="FIGURE 1" captionStartId="2.[151,250,858,883]" captionTargetBox="[162,1423,526,831]" captionTargetId="figure@2.[151,1436,517,839]" captionTargetPageId="2" captionText="FIGURE 1. Cratera obsidiana sp. nov.: dorsal aspect and eye pattern of fixed specimen (paratype MZU PL. 272)." httpUri="https://zenodo.org/record/3386953/files/figure.png" pageId="1" pageNumber="536">Fig. 1</figureCitation>
). Towards the posterior tip, the eyes remain dorsal but become less numerous.
</paragraph>
<paragraph id="8B923694FF8EFFE6FF2EFF0CFE27FF2E" blockId="2.[151,1437,152,883]" pageId="2" pageNumber="537">
<emphasis id="B959EA86FF8EFFE6FF2EFF0CFCD5FF66" bold="true" box="[199,797,152,179]" pageId="2" pageNumber="537">Sensory organs, epidermis and body musculature.</emphasis>
Sensory pits (
<figureCitation id="13162A11FF8EFFE6FC57FF0CFBC9FF66" box="[958,1025,152,179]" captionStart="FIGURES 2–8" captionStartId="4.[151,264,1934,1959]" captionTargetBox="[217,1387,166,1915]" captionTargetId="figure@4.[199,1387,166,1915]" captionTargetPageId="4" captionText="FIGURES 2–8. Cratera obsidiana sp. nov., holotype in transverse (2–6) and sagittal sections with the anterior end to the left (7, 8): (2) cephalic region; (3) detail of body-margin in pre-pharyngeal region; (4) pre-pharyngeal region; (5) detail of the dorsal surface in the pre-pharyngeal region; (6) detail of the ventral surface in the pre-pharyngeal region; (7) pharynx; (8) ovary." httpUri="https://zenodo.org/record/3386955/files/figure.png" pageId="2" pageNumber="537">Fig. 2</figureCitation>
) as simple invaginations (
<quantity id="4CD59B71FF8EFFE6FACEFF0DFA54FF66" box="[1319,1436,153,179]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="2" pageNumber="537" unit="mm" value="25.0" valueMax="30.0" valueMin="20.0">20–30 µm</quantity>
deep) contour anterior tip and occur ventromarginally in one row on each side of the body in the cephalic region (about 7% of the body length).
</paragraph>
<paragraph id="8B923694FF8EFFE6FF2EFE90FE0CFE22" blockId="2.[151,1437,152,883]" pageId="2" pageNumber="537">
The creeping sole occupies the whole body width (
<tableCitation id="C6AF032FFF8EFFE6FCFBFE90FCA0FECA" box="[786,872,260,287]" captionStart="TABLE 1" captionStartId="2.[151,239,931,955]" captionTargetPageId="2" captionText="TABLE 1. Measurements, in mm, of specimens of Cratera obsidiana sp. nov. –: not measured; *: after fixation; DG: distance of gonopore from anterior end; DM: distance of mouth from anterior end; DMG: distance between mouth and gonopore; DPVP: distance between prostatic vesicle and pharyngeal pouch. The numbers given in parentheses represent the position relative to body length." httpUri="http://table.plazi.org/id/DF52661CFF8EFFE6FF7EFC37FDDBFBF2" pageId="2" pageNumber="537" tableUuid="DF52661CFF8EFFE6FF7EFC37FDDBFBF2">Table 1</tableCitation>
). Three
<typeStatus id="54968836FF8EFFE6FC25FE91FBCFFECA" box="[972,1031,261,287]" pageId="2" pageNumber="537">types</typeStatus>
of glands discharge through whole epidermis of pre-pharyngeal region: rhabditogen glands with xanthophil secretion (ventrally with smaller rhabdites) and cyanophil glands with amorphous secretion, besides scarce xanthophil glands with coarse granular secretion (
<figureCitation id="13162A11FF8EFFE6FF76FEE4FEC1FE5E" box="[159,265,368,395]" captionStart="FIGURES 2–8" captionStartId="4.[151,264,1934,1959]" captionTargetBox="[217,1387,166,1915]" captionTargetId="figure@4.[199,1387,166,1915]" captionTargetPageId="4" captionText="FIGURES 2–8. Cratera obsidiana sp. nov., holotype in transverse (2–6) and sagittal sections with the anterior end to the left (7, 8): (2) cephalic region; (3) detail of body-margin in pre-pharyngeal region; (4) pre-pharyngeal region; (5) detail of the dorsal surface in the pre-pharyngeal region; (6) detail of the ventral surface in the pre-pharyngeal region; (7) pharynx; (8) ovary." httpUri="https://zenodo.org/record/3386955/files/figure.png" pageId="2" pageNumber="537">Figs. 3, 6</figureCitation>
). The glandular margin (
<figureCitation id="13162A11FF8EFFE6FDF5FEE4FD4DFE5E" box="[540,645,368,395]" captionStart="FIGURES 2–8" captionStartId="4.[151,264,1934,1959]" captionTargetBox="[217,1387,166,1915]" captionTargetId="figure@4.[199,1387,166,1915]" captionTargetPageId="4" captionText="FIGURES 2–8. Cratera obsidiana sp. nov., holotype in transverse (2–6) and sagittal sections with the anterior end to the left (7, 8): (2) cephalic region; (3) detail of body-margin in pre-pharyngeal region; (4) pre-pharyngeal region; (5) detail of the dorsal surface in the pre-pharyngeal region; (6) detail of the ventral surface in the pre-pharyngeal region; (7) pharynx; (8) ovary." httpUri="https://zenodo.org/record/3386955/files/figure.png" pageId="2" pageNumber="537">Figs. 3, 4</figureCitation>
) is conspicuous after the cephalic region (about 8% of the body length) and receives the openings of two
<typeStatus id="54968836FF8EFFE6FDEEFE01FD8AFE7A" box="[519,578,405,431]" pageId="2" pageNumber="537">types</typeStatus>
of glands: xanthophil cells with coarse granular secretion and cyanophil cells with amorphous secretions. Glands discharging through the anterior tip of the body are similar to those of the prepharyngeal region (
<figureCitation id="13162A11FF8EFFE6FE98FE48FE7EFE22" box="[369,438,476,503]" captionStart="FIGURES 2–8" captionStartId="4.[151,264,1934,1959]" captionTargetBox="[217,1387,166,1915]" captionTargetId="figure@4.[199,1387,166,1915]" captionTargetPageId="4" captionText="FIGURES 2–8. Cratera obsidiana sp. nov., holotype in transverse (2–6) and sagittal sections with the anterior end to the left (7, 8): (2) cephalic region; (3) detail of body-margin in pre-pharyngeal region; (4) pre-pharyngeal region; (5) detail of the dorsal surface in the pre-pharyngeal region; (6) detail of the ventral surface in the pre-pharyngeal region; (7) pharynx; (8) ovary." httpUri="https://zenodo.org/record/3386955/files/figure.png" pageId="2" pageNumber="537">Fig. 3</figureCitation>
).
</paragraph>
<caption id="DF52661CFF8EFFE6FF7EFCCEFAECFCA6" box="[151,1316,858,883]" httpUri="https://zenodo.org/record/3386953/files/figure.png" pageId="2" pageNumber="537" startId="2.[151,250,858,883]" targetBox="[162,1423,526,831]" targetPageId="2">
<paragraph id="8B923694FF8EFFE6FF7EFCCEFAECFCA6" blockId="2.[151,1437,152,883]" box="[151,1316,858,883]" pageId="2" pageNumber="537">
<emphasis id="B959EA86FF8EFFE6FF7EFCCEFEDBFCA6" bold="true" box="[151,275,858,883]" pageId="2" pageNumber="537">FIGURE 1.</emphasis>
<taxonomicName id="4C2D4D17FF8EFFE6FEF0FCCFFE1CFCA6" authority="Boll &amp; Amaral &amp; Leal-Zanchet, 2019" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[281,468,859,883]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="2" pageNumber="537" phylum="Platyhelminthes" rank="species" species="obsidiana" status="sp. nov.">
<emphasis id="B959EA86FF8EFFE6FEF0FCCFFE1CFCA6" box="[281,468,859,883]" italics="true" pageId="2" pageNumber="537">Cratera obsidiana</emphasis>
</taxonomicName>
<emphasis id="B959EA86FF8EFFE6FE33FCCFFDE3FCA6" bold="true" box="[474,555,859,883]" pageId="2" pageNumber="537">
<taxonomicNameLabel id="A26A57FDFF8EFFE6FE33FCCFFDE3FCA6" box="[474,555,859,883]" pageId="2" pageNumber="537" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: dorsal aspect and eye pattern of fixed specimen (paratype MZU PL. 272).
</paragraph>
</caption>
<caption id="DF52661CFF8EFFE6FF7EFC37FDDBFBF2" ID-Table-UUID="DF52661CFF8EFFE6FF7EFC37FDDBFBF2" httpUri="http://table.plazi.org/id/DF52661CFF8EFFE6FF7EFC37FDDBFBF2" pageId="2" pageNumber="537" targetBox="[166,1406,1074,1965]" targetIsTable="true" targetPageId="2">
<paragraph id="8B923694FF8EFFE6FF7EFC37FDDBFBF2" blockId="2.[151,1437,931,1063]" pageId="2" pageNumber="537">
<emphasis id="B959EA86FF8EFFE6FF7EFC37FEC4FC6E" bold="true" box="[151,268,931,955]" pageId="2" pageNumber="537">TABLE 1.</emphasis>
Measurements, in mm, of specimens of
<taxonomicName id="4C2D4D17FF8EFFE6FD20FC37FC47FC6E" authority="Boll &amp; Amaral &amp; Leal-Zanchet, 2019" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[713,911,931,955]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="2" pageNumber="537" phylum="Platyhelminthes" rank="species" species="obsidiana" status="sp. nov.">
<emphasis id="B959EA86FF8EFFE6FD20FC37FC47FC6E" box="[713,911,931,955]" italics="true" pageId="2" pageNumber="537">Cratera obsidiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A57FDFF8EFFE6FC7EFC37FC38FC6F" box="[919,1008,931,955]" pageId="2" pageNumber="537" rank="species">
<emphasis id="B959EA86FF8EFFE6FC7EFC37FC20FC6E" bold="true" box="[919,1000,931,955]" pageId="2" pageNumber="537">sp. nov</emphasis>
.
</taxonomicNameLabel>
–: not measured; *: after fixation; DG: distance of gonopore from anterior end; DM: distance of mouth from anterior end; DMG: distance between mouth and gonopore; DPVP: distance between prostatic vesicle and pharyngeal pouch. The numbers given in parentheses represent the position relative to body length.
