525 lines
69 KiB
XML
525 lines
69 KiB
XML
<document id="025C4724D3473E3CE62A57354B0540CA" ID-CLB-Dataset="304981" ID-DOI="10.11646/zootaxa.5536.2.1" ID-GBIF-Dataset="903c9627-28ff-4bfb-9c49-d911e9673c71" ID-ISSN="1175-5326" ID-Zenodo-Dep="14033618" ID-ZooBank="5F1FBB59-9C69-4E66-9B0D-69A015F30EAA" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1730712964139" checkinUser="plazi" docAuthor="ALEKSEEV, VITALII, MCKELLAR, RYAN C. & BUKEJS, ANDRIS" docDate="2024" docId="03F4DF38FFF16654FF506B9BB912571A" docLanguage="en" docName="zootaxa.5536.2.1.pdf" docOrigin="Zootaxa 5536 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5536.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helioctamenus groehni Alekseev et Bukejs 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="223" masterDocId="FFCDA740FFE06643FFC76A41BB4B5607" masterDocTitle="A revision and addition to Zopheridae (Coleoptera: Tenebrionoidea) in Baltic amber: possible connections between modern Holarctic distributions and Eocene ‘ amber forests’" masterLastPageNumber="247" masterPageNumber="201" pageNumber="218" updateTime="1730817957888" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="648473200E2CA6F0C8E482FB88DA7ED8">A revision and addition to Zopheridae (Coleoptera: Tenebrionoidea) in Baltic amber: possible connections between modern Holarctic distributions and Eocene ‘ amber forests’</mods:title>
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<mods:name id="A04812F8814A522F3E626559CB6DEF28" type="personal">
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<mods:roleTerm id="66FA1ED32C777CF849DD058C209398ED">Author</mods:roleTerm>
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<mods:namePart id="EA478AD395199DC1CBABBFB66AE59A2E">ALEKSEEV, VITALII</mods:namePart>
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<mods:nameIdentifier id="F736D71FA8527D7026DB9D1221F6EFC0" type="ORCID">0000-0003-4390-5443</mods:nameIdentifier>
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<mods:affiliation id="3793469F676726C9B68E55CFD94234D2">Immanuel Kant Baltic Federal University, Nevskogo 14, Kaliningrad 236016, Russia.</mods:affiliation>
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<mods:nameIdentifier id="54AB67854FB0CA52CA0064C1F6BA827A" type="email">alekseew0802@yahoo.com</mods:nameIdentifier>
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<mods:name id="F86FABBCF18CD3F55AEFEA05B4521C39" type="personal">
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<mods:roleTerm id="EE091425806C117EDEBDB795D5A1E6D9">Author</mods:roleTerm>
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<mods:namePart id="81383C4485D0C862F016C89B14A553C2">MCKELLAR, RYAN C.</mods:namePart>
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<mods:nameIdentifier id="34FC42DDFEB1F5ED759F1A3726283250" type="ORCID">0000-0002-7451-4368</mods:nameIdentifier>
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<mods:affiliation id="66B7EFA5F900DC3F72E2936FBC93611A">Royal Saskatchewan Museum, 2445 Albert St., Regina, SK, S 4 P 4 W 7, Canada. & Biology Department, University of Regina, Regina, SK, S 4 S 0 A 2, Canada; & Department of Ecology & Evolutionary Biology, University of Kansas, Lawrence, Kansas, 66045, USA;</mods:affiliation>
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<mods:nameIdentifier id="0B62870AAAFDD613D977CB0E1878F81D" type="email">ryan.mckellar@gov.sk.ca</mods:nameIdentifier>
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</mods:role>
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<mods:namePart id="019226464FD5B0B5BB65F2E1B93FB6A3">BUKEJS, ANDRIS</mods:namePart>
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<mods:nameIdentifier id="45D3FD8BC36B78DA0CDEB4F1618D79EF" type="ORCID">0000-0001-7165-3023</mods:nameIdentifier>
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<mods:affiliation id="7D46315F42A0AD811043841D3B6378F2">Institute of Life Sciences and Technologies, Daugavpils University, Vienības 13, Daugavpils LV- 5401, Latvia.</mods:affiliation>
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<mods:nameIdentifier id="D150C4E9C4A3514A6C3F0E060443C004" type="email">carabidae@inbox.lv</mods:nameIdentifier>
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<mods:title id="E11E239DEE95964C2B03086C8F7AD749">Zootaxa</mods:title>
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<mods:part id="2C233AA96E841E3CE0351428A01D246D">
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<mods:date id="ED2C1346C502CC7551CE76D04A9A5325">2024</mods:date>
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<mods:number id="4A2DDA0BB37E816B6C658821A132A924">2024-11-04</mods:number>
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<mods:number id="D94B4872560F853496990BAA2E25DA53">5536</mods:number>
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<mods:url id="F79C351FDF7FC7BE349D1482E99F1241">http://dx.doi.org/10.11646/zootaxa.5536.2.1</mods:url>
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<mods:classification id="D47E629A6E806D3756DCB17201C6A15A">journal article</mods:classification>
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<mods:identifier id="C8A496281C297937333E4D0E1FA42578" type="Zenodo-Dep">14033618</mods:identifier>
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<treatment id="03F4DF38FFF16654FF506B9BB912571A" ID-DOI="http://doi.org/10.5281/zenodo.14040782" ID-Zenodo-Dep="14040782" LSID="urn:lsid:plazi:treatment:03F4DF38FFF16654FF506B9BB912571A" httpUri="http://treatment.plazi.org/id/03F4DF38FFF16654FF506B9BB912571A" lastPageId="23" lastPageNumber="223" pageId="17" pageNumber="218">
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<subSubSection id="C3473DA5FFF16652FF506B9BB84157F2" box="[151,778,474,501]" pageId="17" pageNumber="218" type="nomenclature">
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<paragraph id="8BE26E2EFFF16652FF506B9BB84157F2" blockId="17.[151,778,474,537]" box="[151,778,474,501]" pageId="17" pageNumber="218">
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<heading id="D0AAD942FFF16652FF506B9BB84157F2" bold="true" box="[151,778,474,501]" fontSize="11" level="1" pageId="17" pageNumber="218" reason="1">
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<emphasis id="B929B23CFFF16652FF506B9BB84157F2" bold="true" box="[151,778,474,501]" pageId="17" pageNumber="218">
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<taxonomicName id="4C5D15ADFFF16652FF506B9BB9EE57F3" authority="Alekseev et Bukejs" authorityName="Alekseev et Bukejs" authorityYear="2024" box="[151,677,474,501]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="groehni" status="sp. nov.">
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<emphasis id="B929B23CFFF16652FF506B9BBAFF57F2" bold="true" box="[151,436,474,501]" italics="true" pageId="17" pageNumber="218">Helioctamenus groehni</emphasis>
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Alekseev et Bukejs
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</taxonomicName>
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<taxonomicNameLabel id="A21A0F47FFF16652FD6A6B9AB84157F2" box="[685,778,475,501]" pageId="17" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3473DA5FFF16652FF506BBEBA6C541E" box="[151,295,511,537]" pageId="17" pageNumber="218" type="description">
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<paragraph id="8BE26E2EFFF16652FF506BBEBA6C541E" blockId="17.[151,778,474,537]" box="[151,295,511,537]" pageId="17" pageNumber="218">
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(
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<figureCitation id="136672ABFFF16652FF586BBEBA54541E" box="[159,287,511,537]" captionStart-0="FIGURES 17–19" captionStart-1="FIGURES 20–21" captionStart-2="FIGURES 22–24" captionStart-3="FIGURES 25–26" captionStartId-0="19.[151,264,1935,1959]" captionStartId-1="20.[151,264,1805,1829]" captionStartId-2="21.[151,264,1869,1893]" captionStartId-3="22.[151,264,1430,1454]" captionTargetBox-0="[156,1431,179,1908]" captionTargetBox-1="[151,1435,198,1779]" captionTargetBox-2="[151,1435,179,1843]" captionTargetBox-3="[151,1435,179,1404]" captionTargetId-0="figure-17@19.[156,1431,179,1908]" captionTargetId-1="figure-21@20.[151,1435,198,1779]" captionTargetId-2="figure-17@21.[151,1435,179,1843]" captionTargetId-3="figure-21@22.[151,1435,179,1404]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="21" captionTargetPageId-3="22" captionText-0="FIGURES 17–19. Helioctamenus groehni sp. nov., holotype, No. 5209 [GPIH]: 17—habitus, dorsal view; 18—habitus, ventral view; 19—habitus, dorso-lateral view. Scale bar = 0.5 mm." captionText-1="FIGURES 20–21. Helioctamenus groehni sp. nov., holotype, No. 5209 [GPIH]: 20—habitus, left lateral view; 21—details of anterior part of body, dorso-lateral view.Abbreviations: a1–a11—antennomeres 1–11, respectively. Scale bars = 1.0 mm for Fig. 20; 0.25 mm for Fig. 21." captionText-2="FIGURES 22–24. Helioctamenus groehni sp. nov., paratype, No. 5217 [GPIH]: 22—habitus, dorsal view; 23—habitus, ventral view; 24—habitus, left lateral view. Scale bar = 0.5 mm." captionText-3="FIGURES 25–26. Helioctamenus groehni sp. nov., paratype, KA-CLP-001 [CKVA]: 25—habitus, dorsal view; 26—habitus, ventral view. Scale bar = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14033638" figureDoi-1="http://doi.org/10.5281/zenodo.14033646" figureDoi-2="http://doi.org/10.5281/zenodo.14033650" figureDoi-3="http://doi.org/10.5281/zenodo.14033660" httpUri-0="https://zenodo.org/record/14033638/files/figure.png" httpUri-1="https://zenodo.org/record/14033646/files/figure.png" httpUri-2="https://zenodo.org/record/14033650/files/figure.png" httpUri-3="https://zenodo.org/record/14033660/files/figure.png" pageId="17" pageNumber="218">Figs 17–26</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3473DA5FFF16652FF506806BED8525E" pageId="17" pageNumber="218" type="materials_examined">
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||
<paragraph id="8BE26E2EFFF16652FF506806BFBB54CA" blockId="17.[151,1437,582,2013]" pageId="17" pageNumber="218">
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<emphasis id="B929B23CFFF16652FF506806BA0D5467" bold="true" box="[151,326,582,609]" pageId="17" pageNumber="218">Type material.</emphasis>
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<materialsCitation id="3B356473FFF16652FE8A6806BE525482" collectionCode="GPIH, CCGG" country="Russia" determinerName="Alekseev et Bukejs" location="Yantarny" municipality="Sambian" pageId="17" pageNumber="218" specimenCode="5209, 8706, JDC-12544" specimenCount="1" specimenCount-adult="1" stateProvince="Kaliningrad Region" typeStatus="holotype">
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<typeStatus id="54E6D08CFFF16652FE8A6806BAF15466" box="[333,442,583,609]" pageId="17" pageNumber="218" type="holotype">Holotype</typeStatus>
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: No.