</paragraph>
</caption>
<paragraph id="8B923694FF8EFFE6FD97FBA7FAC2F878" pageId="2" pageNumber="537">
<table id="F92DC434FF8E001BFF4FFBA6FAB6F878" box="[166,1406,1074,1965]" gridcols="5" gridrows="20" pageId="2" pageNumber="537">
<tr id="351D34D6FF8E001BFF4FFBA6FAB6FB9E" box="[166,1406,1074,1099]" gridrow="0" pageId="2" pageNumber="537" rowspan-0="1">
<th id="76CC5DAAFF8E001BFD97FBA6FCF6FB9E" box="[638,830,1074,1099]" gridcol="1" gridrow="0" pageId="2" pageNumber="537">Holotype MZUSP</th>
<th id="76CC5DAAFF8E001BFC8AFBA6FC3FFB9E" box="[867,1015,1074,1099]" gridcol="2" gridrow="0" pageId="2" pageNumber="537">Paratype</th>
<th id="76CC5DAAFF8E001BFBCFFBA6FB73FB9E" box="[1062,1211,1074,1099]" gridcol="3" gridrow="0" pageId="2" pageNumber="537">Paratype</th>
<th id="76CC5DAAFF8E001BFB03FBA6FAB6FB9E" box="[1258,1406,1074,1099]" gridcol="4" gridrow="0" pageId="2" pageNumber="537">
<emphasis id="B959EA86FF8EFFE6FB03FBA7FA84FB9E" bold="true" box="[1258,1356,1075,1099]" pageId="2" pageNumber="537">Paratype</emphasis>
</th>
</tr>
<tr id="351D34D6FF8E001BFF4FFBC2FAB6FBBA" box="[166,1406,1110,1135]" gridrow="1" pageId="2" pageNumber="537" rowspan-0="1">
<td id="76CC5DAAFF8E001BFD97FBC2FCF6FBBA" box="[638,830,1110,1135]" gridcol="1" gridrow="1" pageId="2" pageNumber="537">PL.2158</td>
<td id="76CC5DAAFF8E001BFC8AFBC2FC3FFBBA" box="[867,1015,1110,1135]" gridcol="2" gridrow="1" pageId="2" pageNumber="537">MZU PL. 270</td>
<td id="76CC5DAAFF8E001BFBCFFBC2FB73FBBA" box="[1062,1211,1110,1135]" gridcol="3" gridrow="1" pageId="2" pageNumber="537">MZU PL. 271</td>
<td id="76CC5DAAFF8E001BFB03FBC2FAB6FBBA" box="[1258,1406,1110,1135]" gridcol="4" gridrow="1" pageId="2" pageNumber="537">MZU PL. 272</td>
</tr>
<tr id="351D34D6FF8E001BFF4FFBE3FAB6FB45" box="[166,1406,1143,1168]" gridrow="2" pageId="2" pageNumber="537">
<th id="76CC5DAAFF8E001BFF4FFBE3FE1EFB45" box="[166,470,1143,1168]" gridcol="0" gridrow="2" pageId="2" pageNumber="537">Maximum length in extension</th>
<td id="76CC5DAAFF8E001BFD97FBE3FCF6FB45" box="[638,830,1143,1168]" gridcol="1" gridrow="2" pageId="2" pageNumber="537">35</td>
<td id="76CC5DAAFF8E001BFC8AFBE3FC3FFB45" box="[867,1015,1143,1168]" gridcol="2" gridrow="2" pageId="2" pageNumber="537">35</td>
<td id="76CC5DAAFF8E001BFBCFFBE3FB73FB45" box="[1062,1211,1143,1168]" gridcol="3" gridrow="2" pageId="2" pageNumber="537">40</td>
<td id="76CC5DAAFF8E001BFB03FBE3FAB6FB45" box="[1258,1406,1143,1168]" gridcol="4" gridrow="2" pageId="2" pageNumber="537">40</td>
</tr>
<tr id="351D34D6FF8E001BFF4FFB09FAB6FB63" box="[166,1406,1181,1206]" gridrow="3" pageId="2" pageNumber="537">
<th id="76CC5DAAFF8E001BFF4FFB09FE1EFB63" box="[166,470,1181,1206]" gridcol="0" gridrow="3" pageId="2" pageNumber="537">Maximum width in extension</th>
<td id="76CC5DAAFF8E001BFD97FB09FCF6FB63" box="[638,830,1181,1206]" gridcol="1" gridrow="3" pageId="2" pageNumber="537">4</td>
<td id="76CC5DAAFF8E001BFC8AFB09FC3FFB63" box="[867,1015,1181,1206]" gridcol="2" gridrow="3" pageId="2" pageNumber="537">3</td>
<td id="76CC5DAAFF8E001BFBCFFB09FB73FB63" box="[1062,1211,1181,1206]" gridcol="3" gridrow="3" pageId="2" pageNumber="537">3</td>
<td id="76CC5DAAFF8E001BFB03FB09FAB6FB63" box="[1258,1406,1181,1206]" gridcol="4" gridrow="3" pageId="2" pageNumber="537">2</td>
</tr>
<tr id="351D34D6FF8E001BFF4FFB56FAB6FB0E" box="[166,1406,1218,1243]" gridrow="4" pageId="2" pageNumber="537">
<th id="76CC5DAAFF8E001BFF4FFB56FE1EFB0E" box="[166,470,1218,1243]" gridcol="0" gridrow="4" pageId="2" pageNumber="537">Length at rest</th>
<td id="76CC5DAAFF8E001BFD97FB56FCF6FB0E" box="[638,830,1218,1243]" gridcol="1" gridrow="4" pageId="2" pageNumber="537">20</td>
<td id="76CC5DAAFF8E001BFC8AFB56FC3FFB0E" box="[867,1015,1218,1243]" gridcol="2" gridrow="4" pageId="2" pageNumber="537">30</td>
<td id="76CC5DAAFF8E001BFBCFFB56FB73FB0E" box="[1062,1211,1218,1243]" gridcol="3" gridrow="4" pageId="2" pageNumber="537">30</td>
<td id="76CC5DAAFF8E001BFB03FB56FAB6FB0E" box="[1258,1406,1218,1243]" gridcol="4" gridrow="4" pageId="2" pageNumber="537">20</td>
</tr>
<tr id="351D34D6FF8E001BFF4FFB7CFAB6FAD4" box="[166,1406,1256,1281]" gridrow="5" pageId="2" pageNumber="537">
<th id="76CC5DAAFF8E001BFF4FFB7CFE1EFAD4" box="[166,470,1256,1281]" gridcol="0" gridrow="5" pageId="2" pageNumber="537">Width at rest</th>
<td id="76CC5DAAFF8E001BFD97FB7CFCF6FAD4" box="[638,830,1256,1281]" gridcol="1" gridrow="5" pageId="2" pageNumber="537">5</td>
<td id="76CC5DAAFF8E001BFC8AFB7CFC3FFAD4" box="[867,1015,1256,1281]" gridcol="2" gridrow="5" pageId="2" pageNumber="537">4</td>
<td id="76CC5DAAFF8E001BFBCFFB7CFB73FAD4" box="[1062,1211,1256,1281]" gridcol="3" gridrow="5" pageId="2" pageNumber="537">4</td>
<td id="76CC5DAAFF8E001BFB03FB7CFAB6FAD4" box="[1258,1406,1256,1281]" gridcol="4" gridrow="5" pageId="2" pageNumber="537">3</td>
</tr>
<tr id="351D34D6FF8E001BFF4FFA9AFAB6FAF2" box="[166,1406,1294,1319]" gridrow="6" pageId="2" pageNumber="537">
<th id="76CC5DAAFF8E001BFF4FFA9AFE1EFAF2" box="[166,470,1294,1319]" gridcol="0" gridrow="6" pageId="2" pageNumber="537">Length*</th>
<td id="76CC5DAAFF8E001BFD97FA9AFCF6FAF2" box="[638,830,1294,1319]" gridcol="1" gridrow="6" pageId="2" pageNumber="537">34</td>
<td id="76CC5DAAFF8E001BFC8AFA9AFC3FFAF2" box="[867,1015,1294,1319]" gridcol="2" gridrow="6" pageId="2" pageNumber="537">38</td>
<td id="76CC5DAAFF8E001BFBCFFA9AFB73FAF2" box="[1062,1211,1294,1319]" gridcol="3" gridrow="6" pageId="2" pageNumber="537">36</td>
<td id="76CC5DAAFF8E001BFB03FA9AFAB6FAF2" box="[1258,1406,1294,1319]" gridcol="4" gridrow="6" pageId="2" pageNumber="537">26</td>
</tr>
<tr id="351D34D6FF8E001BFF4FFAA0FAB6FA98" box="[166,1406,1332,1357]" gridrow="7" pageId="2" pageNumber="537">
<th id="76CC5DAAFF8E001BFF4FFAA0FE1EFA98" box="[166,470,1332,1357]" gridcol="0" gridrow="7" pageId="2" pageNumber="537">Width*</th>
<td id="76CC5DAAFF8E001BFD97FAA0FCF6FA98" box="[638,830,1332,1357]" gridcol="1" gridrow="7" pageId="2" pageNumber="537">5</td>
<td id="76CC5DAAFF8E001BFC8AFAA0FC3FFA98" box="[867,1015,1332,1357]" gridcol="2" gridrow="7" pageId="2" pageNumber="537">3</td>
<td id="76CC5DAAFF8E001BFBCFFAA0FB73FA98" box="[1062,1211,1332,1357]" gridcol="3" gridrow="7" pageId="2" pageNumber="537">5</td>
<td id="76CC5DAAFF8E001BFB03FAA0FAB6FA98" box="[1258,1406,1332,1357]" gridcol="4" gridrow="7" pageId="2" pageNumber="537">3</td>
</tr>
<tr id="351D34D6FF8E001BFF4FFACEFAB6FAA7" box="[166,1406,1370,1394]" gridrow="8" pageId="2" pageNumber="537">
<th id="76CC5DAAFF8E001BFF4FFACEFE1EFAA7" box="[166,470,1370,1394]" gridcol="0" gridrow="8" pageId="2" pageNumber="537">DM*</th>
<td id="76CC5DAAFF8E001BFD97FACEFCF6FAA7" box="[638,830,1370,1394]" gridcol="1" gridrow="8" pageId="2" pageNumber="537">24</td>
<td id="76CC5DAAFF8E001BFC8AFACEFC3FFAA7" box="[867,1015,1370,1394]" gridcol="2" gridrow="8" pageId="2" pageNumber="537">30</td>
<td id="76CC5DAAFF8E001BFBCFFACEFB73FAA7" box="[1062,1211,1370,1394]" gridcol="3" gridrow="8" pageId="2" pageNumber="537">27</td>
<td id="76CC5DAAFF8E001BFB03FACEFAB6FAA7" box="[1258,1406,1370,1394]" gridcol="4" gridrow="8" pageId="2" pageNumber="537">19</td>
</tr>
<tr id="351D34D6FF8E001BFF4FFAEAFAB6FA62" box="[166,1406,1406,1463]" gridrow="9" pageId="2" pageNumber="537">
<th id="76CC5DAAFF8E001BFF4FFAEAFE1EFA62" box="[166,470,1406,1463]" gridcol="0" gridrow="9" pageId="2" pageNumber="537">DG*</th>
<td id="76CC5DAAFF8E001BFD97FAEAFCF6FA62" box="[638,830,1406,1463]" gridcol="1" gridrow="9" pageId="2" pageNumber="537">(70.5) 29</td>
<td id="76CC5DAAFF8E001BFC8AFAEAFC3FFA62" box="[867,1015,1406,1463]" gridcol="2" gridrow="9" pageId="2" pageNumber="537">(79) 33</td>
<td id="76CC5DAAFF8E001BFBCFFAEAFB73FA62" box="[1062,1211,1406,1463]" gridcol="3" gridrow="9" pageId="2" pageNumber="537">(75) 32</td>
<td id="76CC5DAAFF8E001BFB03FAEAFAB6FA62" box="[1258,1406,1406,1463]" gridcol="4" gridrow="9" pageId="2" pageNumber="537">23</td>
</tr>
<tr id="351D34D6FF8E001BFF4FFA56FAB6FA29" box="[166,1406,1474,1532]" gridrow="10" pageId="2" pageNumber="537">
<th id="76CC5DAAFF8E001BFF4FFA56FE1EFA29" box="[166,470,1474,1532]" gridcol="0" gridrow="10" pageId="2" pageNumber="537">DMG*</th>
<td id="76CC5DAAFF8E001BFD97FA56FCF6FA29" box="[638,830,1474,1532]" gridcol="1" gridrow="10" pageId="2" pageNumber="537">(85) 5</td>
<td id="76CC5DAAFF8E001BFC8AFA56FC3FFA29" box="[867,1015,1474,1532]" gridcol="2" gridrow="10" pageId="2" pageNumber="537">(87) 3</td>
<td id="76CC5DAAFF8E001BFBCFFA56FB73FA29" box="[1062,1211,1474,1532]" gridcol="3" gridrow="10" pageId="2" pageNumber="537">(89) 5</td>
<td id="76CC5DAAFF8E001BFB03FA56FAB6FA29" box="[1258,1406,1474,1532]" gridcol="4" gridrow="10" pageId="2" pageNumber="537">4</td>
</tr>
<tr id="351D34D6FF8E001BFF4FF99DFAB6F9F7" box="[166,1406,1545,1570]" gridrow="11" pageId="2" pageNumber="537">
<th id="76CC5DAAFF8E001BFF4FF99DFE1EF9F7" box="[166,470,1545,1570]" gridcol="0" gridrow="11" pageId="2" pageNumber="537">DPVP*</th>
<td id="76CC5DAAFF8E001BFD97F99DFCF6F9F7" box="[638,830,1545,1570]" gridcol="1" gridrow="11" pageId="2" pageNumber="537">2.6</td>
<td id="76CC5DAAFF8E001BFC8AF99DFC3FF9F7" box="[867,1015,1545,1570]" gridcol="2" gridrow="11" pageId="2" pageNumber="537">–</td>
<td id="76CC5DAAFF8E001BFBCFF99DFB73F9F7" box="[1062,1211,1545,1570]" gridcol="3" gridrow="11" pageId="2" pageNumber="537">2.4</td>
<td id="76CC5DAAFF8E001BFB03F99DFAB6F9F7" box="[1258,1406,1545,1570]" gridcol="4" gridrow="11" pageId="2" pageNumber="537">–</td>
</tr>
<tr id="351D34D6FF8E001BFF4FF9BBFAB6F99D" box="[166,1406,1583,1608]" gridrow="12" pageId="2" pageNumber="537">
<th id="76CC5DAAFF8E001BFF4FF9BBFE1EF99D" box="[166,470,1583,1608]" gridcol="0" gridrow="12" pageId="2" pageNumber="537">Creeping sole %</th>
<td id="76CC5DAAFF8E001BFD97F9BBFCF6F99D" box="[638,830,1583,1608]" gridcol="1" gridrow="12" pageId="2" pageNumber="537">100</td>
<td id="76CC5DAAFF8E001BFC8AF9BBFC3FF99D" box="[867,1015,1583,1608]" gridcol="2" gridrow="12" pageId="2" pageNumber="537">–</td>
<td id="76CC5DAAFF8E001BFBCFF9BBFB73F99D" box="[1062,1211,1583,1608]" gridcol="3" gridrow="12" pageId="2" pageNumber="537">100</td>
<td id="76CC5DAAFF8E001BFB03F9BBFAB6F99D" box="[1258,1406,1583,1608]" gridcol="4" gridrow="12" pageId="2" pageNumber="537">–</td>
</tr>
<tr id="351D34D6FF8E001BFF4FF9C0FAB6F9B8" box="[166,1406,1620,1645]" gridrow="13" pageId="2" pageNumber="537">
<th id="76CC5DAAFF8E001BFF4FF9C0FE1EF9B8" box="[166,470,1620,1645]" gridcol="0" gridrow="13" pageId="2" pageNumber="537">Ovaries</th>
<td id="76CC5DAAFF8E001BFD97F9C0FCF6F9B8" box="[638,830,1620,1645]" gridcol="1" gridrow="13" pageId="2" pageNumber="537">9</td>
<td id="76CC5DAAFF8E001BFC8AF9C0FC3FF9B8" box="[867,1015,1620,1645]" gridcol="2" gridrow="13" pageId="2" pageNumber="537">–</td>
<td id="76CC5DAAFF8E001BFBCFF9C0FB73F9B8" box="[1062,1211,1620,1645]" gridcol="3" gridrow="13" pageId="2" pageNumber="537">9</td>
<td id="76CC5DAAFF8E001BFB03F9C0FAB6F9B8" box="[1258,1406,1620,1645]" gridcol="4" gridrow="13" pageId="2" pageNumber="537">–</td>
</tr>
<tr id="351D34D6FF8E001BFF4FF9ECFAB6F967" box="[166,1406,1656,1714]" gridrow="14" pageId="2" pageNumber="537">
<th id="76CC5DAAFF8E001BFF4FF9ECFE1EF967" box="[166,470,1656,1714]" gridcol="0" gridrow="14" pageId="2" pageNumber="537">Anteriormost testes</th>
<td id="76CC5DAAFF8E001BFD97F9ECFCF6F967" box="[638,830,1656,1714]" gridcol="1" gridrow="14" pageId="2" pageNumber="537">(26) 9</td>
<td id="76CC5DAAFF8E001BFC8AF9ECFC3FF967" box="[867,1015,1656,1714]" gridcol="2" gridrow="14" pageId="2" pageNumber="537">–</td>
<td id="76CC5DAAFF8E001BFBCFF9ECFB73F967" box="[1062,1211,1656,1714]" gridcol="3" gridrow="14" pageId="2" pageNumber="537">(25) 10.5</td>
<td id="76CC5DAAFF8E001BFB03F9ECFAB6F967" box="[1258,1406,1656,1714]" gridcol="4" gridrow="14" pageId="2" pageNumber="537">–</td>
</tr>
<tr id="351D34D6FF8E001BFF4FF929FAB6F922" box="[166,1406,1725,1783]" gridrow="15" pageId="2" pageNumber="537">
<th id="76CC5DAAFF8E001BFF4FF929FE1EF922" box="[166,470,1725,1783]" gridcol="0" gridrow="15" pageId="2" pageNumber="537">Posteriormost testes</th>
<td id="76CC5DAAFF8E001BFD97F929FCF6F922" box="[638,830,1725,1783]" gridcol="1" gridrow="15" pageId="2" pageNumber="537">(26) 21</td>
<td id="76CC5DAAFF8E001BFC8AF929FC3FF922" box="[867,1015,1725,1783]" gridcol="2" gridrow="15" pageId="2" pageNumber="537">–</td>
<td id="76CC5DAAFF8E001BFBCFF929FB73F922" box="[1062,1211,1725,1783]" gridcol="3" gridrow="15" pageId="2" pageNumber="537">(29) 19</td>
<td id="76CC5DAAFF8E001BFB03F929FAB6F922" box="[1258,1406,1725,1783]" gridcol="4" gridrow="15" pageId="2" pageNumber="537">–</td>
</tr>
<tr id="351D34D6FF8E001BFF4FF896FAB6F8EE" box="[166,1406,1794,1851]" gridrow="16" pageId="2" pageNumber="537">
<th id="76CC5DAAFF8E001BFF4FF896FE1EF8EE" box="[166,470,1794,1851]" gridcol="0" gridrow="16" pageId="2" pageNumber="537">Prostatic vesicle</th>
<td id="76CC5DAAFF8E001BFD97F896FCF6F8EE" box="[638,830,1794,1851]" gridcol="1" gridrow="16" pageId="2" pageNumber="537">(62) 0.3</td>
<td id="76CC5DAAFF8E001BFC8AF896FC3FF8EE" box="[867,1015,1794,1851]" gridcol="2" gridrow="16" pageId="2" pageNumber="537">0.2</td>
<td id="76CC5DAAFF8E001BFBCFF896FB73F8EE" box="[1062,1211,1794,1851]" gridcol="3" gridrow="16" pageId="2" pageNumber="537">(53) 0.3</td>
<td id="76CC5DAAFF8E001BFB03F896FAB6F8EE" box="[1258,1406,1794,1851]" gridcol="4" gridrow="16" pageId="2" pageNumber="537">0.15</td>
</tr>
<tr id="351D34D6FF8E001BFF4FF8DCFAB6F8B4" box="[166,1406,1864,1889]" gridrow="17" pageId="2" pageNumber="537">
<th id="76CC5DAAFF8E001BFF4FF8DCFE1EF8B4" box="[166,470,1864,1889]" gridcol="0" gridrow="17" pageId="2" pageNumber="537">Penis papilla</th>
<td id="76CC5DAAFF8E001BFD97F8DCFCF6F8B4" box="[638,830,1864,1889]" gridcol="1" gridrow="17" pageId="2" pageNumber="537">0.7</td>
<td id="76CC5DAAFF8E001BFC8AF8DCFC3FF8B4" box="[867,1015,1864,1889]" gridcol="2" gridrow="17" pageId="2" pageNumber="537">0.4</td>
<td id="76CC5DAAFF8E001BFBCFF8DCFB73F8B4" box="[1062,1211,1864,1889]" gridcol="3" gridrow="17" pageId="2" pageNumber="537">0.5</td>
<td id="76CC5DAAFF8E001BFB03F8DCFAB6F8B4" box="[1258,1406,1864,1889]" gridcol="4" gridrow="17" pageId="2" pageNumber="537">0.6</td>
</tr>
<tr id="351D34D6FF8E001BFF4FF8FAFAB6F852" box="[166,1406,1902,1927]" gridrow="18" pageId="2" pageNumber="537">
<th id="76CC5DAAFF8E001BFF4FF8FAFE1EF852" box="[166,470,1902,1927]" gridcol="0" gridrow="18" pageId="2" pageNumber="537">Male atrium</th>
<td id="76CC5DAAFF8E001BFD97F8FAFCF6F852" box="[638,830,1902,1927]" gridcol="1" gridrow="18" pageId="2" pageNumber="537">0.7</td>
<td id="76CC5DAAFF8E001BFC8AF8FAFC3FF852" box="[867,1015,1902,1927]" gridcol="2" gridrow="18" pageId="2" pageNumber="537">0.5</td>
<td id="76CC5DAAFF8E001BFBCFF8FAFB73F852" box="[1062,1211,1902,1927]" gridcol="3" gridrow="18" pageId="2" pageNumber="537">0.7</td>
<td id="76CC5DAAFF8E001BFB03F8FAFAB6F852" box="[1258,1406,1902,1927]" gridcol="4" gridrow="18" pageId="2" pageNumber="537">0.5</td>
</tr>
<tr id="351D34D6FF8E001BFF4FF800FAB6F878" box="[166,1406,1940,1965]" gridrow="19" pageId="2" pageNumber="537">
<th id="76CC5DAAFF8E001BFF4FF800FE1EF878" box="[166,470,1940,1965]" gridcol="0" gridrow="19" pageId="2" pageNumber="537">Female atrium</th>
<td id="76CC5DAAFF8E001BFD97F800FCF6F878" box="[638,830,1940,1965]" gridcol="1" gridrow="19" pageId="2" pageNumber="537">0.4</td>
<td id="76CC5DAAFF8E001BFC8AF800FC3FF878" box="[867,1015,1940,1965]" gridcol="2" gridrow="19" pageId="2" pageNumber="537">0.3</td>
<td id="76CC5DAAFF8E001BFBCFF800FB73F878" box="[1062,1211,1940,1965]" gridcol="3" gridrow="19" pageId="2" pageNumber="537">0.4</td>
<td id="76CC5DAAFF8E001BFB03F800FAB6F878" box="[1258,1406,1940,1965]" gridcol="4" gridrow="19" pageId="2" pageNumber="537">0.2</td>
</tr>
</table>
</paragraph>
<paragraph id="8B923694FF8FFFE7FF2EFF0DFB51FE96" blockId="3.[151,1437,153,431]" pageId="3" pageNumber="538">
Cutaneous musculature with the usual three layers: circular, oblique and longitudinal layers, the latter with thick bundles (
<figureCitation id="13162A11FF8FFFE7FF15FF29FEADFF02" box="[252,357,188,215]" captionStart="FIGURES 2–8" captionStartId="4.[151,264,1934,1959]" captionTargetBox="[217,1387,166,1915]" captionTargetId="figure@4.[199,1387,166,1915]" captionTargetPageId="4" captionText="FIGURES 2–8. Cratera obsidiana sp. nov., holotype in transverse (2–6) and sagittal sections with the anterior end to the left (7, 8): (2) cephalic region; (3) detail of body-margin in pre-pharyngeal region; (4) pre-pharyngeal region; (5) detail of the dorsal surface in the pre-pharyngeal region; (6) detail of the ventral surface in the pre-pharyngeal region; (7) pharynx; (8) ovary." httpUri="https://zenodo.org/record/3386955/files/figure.png" pageId="3" pageNumber="538">Figs. 5, 6</figureCitation>
,
<tableCitation id="C6AF032FFF8FFFE7FE99FF29FE0CFF02" box="[368,452,189,215]" captionStart="TABLE 2" captionStartId="3.[151,239,478,502]" captionTargetPageId="3" captionText="TABLE 2. Body height and cutaneous musculature in the median region of a transversal section of the pre-pharyngeal region, in micrometers, and ratio of the height of cutaneous musculature to the height of the body (mc:h index) of specimens of Cratera obsidiana sp. nov." httpUri="http://table.plazi.org/id/DF52661CFF8FFFE7FF7EFE4AFDDCFDEA" pageId="3" pageNumber="538" tableUuid="DF52661CFF8FFFE7FF7EFE4AFDDCFDEA">Table 2</tableCitation>
). Cutaneous musculature thinner in the pre-pharyngeal region than in the cephalic region of the body, but gradually diminishing its thickness towards anterior tip. Ventral musculature with similar thickness or slightly thinner than dorsal musculature at the sagittal plane (
<tableCitation id="C6AF032FFF8FFFE7FCB6FE91FC7DFECA" box="[863,949,261,287]" captionStart="TABLE 2" captionStartId="3.[151,239,478,502]" captionTargetPageId="3" captionText="TABLE 2. Body height and cutaneous musculature in the median region of a transversal section of the pre-pharyngeal region, in micrometers, and ratio of the height of cutaneous musculature to the height of the body (mc:h index) of specimens of Cratera obsidiana sp. nov." httpUri="http://table.plazi.org/id/DF52661CFF8FFFE7FF7EFE4AFDDCFDEA" pageId="3" pageNumber="538" tableUuid="DF52661CFF8FFFE7FF7EFE4AFDDCFDEA">Table 2</tableCitation>
) between four and seven times thicker than the epidermis, becoming progressively thinner toward body margins. Mc:h 7–8% (
<tableCitation id="C6AF032FFF8FFFE7FBDDFEBDFB43FE96" box="[1076,1163,297,323]" captionStart="TABLE 2" captionStartId="3.[151,239,478,502]" captionTargetPageId="3" captionText="TABLE 2. Body height and cutaneous musculature in the median region of a transversal section of the pre-pharyngeal region, in micrometers, and ratio of the height of cutaneous musculature to the height of the body (mc:h index) of specimens of Cratera obsidiana sp. nov." httpUri="http://table.plazi.org/id/DF52661CFF8FFFE7FF7EFE4AFDDCFDEA" pageId="3" pageNumber="538" tableUuid="DF52661CFF8FFFE7FF7EFE4AFDDCFDEA">Table 2</tableCitation>
).
</paragraph>
<paragraph id="8B923694FF8FFFE7FF2EFED9FF2EFE7A" blockId="3.[151,1437,153,431]" pageId="3" pageNumber="538">
Mesenchymal musculature (
<figureCitation id="13162A11FF8FFFE7FDEDFED9FD8FFEB2" box="[516,583,333,359]" captionStart="FIGURES 2–8" captionStartId="4.[151,264,1934,1959]" captionTargetBox="[217,1387,166,1915]" captionTargetId="figure@4.[199,1387,166,1915]" captionTargetPageId="4" captionText="FIGURES 2–8. Cratera obsidiana sp. nov., holotype in transverse (2–6) and sagittal sections with the anterior end to the left (7, 8): (2) cephalic region; (3) detail of body-margin in pre-pharyngeal region; (4) pre-pharyngeal region; (5) detail of the dorsal surface in the pre-pharyngeal region; (6) detail of the ventral surface in the pre-pharyngeal region; (7) pharynx; (8) ovary." httpUri="https://zenodo.org/record/3386955/files/figure.png" pageId="3" pageNumber="538">Fig. 4</figureCitation>
) well developed, with a similar arrangement to that described for other species of the genus. At the cephalic region, the mesenchymal musculature is less developed than in the pre-pharyngeal region.