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<collectionCode id="ED4CF6EBFFF16652FE316806B97C5466" box="[502,567,583,609]" collectionName="GPIH" country="Germany" httpUri="http://grbio.org/cool/8qxf-pkip" name="Geologisch-Palaeontologiches Institut der Universitt Haemburg" pageId="17" pageNumber="218">GPIH</collectionCode>
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no.
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<specimenCode id="DBFBC655FFF16652FDAE6807B9EF5466" box="[617,676,582,609]" pageId="17" pageNumber="218">5209</specimenCode>
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,
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<collectionCode id="ED4CF6EBFFF16652FD776806B9B55466" box="[688,766,583,609]" collectionName="CCGG" pageId="17" pageNumber="218">CCGG</collectionCode>
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no.
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<specimenCode id="DBFBC655FFF16652FCF76806B8235466" box="[816,872,583,609]" pageId="17" pageNumber="218">8706</specimenCode>
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(ex coll. Jonas Damzen
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<specimenCode id="DBFBC655FFF16652FBB96807BE4E5466" box="[1150,1285,582,609]" pageId="17" pageNumber="218">JDC-12544</specimenCode>
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); “Holotype /
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<taxonomicName id="4C5D15ADFFF16652FF50682ABAD05482" authorityName="Alekseev & Bukejs" authorityYear="2024" box="[151,411,619,645]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="groehni" status="sp. nov.">
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<emphasis id="B929B23CFFF16652FF50682ABAD05482" box="[151,411,619,645]" italics="true" pageId="17" pageNumber="218">Helioctamenus groehni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21A0F47FFF16652FE66682ABABC5482" box="[417,503,619,645]" pageId="17" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
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/
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<determinerName id="7E3E82EAFFF16652FDCE682AB9AB5482" box="[521,736,619,645]" pageId="17" pageNumber="218">Alekseev et Bukejs</determinerName>
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des. 2024” [red printed label];
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<specimenCount id="9D5BA5A7FFF16652FBFE682ABF3F5482" box="[1081,1140,619,645]" pageId="17" pageNumber="218" type="adult">adult</specimenCount>
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, sex unknown.
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</materialsCitation>
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A complete beetle included in a transparent, yellow amber piece with dimensions of 20×
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<quantity id="4CA5C3CBFFF16652FC3E68CEBF7154AE" box="[1017,1082,655,681]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="218" unit="mm" value="7.0">7 mm</quantity>
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and a maximum thickness of
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<quantity id="4CA5C3CBFFF16652FA4968CEBB8E54CA" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="218" unit="mm" value="3.0">3 mm</quantity>
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, preserved without supplementary fixation.
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<collectorName id="26A80BF8FFF16652FD7568F2B81F54CA" box="[690,852,691,717]" pageId="17" pageNumber="218">Syninclusions</collectorName>
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||
: several stellate
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||
<taxonomicName id="4C5D15ADFFF16652FBCC68F2BF3854CA" authorityName="Dumortier" authorityYear="1829" box="[1035,1139,691,717]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="17" pageNumber="218" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
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trichomes.
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</paragraph>
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||
<paragraph id="8BE26E2EFFF16652FF006896BF08555A" blockId="17.[151,1437,582,2013]" pageId="17" pageNumber="218">
|
||
<materialsCitation id="3B356473FFF16652FF006896BF215512" collectionCode="GPIH, CCGG" country="Russia" determinerName="Alekseev et Bukejs" location="Yantarny" municipality="Sambian" pageId="17" pageNumber="218" specimenCode="5217, 8707, JDC-13157" specimenCount="1" specimenCount-adult="1" stateProvince="Kaliningrad Region" typeStatus="paratype">
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<typeStatus id="54E6D08CFFF16652FF006896BA6654F6" box="[199,301,727,753]" pageId="17" pageNumber="218" type="paratype">Paratype</typeStatus>
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||
: No.
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||
<collectionCode id="ED4CF6EBFFF16652FEA16896BAEC54F6" box="[358,423,727,753]" collectionName="GPIH" country="Germany" httpUri="http://grbio.org/cool/8qxf-pkip" name="Geologisch-Palaeontologiches Institut der Universitt Haemburg" pageId="17" pageNumber="218">GPIH</collectionCode>
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no.
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<specimenCode id="DBFBC655FFF16652FE126897B95B54F6" box="[469,528,726,753]" pageId="17" pageNumber="218">5217</specimenCode>
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,
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<collectionCode id="ED4CF6EBFFF16652FDDD6896B92354F6" box="[538,616,727,753]" collectionName="CCGG" pageId="17" pageNumber="218">CCGG</collectionCode>
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no.
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<specimenCode id="DBFBC655FFF16652FD516896B98554F6" box="[662,718,727,753]" pageId="17" pageNumber="218">8707</specimenCode>
|
||
(ex coll. Jonas Damzen
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<specimenCode id="DBFBC655FFF16652FC1C6897BF2B54F6" box="[987,1120,726,753]" pageId="17" pageNumber="218">JDC-13157</specimenCode>
|
||
); “Paratype /
|
||
<taxonomicName id="4C5D15ADFFF16652FB336896BBA65512" authorityName="Alekseev & Bukejs" authorityYear="2024" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="groehni" status="sp. nov.">
|
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<emphasis id="B929B23CFFF16652FB336896BBA65512" italics="true" pageId="17" pageNumber="218">Helioctamenus groehni</emphasis>
|
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</taxonomicName>
|
||
<taxonomicNameLabel id="A21A0F47FFF16652FF3468BABA025512" box="[243,329,763,789]" pageId="17" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
|
||
/
|
||
<determinerName id="7E3E82EAFFF16652FE9C68BAB97A5512" box="[347,561,763,789]" pageId="17" pageNumber="218">Alekseev et Bukejs</determinerName>
|
||
des. 2024” [red printed label];
|
||
<specimenCount id="9D5BA5A7FFF16652FC4D68BAB88E5512" box="[906,965,763,789]" pageId="17" pageNumber="218" type="adult">adult</specimenCount>
|
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, sex unknown.
|
||
</materialsCitation>
|
||
A complete beetle included in a transparent, yellow amber piece with dimensions of 17×
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<quantity id="4CA5C3CBFFF16652FC80695EB8C2553E" box="[839,905,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="17" pageNumber="218" unit="mm" value="6.0">6 mm</quantity>
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and a maximum thickness of
|
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<quantity id="4CA5C3CBFFF16652FB18695EBE68553E" box="[1247,1315,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="218" unit="mm" value="3.0">3 mm</quantity>
|
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, preserved without supplementary fixation.
|
||
<collectorName id="26A80BF8FFF16652FDC26902B9EC555A" box="[517,679,835,861]" pageId="17" pageNumber="218">Syninclusions</collectorName>
|
||
: several stellate
|
||
<taxonomicName id="4C5D15ADFFF16652FC9A6902B88E555A" authorityName="Dumortier" authorityYear="1829" box="[861,965,835,861]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="17" pageNumber="218" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
|
||
trichomes.