</paragraph>
<caption id="DF52661CFF8FFFE7FF7EFE4AFDDCFDEA" ID-Table-UUID="DF52661CFF8FFFE7FF7EFE4AFDDCFDEA" httpUri="http://table.plazi.org/id/DF52661CFF8FFFE7FF7EFE4AFDDCFDEA" pageId="3" pageNumber="538" targetBox="[166,1163,587,869]" targetIsTable="true" targetPageId="3">
<paragraph id="8B923694FF8FFFE7FF7EFE4AFDDCFDEA" blockId="3.[151,1436,478,575]" pageId="3" pageNumber="538">
<emphasis id="B959EA86FF8FFFE7FF7EFE4AFEC3FE23" bold="true" box="[151,267,478,502]" pageId="3" pageNumber="538">TABLE 2.</emphasis>
Body height and cutaneous musculature in the median region of a transversal section of the pre-pharyngeal region, in micrometers, and ratio of the height of cutaneous musculature to the height of the body (mc:h index) of specimens of
<taxonomicName id="4C2D4D17FF8FFFE7FF1AFDB3FE7FFDEA" authority="Boll &amp; Amaral &amp; Leal-Zanchet, 2019" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[243,439,551,575]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="3" pageNumber="538" phylum="Platyhelminthes" rank="species" species="obsidiana" status="sp. nov.">
<emphasis id="B959EA86FF8FFFE7FF1AFDB3FE7FFDEA" box="[243,439,551,575]" italics="true" pageId="3" pageNumber="538">Cratera obsidiana</emphasis>
</taxonomicName>
<emphasis id="B959EA86FF8FFFE7FE57FDB3FDDCFDEA" bold="true" box="[446,532,551,575]" pageId="3" pageNumber="538">
<taxonomicNameLabel id="A26A57FDFF8FFFE7FE57FDB3FDDCFDEA" box="[446,532,551,575]" pageId="3" pageNumber="538" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</caption>
<paragraph id="8B923694FF8FFFE7FDF4FDDFFBCBFCB1" pageId="3" pageNumber="538">
<table id="F92DC434FF8F001BFF4FFDDFFB43FCB0" box="[166,1163,587,869]" gridcols="3" gridrows="8" pageId="3" pageNumber="538">
<tr id="351D34D6FF8F001BFF4FFDDFFB43FDB6" box="[166,1163,587,611]" gridrow="0" pageId="3" pageNumber="538" rowspan-0="1">
<th id="76CC5DAAFF8F001BFDF4FDDFFD1DFDB6" box="[541,725,587,611]" gridcol="1" gridrow="0" pageId="3" pageNumber="538">Holotype</th>
<th id="76CC5DAAFF8F001BFC1FFDDFFB43FDB6" box="[1014,1163,587,611]" gridcol="2" gridrow="0" pageId="3" pageNumber="538">
<emphasis id="B959EA86FF8FFFE7FC1FFDDFFB90FDB6" bold="true" box="[1014,1112,587,611]" pageId="3" pageNumber="538">Paratype</emphasis>
</th>
</tr>
<tr id="351D34D6FF8F001BFF4FFDFAFB43FD52" box="[166,1163,622,647]" gridrow="1" pageId="3" pageNumber="538" rowspan-0="1">
<td id="76CC5DAAFF8F001BFDF4FDFAFD1DFD52" box="[541,725,622,647]" gridcol="1" gridrow="1" pageId="3" pageNumber="538">MZUSP PL.2158</td>
<td id="76CC5DAAFF8F001BFC1FFDFAFB43FD52" box="[1014,1163,622,647]" gridcol="2" gridrow="1" pageId="3" pageNumber="538">MZU PL. 270</td>
</tr>
<tr id="351D34D6FF8F001BFF4FFD1BFB43FD7D" box="[166,1163,655,680]" gridrow="2" pageId="3" pageNumber="538">
<th id="76CC5DAAFF8F001BFF4FFD1BFEB9FD7D" box="[166,369,655,680]" gridcol="0" gridrow="2" pageId="3" pageNumber="538">Dorsal epidermis</th>
<td id="76CC5DAAFF8F001BFDF4FD1BFD1DFD7D" box="[541,725,655,680]" gridcol="1" gridrow="2" pageId="3" pageNumber="538">9</td>
<td id="76CC5DAAFF8F001BFC1FFD1BFB43FD7D" box="[1014,1163,655,680]" gridcol="2" gridrow="2" pageId="3" pageNumber="538">12</td>
</tr>
<tr id="351D34D6FF8F001BFF4FFD21FB43FD1B" box="[166,1163,693,718]" gridrow="3" pageId="3" pageNumber="538">
<th id="76CC5DAAFF8F001BFF4FFD21FEB9FD1B" box="[166,369,693,718]" gridcol="0" gridrow="3" pageId="3" pageNumber="538">Ventral epidermis</th>
<td id="76CC5DAAFF8F001BFDF4FD21FD1DFD1B" box="[541,725,693,718]" gridcol="1" gridrow="3" pageId="3" pageNumber="538">9</td>
<td id="76CC5DAAFF8F001BFC1FFD21FB43FD1B" box="[1014,1163,693,718]" gridcol="2" gridrow="3" pageId="3" pageNumber="538">12</td>
</tr>
<tr id="351D34D6FF8F001BFF4FFD4FFB43FD21" box="[166,1163,731,756]" gridrow="4" pageId="3" pageNumber="538">
<th id="76CC5DAAFF8F001BFF4FFD4FFEB9FD21" box="[166,369,731,756]" gridcol="0" gridrow="4" pageId="3" pageNumber="538">Dorsal musculature</th>
<td id="76CC5DAAFF8F001BFDF4FD4FFD1DFD21" box="[541,725,731,756]" gridcol="1" gridrow="4" pageId="3" pageNumber="538">63</td>
<td id="76CC5DAAFF8F001BFC1FFD4FFB43FD21" box="[1014,1163,731,756]" gridcol="2" gridrow="4" pageId="3" pageNumber="538">45</td>
</tr>
<tr id="351D34D6FF8F001BFF4FFC94FB43FCCC" box="[166,1163,768,793]" gridrow="5" pageId="3" pageNumber="538">
<th id="76CC5DAAFF8F001BFF4FFC94FEB9FCCC" box="[166,369,768,793]" gridcol="0" gridrow="5" pageId="3" pageNumber="538">Ventral musculature</th>
<td id="76CC5DAAFF8F001BFDF4FC94FD1DFCCC" box="[541,725,768,793]" gridcol="1" gridrow="5" pageId="3" pageNumber="538">55</td>
<td id="76CC5DAAFF8F001BFC1FFC94FB43FCCC" box="[1014,1163,768,793]" gridcol="2" gridrow="5" pageId="3" pageNumber="538">45</td>
</tr>
<tr id="351D34D6FF8F001BFF4FFCB2FB43FCEA" box="[166,1163,806,831]" gridrow="6" pageId="3" pageNumber="538">
<th id="76CC5DAAFF8F001BFF4FFCB2FEB9FCEA" box="[166,369,806,831]" gridcol="0" gridrow="6" pageId="3" pageNumber="538">Body height</th>
<td id="76CC5DAAFF8F001BFDF4FCB2FD1DFCEA" box="[541,725,806,831]" gridcol="1" gridrow="6" pageId="3" pageNumber="538">875</td>
<td id="76CC5DAAFF8F001BFC1FFCB2FB43FCEA" box="[1014,1163,806,831]" gridcol="2" gridrow="6" pageId="3" pageNumber="538">1225</td>
</tr>
<tr id="351D34D6FF8F001BFF4FFCD8FB43FCB0" box="[166,1163,844,869]" gridrow="7" pageId="3" pageNumber="538">
<th id="76CC5DAAFF8F001BFF4FFCD8FEB9FCB0" box="[166,369,844,869]" gridcol="0" gridrow="7" pageId="3" pageNumber="538">Mc:h (%)</th>
<td id="76CC5DAAFF8F001BFDF4FCD8FD1DFCB0" box="[541,725,844,869]" gridcol="1" gridrow="7" pageId="3" pageNumber="538">8</td>
<td id="76CC5DAAFF8F001BFC1FFCD8FB43FCB0" box="[1014,1163,844,869]" gridcol="2" gridrow="7" pageId="3" pageNumber="538">7</td>
</tr>
</table>
</paragraph>
<paragraph id="8B923694FF8FFFE7FF2EFC02FAF3FC01" blockId="3.[151,1437,918,2024]" pageId="3" pageNumber="538">
<emphasis id="B959EA86FF8FFFE7FF2EFC02FEFDFC65" bold="true" box="[199,309,918,944]" pageId="3" pageNumber="538">Pharynx.</emphasis>
The pharynx is cylindrical, about 6% of body length, with folded walls and dorsal insertion posteriorly shifted. The mouth is located in the median third of the pharyngeal pouch (
<figureCitation id="13162A11FF8FFFE7FBFBFC2EFB9EFC01" box="[1042,1110,954,980]" captionStart="FIGURES 2–8" captionStartId="4.[151,264,1934,1959]" captionTargetBox="[217,1387,166,1915]" captionTargetId="figure@4.[199,1387,166,1915]" captionTargetPageId="4" captionText="FIGURES 2–8. Cratera obsidiana sp. nov., holotype in transverse (2–6) and sagittal sections with the anterior end to the left (7, 8): (2) cephalic region; (3) detail of body-margin in pre-pharyngeal region; (4) pre-pharyngeal region; (5) detail of the dorsal surface in the pre-pharyngeal region; (6) detail of the ventral surface in the pre-pharyngeal region; (7) pharynx; (8) ovary." httpUri="https://zenodo.org/record/3386955/files/figure.png" pageId="3" pageNumber="538">Fig. 7</figureCitation>
). Esophagus absent.
</paragraph>
<paragraph id="8B923694FF8FFFE7FF2EFC49FA48FA51" blockId="3.[151,1437,918,2024]" pageId="3" pageNumber="538">
<emphasis id="B959EA86FF8FFFE7FF2EFC49FE02FC2D" bold="true" box="[199,458,989,1016]" pageId="3" pageNumber="538">Reproductive organs.</emphasis>
The testes are arranged in one irregular row on either side of the body, beneath the dorsal transverse mesenchymal muscles (
<figureCitation id="13162A11FF8FFFE7FD8CFB96FD18FBC9" box="[613,720,1025,1052]" captionStart="FIGURES 2–8" captionStartId="4.[151,264,1934,1959]" captionTargetBox="[217,1387,166,1915]" captionTargetId="figure@4.[199,1387,166,1915]" captionTargetPageId="4" captionText="FIGURES 2–8. Cratera obsidiana sp. nov., holotype in transverse (2–6) and sagittal sections with the anterior end to the left (7, 8): (2) cephalic region; (3) detail of body-margin in pre-pharyngeal region; (4) pre-pharyngeal region; (5) detail of the dorsal surface in the pre-pharyngeal region; (6) detail of the ventral surface in the pre-pharyngeal region; (7) pharynx; (8) ovary." httpUri="https://zenodo.org/record/3386955/files/figure.png" pageId="3" pageNumber="538">Figs. 4, 5</figureCitation>
). They begin in the anterior third of the body and extend to near the root of the pharynx. Sperm ducts dorsal to ovovitelline ducts in pre-pharyngeal region (
<figureCitation id="13162A11FF8FFFE7FB71FBB2FB15FB94" box="[1176,1245,1062,1089]" captionStart="FIGURES 2–8" captionStartId="4.[151,264,1934,1959]" captionTargetBox="[217,1387,166,1915]" captionTargetId="figure@4.[199,1387,166,1915]" captionTargetPageId="4" captionText="FIGURES 2–8. Cratera obsidiana sp. nov., holotype in transverse (2–6) and sagittal sections with the anterior end to the left (7, 8): (2) cephalic region; (3) detail of body-margin in pre-pharyngeal region; (4) pre-pharyngeal region; (5) detail of the dorsal surface in the pre-pharyngeal region; (6) detail of the ventral surface in the pre-pharyngeal region; (7) pharynx; (8) ovary." httpUri="https://zenodo.org/record/3386955/files/figure.png" pageId="3" pageNumber="538">Fig. 6</figureCitation>
). Distally, sperm ducts—forming spermiducal vesicles—enter laterally into the proximal part of the prostatic vesicle (
<figureCitation id="13162A11FF8FFFE7FAF8FBDEFA47FBB1" box="[1297,1423,1098,1124]" captionStart="FIGURES 9–10" captionStartId="5.[151,264,1964,1989]" captionTargetBox="[176,1436,151,1944]" captionTargetId="figure@5.[151,1436,151,1944]" captionTargetPageId="5" captionText="FIGURES 9–10. Cratera obsidiana sp. nov., composite reconstruction of the copulatory apparatus: (9) holotype in sagittal sections; (10) paratype MZU PL. 271 in horizontal sections. Anterior end to the left." httpUri="https://zenodo.org/record/3386957/files/figure.png" pageId="3" pageNumber="538">Figs. 9, 10</figureCitation>
). Extrabulbar prostatic vesicle, tubular and unpaired, located near the common muscle coat. The proximal portion is laterally expanded (T-shaped) and slightly dorsally oriented (
<figureCitation id="13162A11FF8FFFE7FCA6FB06FC04FB79" box="[847,972,1170,1196]" captionStart-0="FIGURES 9–10" captionStart-1="FIGURES 11–12" captionStartId-0="5.[151,264,1964,1989]" captionStartId-1="6.[151,264,1964,1989]" captionTargetBox-0="[176,1436,151,1944]" captionTargetBox-1="[196,1392,188,1945]" captionTargetId-0="figure@5.[151,1436,151,1944]" captionTargetId-1="figure@6.[196,1392,166,1945]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 9–10. Cratera obsidiana sp. nov., composite reconstruction of the copulatory apparatus: (9) holotype in sagittal sections; (10) paratype MZU PL. 271 in horizontal sections. Anterior end to the left." captionText-1="FIGURES 11–12. Cratera obsidiana sp. nov.: (11) copulatory apparatus of the holotype in sagittal section; (12) copulatory apparatus of the paratype MZU PL. 271 in horizontal section." httpUri-0="https://zenodo.org/record/3386957/files/figure.png" httpUri-1="https://zenodo.org/record/3386959/files/figure.png" pageId="3" pageNumber="538">Figs. 9–12</figureCitation>
); it is located at about the same distance from the ventral and dorsal epidermis. Its distal portion penetrates the common muscle coat, becoming sinuous, and continues inside the penis papilla as an ejaculatory duct. This duct is almost straight and opens through an expansion at the tip of the penis papilla (
<figureCitation id="13162A11FF8FFFE7FDCEFB6AFDBFFACD" box="[551,631,1278,1304]" captionStart="FIGURES 9–10" captionStartId="5.[151,264,1964,1989]" captionTargetBox="[176,1436,151,1944]" captionTargetId="figure@5.[151,1436,151,1944]" captionTargetPageId="5" captionText="FIGURES 9–10. Cratera obsidiana sp. nov., composite reconstruction of the copulatory apparatus: (9) holotype in sagittal sections; (10) paratype MZU PL. 271 in horizontal sections. Anterior end to the left." httpUri="https://zenodo.org/record/3386957/files/figure.png" pageId="3" pageNumber="538">Figs. 9</figureCitation>
,