|
||
</paragraph>
|
||
<paragraph id="8BE26E2EFFF16652FF006926BFAB55EA" blockId="17.[151,1437,582,2013]" pageId="17" pageNumber="218">
|
||
<materialsCitation id="3B356473FFF16652FF006926B99555A2" collectionCode="CKVA" country="Russia" determinerName="Alekseev et Bukejs" location="Yantarny" municipality="Sambian" pageId="17" pageNumber="218" specimenCode="KA-CLP-001" specimenCount="1" specimenCount-adult="1" stateProvince="Kaliningrad Region" typeStatus="paratype">
|
||
<typeStatus id="54E6D08CFFF16652FF006926BA665586" box="[199,301,871,897]" pageId="17" pageNumber="218" type="paratype">Paratype</typeStatus>
|
||
: No.
|
||
<specimenCode id="DBFBC655FFF16652FEAC6926B9485586" box="[363,515,871,897]" pageId="17" pageNumber="218">KA-CLP-001</specimenCode>
|
||
[
|
||
<collectionCode id="ED4CF6EBFFF16652FDD56926B92C5586" box="[530,615,871,897]" collectionName="CKVA" pageId="17" pageNumber="218">CKVA</collectionCode>
|
||
]; “Paratype /
|
||
<taxonomicName id="4C5D15ADFFF16652FCC76926B8E35586" baseAuthorityName="Slipinski & Schuh" baseAuthorityYear="2008" box="[768,936,871,897]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B929B23CFFF16652FCC76926B8E35586" box="[768,936,871,897]" italics="true" pageId="17" pageNumber="218">Helioctamenus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C5D15ADFFF16652FC066926BF5C5586" authorityName="Alekseev & Bukejs" authorityYear="2024" box="[961,1047,871,897]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="groehni" status="sp. nov.">
|
||
<emphasis id="B929B23CFFF16652FC066926BF5C5586" box="[961,1047,871,897]" italics="true" pageId="17" pageNumber="218">groehni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A0F47FFF16652FBD86926BF3C5586" box="[1055,1143,871,897]" pageId="17" pageNumber="218" rank="species">sp. nov.</taxonomicNameLabel>
|
||
/
|
||
<determinerName id="7E3E82EAFFF16652FB4A6926BE235586" box="[1165,1384,871,897]" pageId="17" pageNumber="218">Alekseev et Bukejs</determinerName>
|
||
des. 2024” [red handwritten label];
|
||
<specimenCount id="9D5BA5A7FFF16652FE3E69CAB97F55A2" box="[505,564,907,933]" pageId="17" pageNumber="218" type="adult">adult</specimenCount>
|
||
, sex unknown.
|
||
</materialsCitation>
|
||
A complete beetle with partially exposed metathoracic wings included in a transparent, yellow amber piece with dimensions of 26×
|
||
<quantity id="4CA5C3CBFFF16652FC7769EEB8B555CE" box="[944,1022,943,969]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="17" pageNumber="218" unit="mm" value="18.0">18 mm</quantity>
|
||
and a maximum thickness of
|
||
<quantity id="4CA5C3CBFFF16652FA9369EEBED355CE" box="[1364,1432,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="218" unit="mm" value="7.0">7 mm</quantity>
|
||
, preserved without supplementary fixation. Syninclusions:
|
||
<specimenCount id="9D5BA5A7FFF16652FCE36992B8F555EA" box="[804,958,979,1005]" count="1" pageId="17" pageNumber="218" type="generic">one specimen</specimenCount>
|
||
of Nematocera (
|
||
<taxonomicName id="4C5D15ADFFF16652FBBE6992BF9F55EA" box="[1145,1236,979,1005]" class="Insecta" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE26E2EFFF16652FF0069B6BAEF5232" blockId="17.[151,1437,582,2013]" pageId="17" pageNumber="218">
|
||
<emphasis id="B929B23CFFF16652FF0069B6BA385216" bold="true" box="[199,371,1015,1041]" pageId="17" pageNumber="218">Type stratum.</emphasis>
|
||
Baltic amber from Eocene amber-bearing Blaue Erde deposits; estimated age: middle–late Eocene (Standke 1998).
|
||
</paragraph>
|
||
<paragraph id="8BE26E2EFFF16652FF006E7EBED8525E" blockId="17.[151,1437,582,2013]" box="[199,1427,1086,1113]" pageId="17" pageNumber="218">
|
||
<emphasis id="B929B23CFFF16652FF006E7EBA13525F" bold="true" box="[199,344,1086,1113]" pageId="17" pageNumber="218">Type locality.</emphasis>
|
||
<location id="8E8238F5FFF16652FE996E7EBAF0525E" LSID="urn:lsid:plazi:treatment:03F4DF38FFF16654FF506B9BB912571A:8E8238F5FFF16652FE996E7EBAF0525E" box="[350,443,1087,1113]" country="Russia" municipality="Sambian" name="Yantarny" pageId="17" pageNumber="218" stateProvince="Kaliningrad Region">Yantarny</location>
|
||
settlement (formerly
|
||
<location id="8E8238F5FFF16652FD5C6E7EB852525E" LSID="urn:lsid:plazi:treatment:03F4DF38FFF16654FF506B9BB912571A:8E8238F5FFF16652FD5C6E7EB852525E" box="[667,793,1087,1113]" country="Russia" municipality="Sambian" name="Palmnicken" pageId="17" pageNumber="218" stateProvince="Kaliningrad Region">Palmnicken</location>
|
||
),
|
||
<collectingMunicipality id="6B86F454FFF16652FCED6E7EB8CF525E" box="[810,900,1087,1113]" pageId="17" pageNumber="218">Sambian</collectingMunicipality>
|
||
(Samland)
|
||
<collectingMunicipality id="6B86F454FFF16652FC3A6E7EBF2F525E" box="[1021,1124,1087,1113]" pageId="17" pageNumber="218">Peninsula</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="4999A0CCFFF16652FBA96E7EBE77525E" box="[1134,1340,1087,1113]" country="Russia" name="Kaliningrad" pageId="17" pageNumber="218">Kaliningrad Region</collectingRegion>
|
||
,
|
||
<collectingCountry id="F34A2EBEFFF16652FA806E7EBEC4525E" box="[1351,1423,1087,1113]" name="Russia" pageId="17" pageNumber="218">Russia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3473DA5FFF16651FF006E23BA805025" lastPageId="18" lastPageNumber="219" pageId="17" pageNumber="218" type="description">
|
||
<paragraph id="8BE26E2EFFF16652FF006E23BAA952EE" blockId="17.[151,1437,582,2013]" pageId="17" pageNumber="218">
|
||
<emphasis id="B929B23CFFF16652FF006E23BA95527A" bold="true" box="[199,478,1122,1149]" pageId="17" pageNumber="218">
|
||
Description of
|
||
<typeStatus id="54E6D08CFFF16652FEB06E22BA95527A" box="[375,478,1123,1149]" pageId="17" pageNumber="218" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
. Measurements: total body length (including visible part of head)
|
||
<quantity id="4CA5C3CBFFF16652FB006E22BE54527A" box="[1223,1311,1123,1149]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="17" pageNumber="218" unit="mm" value="3.1">3.1 mm</quantity>
|
||
, maximum body width (across elytra)
|
||
<quantity id="4CA5C3CBFFF16652FE0B6EC7B97E52A6" box="[460,565,1158,1185]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="17" pageNumber="218" unit="mm" value="1.15">1.15 mm</quantity>
|
||
; head length
|
||
<quantity id="4CA5C3CBFFF16652FD176EC7B86152A6" box="[720,810,1158,1185]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="218" unit="mm" value="0.5">0.5 mm</quantity>
|
||
, head maximum width (across eyes)
|
||
<quantity id="4CA5C3CBFFF16652FB116EC6BE7A52A6" box="[1238,1329,1159,1185]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="218" unit="mm" value="0.7">0.7 mm</quantity>
|
||
; pronotal length
|
||
<quantity id="4CA5C3CBFFF16652FF236EEABA7052C2" box="[228,315,1195,1221]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="17" pageNumber="218" unit="mm" value="0.6">0.6 mm</quantity>
|
||
, maximum pronotal width
|
||
<quantity id="4CA5C3CBFFF16652FDAE6EEAB98A52C2" box="[617,705,1195,1221]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="218" unit="mm" value="0.7">0.7 mm</quantity>
|
||
, basal pronotal width
|
||
<quantity id="4CA5C3CBFFF16652FC716EEABF4452C2" box="[950,1039,1195,1221]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="17" pageNumber="218" unit="mm" value="0.6">0.6 mm</quantity>
|
||
; elytral length (along elytral suture, including scutellum) 2.0 mm.
|
||
</paragraph>
|
||
<paragraph id="8BE26E2EFFF16652FF006EB2BF805352" blockId="17.[151,1437,582,2013]" pageId="17" pageNumber="218">Body subparallel-sided, elongate, about 3.3× as long as wide, weakly convex dorsally and ventrally, unicolorous dark brown. Pubescence: head and pronotum with short, strongly curved, decumbent, narrow setae; elytra with short, slightly curved, semierect, narrow setae arranged in regular rows and not forming pattern.</paragraph>
|
||
<paragraph id="8BE26E2EFFF16652FF006F1EBF9E514A" blockId="17.[151,1437,582,2013]" pageId="17" pageNumber="218">Head prognathous, transverse; densely granulose dorsally, with granules round, about as wide as diameter of eye facet, and each granule bearing seta; covered with coarse punctures ventrally. Anterior margin of clypeus widely rounded. Compound eyes hemispherical, strongly protruding, with coarse facets, without interfacetal setae. Antennal insertions concealed dorsally, distant from eyes; antennal grooves ventrad eye indistinct. Antennae short, extending to about midlength of pronotum; sparsely covered with short setae; 11-segmented without distinctly separated antennal club; scape invisible from above, cylindrical; pedicel cylindrical, slightly elongate, 1.15× as long as wide, about as wide as scape; antennomere 3 conical, slightly dilated apically, elongate, about 1.4× as long as wide, about 1.35× as long as pedicel; antennomere 4 subtrapezoidal, 1.2× as long as wide, about 0.7× as long as antennomere 3; antennomeres 5–8 subquadratic, nearly as long as wide; antennomere 9 subtrapezoidal, slightly dilated apically, 1.3× as wide as long, slightly wider than antennomere 8; antennomere 10 trapezoidal, dilated apically, transverse, 1.7× as wide long, distinctly wider than antennomere 9; antennomere 11 circular, as long as wide, with widely rounded apex, 0.7× as wide as antennomere 10. Maxillary palp with four palpomeres; palpomere 1 smallest, subtriangular; palpomeres 2 and 3 subquadratic, about as long as wide; palpomere 4 elongate oval, truncate, about 1.9× as long as palpomere 3. Terminal labial palpomere triangular, acute apically.</paragraph>
|
||
<paragraph id="8BE26E2EFFF16651FF006D16B8CF5738" blockId="17.[151,1437,582,2013]" lastBlockId="18.[151,1437,151,2036]" lastPageId="18" lastPageNumber="219" pageId="17" pageNumber="218">Pronotum subquadrate, slightly transverse, 1.16× as wide as long, widest at anterior angles, weakly narrowed posteriad, maximum pronotal width / basal pronotal width = 1.16; pronotal disc convex, uneven, longitudinally impressed medially, sloped at lateral sides; pronotal surface densely granulose, with distance between granules smaller than diameter of one granule, and each round granule bearing seta. Pronotal lateral margins subparallel, finely denticulate; basal margin widely rounded medially; anterior margin with weakly rounded middle lobe. Anterior pronotal angles distinct, rectangular; posterior angles obtuse. Prohypomera concave, densely punctate. Prosternum convex, densely covered with coarse punctuation. Intercoxal prosternal process elongate, extending slightly beyond posterior margin of procoxae, with subparallel lateral margins, rounded apically, rather wide, about as wide as diameter of procoxa. Procoxal cavities open posteriorly.</paragraph>
|
||
<paragraph id="8BE26E2EFFF26651FF006B08B99F5764" blockId="18.[151,1437,151,2036]" box="[199,724,329,355]" pageId="18" pageNumber="219">Scutellar shield distinct, small, almost circular.</paragraph>
|
||
<paragraph id="8BE26E2EFFF26651FF006B2CBF035486" blockId="18.[151,1437,151,2036]" pageId="18" pageNumber="219">Elytra almost parallel-sided within anterior two-thirds of their lengths, tapered apically, about 1.7× as long as wide combined, convex, without carinae, distinctly wider than pronotum basally, 3.3× as long as pronotum. Humeri well-developed; humeral angles rounded. Elytra striate-punctate. Scutellary striole absent. Elytral striae with rows of dense, elongate granules; intervals flat, with micropunctation. Epipleura well-developed, widest in basal half, slightly narrowing posteriad, punctate, reaching elytral apices. Mesocoxal cavities closed. Metanepisternum narrow, about 7.0× as long as wide, punctate, with lateral margins slightly emarginate. Metaventrite slightly convex, distinctly longer than abdominal ventrite 1, densely covered with coarse punctation, with distance between punctures distinctly smaller than diameter of one puncture; discrimen distinct in posterior half.</paragraph>
|
||
<paragraph id="8BE26E2EFFF26651FF0068CAB9FF557C" blockId="18.[151,1437,151,2036]" pageId="18" pageNumber="219">Legs rather short and robust, finely punctate and sparsely setose. Procoxa small, nearly rounded, separated by about one diameter of procoxa; mesocoxae hemispherical, narrowly separated by about 0.2× diameter of mesocoxa; metacoxae widely oval, transverse, narrowly separated by about 0.5× longitudinal diameter of metacoxa. Femora nearly spindle-shaped, slightly flattened, widened medially. Tibiae straight; tibiae and femora subequal in length. Tarsal formula 4-4-4; tarsomeres simple, non-lobed, with fine setation ventrally; tarsomeres 1–3 subequal in length and shape, cylindrical; tarsomere 4 longest, about as long as tarsomeres 1–3 combined, slightly curved. Claws simple, large, about 0.3× as long as tarsomere 4.</paragraph>
|
||
<paragraph id="8BE26E2EFFF26651FF0069C4BF5E520C" blockId="18.[151,1437,151,2036]" pageId="18" pageNumber="219">Abdomen with five visible ventrites; densely covered with fine punctuation, distance between punctures equal to 0.5–1.0× diameter of one puncture; abdominal sutures distinct throughout length. Relative length ratios of ventrites 1–5 equal to 22:17:14:10:13. Abdominal ventrite 1 with triangular, apically rounded intercoxal process. Abdominal ventrite 5 with widely rounded posterior margin, weakly concave towards apex.</paragraph>
|
||
<paragraph id="8BE26E2EFFF26651FF006E55BABC534B" blockId="18.[151,1437,151,2036]" pageId="18" pageNumber="219">
|
||
<emphasis id="B929B23CFFF26651FF006E55BA705229" bold="true" box="[199,315,1044,1070]" pageId="18" pageNumber="219">
|
||
<typeStatus id="54E6D08CFFF26651FF006E55BA7D5229" box="[199,310,1044,1070]" pageId="18" pageNumber="219" type="paratype">Paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
No. GPIH no. 5217, CCGG no. 8707 (
|
||
<figureCitation id="136672ABFFF26651FCCE6E55B8DA5229" box="[777,913,1044,1070]" captionStart="FIGURES 22–24" captionStartId="21.[151,264,1869,1893]" captionTargetBox="[151,1435,179,1843]" captionTargetId="figure-17@21.[151,1435,179,1843]" captionTargetPageId="21" captionText="FIGURES 22–24. Helioctamenus groehni sp. nov., paratype, No. 5217 [GPIH]: 22—habitus, dorsal view; 23—habitus, ventral view; 24—habitus, left lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14033650" httpUri="https://zenodo.org/record/14033650/files/figure.png" pageId="18" pageNumber="219">Figs 22–24</figureCitation>
|
||
). Measurements: total body length (including visible part of head)
|
||
<quantity id="4CA5C3CBFFF26651FE436E79BA965255" box="[388,477,1080,1106]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="18" pageNumber="219" unit="mm" value="3.1">3.1 mm</quantity>
|
||
, maximum body width (across elytra)
|
||
<quantity id="4CA5C3CBFFF26651FC526E79B8A45255" box="[917,1007,1080,1106]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="18" pageNumber="219" unit="mm" value="1.1">1.1 mm</quantity>
|
||
; head length
|
||
<quantity id="4CA5C3CBFFF26651FB406E79BFAA5255" box="[1159,1249,1080,1106]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="219" unit="mm" value="0.3">0.3 mm</quantity>
|
||
, head maximum width (across eyes)
|
||
<quantity id="4CA5C3CBFFF26651FE446E1ABAAB5271" box="[387,480,1115,1142]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="18" pageNumber="219" unit="mm" value="0.6">0.6 mm</quantity>
|
||
; pronotal length
|
||
<quantity id="4CA5C3CBFFF26651FD6F6E1DB84F5271" box="[680,772,1116,1142]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="18" pageNumber="219" unit="mm" value="0.7">0.7 mm</quantity>
|
||
, maximum pronotal width
|
||
<quantity id="4CA5C3CBFFF26651FB836E1DBFEB5271" box="[1092,1184,1116,1142]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="18" pageNumber="219" unit="mm" value="0.8">0.8 mm</quantity>
|
||
, basal pronotal width
|
||
<quantity id="4CA5C3CBFFF26651FF506E3EBBB4529D" box="[151,255,1151,1178]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="18" pageNumber="219" unit="mm" value="0.75">0.75 mm</quantity>
|
||
; elytral length (along elytral suture, including scutellum)
|
||
<quantity id="4CA5C3CBFFF26651FC4A6E3EB8AC529D" box="[909,999,1151,1178]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="18" pageNumber="219" unit="mm" value="2.1">2.1 mm</quantity>
|
||
. Elytral surface of
|
||
<typeStatus id="54E6D08CFFF26651FB076EC1BE6B529D" box="[1216,1312,1152,1178]" pageId="18" pageNumber="219" type="paratype">paratype</typeStatus>
|
||
is visually represented by coarse, apparently irregular, and generally homogenous sculpture, which consists of rounded forms of variable size—this significantly differs from striate-punctate elytra of the
|
||
<typeStatus id="54E6D08CFFF26651FBD06E86BF3752E6" box="[1047,1148,1223,1249]" pageId="18" pageNumber="219" type="holotype">holotype</typeStatus>
|
||
, which are covered with regularly arranged elongate granules. We interpreted this apparent difference as probable contamination covering beetle dorsum (i.e., wood dust and air covering this region within resin), and not natural elements of dorsal sculpture with species-level significance.