<figureCitation id="13162A11FF8FFFE7FD6CFB6AFD6AFACD" box="[645,674,1278,1304]" captionStart="FIGURES 11–12" captionStartId="6.[151,264,1964,1989]" captionTargetBox="[196,1392,188,1945]" captionTargetId="figure@6.[196,1392,166,1945]" captionTargetPageId="6" captionText="FIGURES 11–12. Cratera obsidiana sp. nov.: (11) copulatory apparatus of the holotype in sagittal section; (12) copulatory apparatus of the paratype MZU PL. 271 in horizontal section." httpUri="https://zenodo.org/record/3386959/files/figure.png" pageId="3" pageNumber="538">11</figureCitation>
). The male atrium is almost unfolded and occupied by the conical, almost symmetric penis papilla, which has a dorsal fold. The dorsal wall of the male atrium receives the openings of abundant cyanophil glands (
<figureCitation id="13162A11FF8FFFE7FE04FAD2FDAFFAB5" box="[493,615,1350,1376]" captionStart-0="FIGURES 9–10" captionStart-1="FIGURES 11–12" captionStartId-0="5.[151,264,1964,1989]" captionStartId-1="6.[151,264,1964,1989]" captionTargetBox-0="[176,1436,151,1944]" captionTargetBox-1="[196,1392,188,1945]" captionTargetId-0="figure@5.[151,1436,151,1944]" captionTargetId-1="figure@6.[196,1392,166,1945]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 9–10. Cratera obsidiana sp. nov., composite reconstruction of the copulatory apparatus: (9) holotype in sagittal sections; (10) paratype MZU PL. 271 in horizontal sections. Anterior end to the left." captionText-1="FIGURES 11–12. Cratera obsidiana sp. nov.: (11) copulatory apparatus of the holotype in sagittal section; (12) copulatory apparatus of the paratype MZU PL. 271 in horizontal section." httpUri-0="https://zenodo.org/record/3386957/files/figure.png" httpUri-1="https://zenodo.org/record/3386959/files/figure.png" pageId="3" pageNumber="538">Figs. 9–12</figureCitation>
). Histological aspects of the epithelium, musculature and secretions of the prostatic vesicle, penis papilla, ejaculatory duct and male atrium are similar to those of other species of
<taxonomicName id="4C2D4D17FF8FFFE7FAC9FAFEFAB1FA51" authorityName="Carbayo, Alvarez-Presas, Olivares, Marques, Froehlich &amp; Riutort" authorityYear="2013" box="[1312,1401,1386,1412]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="3" pageNumber="538" phylum="Platyhelminthes" rank="genus">
<emphasis id="B959EA86FF8FFFE7FAC9FAFEFAB1FA51" box="[1312,1401,1386,1412]" italics="true" pageId="3" pageNumber="538">Cratera</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B923694FF8FFFE7FF2EFA1AFE66F939" blockId="3.[151,1437,918,2024]" pageId="3" pageNumber="538">
Vitelline follicles, situated between intestinal branches, well-developed in all studied specimens (
<figureCitation id="13162A11FF8FFFE7FAEEFA1AFA46FA7D" box="[1287,1422,1422,1448]" captionStart="FIGURES 2–8" captionStartId="4.[151,264,1934,1959]" captionTargetBox="[217,1387,166,1915]" captionTargetId="figure@4.[199,1387,166,1915]" captionTargetPageId="4" captionText="FIGURES 2–8. Cratera obsidiana sp. nov., holotype in transverse (2–6) and sagittal sections with the anterior end to the left (7, 8): (2) cephalic region; (3) detail of body-margin in pre-pharyngeal region; (4) pre-pharyngeal region; (5) detail of the dorsal surface in the pre-pharyngeal region; (6) detail of the ventral surface in the pre-pharyngeal region; (7) pharynx; (8) ovary." httpUri="https://zenodo.org/record/3386955/files/figure.png" pageId="3" pageNumber="538">Figs. 4, 6, 8</figureCitation>
). Ovaries ovoid, about twice longer than wide, measuring 0.3 mm in their antero-posterior axis. They are located dorsally to the ventral nerve plate at about the same transversal level as the anteriormost testes, in the anterior third of the body (
<tableCitation id="C6AF032FFF8FFFE7FEEEFA6EFE93F9C1" box="[263,347,1530,1556]" captionStart="TABLE 1" captionStartId="2.[151,239,931,955]" captionTargetPageId="2" captionText="TABLE 1. Measurements, in mm, of specimens of Cratera obsidiana sp. nov. –: not measured; *: after fixation; DG: distance of gonopore from anterior end; DM: distance of mouth from anterior end; DMG: distance between mouth and gonopore; DPVP: distance between prostatic vesicle and pharyngeal pouch. The numbers given in parentheses represent the position relative to body length." httpUri="http://table.plazi.org/id/DF52661CFF8EFFE6FF7EFC37FDDBFBF2" pageId="3" pageNumber="538" tableUuid="DF52661CFF8EFFE6FF7EFC37FDDBFBF2">Table 1</tableCitation>
,
<figureCitation id="13162A11FF8FFFE7FE80FA6EFE64F9C1" box="[361,428,1530,1556]" captionStart="FIGURES 2–8" captionStartId="4.[151,264,1934,1959]" captionTargetBox="[217,1387,166,1915]" captionTargetId="figure@4.[199,1387,166,1915]" captionTargetPageId="4" captionText="FIGURES 2–8. Cratera obsidiana sp. nov., holotype in transverse (2–6) and sagittal sections with the anterior end to the left (7, 8): (2) cephalic region; (3) detail of body-margin in pre-pharyngeal region; (4) pre-pharyngeal region; (5) detail of the dorsal surface in the pre-pharyngeal region; (6) detail of the ventral surface in the pre-pharyngeal region; (7) pharynx; (8) ovary." httpUri="https://zenodo.org/record/3386955/files/figure.png" pageId="3" pageNumber="538">Fig. 8</figureCitation>
). Ovovitelline ducts emerge dorsally from the median third of the ovaries and run posteriorly immediately above the nerve plate. Laterally to the female atrium, the ovovitelline ducts ascend dorsomedially and unite dorsally to the proximal third of the female canal to form the common glandular ovovitelline duct (
<figureCitation id="13162A11FF8FFFE7FA8DF9D6FF1AF955" captionStart-0="FIGURES 9–10" captionStart-1="FIGURES 11–12" captionStartId-0="5.[151,264,1964,1989]" captionStartId-1="6.[151,264,1964,1989]" captionTargetBox-0="[176,1436,151,1944]" captionTargetBox-1="[196,1392,188,1945]" captionTargetId-0="figure@5.[151,1436,151,1944]" captionTargetId-1="figure@6.[196,1392,166,1945]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 9–10. Cratera obsidiana sp. nov., composite reconstruction of the copulatory apparatus: (9) holotype in sagittal sections; (10) paratype MZU PL. 271 in horizontal sections. Anterior end to the left." captionText-1="FIGURES 11–12. Cratera obsidiana sp. nov.: (11) copulatory apparatus of the holotype in sagittal section; (12) copulatory apparatus of the paratype MZU PL. 271 in horizontal section." httpUri-0="https://zenodo.org/record/3386957/files/figure.png" httpUri-1="https://zenodo.org/record/3386959/files/figure.png" pageId="3" pageNumber="538">Figs. 9–11</figureCitation>
). The female genital duct is dorso-anteriorly curved (
<figureCitation id="13162A11FF8FFFE7FCDDF9F2FC79F955" box="[820,945,1638,1664]" captionStart-0="FIGURES 9–10" captionStart-1="FIGURES 11–12" captionStartId-0="5.[151,264,1964,1989]" captionStartId-1="6.[151,264,1964,1989]" captionTargetBox-0="[176,1436,151,1944]" captionTargetBox-1="[196,1392,188,1945]" captionTargetId-0="figure@5.[151,1436,151,1944]" captionTargetId-1="figure@6.[196,1392,166,1945]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 9–10. Cratera obsidiana sp. nov., composite reconstruction of the copulatory apparatus: (9) holotype in sagittal sections; (10) paratype MZU PL. 271 in horizontal sections. Anterior end to the left." captionText-1="FIGURES 11–12. Cratera obsidiana sp. nov.: (11) copulatory apparatus of the holotype in sagittal section; (12) copulatory apparatus of the paratype MZU PL. 271 in horizontal section." httpUri-0="https://zenodo.org/record/3386957/files/figure.png" httpUri-1="https://zenodo.org/record/3386959/files/figure.png" pageId="3" pageNumber="538">Figs. 9–12</figureCitation>
). The female atrium is funnel-shaped, with small lateral folds, and has about half the length of the male atrium (
<tableCitation id="C6AF032FFF8FFFE7FC60F91EFC15F971" box="[905,989,1674,1700]" captionStart="TABLE 1" captionStartId="2.[151,239,931,955]" captionTargetPageId="2" captionText="TABLE 1. Measurements, in mm, of specimens of Cratera obsidiana sp. nov. –: not measured; *: after fixation; DG: distance of gonopore from anterior end; DM: distance of mouth from anterior end; DMG: distance between mouth and gonopore; DPVP: distance between prostatic vesicle and pharyngeal pouch. The numbers given in parentheses represent the position relative to body length." httpUri="http://table.plazi.org/id/DF52661CFF8EFFE6FF7EFC37FDDBFBF2" pageId="3" pageNumber="538" tableUuid="DF52661CFF8EFFE6FF7EFC37FDDBFBF2">Table 1</tableCitation>
,
<figureCitation id="13162A11FF8FFFE7FC03F91EFBACF971" box="[1002,1124,1674,1700]" captionStart-0="FIGURES 9–10" captionStart-1="FIGURES 11–12" captionStartId-0="5.[151,264,1964,1989]" captionStartId-1="6.[151,264,1964,1989]" captionTargetBox-0="[176,1436,151,1944]" captionTargetBox-1="[196,1392,188,1945]" captionTargetId-0="figure@5.[151,1436,151,1944]" captionTargetId-1="figure@6.[196,1392,166,1945]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 9–10. Cratera obsidiana sp. nov., composite reconstruction of the copulatory apparatus: (9) holotype in sagittal sections; (10) paratype MZU PL. 271 in horizontal sections. Anterior end to the left." captionText-1="FIGURES 11–12. Cratera obsidiana sp. nov.: (11) copulatory apparatus of the holotype in sagittal section; (12) copulatory apparatus of the paratype MZU PL. 271 in horizontal section." httpUri-0="https://zenodo.org/record/3386957/files/figure.png" httpUri-1="https://zenodo.org/record/3386959/files/figure.png" pageId="3" pageNumber="538">Figs. 9–12</figureCitation>
). Histological aspects of the epithelium, musculature and secretions of the ovovitelline ducts and female canal and atrium are similar to those of other species of
<taxonomicName id="4C2D4D17FF8FFFE7FEA7F946FE6FF939" authorityName="Carbayo, Alvarez-Presas, Olivares, Marques, Froehlich &amp; Riutort" authorityYear="2013" box="[334,423,1746,1772]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="3" pageNumber="538" phylum="Platyhelminthes" rank="genus">
<emphasis id="B959EA86FF8FFFE7FEA7F946FE6FF939" box="[334,423,1746,1772]" italics="true" pageId="3" pageNumber="538">Cratera</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B923694FF8FFFE7FF2EF962FE71F8A9" blockId="3.[151,1437,918,2024]" pageId="3" pageNumber="538">
Gonopore canal vertical at the sagittal plane. Male and female atria with ample communication, without separating folds (
<figureCitation id="13162A11FF8FFFE7FECEF88EFE6AF8E1" box="[295,418,1818,1844]" captionStart-0="FIGURES 9–10" captionStart-1="FIGURES 11–12" captionStartId-0="5.[151,264,1964,1989]" captionStartId-1="6.[151,264,1964,1989]" captionTargetBox-0="[176,1436,151,1944]" captionTargetBox-1="[196,1392,188,1945]" captionTargetId-0="figure@5.[151,1436,151,1944]" captionTargetId-1="figure@6.[196,1392,166,1945]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 9–10. Cratera obsidiana sp. nov., composite reconstruction of the copulatory apparatus: (9) holotype in sagittal sections; (10) paratype MZU PL. 271 in horizontal sections. Anterior end to the left." captionText-1="FIGURES 11–12. Cratera obsidiana sp. nov.: (11) copulatory apparatus of the holotype in sagittal section; (12) copulatory apparatus of the paratype MZU PL. 271 in horizontal section." httpUri-0="https://zenodo.org/record/3386957/files/figure.png" httpUri-1="https://zenodo.org/record/3386959/files/figure.png" pageId="3" pageNumber="538">Figs. 9–12</figureCitation>
), showing well developed, continuous muscle coats with longitudinal, oblique and circular fibers (
<figureCitation id="13162A11FF8FFFE7FF0CF8AAFEA2F88D" box="[229,362,1854,1880]" captionStart="FIGURES 11–12" captionStartId="6.[151,264,1964,1989]" captionTargetBox="[196,1392,188,1945]" captionTargetId="figure@6.[196,1392,166,1945]" captionTargetPageId="6" captionText="FIGURES 11–12. Cratera obsidiana sp. nov.: (11) copulatory apparatus of the holotype in sagittal section; (12) copulatory apparatus of the paratype MZU PL. 271 in horizontal section." httpUri="https://zenodo.org/record/3386959/files/figure.png" pageId="3" pageNumber="538">Figs. 11, 12</figureCitation>
). The common muscle coat is thicker entally and along the dorsal wall of the male atrium, thinner around the female atrium.