|
||
</paragraph>
|
||
<paragraph id="8BE26E2EFFF26651FF006F17BA805025" blockId="18.[151,1437,151,2036]" pageId="18" pageNumber="219">
|
||
<emphasis id="B929B23CFFF26651FF006F17BA705377" bold="true" box="[199,315,1366,1392]" pageId="18" pageNumber="219">
|
||
<typeStatus id="54E6D08CFFF26651FF006F17BA7D5377" box="[199,310,1366,1392]" pageId="18" pageNumber="219" type="paratype">Paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
No. KA-CLP-001 [CKVA] (
|
||
<figureCitation id="136672ABFFF26651FD406F14B8465377" box="[647,781,1365,1392]" captionStart="FIGURES 25–26" captionStartId="22.[151,264,1430,1454]" captionTargetBox="[151,1435,179,1404]" captionTargetId="figure-21@22.[151,1435,179,1404]" captionTargetPageId="22" captionText="FIGURES 25–26. Helioctamenus groehni sp. nov., paratype, KA-CLP-001 [CKVA]: 25—habitus, dorsal view; 26—habitus, ventral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14033660" httpUri="https://zenodo.org/record/14033660/files/figure.png" pageId="18" pageNumber="219">Figs 25–26</figureCitation>
|
||
). Measurements: total body length (including visible part of head)
|
||
<quantity id="4CA5C3CBFFF26651FF3D6F38BA195393" box="[250,338,1401,1428]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="18" pageNumber="219" unit="mm" value="3.1">3.1 mm</quantity>
|
||
, maximum body width (across elytra)
|
||
<quantity id="4CA5C3CBFFF26651FCC56F38B8105393" box="[770,859,1401,1428]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="18" pageNumber="219" unit="mm" value="1.2">1.2 mm</quantity>
|
||
; head length
|
||
<quantity id="4CA5C3CBFFF26651FC286F38BF0C5393" box="[1007,1095,1401,1428]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="219" unit="mm" value="0.2">0.2 mm</quantity>
|
||
, head maximum width (across eyes)
|
||
<quantity id="4CA5C3CBFFF26651FF1C6FDCBA7D53BF" box="[219,310,1437,1464]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="18" pageNumber="219" unit="mm" value="0.7">0.7 mm</quantity>
|
||
; pronotal length
|
||
<quantity id="4CA5C3CBFFF26651FE306FDCB91A53BF" box="[503,593,1437,1464]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="18" pageNumber="219" unit="mm" value="0.8">0.8 mm</quantity>
|
||
, maximum pronotal width
|
||
<quantity id="4CA5C3CBFFF26651FC406FDCB8AA53BF" box="[903,993,1437,1464]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="18" pageNumber="219" unit="mm" value="0.8">0.8 mm</quantity>
|
||
, basal pronotal width
|
||
<quantity id="4CA5C3CBFFF26651FB186FDCBE0353BF" box="[1247,1352,1437,1464]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="18" pageNumber="219" unit="mm" value="0.75">0.75 mm</quantity>
|
||
; elytral length (along elytral suture, including scutellum)
|
||
<quantity id="4CA5C3CBFFF26651FD7D6F80B85953DC" box="[698,786,1473,1499]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="18" pageNumber="219" unit="mm" value="2.1">2.1 mm</quantity>
|
||
. Discrimen distinct in posterior two-thirds of metaventrite. Metathoracic wings well developed. Otherwise, this specimen is similar in all visible morphological characters to
|
||
<typeStatus id="54E6D08CFFF26651FF506C49BBB25025" box="[151,249,1544,1570]" pageId="18" pageNumber="219" type="holotype">holotype</typeStatus>
|
||
No. 5209 [GPIH].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3473DA5FFF26651FF006C6DBF3F51F3" pageId="18" pageNumber="219" type="diagnosis">
|
||
<paragraph id="8BE26E2EFFF26651FF006C6DB9F0511B" blockId="18.[151,1437,151,2036]" pageId="18" pageNumber="219">
|
||
<emphasis id="B929B23CFFF26651FF006C6DBA9A5041" bold="true" box="[199,465,1580,1606]" pageId="18" pageNumber="219">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C5D15ADFFF26651FE1C6C6DB9A85041" authorityName="Alekseev & Bukejs" authorityYear="2024" box="[475,739,1580,1606]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="groehni" status="sp. nov.">
|
||
<emphasis id="B929B23CFFF26651FE1C6C6DB9A85041" box="[475,739,1580,1606]" italics="true" pageId="18" pageNumber="219">Helioctamenus groehni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B929B23CFFF26651FD2A6C6DB8015041" bold="true" box="[749,842,1580,1606]" pageId="18" pageNumber="219">
|
||
<taxonomicNameLabel id="A21A0F47FFF26651FD2A6C6DB8015041" box="[749,842,1580,1606]" pageId="18" pageNumber="219" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from all extant congeners in possessing the following combination of characters: antennae distinctly 11-segmented with antennomere 11 well-developed, not fused with previous antennomere, and as long as antennomere 10; elytral intervals flat, not carinate; humeri well-developed; metathoracic wings well developed; pronotum distinctly narrower than elytral base; and scutellar shield distinct. The elongate body of the new species resembles such species as
|
||
<taxonomicName id="4C5D15ADFFF26651FC286CFDBFBB50D2" authority="Dajoz, 1971" authorityName="Dajoz" authorityYear="1971" box="[1007,1264,1723,1749]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="bedeli">
|
||
<emphasis id="B929B23CFFF26651FC286CFDBF1350D2" box="[1007,1112,1723,1749]" italics="true" pageId="18" pageNumber="219">H. bedeli</emphasis>
|
||
<bibRefCitation id="EFCC13DFFFF26651FBA56CFABFBB50D2" author="Dajoz, R." box="[1122,1264,1723,1749]" pageId="18" pageNumber="219" pagination="83 - 101" refId="ref25392" refString="Dajoz, R. (1971) Coleopteres Colydiidae nouveaux ou peu connus de la region palearctique. L'Entomologiste, XXVII (4 - 5), 83 - 101." type="book chapter" year="1971">Dajoz, 1971</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5D15ADFFF26651FB386CFDBA7950FE" authority="(Bedel, 1891)" baseAuthorityName="Bedel" baseAuthorityYear="1891" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="fumicornis">
|
||
<emphasis id="B929B23CFFF26651FB386CFDBED750D2" box="[1279,1436,1723,1749]" italics="true" pageId="18" pageNumber="219">H. fumicornis</emphasis>
|
||
(Bedel, 1891)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C5D15ADFFF26651FEAC6C9EB9E150FE" authority="Oromi, 1984" authorityName="Oromi" authorityYear="1984" box="[363,682,1759,1785]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="occidentalis">
|
||
<emphasis id="B929B23CFFF26651FEAC6C9EB95850FE" box="[363,531,1759,1785]" italics="true" pageId="18" pageNumber="219">H. occidentalis</emphasis>
|
||
<bibRefCitation id="EFCC13DFFFF26651FDDE6C9EB9E150FE" author="Oromi, P." box="[537,682,1759,1785]" pageId="18" pageNumber="219" pagination="225 - 232" refId="ref27145" refString="Oromi, P. (1984) Helioctamenus occidentalis, un nuevo Colydiidae de Canarias (Coleoptera). Vieraea, 13 (1 - 2), 225 - 232." type="journal article" year="1984">Oromí, 1984</bibRefCitation>
|
||
</taxonomicName>
|
||
, but distinctly differs from shorter and more oval bodies of
|
||
<taxonomicName id="4C5D15ADFFF26651FAFF6C9EBA01511B" authority="(Espanol, 1948)" baseAuthorityName="Espanol" baseAuthorityYear="1948" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="gineri">
|
||
<emphasis id="B929B23CFFF26651FAFF6C9EBED750FF" box="[1336,1436,1759,1784]" italics="true" pageId="18" pageNumber="219">H. gineri</emphasis>
|
||
(
|
||
<bibRefCitation id="EFCC13DFFFF26651FF586D43BA09511B" author="Espanol, F." box="[159,322,1794,1820]" pageId="18" pageNumber="219" pagination="311 - 315" refId="ref25710" refString="Espanol, F. (1948) Notas sobre Coleopteros africanos: I. Un nuevo " Acotulus " (" Colydiidae ") del Sahara espanol. Eos, XXIV (3), 311 - 315." type="journal article" year="1948">Español, 1948</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5D15ADFFF26651FE476D42B9FC511B" authority="Reitter, 1903" authorityName="Reitter" authorityYear="1903" box="[384,695,1794,1820]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="lusitanicus">
|
||
<emphasis id="B929B23CFFF26651FE476D42B957511B" box="[384,540,1794,1820]" italics="true" pageId="18" pageNumber="219">H. lusitanicus</emphasis>
|
||
<bibRefCitation id="EFCC13DFFFF26651FDE46D43B9FC511B" author="Reitter, E." box="[547,695,1794,1820]" pageId="18" pageNumber="219" pagination="173 - 178" refId="ref27539" refString="Reitter, E. (1903) Siebzehnter Beitrag zur Coleopterenfauna von Europa und den angrenzenden Landern. Wiener Entomologische Zeitung, XXII (VI), 173 - 178." type="book chapter" year="1903">Reitter, 1903</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE26E2EFFF26651FF006D67BF3F51F3" blockId="18.[151,1437,151,2036]" pageId="18" pageNumber="219">
|
||
Among the zopherids described from Baltic amber, the new species is the first described representative of the tribe
|
||
<taxonomicName id="4C5D15ADFFF26651FF156D0BBA035163" authorityName="Billberg" authorityYear="1820" box="[210,328,1866,1892]" class="Insecta" family="Zopheridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="tribe" tribe="Sarrotriini">Sarrotriini</taxonomicName>
|
||
, thus
|
||
<taxonomicName id="4C5D15ADFFF26651FE4F6D0AB94A5163" authorityName="Alekseev & Bukejs" authorityYear="2024" box="[392,513,1866,1892]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="groehni" status="sp. nov.">
|
||
<emphasis id="B929B23CFFF26651FE4F6D0AB94A5163" box="[392,513,1866,1892]" italics="true" pageId="18" pageNumber="219">H. groehni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21A0F47FFF26651FDCF6D0BB9285163" box="[520,611,1866,1892]" pageId="18" pageNumber="219" rank="species">
|
||
<emphasis id="B929B23CFFF26651FDCF6D0BB9175163" bold="true" box="[520,604,1866,1892]" pageId="18" pageNumber="219">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
may be distinguished by the set of the tribal level characters that include: antennae setose, 11-segmented, without distinctly separated antennal club; wide prosternal process; and intercoxal abdominal process triangular, rounded apically. The new species may be confused only with the apparently similar
|
||
<taxonomicName id="4C5D15ADFFF26651FF506DF4B92651C8" authority="Alekseev et Bukejs" authorityName="Alekseev et Bukejs" authorityYear="2016" box="[151,621,1973,2000]" class="Insecta" family="Zopheridae" genus="Xylolaemus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="legalovi">
|
||
<emphasis id="B929B23CFFF26651FF506DF4BA3451C8" box="[151,383,1973,1999]" italics="true" pageId="18" pageNumber="219">Xylolaemus legalovi</emphasis>
|
||
Alekseev et Bukejs
|
||
</taxonomicName>
|
||
, but differs from it in the non-explanate lateral pronotal sides, elytral intervals flat and elytra without additional setose patches consisting of lanceolate scales.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF223EA6FFF36650FF506DCEB9A551CD" ID-DOI="http://doi.org/10.5281/zenodo.14033638" ID-Zenodo-Dep="14033638" httpUri="https://zenodo.org/record/14033638/files/figure.png" pageId="19" pageNumber="220" startId="19.[151,264,1935,1959]" targetBox="[156,1431,179,1908]" targetPageId="19" targetType="figure">
|
||
<paragraph id="8BE26E2EFFF36650FF506DCEB9A551CD" blockId="19.[151,1436,1935,1995]" pageId="19" pageNumber="220">
|
||
<emphasis id="B929B23CFFF36650FF506DCEBA1851A0" bold="true" box="[151,339,1935,1959]" pageId="19" pageNumber="220">FIGURES 17–19.</emphasis>
|
||
<taxonomicName id="4C5D15ADFFF36650FE9F6DCEB90F51A0" authorityName="Alekseev & Bukejs" authorityYear="2024" box="[344,580,1935,1959]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="220" phylum="Arthropoda" rank="species" species="groehni" status="sp. nov.">
|
||
<emphasis id="B929B23CFFF36650FE9F6DCEB90F51A0" box="[344,580,1935,1959]" italics="true" pageId="19" pageNumber="220">Helioctamenus groehni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B929B23CFFF36650FD8E6DCEB9D251A0" bold="true" box="[585,665,1935,1959]" pageId="19" pageNumber="220">
|
||
<taxonomicNameLabel id="A21A0F47FFF36650FD8E6DCEB9D251A0" box="[585,665,1935,1959]" pageId="19" pageNumber="220" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, No. 5209 [GPIH]: 17—habitus, dorsal view; 18—habitus, ventral view; 19—habitus, dorso-lateral view. Scale bar = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF223EA6FFF46657FF506D4CBADA516A" ID-DOI="http://doi.org/10.5281/zenodo.14033646" ID-Zenodo-Dep="14033646" httpUri="https://zenodo.org/record/14033646/files/figure.png" pageId="20" pageNumber="221" startId="20.[151,264,1805,1829]" targetBox="[151,1435,198,1779]" targetPageId="20" targetType="figure">
|
||
<paragraph id="8BE26E2EFFF46657FF506D4CBADA516A" blockId="20.[151,1437,1805,1901]" pageId="20" pageNumber="221">
|
||
<emphasis id="B929B23CFFF46657FF506D4CBA1E5122" bold="true" box="[151,341,1805,1829]" pageId="20" pageNumber="221">FIGURES 20–21.</emphasis>
|
||
<taxonomicName id="4C5D15ADFFF46657FE9C6D4CB9035122" authorityName="Alekseev & Bukejs" authorityYear="2024" box="[347,584,1805,1829]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="groehni" status="sp. nov.">
|
||
<emphasis id="B929B23CFFF46657FE9C6D4CB9035122" box="[347,584,1805,1829]" italics="true" pageId="20" pageNumber="221">Helioctamenus groehni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B929B23CFFF46657FD886D4CB9EB5122" bold="true" box="[591,672,1805,1829]" pageId="20" pageNumber="221">
|
||
<taxonomicNameLabel id="A21A0F47FFF46657FD886D4CB9EB5122" box="[591,672,1805,1829]" pageId="20" pageNumber="221" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, No. 5209 [GPIH]: 20—habitus, left lateral view; 21—details of anterior part of body, dorso-lateral view. Abbreviations: a1–a11—antennomeres 1–11, respectively. Scale bars = 1.0 mm for Fig. 20; 0.25 mm for Fig. 21.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF223EA6FFF56656FF506D0CB99E518E" ID-DOI="http://doi.org/10.5281/zenodo.14033650" ID-Zenodo-Dep="14033650" httpUri="https://zenodo.org/record/14033650/files/figure.png" pageId="21" pageNumber="222" startId="21.[151,264,1869,1893]" targetBox="[151,1435,179,1843]" targetPageId="21" targetType="figure">
|
||
<paragraph id="8BE26E2EFFF56656FF506D0CB99E518E" blockId="21.[151,1436,1869,1930]" pageId="21" pageNumber="222">
|
||
<emphasis id="B929B23CFFF56656FF506D0CBA1F5162" bold="true" box="[151,340,1869,1893]" pageId="21" pageNumber="222">FIGURES 22–24.</emphasis>
|
||
<taxonomicName id="4C5D15ADFFF56656FE9E6D0FB90F5161" authorityName="Alekseev & Bukejs" authorityYear="2024" box="[345,580,1870,1894]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="222" phylum="Arthropoda" rank="species" species="groehni" status="sp. nov.">
|
||
<emphasis id="B929B23CFFF56656FE9E6D0FB90F5161" box="[345,580,1870,1894]" italics="true" pageId="21" pageNumber="222">Helioctamenus groehni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B929B23CFFF56656FD8D6D0FB9D15161" bold="true" box="[586,666,1870,1894]" pageId="21" pageNumber="222">
|
||
<taxonomicNameLabel id="A21A0F47FFF56656FD8D6D0FB9D15161" box="[586,666,1870,1894]" pageId="21" pageNumber="222" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype, No. 5217 [GPIH]: 22—habitus, dorsal view; 23—habitus, ventral view; 24—habitus, left lateral view. Scale bar = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF223EA6FFF66655FF506FD7BAA653D5" ID-DOI="http://doi.org/10.5281/zenodo.14033660" ID-Zenodo-Dep="14033660" httpUri="https://zenodo.org/record/14033660/files/figure.png" pageId="22" pageNumber="223" startId="22.[151,264,1430,1454]" targetBox="[151,1435,179,1404]" targetPageId="22" targetType="figure">
|
||
<paragraph id="8BE26E2EFFF66655FF506FD7BAA653D5" blockId="22.[151,1436,1430,1490]" pageId="22" pageNumber="223">
|
||
<emphasis id="B929B23CFFF66655FF506FD7BA1E53A9" bold="true" box="[151,341,1430,1454]" pageId="22" pageNumber="223">FIGURES 25–26.</emphasis>
|
||
<taxonomicName id="4C5D15ADFFF66655FE9B6FD7B90153A9" authorityName="Alekseev & Bukejs" authorityYear="2024" box="[348,586,1430,1454]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="groehni" status="sp. nov.">
|
||
<emphasis id="B929B23CFFF66655FE9B6FD7B90153A9" box="[348,586,1430,1454]" italics="true" pageId="22" pageNumber="223">Helioctamenus groehni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B929B23CFFF66655FD966FD6B9E853A8" bold="true" box="[593,675,1431,1455]" pageId="22" pageNumber="223">