</paragraph>
<paragraph id="8B923694FF8FFFE7FF2EF811FB27F83D" blockId="3.[151,1437,918,2024]" pageId="3" pageNumber="538">
<emphasis id="B959EA86FF8FFFE7FF2EF811FE26F875" bold="true" box="[199,494,1925,1952]" pageId="3" pageNumber="538">Comparative discussion.</emphasis>
The presence of an expanded ejaculatory cavity at the tip of the penis papilla and cyanophil glands discharging through the roof of the male atrium, as well as other characters, such as dorsal monolobated eyes, support the inclusion of
<taxonomicName id="4C2D4D17FF8FFFE7FDA8F85AFCC6F83D" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[577,782,1998,2024]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="3" pageNumber="538" phylum="Platyhelminthes" rank="species" species="obsidiana">
<emphasis id="B959EA86FF8FFFE7FDA8F85AFCC6F83D" box="[577,782,1998,2024]" italics="true" pageId="3" pageNumber="538">Cratera obsidiana</emphasis>
</taxonomicName>
in the genus
<taxonomicName id="4C2D4D17FF8FFFE7FC4DF85AFB23F83D" authority="Carbayo et al., 2013" authorityName="Carbayo" authorityYear="2013" box="[932,1259,1998,2024]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="3" pageNumber="538" phylum="Platyhelminthes" rank="genus">
<emphasis id="B959EA86FF8FFFE7FC4DF85AFC35F83D" box="[932,1021,1998,2024]" italics="true" pageId="3" pageNumber="538">Cratera</emphasis>
<bibRefCitation id="EFBC4B65FF8FFFE7FBEDF85AFB23F83D" author="Carbayo, F. &amp; Alvarez-Presas, M. &amp; Olivares, C. T. &amp; Marques, F. P. L. &amp; Froehlich, E. M. &amp; Riutort, M." box="[1028,1259,1998,2024]" pageId="3" pageNumber="538" pagination="508 - 528" refId="ref10548" refString="Carbayo, F., Alvarez-Presas, M., Olivares, C. T., Marques, F. P. L., Froehlich, E. M. &amp; Riutort, M. (2013) Molecular phylogeny of Geoplaninae (Platyhelminthes) challenges current classification: proposal of taxonomic actions. Zoologica Scripta, 42, 508 - 528. https: // doi. org / 10.1111 / zsc. 1201 9" type="journal article" year="2013">
Carbayo
<emphasis id="B959EA86FF8FFFE7FB83F85BFB53F83D" box="[1130,1179,1998,2024]" italics="true" pageId="3" pageNumber="538">et al</emphasis>
., 2013
</bibRefCitation>
</taxonomicName>
.
</paragraph>
<caption id="DF52661CFF88FFE0FF7EF81AFAA0F825" httpUri="https://zenodo.org/record/3386955/files/figure.png" pageId="4" pageNumber="539" startId="4.[151,264,1934,1959]" targetBox="[217,1387,166,1915]" targetPageId="4">
<paragraph id="8B923694FF88FFE0FF7EF81AFAA0F825" blockId="4.[151,1436,1934,2032]" pageId="4" pageNumber="539">
<emphasis id="B959EA86FF88FFE0FF7EF81AFEF3F872" bold="true" box="[151,315,1934,1959]" pageId="4" pageNumber="539">FIGURES 2–8.</emphasis>
<taxonomicName id="4C2D4D17FF88FFE0FEABF81BFE35F872" authority="Boll &amp; Amaral &amp; Leal-Zanchet, 2019" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[322,509,1935,1959]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="4" pageNumber="539" phylum="Platyhelminthes" rank="species" species="obsidiana" status="sp. nov.">
<emphasis id="B959EA86FF88FFE0FEABF81BFE35F872" box="[322,509,1935,1959]" italics="true" pageId="4" pageNumber="539">Cratera obsidiana</emphasis>
</taxonomicName>
<emphasis id="B959EA86FF88FFE0FDEDF81BFD9EF872" bold="true" box="[516,598,1935,1959]" pageId="4" pageNumber="539">
<taxonomicNameLabel id="A26A57FDFF88FFE0FDEDF81BFD9EF872" box="[516,598,1935,1959]" pageId="4" pageNumber="539" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype in transverse (2–6) and sagittal sections with the anterior end to the left (7, 8): (2) cephalic region; (3) detail of body-margin in pre-pharyngeal region; (4) pre-pharyngeal region; (5) detail of the dorsal surface in the pre-pharyngeal region; (6) detail of the ventral surface in the pre-pharyngeal region; (7) pharynx; (8) ovary.
</paragraph>
</caption>
<caption id="DF52661CFF89FFE1FF7EF838FC05F83C" httpUri="https://zenodo.org/record/3386957/files/figure.png" pageId="5" pageNumber="540" startId="5.[151,264,1964,1989]" targetBox="[176,1436,151,1944]" targetPageId="5">
<paragraph id="8B923694FF89FFE1FF7EF838FC05F83C" blockId="5.[151,1436,1964,2025]" pageId="5" pageNumber="540">
<emphasis id="B959EA86FF89FFE1FF7EF838FE8FF810" bold="true" box="[151,327,1964,1989]" pageId="5" pageNumber="540">FIGURES 9–10.</emphasis>
<taxonomicName id="4C2D4D17FF89FFE1FEA5F839FDCEF811" authority="Boll &amp; Amaral &amp; Leal-Zanchet, 2019" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[332,518,1964,1988]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="5" pageNumber="540" phylum="Platyhelminthes" rank="species" species="obsidiana" status="sp. nov.">
<emphasis id="B959EA86FF89FFE1FEA5F839FDCEF811" box="[332,518,1964,1988]" italics="true" pageId="5" pageNumber="540">Cratera obsidiana</emphasis>
</taxonomicName>
<emphasis id="B959EA86FF89FFE1FDE2F839FD93F810" bold="true" box="[523,603,1965,1989]" pageId="5" pageNumber="540">
<taxonomicNameLabel id="A26A57FDFF89FFE1FDE2F839FD93F810" box="[523,603,1965,1989]" pageId="5" pageNumber="540" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, composite reconstruction of the copulatory apparatus: (9) holotype in sagittal sections; (10) paratype MZU PL. 271 in horizontal sections. Anterior end to the left.
</paragraph>
</caption>
<caption id="DF52661CFF8AFFE2FF7EF838FCC0F83C" httpUri="https://zenodo.org/record/3386959/files/figure.png" pageId="6" pageNumber="541" startId="6.[151,264,1964,1989]" targetBox="[196,1392,188,1945]" targetPageId="6">
<paragraph id="8B923694FF8AFFE2FF7EF838FCC0F83C" blockId="6.[151,1436,1964,2026]" pageId="6" pageNumber="541">
<emphasis id="B959EA86FF8AFFE2FF7EF838FE86F810" bold="true" box="[151,334,1964,1989]" pageId="6" pageNumber="541">FIGURES 11–12</emphasis>
.
<taxonomicName id="4C2D4D17FF8AFFE2FEB4F83AFDD1F810" authority="Boll &amp; Amaral &amp; Leal-Zanchet, 2019" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[349,537,1965,1989]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="6" pageNumber="541" phylum="Platyhelminthes" rank="species" species="obsidiana" status="sp. nov.">
<emphasis id="B959EA86FF8AFFE2FEB4F83AFDD1F810" box="[349,537,1965,1989]" italics="true" pageId="6" pageNumber="541">Cratera obsidiana</emphasis>
</taxonomicName>
<emphasis id="B959EA86FF8AFFE2FDC8F839FDBCF810" bold="true" box="[545,628,1965,1989]" pageId="6" pageNumber="541">
<taxonomicNameLabel id="A26A57FDFF8AFFE2FDC8F839FDBCF810" box="[545,628,1965,1989]" pageId="6" pageNumber="541" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: (11) copulatory apparatus of the holotype in sagittal section; (12) copulatory apparatus of the paratype MZU PL. 271 in horizontal section.
</paragraph>
</caption>
<paragraph id="8B923694FF8BFFE3FF2EFF0DFB2BFE5E" blockId="7.[151,1437,152,971]" pageId="7" pageNumber="542">
The new species shares similarities with eleven species of the genus,
<taxonomicName id="4C2D4D17FF8BFFE3FC23FF0DFB19FF66" authority="(Marcus, 1951)" authorityName="Marcus" authorityYear="1951" baseAuthorityName="Marcus" baseAuthorityYear="1951" box="[970,1233,152,179]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="hina">
<emphasis id="B959EA86FF8BFFE3FC23FF0DFBD5FF66" box="[970,1053,153,179]" italics="true" pageId="7" pageNumber="542">C. hina</emphasis>
(Marcus, 1951)
</taxonomicName>
,
<taxonomicName id="4C2D4D17FF8BFFE3FB34FF0DFE9BFF02" authority="(E. M. Froehlich, 1955)" authorityName="E. M. Froehlich" authorityYear="1955" baseAuthorityName="E. M. Froehlich" baseAuthorityYear="1955" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="crioula">
<emphasis id="B959EA86FF8BFFE3FB34FF0DFA86FF66" box="[1245,1358,153,179]" italics="true" pageId="7" pageNumber="542">C. crioula</emphasis>
(E. M. Froehlich, 1955)
</taxonomicName>
,
<taxonomicName id="4C2D4D17FF8BFFE3FEB7FF29FDBEFF02" authority="(Froehlich, 1956)" authorityName="Froehlich" authorityYear="1956" baseAuthorityName="Froehlich" baseAuthorityYear="1956" box="[350,630,188,215]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="joia">
<emphasis id="B959EA86FF8BFFE3FEB7FF29FE62FF02" box="[350,426,189,215]" italics="true" pageId="7" pageNumber="542">C. joia</emphasis>
(Froehlich, 1956)
</taxonomicName>
,
<taxonomicName id="4C2D4D17FF8BFFE3FD68FF29FC0FFF02" authority="Carbayo, 2015" authorityName="Carbayo" authorityYear="2015" box="[641,967,188,215]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="anamariae">
<emphasis id="B959EA86FF8BFFE3FD68FF29FCD2FF02" box="[641,794,189,215]" italics="true" pageId="7" pageNumber="542">C. anamariae</emphasis>
<bibRefCitation id="EFBC4B65FF8BFFE3FCC9FF29FC0FFF02" author="Carbayo, F. &amp; Almeida, A. L." box="[800,967,188,215]" pageId="7" pageNumber="542" pagination="27 - 40" refId="ref10482" refString="Carbayo, F. &amp; Almeida, A. L. (2015) Anatomical deviation of male organs of land planarians from Rio de Janeiro, Brazil, with description of two new species of Cratera (Platyhelminthes, Tricladida). Zootaxa, 3931 (1), 27 - 40. https: // doi. org / 10.11646 / zootaxa. 3931.1.2" type="journal article" year="2015">Carbayo, 2015</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2D4D17FF8BFFE3FC3BFF29FB35FF02" authority="Rossi et al., 2015" authorityName="Rossi" authorityYear="2015" box="[978,1277,188,215]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="ochra">
<emphasis id="B959EA86FF8BFFE3FC3BFF29FBFBFF02" box="[978,1075,189,215]" italics="true" pageId="7" pageNumber="542">C. ochra</emphasis>
Rossi
<emphasis id="B959EA86FF8BFFE3FB97FF2AFB66FF02" box="[1150,1198,189,215]" italics="true" pageId="7" pageNumber="542">et al</emphasis>
., 2015
</taxonomicName>
,
<taxonomicName id="4C2D4D17FF8BFFE3FAE1FF29FE28FF2E" authority="Negrete &amp; Brusa, 2016" authorityName="Negrete &amp; Brusa" authorityYear="2016" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="viridimaculata">
<emphasis id="B959EA86FF8BFFE3FAE1FF29FF19FF2E" italics="true" pageId="7" pageNumber="542">C. viridimaculata</emphasis>
<bibRefCitation id="EFBC4B65FF8BFFE3FF31FF75FE28FF2E" author="Negrete, L. &amp; Brusa, F." box="[216,480,225,251]" pageId="7" pageNumber="542" pagination="1 - 12" refId="ref11443" refString="Negrete, L. &amp; Brusa, F. (2016) First report of the genus Cratera (Platyhelminthes, Geoplanidae) in Argentina, with description of a new species and comments on the species of the genus. ZooKeys, 610, 1 - 12. https: // doi. org / 10.3897 / zookeys. 610.946 5" type="journal article" year="2016">Negrete &amp; Brusa, 2016</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2D4D17FF8BFFE3FE05FF75FC23FF2E" authority="Rossi &amp; Leal-Zanchet, 2016" authorityName="Rossi &amp; Leal-Zanchet" authorityYear="2016" box="[492,1003,225,251]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="cryptolineata">
<emphasis id="B959EA86FF8BFFE3FE05FF75FD6AFF2E" box="[492,674,225,251]" italics="true" pageId="7" pageNumber="542">C. cryptolineata</emphasis>
<bibRefCitation id="EFBC4B65FF8BFFE3FD40FF75FC23FF2E" author="Rossi, I. &amp; Amaral, S. V. &amp; Ribeiro, G. G. &amp; Cauduro, G. P. &amp; Fick, I. &amp; Valiati, V. H. &amp; Leal-Zanchet, A. M." box="[681,1003,225,251]" pageId="7" pageNumber="542" pagination="787 - 815" refId="ref11524" refString="Rossi, I., Amaral, S. V., Ribeiro, G. G., Cauduro, G. P., Fick, I., Valiati, V. H. &amp; Leal-Zanchet, A. M. (2016) Two new Geoplaninae species (Platyhelminthes: Continenticola) from Southern Brazil based on an integrative taxonomic approach. Journal of Natural History, 50, 787 - 815. https: // doi. org / 10.1080 / 00222933.2015.108405 7" type="journal article" year="2016">Rossi &amp; Leal-Zanchet, 2016</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2D4D17FF8BFFE3FC1EFF75FEC6FECA" authority="Rossi &amp; Leal-Zanchet, 2016" authorityName="Rossi &amp; Leal-Zanchet" authorityYear="2016" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="nigrimarginata">
<emphasis id="B959EA86FF8BFFE3FC1EFF75FB0AFF2E" box="[1015,1218,225,251]" italics="true" pageId="7" pageNumber="542">C. nigrimarginata</emphasis>
<bibRefCitation id="EFBC4B65FF8BFFE3FB20FF75FEC6FECA" author="Rossi, I. &amp; Amaral, S. V. &amp; Ribeiro, G. G. &amp; Cauduro, G. P. &amp; Fick, I. &amp; Valiati, V. H. &amp; Leal-Zanchet, A. M." pageId="7" pageNumber="542" pagination="787 - 815" refId="ref11524" refString="Rossi, I., Amaral, S. V., Ribeiro, G. G., Cauduro, G. P., Fick, I., Valiati, V. H. &amp; Leal-Zanchet, A. M. (2016) Two new Geoplaninae species (Platyhelminthes: Continenticola) from Southern Brazil based on an integrative taxonomic approach. Journal of Natural History, 50, 787 - 815. https: // doi. org / 10.1080 / 00222933.2015.108405 7" type="journal article" year="2016">Rossi &amp; Leal-Zanchet, 2016</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2D4D17FF8BFFE3FEF0FE91FCEEFECA" authority="Rossi &amp; Leal-Zanchet, 2016" authorityName="Rossi &amp; Leal-Zanchet" authorityYear="2016" box="[281,806,261,287]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="aureomaculata">
<emphasis id="B959EA86FF8BFFE3FEF0FE91FE2AFECA" box="[281,482,261,287]" italics="true" pageId="7" pageNumber="542">C. aureomaculata</emphasis>
<bibRefCitation id="EFBC4B65FF8BFFE3FE01FE91FCEEFECA" author="Rossi, I. &amp; Amaral, S. V. &amp; Ribeiro, G. G. &amp; Cauduro, G. P. &amp; Fick, I. &amp; Valiati, V. H. &amp; Leal-Zanchet, A. M." box="[488,806,261,287]" pageId="7" pageNumber="542" pagination="787 - 815" refId="ref11524" refString="Rossi, I., Amaral, S. V., Ribeiro, G. G., Cauduro, G. P., Fick, I., Valiati, V. H. &amp; Leal-Zanchet, A. M. (2016) Two new Geoplaninae species (Platyhelminthes: Continenticola) from Southern Brazil based on an integrative taxonomic approach. Journal of Natural History, 50, 787 - 815. https: // doi. org / 10.1080 / 00222933.2015.108405 7" type="journal article" year="2016">Rossi &amp; Leal-Zanchet, 2016</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2D4D17FF8BFFE3FCD8FE91FB38FECA" authority="Lago-Barcia &amp; Carbayo, 2018" authorityName="Lago-Barcia &amp; Carbayo" authorityYear="2018" box="[817,1264,261,287]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="picuia">
<emphasis id="B959EA86FF8BFFE3FCD8FE91FC5FFECA" box="[817,919,261,287]" italics="true" pageId="7" pageNumber="542">C. picuia</emphasis>
<bibRefCitation id="EFBC4B65FF8BFFE3FC74FE91FB38FECA" author="Lago-Barcia, D. &amp; Carbayo, F." box="[925,1264,261,287]" pageId="7" pageNumber="542" pagination="517 - 542" refId="ref11110" refString="Lago-Barcia, D. &amp; Carbayo, F. (2018) Taxonomic revision of four species of the Neotropical land planarian genera Cratera and Geoplana (Platyhelminthes, Tricladida) with a description of two new species and an emendation of Cratera. Zootaxa, 4500, 517 - 542. https: // doi. org / 10.11646 / zootaxa. 4500.4. 3" type="journal article" year="2018">Lago-Barcia &amp; Carbayo, 2018</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C2D4D17FF8BFFE3FACCFE91FE24FE96" authority="Lago-Barcia &amp; Carbayo, 2018" authorityName="Lago-Barcia &amp; Carbayo" authorityYear="2018" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="arucuia">
<emphasis id="B959EA86FF8BFFE3FACCFE91FA54FECA" box="[1317,1436,261,287]" italics="true" pageId="7" pageNumber="542">C. arucuia</emphasis>
<bibRefCitation id="EFBC4B65FF8BFFE3FF7EFEBDFE24FE96" author="Lago-Barcia, D. &amp; Carbayo, F." box="[151,492,297,323]" pageId="7" pageNumber="542" pagination="517 - 542" refId="ref11110" refString="Lago-Barcia, D. &amp; Carbayo, F. (2018) Taxonomic revision of four species of the Neotropical land planarian genera Cratera and Geoplana (Platyhelminthes, Tricladida) with a description of two new species and an emendation of Cratera. Zootaxa, 4500, 517 - 542. https: // doi. org / 10.11646 / zootaxa. 4500.4. 3" type="journal article" year="2018">Lago-Barcia &amp; Carbayo, 2018</bibRefCitation>
</taxonomicName>
, showing eyes spreading over the dorsal surface and a cylindrical pharynx (
<bibRefCitation id="EFBC4B65FF8BFFE3FAC6FEBDFF17FEB2" author="Froehlich, C. G." pageId="7" pageNumber="542" pagination="195 - 279" refId="ref10802" refString="Froehlich, C. G. (1954 a) Sobre Morfologia e Taxonomia das Geoplanidae. Boletim da Faculdade de Filosofia, Ciencias e Letras da Universidade de Sao Paulo, Serie Zoologia, 19, 195 - 279. https: // doi. org / 10.11606 / issn. 2526 - 3382. bffclzoologia. 1954.12009 2" type="journal article" year="1954">Froehlich 1954a</bibRefCitation>
,
<bibRefCitation id="EFBC4B65FF8BFFE3FF00FED8FEFBFEB2" author="Froehlich, C. G." box="[233,307,332,359]" pageId="7" pageNumber="542" pagination="313 - 344" refId="ref10885" refString="Froehlich, C. G. (1956 b) Tricladida Terricola das regioes de Teresopolis e Ubatuba. Papeis Avulsos do Departamento de Zoologia Secretaria de Agricultura Sao Paulo, 12, 313 - 344." type="journal article" year="1956">1956b</bibRefCitation>
;
<bibRefCitation id="EFBC4B65FF8BFFE3FED7FED9FDA9FEB2" author="Carbayo, F. &amp; Almeida, A. L." box="[318,609,332,359]" pageId="7" pageNumber="542" pagination="27 - 40" refId="ref10482" refString="Carbayo, F. &amp; Almeida, A. L. (2015) Anatomical deviation of male organs of land planarians from Rio de Janeiro, Brazil, with description of two new species of Cratera (Platyhelminthes, Tricladida). Zootaxa, 3931 (1), 27 - 40. https: // doi. org / 10.11646 / zootaxa. 3931.1.2" type="journal article" year="2015">Carbayo &amp; Almeida 2015</bibRefCitation>
;
<bibRefCitation id="EFBC4B65FF8BFFE3FD84FED9FCA4FEB2" author="Negrete, L. &amp; Brusa, F." box="[621,876,333,359]" pageId="7" pageNumber="542" pagination="1 - 12" refId="ref11443" refString="Negrete, L. &amp; Brusa, F. (2016) First report of the genus Cratera (Platyhelminthes, Geoplanidae) in Argentina, with description of a new species and comments on the species of the genus. ZooKeys, 610, 1 - 12. https: // doi. org / 10.3897 / zookeys. 610.946 5" type="journal article" year="2016">Negrete &amp; Brusa 2016</bibRefCitation>
;
<bibRefCitation id="EFBC4B65FF8BFFE3FC9FFED9FBFDFEB2" author="Rossi, I. &amp; Amaral, S. V. &amp; Ribeiro, G. G. &amp; Cauduro, G. P. &amp; Fick, I. &amp; Valiati, V. H. &amp; Leal-Zanchet, A. M." box="[886,1077,333,359]" pageId="7" pageNumber="542" pagination="787 - 815" refId="ref11524" refString="Rossi, I., Amaral, S. V., Ribeiro, G. G., Cauduro, G. P., Fick, I., Valiati, V. H. &amp; Leal-Zanchet, A. M. (2016) Two new Geoplaninae species (Platyhelminthes: Continenticola) from Southern Brazil based on an integrative taxonomic approach. Journal of Natural History, 50, 787 - 815. https: // doi. org / 10.1080 / 00222933.2015.108405 7" type="journal article" year="2016">
Rossi
<emphasis id="B959EA86FF8BFFE3FC52FEDAFC3BFEB2" box="[955,1011,333,359]" italics="true" pageId="7" pageNumber="542">et al.</emphasis>
2016
</bibRefCitation>
;
<bibRefCitation id="EFBC4B65FF8BFFE3FBA9FED9FA59FEB2" author="Lago-Barcia, D. &amp; Carbayo, F." box="[1088,1425,333,359]" pageId="7" pageNumber="542" pagination="517 - 542" refId="ref11110" refString="Lago-Barcia, D. &amp; Carbayo, F. (2018) Taxonomic revision of four species of the Neotropical land planarian genera Cratera and Geoplana (Platyhelminthes, Tricladida) with a description of two new species and an emendation of Cratera. Zootaxa, 4500, 517 - 542. https: // doi. org / 10.11646 / zootaxa. 4500.4. 3" type="journal article" year="2018">Lago-Barcia &amp; Carbayo 2018</bibRefCitation>
). It differs from these species, however, by the color pattern and details of the copulatory apparatus.