|
||
<taxonomicNameLabel id="A21A0F47FFF66655FD966FD6B9E853A8" box="[593,675,1431,1455]" pageId="22" pageNumber="223" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype, KA-CLP-001 [CKVA]: 25—habitus, dorsal view; 26—habitus, ventral view. Scale bar = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3473DA5FFF66655FF006C41B82F5038" pageId="22" pageNumber="223" type="etymology">
|
||
<paragraph id="8BE26E2EFFF66655FF006C41B82F5038" blockId="22.[151,1437,1536,2031]" pageId="22" pageNumber="223">
|
||
<emphasis id="B929B23CFFF66655FF006C41BAD7501D" bold="true" box="[199,412,1536,1562]" pageId="22" pageNumber="223">Derivatio nominis</emphasis>
|
||
. The specific epithet is a patronym; the new species is named in honor of Mr. Carsten Gröhn (Glinde,
|
||
<collectingCountry id="F34A2EBEFFF66655FF3C6C64BA235038" box="[251,360,1573,1599]" name="Germany" pageId="22" pageNumber="223">Germany</collectingCountry>
|
||
), an enthusiast and specialist in Baltic amber.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3473DA5FFF66655FF006C09B97B513C" pageId="22" pageNumber="223" type="distribution">
|
||
<paragraph id="8BE26E2EFFF66655FF006C09B97B513C" blockId="22.[151,1437,1536,2031]" pageId="22" pageNumber="223">
|
||
<emphasis id="B929B23CFFF66655FF006C09BAB95064" bold="true" box="[199,498,1608,1635]" pageId="22" pageNumber="223">Distribution of congeners</emphasis>
|
||
. The genus
|
||
<taxonomicName id="4C5D15ADFFF66655FDB56C08B8515064" baseAuthorityName="Slipinski & Schuh" baseAuthorityYear="2008" box="[626,794,1609,1635]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B929B23CFFF66655FDB56C08B8515064" box="[626,794,1609,1635]" italics="true" pageId="22" pageNumber="223">Helioctamenus</emphasis>
|
||
</taxonomicName>
|
||
is currently distributed in the Mediterranean region (
|
||
<bibRefCitation id="EFCC13DFFFF66655FA906C08BB995080" author="Dajoz, R." pageId="22" pageNumber="223" pagination="83 - 101" refId="ref25392" refString="Dajoz, R. (1971) Coleopteres Colydiidae nouveaux ou peu connus de la region palearctique. L'Entomologiste, XXVII (4 - 5), 83 - 101." type="book chapter" year="1971">Dajoz 1971</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFCC13DFFFF66655FF186C2FBABF5080" author="Slipinski, S. A. & Schuh, R." box="[223,500,1645,1673]" pageId="22" pageNumber="223" refId="ref28217" refString="Slipinski, S. A. & Schuh, R. (2008) Zopheridae. In: Lobl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 5. Apollo Books, Stenstrup, pp. 78 - 87." type="book" year="2008">Ślipiński & Schuh 2008</bibRefCitation>
|
||
), namely in the Iberian Peninsula (3 species), Baleares (1 species), Canary Islands (2 species), and North Africa (4 species). In total, ten species are known:
|
||
<taxonomicName id="4C5D15ADFFF66655FC2A6CD3BFA550AC" authority="Dajoz, 1971" authorityName="Dajoz" authorityYear="1971" box="[1005,1262,1681,1707]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="bedeli">
|
||
<emphasis id="B929B23CFFF66655FC2A6CD3BF1D50AC" box="[1005,1110,1681,1707]" italics="true" pageId="22" pageNumber="223">H. bedeli</emphasis>
|
||
<bibRefCitation id="EFCC13DFFFF66655FBA76CD0BFA550AC" author="Dajoz, R." box="[1120,1262,1681,1707]" pageId="22" pageNumber="223" pagination="83 - 101" refId="ref25392" refString="Dajoz, R. (1971) Coleopteres Colydiidae nouveaux ou peu connus de la region palearctique. L'Entomologiste, XXVII (4 - 5), 83 - 101." type="book chapter" year="1971">Dajoz, 1971</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C5D15ADFFF66655FB3A6CD3BA4E50C8" authority="Pic, 1922" authorityName="Pic" authorityYear="1922" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="curticornis">
|
||
<emphasis id="B929B23CFFF66655FB3A6CD3BED750AC" box="[1277,1436,1682,1707]" italics="true" pageId="22" pageNumber="223">H. curticornis</emphasis>
|
||
<bibRefCitation id="EFCC13DFFFF66655FF506CF4BA4E50C8" author="Pic, M." box="[151,261,1717,1743]" pageId="22" pageNumber="223" pagination="21 - 23" refId="ref27356" refString="Pic, M. (1922) Notes diverses, descriptions et diagnoses (suite). L'Echange, Revue Linneenne, 38 (408), 21 - 23." type="journal article" year="1922">Pic, 1922</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C5D15ADFFF66655FED66CF7B96750D7" authority="Cobos, 1950" authorityName="Cobos" authorityYear="1950" box="[273,556,1717,1744]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="espanoli">
|
||
<emphasis id="B929B23CFFF66655FED66CF7BAD850C8" box="[273,403,1717,1743]" italics="true" pageId="22" pageNumber="223">H. espanoli</emphasis>
|
||
Cobos, 1950
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C5D15ADFFF66655FDFE6CF7B82050D7" authority="Cobos, 1965" authorityName="Cobos" authorityYear="1965" box="[569,875,1717,1744]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="fernandezi">
|
||
<emphasis id="B929B23CFFF66655FDFE6CF7B99950C8" box="[569,722,1717,1743]" italics="true" pageId="22" pageNumber="223">H. fernandezi</emphasis>
|
||
Cobos, 1965
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C5D15ADFFF66655FCB06CF7BFF350C8" authority="(Bedel, 1891)" baseAuthorityName="Bedel" baseAuthorityYear="1891" box="[887,1208,1717,1743]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="fumicornis">
|
||
<emphasis id="B929B23CFFF66655FCB06CF7BF5950C8" box="[887,1042,1717,1743]" italics="true" pageId="22" pageNumber="223">H. fumicornis</emphasis>
|
||
(Bedel, 1891)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C5D15ADFFF66655FB036CF7BB9050F4" authority="(Espanol, 1948)" baseAuthorityName="Espanol" baseAuthorityYear="1948" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="gineri">
|
||
<emphasis id="B929B23CFFF66655FB036CF7BE6150C8" box="[1220,1322,1718,1743]" italics="true" pageId="22" pageNumber="223">H. gineri</emphasis>
|
||
(
|
||
<bibRefCitation id="EFCC13DFFFF66655FAFE6CF4BB9850F4" author="Espanol, F." pageId="22" pageNumber="223" pagination="311 - 315" refId="ref25710" refString="Espanol, F. (1948) Notas sobre Coleopteros africanos: I. Un nuevo " Acotulus " (" Colydiidae ") del Sahara espanol. Eos, XXIV (3), 311 - 315." type="journal article" year="1948">Español, 1948</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C5D15ADFFF66655FF206C9BB92350F4" authority="Schaufuss, 1882" authorityName="Schaufuss" authorityYear="1882" box="[231,616,1753,1779]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="hippopotamus">
|
||
<emphasis id="B929B23CFFF66655FF206C9BBAE350F4" box="[231,424,1753,1779]" italics="true" pageId="22" pageNumber="223">H. hippopotamus</emphasis>
|
||
<bibRefCitation id="EFCC13DFFFF66655FE696C98B92350F4" author="Schaufuss, L. W." box="[430,616,1753,1779]" pageId="22" pageNumber="223" pagination="527 - 560" refId="ref27762" refString="Schaufuss, L. W. (1882) Zoologische Ergebnisse von Excursionen auf den Balearen. III (Addenda und Fortsetzung). Numquam Otiosus, 3, 527 - 560." type="journal article" year="1882">Schaufuss, 1882</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C5D15ADFFF66655FDB36C9BB8E250F4" authority="Reitter, 1903" authorityName="Reitter" authorityYear="1903" box="[628,937,1753,1779]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="lusitanicus">
|
||
<emphasis id="B929B23CFFF66655FDB36C9BB84450F4" box="[628,783,1753,1779]" italics="true" pageId="22" pageNumber="223">H. lusitanicus</emphasis>
|
||
<bibRefCitation id="EFCC13DFFFF66655FCD16C98B8E250F4" author="Reitter, E." box="[790,937,1753,1779]" pageId="22" pageNumber="223" pagination="173 - 178" refId="ref27539" refString="Reitter, E. (1903) Siebzehnter Beitrag zur Coleopterenfauna von Europa und den angrenzenden Landern. Wiener Entomologische Zeitung, XXII (VI), 173 - 178." type="book chapter" year="1903">Reitter, 1903</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C5D15ADFFF66655FC726C9BBFBD50F4" authority="Oromi, 1984" authorityName="Oromi" authorityYear="1984" box="[949,1270,1753,1779]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="occidentalis">