</paragraph>
<paragraph id="8B923694FF8BFFE3FF2EFE01FA83FCC2" blockId="7.[151,1437,152,971]" pageId="7" pageNumber="542">
The color pattern of
<taxonomicName id="4C2D4D17FF8BFFE3FE42FE01FDF1FE7A" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[427,569,405,431]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="obsidiana">
<emphasis id="B959EA86FF8BFFE3FE42FE01FDF1FE7A" box="[427,569,405,431]" italics="true" pageId="7" pageNumber="542">C. obsidiana</emphasis>
</taxonomicName>
, homogeneously black, differs from that of
<taxonomicName id="4C2D4D17FF8BFFE3FBCAFE01FB5CFE7A" baseAuthorityName="E. M. Froehlich" baseAuthorityYear="1955" box="[1059,1172,405,431]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="crioula">
<emphasis id="B959EA86FF8BFFE3FBCAFE01FB5CFE7A" box="[1059,1172,405,431]" italics="true" pageId="7" pageNumber="542">C. crioula</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2D4D17FF8BFFE3FB20FE01FADDFE7A" baseAuthorityName="Froehlich" baseAuthorityYear="1956" box="[1225,1301,405,431]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="joia">
<emphasis id="B959EA86FF8BFFE3FB20FE01FADDFE7A" box="[1225,1301,405,431]" italics="true" pageId="7" pageNumber="542">C. joia</emphasis>
</taxonomicName>
as they also have a black dorsum, but with a median light stripe (
<bibRefCitation id="EFBC4B65FF8BFFE3FD10FE2DFC70FE06" author="Froehlich, E. M." box="[761,952,440,467]" pageId="7" pageNumber="542" pagination="289 - 339" refId="ref10979" refString="Froehlich, E. M. (1954 b) Sobre especies brasileiras do genero Geoplana. Boletim da Faculdade de Filosofia, Ciencias e Letras da Universidade de Sao Paulo, Serie Zoologia, 19, 289 - 339. https: // doi. org / 10.11606 / issn. 2526 - 3382. bffclzoologia. 1954.12009 6" type="journal article" year="1954">Froehlich 1954b</bibRefCitation>
,
<bibRefCitation id="EFBC4B65FF8BFFE3FC2CFE2CFBC6FE06" author="Froehlich, C. G." box="[965,1038,440,467]" pageId="7" pageNumber="542" pagination="173 - 191" refId="ref10862" refString="Froehlich, C. G. (1956 a) Planarias Terrestres do Parana. Dusenia, 7, 173 - 191." type="journal article" year="1956">1956a</bibRefCitation>
). It also can be distinguished from
<taxonomicName id="4C2D4D17FF8BFFE3FF7EFE49FF32FE22" box="[151,250,477,503]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="ochra">
<emphasis id="B959EA86FF8BFFE3FF7EFE49FF32FE22" box="[151,250,477,503]" italics="true" pageId="7" pageNumber="542">C. ochra</emphasis>
</taxonomicName>
, which shows a dorsal yellow-ochre ground color and dispersed grayish or grayish-brown pigmentation constituting two broad dorsal bands (
<bibRefCitation id="EFBC4B65FF8BFFE3FDD2FD95FCC9FDC9" author="Rossi, I. &amp; Amaral, S. V. &amp; Ribeiro, G. G. &amp; Cauduro, G. P. &amp; Fick, I. &amp; Valiati, V. H. &amp; Leal-Zanchet, A. M." box="[571,769,513,540]" pageId="7" pageNumber="542" pagination="787 - 815" refId="ref11524" refString="Rossi, I., Amaral, S. V., Ribeiro, G. G., Cauduro, G. P., Fick, I., Valiati, V. H. &amp; Leal-Zanchet, A. M. (2016) Two new Geoplaninae species (Platyhelminthes: Continenticola) from Southern Brazil based on an integrative taxonomic approach. Journal of Natural History, 50, 787 - 815. https: // doi. org / 10.1080 / 00222933.2015.108405 7" type="journal article" year="2016">
Rossi
<emphasis id="B959EA86FF8BFFE3FD6AFD96FD75FDCE" box="[643,701,513,539]" italics="true" pageId="7" pageNumber="542">et al.</emphasis>
2016
</bibRefCitation>
). The new species also can be distinguished from
<taxonomicName id="4C2D4D17FF8BFFE3FAD2FD95FEC3FDEA" authorityName="Rossi &amp; Leal-Zanchet" authorityYear="2016" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="nigrimarginata">
<emphasis id="B959EA86FF8BFFE3FAD2FD95FEC3FDEA" italics="true" pageId="7" pageNumber="542">C. nigrimarginata</emphasis>
</taxonomicName>
, which has a light-brownish dorsal color bordered by dark margins, and
<taxonomicName id="4C2D4D17FF8BFFE3FBDCFDB1FB35FDEA" authorityName="Rossi &amp; Leal-Zanchet" authorityYear="2016" box="[1077,1277,549,575]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="aureomaculata">
<emphasis id="B959EA86FF8BFFE3FBDCFDB1FB35FDEA" box="[1077,1277,549,575]" italics="true" pageId="7" pageNumber="542">C. aureomaculata</emphasis>
</taxonomicName>
, which shows a yellowish dorsal ground color covered by brownish pigmentation in the cephalic region and blackish pigmentation constituting irregular flecks over the rest of the dorsum (
<bibRefCitation id="EFBC4B65FF8BFFE3FCE7FDF9FB83FD52" author="Rossi, I. &amp; Leal-Zanchet, A." box="[782,1099,621,647]" pageId="7" pageNumber="542" pagination="1 - 32" refId="ref11681" refString="Rossi, I. &amp; Leal-Zanchet, A. (2017) Three new species of Cratera Carbayo et al., 2013 from Araucaria forests with a key to species of the genus (Platyhelminthes, Continenticola). ZooKeys, 643, 1 - 32. https: // doi. org / 10.3897 / zookeys. 643.1109 3" type="journal article" year="2017">Rossi &amp; Leal-Zanchet 2017</bibRefCitation>
). By having a homogeneously pigmented dorsum,
<taxonomicName id="4C2D4D17FF8BFFE3FE9FFD05FDCDFD7E" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[374,517,657,683]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="obsidiana">
<emphasis id="B959EA86FF8BFFE3FE9FFD05FDCDFD7E" box="[374,517,657,683]" italics="true" pageId="7" pageNumber="542">C. obsidiana</emphasis>
</taxonomicName>
can be distinguished from species with striped dorsal surface, such as
<taxonomicName id="4C2D4D17FF8BFFE3FAF5FD05FF71FD1A" authorityName="Carbayo" authorityYear="2015" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="anamariae">
<emphasis id="B959EA86FF8BFFE3FAF5FD05FF71FD1A" italics="true" pageId="7" pageNumber="542">C. anamariae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2D4D17FF8BFFE3FF21FD21FE46FD1A" authorityName="Negrete &amp; Brusa" authorityYear="2016" box="[200,398,693,719]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="viridimaculata">
<emphasis id="B959EA86FF8BFFE3FF21FD21FE46FD1A" box="[200,398,693,719]" italics="true" pageId="7" pageNumber="542">C. viridimaculata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2D4D17FF8BFFE3FE2CFD21FDB3FD1A" authorityName="Rossi &amp; Leal-Zanchet" authorityYear="2016" box="[453,635,693,719]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="cryptolineata">
<emphasis id="B959EA86FF8BFFE3FE2CFD21FDB3FD1A" box="[453,635,693,719]" italics="true" pageId="7" pageNumber="542">C. cryptolineata</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFBC4B65FF8BFFE3FD63FD21FC78FD1A" author="Carbayo, F. &amp; Almeida, A. L." box="[650,944,692,719]" pageId="7" pageNumber="542" pagination="27 - 40" refId="ref10482" refString="Carbayo, F. &amp; Almeida, A. L. (2015) Anatomical deviation of male organs of land planarians from Rio de Janeiro, Brazil, with description of two new species of Cratera (Platyhelminthes, Tricladida). Zootaxa, 3931 (1), 27 - 40. https: // doi. org / 10.11646 / zootaxa. 3931.1.2" type="journal article" year="2015">Carbayo &amp; Almeida 2015</bibRefCitation>
;
<bibRefCitation id="EFBC4B65FF8BFFE3FC54FD21FB76FD1A" author="Negrete, L. &amp; Brusa, F." box="[957,1214,693,719]" pageId="7" pageNumber="542" pagination="1 - 12" refId="ref11443" refString="Negrete, L. &amp; Brusa, F. (2016) First report of the genus Cratera (Platyhelminthes, Geoplanidae) in Argentina, with description of a new species and comments on the species of the genus. ZooKeys, 610, 1 - 12. https: // doi. org / 10.3897 / zookeys. 610.946 5" type="journal article" year="2016">Negrete &amp; Brusa 2016</bibRefCitation>
;
<bibRefCitation id="EFBC4B65FF8BFFE3FB23FD21FEC2FD26" author="Rossi, I. &amp; Leal-Zanchet, A." pageId="7" pageNumber="542" pagination="1 - 32" refId="ref11681" refString="Rossi, I. &amp; Leal-Zanchet, A. (2017) Three new species of Cratera Carbayo et al., 2013 from Araucaria forests with a key to species of the genus (Platyhelminthes, Continenticola). ZooKeys, 643, 1 - 32. https: // doi. org / 10.3897 / zookeys. 643.1109 3" type="journal article" year="2017">Rossi &amp; Leal-Zanchet 2017</bibRefCitation>
). The color pattern of
<taxonomicName id="4C2D4D17FF8BFFE3FDEBFD4DFD9DFD26" baseAuthorityName="Marcus" baseAuthorityYear="1951" box="[514,597,729,755]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="hina">
<emphasis id="B959EA86FF8BFFE3FDEBFD4DFD9DFD26" box="[514,597,729,755]" italics="true" pageId="7" pageNumber="542">C. hina</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2D4D17FF8BFFE3FD8AFD4DFD01FD26" authorityName="Lago-Barcia &amp; Carbayo" authorityYear="2018" box="[611,713,729,755]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="picuia">
<emphasis id="B959EA86FF8BFFE3FD8AFD4DFD01FD26" box="[611,713,729,755]" italics="true" pageId="7" pageNumber="542">C. picuia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2D4D17FF8BFFE3FCE9FD4DFCBFFD26" authorityName="Lago-Barcia &amp; Carbayo" authorityYear="2018" box="[768,887,729,755]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="arucuia">
<emphasis id="B959EA86FF8BFFE3FCE9FD4DFCBFFD26" box="[768,887,729,755]" italics="true" pageId="7" pageNumber="542">C. arucuia</emphasis>
</taxonomicName>
is distinguished from that of
<taxonomicName id="4C2D4D17FF8BFFE3FB28FD4DFA98FD26" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[1217,1360,729,755]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="obsidiana">
<emphasis id="B959EA86FF8BFFE3FB28FD4DFA98FD26" box="[1217,1360,729,755]" italics="true" pageId="7" pageNumber="542">C. obsidiana</emphasis>
</taxonomicName>
by the presence of a light central band followed on each side by a wide dark band (
<bibRefCitation id="EFBC4B65FF8BFFE3FC03FD69FAF7FCC2" author="Lago-Barcia, D. &amp; Carbayo, F." box="[1002,1343,765,791]" pageId="7" pageNumber="542" pagination="517 - 542" refId="ref11110" refString="Lago-Barcia, D. &amp; Carbayo, F. (2018) Taxonomic revision of four species of the Neotropical land planarian genera Cratera and Geoplana (Platyhelminthes, Tricladida) with a description of two new species and an emendation of Cratera. Zootaxa, 4500, 517 - 542. https: // doi. org / 10.11646 / zootaxa. 4500.4. 3" type="journal article" year="2018">Lago-Barcia &amp; Carbayo 2018</bibRefCitation>
).
</paragraph>
<paragraph id="8B923694FF8BFFE3FF2EFCB5FBEBFC1E" blockId="7.[151,1437,152,971]" pageId="7" pageNumber="542">
Regarding the copulatory apparatus,
<taxonomicName id="4C2D4D17FF8BFFE3FD89FCB5FD25FCEE" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[608,749,801,827]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="obsidiana">
<emphasis id="B959EA86FF8BFFE3FD89FCB5FD25FCEE" box="[608,749,801,827]" italics="true" pageId="7" pageNumber="542">C. obsidiana</emphasis>
</taxonomicName>
has a tubular prostatic vesicle with the proximal portion laterally expanded (T-shaped) similar to that of
<taxonomicName id="4C2D4D17FF8BFFE3FD96FCD1FC75FC8A" authority="Rossi et al., 2014" authorityName="Rossi" authorityYear="2014" box="[639,957,837,863]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="steffeni">
<emphasis id="B959EA86FF8BFFE3FD96FCD1FD39FC8A" box="[639,753,837,863]" italics="true" pageId="7" pageNumber="542">C. steffeni</emphasis>
<bibRefCitation id="EFBC4B65FF8BFFE3FD11FCD1FC75FC8A" author="Rossi, I. &amp; Fontoura, M. &amp; Amaral, S. &amp; Leal-Zanchet, A. M." box="[760,957,837,863]" pageId="7" pageNumber="542" pagination="514 - 524" refId="ref11612" refString="Rossi, I., Fontoura, M., Amaral, S. &amp; Leal-Zanchet, A. M. (2014) A new species of land flatworm (Platyhelminthes: Continenticola) from areas of Araucaria Forest in southern Brazil. Zootaxa, 3794 (2), 514 - 524. https: // doi. org / 10.11646 / zootaxa. 3794.4. 2" type="journal article" year="2014">
Rossi
<emphasis id="B959EA86FF8BFFE3FCD7FCD2FCB8FC8A" box="[830,880,837,863]" italics="true" pageId="7" pageNumber="542">et al</emphasis>
., 2014
</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C2D4D17FF8BFFE3FC1CFCD1FB9FFC8A" box="[1013,1111,837,863]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="ochra">
<emphasis id="B959EA86FF8BFFE3FC1CFCD1FB9FFC8A" box="[1013,1111,837,863]" italics="true" pageId="7" pageNumber="542">C. ochra</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFBC4B65FF8BFFE3FB8EFCD1FAE1FC8A" author="Rossi, I. &amp; Fontoura, M. &amp; Amaral, S. &amp; Leal-Zanchet, A. M." box="[1127,1321,837,863]" pageId="7" pageNumber="542" pagination="514 - 524" refId="ref11612" refString="Rossi, I., Fontoura, M., Amaral, S. &amp; Leal-Zanchet, A. M. (2014) A new species of land flatworm (Platyhelminthes: Continenticola) from areas of Araucaria Forest in southern Brazil. Zootaxa, 3794 (2), 514 - 524. https: // doi. org / 10.11646 / zootaxa. 3794.4. 2" type="journal article" year="2014">
Rossi
<emphasis id="B959EA86FF8BFFE3FB47FCD2FB2FFC8A" box="[1198,1255,837,863]" italics="true" pageId="7" pageNumber="542">et al.</emphasis>
2014
</bibRefCitation>
,
<bibRefCitation id="EFBC4B65FF8BFFE3FADCFCD1FABAFCB5" author="Rossi, I. &amp; Amaral, S. V. &amp; Ribeiro, G. G. &amp; Cauduro, G. P. &amp; Fick, I. &amp; Valiati, V. H. &amp; Leal-Zanchet, A. M." box="[1333,1394,837,864]" pageId="7" pageNumber="542" pagination="787 - 815" refId="ref11524" refString="Rossi, I., Amaral, S. V., Ribeiro, G. G., Cauduro, G. P., Fick, I., Valiati, V. H. &amp; Leal-Zanchet, A. M. (2016) Two new Geoplaninae species (Platyhelminthes: Continenticola) from Southern Brazil based on an integrative taxonomic approach. Journal of Natural History, 50, 787 - 815. https: // doi. org / 10.1080 / 00222933.2015.108405 7" type="journal article" year="2016">2016</bibRefCitation>
). In all these three species, the prostatic vesicle is considered to be single, without forked portions, since it just shows a lateral expansion. However,
<taxonomicName id="4C2D4D17FF8BFFE3FE3CFC19FDAAFC72" authorityName="Boll &amp; Amaral &amp; Leal-Zanchet" authorityYear="2019" box="[469,610,909,935]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="species" species="obsidiana">
<emphasis id="B959EA86FF8BFFE3FE3CFC19FDAAFC72" box="[469,610,909,935]" italics="true" pageId="7" pageNumber="542">C. obsidiana</emphasis>
</taxonomicName>
has the proximal portion of the prostatic vesicle slightly dorsally oriented, being such an anatomical feature recorded for the first time in the genus
<taxonomicName id="4C2D4D17FF8BFFE3FC2DFC25FBD5FC1E" box="[964,1053,945,971]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="542" phylum="Platyhelminthes" rank="genus">
<emphasis id="B959EA86FF8BFFE3FC2DFC25FBD5FC1E" box="[964,1053,945,971]" italics="true" pageId="7" pageNumber="542">Cratera</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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