|
||
<emphasis id="B929B23CFFF66655FC726C9BBF1550F4" box="[949,1118,1753,1779]" italics="true" pageId="22" pageNumber="223">H. occidentalis</emphasis>
|
||
<bibRefCitation id="EFCC13DFFFF66655FBA36C98BFBD50F4" author="Oromi, P." box="[1124,1270,1753,1779]" pageId="22" pageNumber="223" pagination="225 - 232" refId="ref27145" refString="Oromi, P. (1984) Helioctamenus occidentalis, un nuevo Colydiidae de Canarias (Coleoptera). Vieraea, 13 (1 - 2), 225 - 232." type="journal article" year="1984">Oromí, 1984</bibRefCitation>
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName id="4C5D15ADFFF66655FAF66C9BBA76511F" authority="Espanol, 1957" authorityName="Espanol" authorityYear="1957" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="pardoi">
|
||
<emphasis id="B929B23CFFF66655FAF66C9BBED650F4" box="[1329,1437,1753,1779]" italics="true" pageId="22" pageNumber="223">H. pardoi</emphasis>
|
||
Español, 1957
|
||
</taxonomicName>
|
||
. The first extinct species described in the current paper confirms the presence of the genus in the Paleogene of the Western Palearctic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3473DA5FFF66654FF006D04B912571A" lastPageId="23" lastPageNumber="224" pageId="22" pageNumber="223" type="biology_ecology">
|
||
<paragraph id="8BE26E2EFFF66654FF006D04B912571A" blockId="22.[151,1437,1536,2031]" lastBlockId="23.[151,1436,151,285]" lastPageId="23" lastPageNumber="224" pageId="22" pageNumber="223">
|
||
<emphasis id="B929B23CFFF66655FF006D04BA825158" bold="true" box="[199,457,1860,1887]" pageId="22" pageNumber="223">Ecology of congeners.</emphasis>
|
||
Recent representatives of
|
||
<taxonomicName id="4C5D15ADFFF66655FD366D04B8D25158" baseAuthorityName="Slipinski & Schuh" baseAuthorityYear="2008" box="[753,921,1861,1887]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B929B23CFFF66655FD366D04B8D25158" box="[753,921,1861,1887]" italics="true" pageId="22" pageNumber="223">Helioctamenus</emphasis>
|
||
</taxonomicName>
|
||
are more or less xerophilous, occurring in dry habitats, under stones or plants (
|
||
<bibRefCitation id="EFCC13DFFFF66655FDCC6D28B9DE5184" author="Dajoz, R." box="[523,661,1897,1923]" pageId="22" pageNumber="223" pagination="83 - 101" refId="ref25392" refString="Dajoz, R. (1971) Coleopteres Colydiidae nouveaux ou peu connus de la region palearctique. L'Entomologiste, XXVII (4 - 5), 83 - 101." type="book chapter" year="1971">Dajoz 1971</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C5D15ADFFF66655FD6D6D28BFF65184" authority="Schaufuss, 1882" authorityName="Schaufuss" authorityYear="1882" box="[682,1213,1897,1923]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="hippopotamus">
|
||
<emphasis id="B929B23CFFF66655FD6D6D28B8B15184" box="[682,1018,1897,1923]" italics="true" pageId="22" pageNumber="223">Helioctamenus hippopotamus</emphasis>
|
||
<bibRefCitation id="EFCC13DFFFF66655FBC46D28BFF65184" author="Schaufuss, L. W." box="[1027,1213,1897,1923]" pageId="22" pageNumber="223" pagination="527 - 560" refId="ref27762" refString="Schaufuss, L. W. (1882) Zoologische Ergebnisse von Excursionen auf den Balearen. III (Addenda und Fortsetzung). Numquam Otiosus, 3, 527 - 560." type="journal article" year="1882">Schaufuss, 1882</bibRefCitation>
|
||
</taxonomicName>
|
||
was collected “sub lapidibus magnus” [under large stones] (
|
||
<bibRefCitation id="EFCC13DFFFF66655FDA26DCCB85551A0" author="Schaufuss, L. W." box="[613,798,1933,1959]" pageId="22" pageNumber="223" pagination="527 - 560" refId="ref27762" refString="Schaufuss, L. W. (1882) Zoologische Ergebnisse von Excursionen auf den Balearen. III (Addenda und Fortsetzung). Numquam Otiosus, 3, 527 - 560." type="journal article" year="1882">Schaufuss 1882</bibRefCitation>
|
||
), and the
|
||
<typeStatus id="54E6D08CFFF66655FC486DCCB8F451A0" box="[911,959,1933,1959]" pageId="22" pageNumber="223">type</typeStatus>
|
||
series of
|
||
<taxonomicName id="4C5D15ADFFF66655FBF66DCFBF9751A0" authorityName="Oromi" authorityYear="1984" box="[1073,1244,1933,1959]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="occidentalis">
|
||
<emphasis id="B929B23CFFF66655FBF66DCFBF9751A0" box="[1073,1244,1933,1959]" italics="true" pageId="22" pageNumber="223">H. occidentalis</emphasis>
|
||
</taxonomicName>
|
||
was sampled on lichens and epiphytic mosses of
|
||
<taxonomicName id="4C5D15ADFFF66655FDC96DF3B82B51CC" authority="(Oromi 1984)" baseAuthorityName="Oromi" baseAuthorityYear="1984" box="[526,864,1969,1995]" class="Arachnida" family="Salticidae" genus="Erica" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="scoparia">
|
||
<emphasis id="B929B23CFFF66655FDC96DF3B9FC51CC" box="[526,695,1970,1995]" italics="true" pageId="22" pageNumber="223">Erica scoparia</emphasis>
|
||
(
|
||
<bibRefCitation id="EFCC13DFFFF66655FD0F6DF0B81351CC" author="Oromi, P." box="[712,856,1969,1995]" pageId="22" pageNumber="223" pagination="225 - 232" refId="ref27145" refString="Oromi, P. (1984) Helioctamenus occidentalis, un nuevo Colydiidae de Canarias (Coleoptera). Vieraea, 13 (1 - 2), 225 - 232." type="journal article" year="1984">Oromí 1984</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. The newly described Eocene
|
||
<taxonomicName id="4C5D15ADFFF66655FB786DF0BE2C51CC" baseAuthorityName="Slipinski & Schuh" baseAuthorityYear="2008" box="[1215,1383,1969,1995]" class="Insecta" family="Colydiinae" genus="Helioctamenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B929B23CFFF66655FB786DF0BE2C51CC" box="[1215,1383,1969,1995]" italics="true" pageId="22" pageNumber="223">Helioctamenus</emphasis>
|
||
</taxonomicName>
|
||
was most likely also associated with a xerophilous community, possibly consisting of epiphytic mosses and lichens on tree bark of trunks, or it could be ground-living, occurring under lichens in sun-exposed microhabitats of the amberiferous forest. Records of different lichens in Baltic amber are known and not rare (e.g.,
|
||
<bibRefCitation id="EFCC13DFFFF76654FB2B6AFABEDC56D2" author="Rikkinen, J." box="[1260,1431,187,213]" pageId="23" pageNumber="224" pagination="1032 - 1036" refId="ref27570" refString="Rikkinen, J. (2003) Calicioid lichens from European Tertiary amber. Mycologia, 95, 1032 - 1036. https: // doi. org / 10.1080 / 15572536.2004.11833019" type="journal article" year="2003">Rikkinen 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFCC13DFFFF76654FF506A9EBACD56FE" author="Rikkinen, J. & Meinke S. K. L. & Grabenhorst, H. & Grohn, C. & Kobbert, M. & Wunderlich, J. & Schmidt, A. R." box="[151,390,223,249]" pageId="23" pageNumber="224" pagination="469 - 479" refId="ref27602" refString="Rikkinen, J., Meinke S. K. L., Grabenhorst, H., Grohn, C., Kobbert, M., Wunderlich, J. & Schmidt, A. R. (2018) Calicioid lichens and fungi in amber - tracing extant lineages back to the Paleogene. Geobios, 51, 469 - 479. https: // doi. org / 10.1016 / j. geobios. 2018.08.009" type="journal article" year="2018">
|
||
Rikkinen
|
||
<emphasis id="B929B23CFFF76654FEC06A9EBA7056FE" box="[263,315,223,249]" italics="true" pageId="23" pageNumber="224">et al</emphasis>
|
||
. 2018
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFCC13DFFFF76654FE536A9EB9ED56FE" author="Kaasalainen, U. & Kukwa, M. & Rikkinen, J. & Schmidt, A. R." box="[404,678,223,249]" pageId="23" pageNumber="224" refId="ref26133" refString="Kaasalainen, U., Kukwa, M., Rikkinen, J. & Schmidt, A. R. (2019) Crustose lichens with lichenicolous fungi from Paleogene amber. Scientific Reports, 9, 10360. https: // doi. org / 10.1038 / s 41598 - 019 - 46692 - w" type="journal volume" year="2019">
|
||
Kaasalainen
|
||
<emphasis id="B929B23CFFF76654FDE16A9EB91156FE" box="[550,602,223,249]" italics="true" pageId="23" pageNumber="224">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
), thus a lichen-related habitat for extinct species may be the most likely analogue among Recent relatives.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |