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<document id="1234CBC4DD73798C5368DCBF5FABBDA3" ID-DOI="10.5281/zenodo.5392079" ID-ISSN="1638-9387" ID-Zenodo-Dep="5392079" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630316032044" checkinUser="marcus" docAuthor="Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul &amp; Spiegel, Didier Van Den" docDate="2007" docId="03F4C378A23A305FFF6E02B6FCCAFAD6" docLanguage="en" docName="Zoosystema.2007.29.1.7-49.pdf" docOrigin="Zoosystema 29 (1)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Glyphiulus beroni Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel 2007, n. sp." docType="treatment" docVersion="3" lastPageNumber="30" masterDocId="FFCDBB00A22F3048FF94075DFFE6FF91" masterDocTitle="Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group" masterLastPageNumber="49" masterPageNumber="7" pageNumber="28" updateTime="1699223690750" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="CC58E336C34536111EFB9AD42E0D01D6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7003509C5E33629D31A07D27598E601D">
<mods:title id="03DD7F258E422A9B655097F34C0060AA">Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group</mods:title>
</mods:titleInfo>
<mods:name id="FA82D0B7D57F0EC4ED2B13D213F9B109" type="personal">
<mods:role id="258B460C93A42B34E0F791DB03A38009">
<mods:roleTerm id="C848064F26C3B39228D698C8603F493E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="39AC0CACC15804376EB91D2B65C233B9">Golovatch, Sergei I.</mods:namePart>
<mods:affiliation id="332FF35ABE3EDA820D8EBBE91E3D4EB0">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071 (Russia) sgol @ orc. ru</mods:affiliation>
<mods:nameIdentifier id="20A46D08FB6FA1BA50342B741113C629" type="email">sgol@orc.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="834A7EB34C914533B4A5BD849A19A0F0" type="personal">
<mods:role id="C096F55B66AAA39951F5883FCD20B864">
<mods:roleTerm id="837CE410BE8D928FF1FFF313269461DE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2D333056B3CD5D03E9F6F0B4EE2CB0C3">Geoffroy, Jean-Jacques</mods:namePart>
<mods:affiliation id="B0BC53D10A4CC9F33F2025031CCF4F8D">Muséum national d’Histoire naturelle, Département Écologie et Gestion de la Biodiversité, USM 306 Réseaux trophiques du sol, 4 avenue du Petit Château, F- 91800 Brunoy (France) geoffroy @ mnhn. fr</mods:affiliation>
<mods:nameIdentifier id="9AAB83BA24B804742F7C92C9CB29AD49" type="email">geoffroy@mnhn.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="F4F45F9CAA853142C81D3292AAA4A462" type="personal">
<mods:role id="2CBE2240443E1924CD61010CCEAACEB9">
<mods:roleTerm id="1E8310DAE12D12ECDB77A8C01876725D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F2F58DDB89C642E84FB8E7FEEBD55160">Mauriès, Jean-Paul</mods:namePart>
<mods:affiliation id="8570C5F81A5158F13DD439566F8FAEA0">Muséum national d’Histoire naturelle, Département Systématique et Évolution, USM 602, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr</mods:affiliation>
<mods:nameIdentifier id="A300674861AD318F6DDD12018068888C" type="email">collmill@mnhn.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="8F313389084221CB229E3EFFA9F86222" type="personal">
<mods:role id="E23487F5536C40A731E386E0ECB2F1C3">
<mods:roleTerm id="021D263351F9B45ACA86BB76D4E3B49A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2F2FA33F3E83E4C21E70D3E56A96B4A3">Spiegel, Didier Van Den</mods:namePart>
<mods:affiliation id="5988E276ECC577B808EC7CCACE86C69F">Musée royal de l’Afrique centrale, Leuvensesteenweg 13, B- 3080 Tervuren (Belgium) didier. van. den. spiegel @ africamuseum. be</mods:affiliation>
</mods:name>
<mods:typeOfResource id="BCA2EBFDE7292724B3ECC981EC98247C">text</mods:typeOfResource>
<mods:relatedItem id="8B773F457A3BD409312856AE888E8FFA" type="host">
<mods:titleInfo id="CE3B7B644BBAB4EDF6283B930DE4189B">
<mods:title id="F100D873436B213E2CBC9CE9F8073727">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="68B6763F7BF346B6D35991365C5E304B">
<mods:date id="424FF5F0AA5808C7BB654BD21BD0971A">2007</mods:date>
<mods:detail id="66102D78EAEB659C8D7C312A85B9578A" type="volume">
<mods:number id="D74604E52AB7FF3D1E138BDFC28EE822">29</mods:number>
</mods:detail>
<mods:detail id="3EE3B6F974CDD2C9872517D2308A2170" type="issue">
<mods:number id="9DCDBF9642EBBB00ADC27D4DF92AC112">1</mods:number>
</mods:detail>
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<mods:start id="76BD6121DE4ECCC91F3E78DD4ACAE598">7</mods:start>
<mods:end id="F25D78F76C55FB0632A9E5E4D4000B7A">49</mods:end>
</mods:extent>
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<mods:classification id="EA72821119B5BF2F74BF329C3B5596C1">journal article</mods:classification>
<mods:identifier id="E7472EADDC37FA8D4315022E7451D6D7" type="DOI">10.5281/zenodo.5392079</mods:identifier>
<mods:identifier id="66027AC07B43621FDE766BC7A5795ABD" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="02DA7D297157E17B31776DED5BBAF80B" type="Zenodo-Dep">5392079</mods:identifier>
</mods:mods>
<treatment id="03F4C378A23A305FFF6E02B6FCCAFAD6" LSID="urn:lsid:plazi:treatment:03F4C378A23A305FFF6E02B6FCCAFAD6" httpUri="http://treatment.plazi.org/id/03F4C378A23A305FFF6E02B6FCCAFAD6" lastPageId="23" lastPageNumber="30" pageId="21" pageNumber="28">
<subSubSection id="C34721E5A23A305DFF6E02B6FE18F994" box="[250,510,1515,1541]" pageId="21" pageNumber="28" type="nomenclature">
<paragraph id="8BE2726EA23A305DFF6E02B6FE18F994" blockId="21.[250,510,1515,1573]" box="[250,510,1515,1541]" pageId="21" pageNumber="28">
<heading id="D0AAC502A23A305DFF6E02B6FE18F994" box="[250,510,1515,1541]" centered="true" fontSize="11" level="2" pageId="21" pageNumber="28" reason="2">
<taxonomicName id="4C5D09EDA23A305DFF6E02B6FE5AF994" ID-CoL="3GGJJ" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[250,444,1515,1541]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="28" phylum="Arthropoda" rank="species" species="beroni" status="sp. nov.">
<emphasis id="B929AE7CA23A305DFF6E02B6FE5AF994" bold="true" box="[250,444,1515,1541]" italics="true" pageId="21" pageNumber="28">Glyphiulus beroni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A1307A23A305DFE5702B6FE18F994" box="[451,510,1515,1541]" pageId="21" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34721E5A23A305DFEA30156FE27F9B4" box="[311,449,1547,1573]" pageId="21" pageNumber="28" type="description">
<paragraph id="8BE2726EA23A305DFEA30156FE27F9B4" blockId="21.[250,510,1515,1573]" box="[311,449,1547,1573]" pageId="21" pageNumber="28">
(
<figureCitation id="13666EEBA23A305DFEAA0156FE76F9B4" box="[318,400,1547,1573]" captionStart="FIG" captionStartId="20.[113,124,1397,1413]" captionTargetBox="[122,1200,215,1354]" captionTargetId="figure-44@20.[391,662,783,974]" captionTargetPageId="20" captionText="FIG. 15. — Glyphiulus beroni n.sp., ♀ paratype:A, anterior part of body, lateral view; B, same, dorsofrontal view;C, middle part of body, dorsal view; D, same, lateral view; E, caudal part of body, dorsal view; F, same, ventral view. Scale bars: A-E, 0.5 mm; F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5392111" httpUri="https://zenodo.org/record/5392111/files/figure.png" pageId="21" pageNumber="28">Figs 15</figureCitation>
;
<figureCitation id="13666EEBA23A305DFE080156FE5EF9B4" box="[412,440,1547,1573]" captionStart="FIG" captionStartId="21.[113,124,1170,1186]" captionTargetBox="[112,1211,214,1130]" captionTargetId="figure-14@21.[112,1211,214,1130]" captionTargetPageId="21" captionText="FIG. 16. — Glyphiulus beroni n. sp., ♂ paratype: A, antenna; B, legs 1, caudal view; C, legs 2, caudal view; D, anterior gonopods, caudal view; E, posterior gonopods, caudal view; F, same, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5392113" httpUri="https://zenodo.org/record/5392113/files/figure.png" pageId="21" pageNumber="28">16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34721E5A23A305DFFE50109FC06FA8E" pageId="21" pageNumber="28" type="materials_examined">
<paragraph id="8BE2726EA23A305DFFE50109FD60F933" blockId="21.[111,647,1620,1698]" pageId="21" pageNumber="28">
<materialsCitation id="3B357833A23A305DFFE50109FF05F933" collectingDate="1989-01-11" collectionCode="NMNHS" collectorName="P. Beron" country="China" county="Jianshui County" location="Cave Ba Guo Dong" pageId="21" pageNumber="28" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
TYPE MATERIAL. —
<emphasis id="B929AE7CA23A305DFEB70109FE82F9FA" bold="true" box="[291,356,1620,1643]" pageId="21" pageNumber="28">
<collectingCountry id="F34A32FEA23A305DFEB70109FE86F9FA" box="[291,352,1620,1643]" name="China" pageId="21" pageNumber="28">China</collectingCountry>
.
</emphasis>
<collectingRegion id="4999BC8CA23A305DFEFC0109FE00F9FA" box="[360,486,1620,1643]" country="China" name="Yunnan" pageId="21" pageNumber="28">Yunnan Prov.</collectingRegion>
,
<collectingCounty id="62830AE2A23A305DFE780109FD65F9FA" box="[492,643,1620,1643]" pageId="21" pageNumber="28">Jianshui County</collectingCounty>
,
<location id="8E8224B5A23A305DFFFB012DFECCF916" LSID="urn:lsid:plazi:treatment:03F4C378A23A305FFF6E02B6FCCAFAD6:8E8224B5A23A305DFFFB012DFECCF916" box="[111,298,1648,1671]" country="China" county="Jianshui County" name="Cave Ba Guo Dong" pageId="21" pageNumber="28" stateProvince="Yunnan">Cave Ba Guo Dong</location>
,
<date id="FFE354AEA23A305DFEA6012DFE75F916" box="[306,403,1648,1671]" pageId="21" pageNumber="28" value="1989-01-11">
<collectingDate id="EFA7AD46A23A305DFEA6012DFE75F916" box="[306,403,1648,1671]" pageId="21" pageNumber="28" value="1989-01-11">11.I.1989</collectingDate>
</date>
, leg.
<collectorName id="26A817B8A23A305DFE55012DFDF6F916" box="[449,528,1648,1671]" pageId="21" pageNumber="28">P. Beron</collectorName>
,
<typeStatus id="54E6CCCCA23A305DFD8D012DFD8BF916" box="[537,621,1648,1671]" pageId="21" pageNumber="28" type="holotype">holotype</typeStatus>
<specimenCount id="9D5BB9E7A23A305DFDE6012FFD60F917" box="[626,646,1650,1670]" pageId="21" pageNumber="28" type="male">♂</specimenCount>
(
<collectionCode id="ED4CEAABA23A305DFFED01D6FF3BF933" box="[121,221,1675,1698]" pageId="21" pageNumber="28">NMNHS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B357833A23A305DFF7901D6FD64F933" box="[237,642,1675,1698]" collectingDate="1989-01-11" collectionCode="NMNHS" collectorName="P. Beron" country="China" county="Jianshui County" isTailEnumeration="true" location="Cave Ba Guo Dong" pageId="21" pageNumber="28" specimenCount="11" specimenCount-female="6" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54E6CCCCA23A305DFF7901D6FEAFF933" box="[237,329,1675,1698]" pageId="21" pageNumber="28" type="paratype">paratypes</typeStatus>
<specimenCount id="9D5BB9E7A23A305DFEDA01D6FE6FF933" box="[334,393,1675,1698]" pageId="21" pageNumber="28" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D5BB9E7A23A305DFE0101D6FE2CF933" box="[405,458,1675,1698]" pageId="21" pageNumber="28" type="female">6 ♀♀</specimenCount>
,
<specimenCount id="9D5BB9E7A23A305DFE4201D6FDEAF933" box="[470,524,1675,1698]" pageId="21" pageNumber="28" type="generic">3 juv.</specimenCount>
(
<collectionCode id="ED4CEAABA23A305DFD8301D6FD9DF933" box="[535,635,1675,1698]" pageId="21" pageNumber="28">NMNHS</collectionCode>
)
</materialsCitation>
;
</paragraph>
<paragraph id="8BE2726EA23A305DFD3103B3FC06FA8E" blockId="21.[677,1210,1262,1311]" pageId="21" pageNumber="28">
<materialsCitation id="3B357833A23A305DFD3103B3FC00FA94" box="[677,998,1262,1285]" collectionCode="MNHN" pageId="21" pageNumber="28" specimenCode="GA 038" specimenCount="4" specimenCount-female="2" specimenCount-male="2">
<specimenCount id="9D5BB9E7A23A305DFD3103B3FD06FA95" box="[677,736,1262,1285]" pageId="21" pageNumber="28" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D5BB9E7A23A305DFD7903B3FCC2FA95" box="[749,804,1262,1285]" pageId="21" pageNumber="28" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="ED4CEAABA23A305DFCA503B3FC63FA94" box="[817,901,1262,1285]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="28">MNHN</collectionCode>
<specimenCode id="DBFBDA15A23A305DFC1803B3FC39FA94" box="[908,991,1262,1285]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="21" pageNumber="28">GA 038</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B357833A23A305DFC6503B3FB53FA94" box="[1009,1205,1262,1285]" collectionCode="ZMUM" pageId="21" pageNumber="28" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D5BB9E7A23A305DFC6503B3FBFFFA95" box="[1009,1049,1262,1285]" pageId="21" pageNumber="28" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D5BB9E7A23A305DFBB203B3FBACFA95" box="[1062,1098,1262,1285]" pageId="21" pageNumber="28" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED4CEAABA23A305DFBCD03B3FB49FA94" box="[1113,1199,1262,1285]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="21" pageNumber="28">ZMUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B357833A23A305DFD310255FC87FA8E" box="[677,865,1288,1311]" collectionCode="ZMUC" pageId="21" pageNumber="28" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D5BB9E7A23A305DFD310255FD2AFA8E" box="[677,716,1288,1311]" pageId="21" pageNumber="28" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D5BB9E7A23A305DFD4C0255FD1AFA8E" box="[728,764,1288,1311]" pageId="21" pageNumber="28" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED4CEAABA23A305DFC9E0255FCBCFA8E" box="[778,858,1288,1311]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="21" pageNumber="28">ZMUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B357833A23A305DFCF80255FC3AFA8E" box="[876,988,1288,1311]" collectionCode="SEM" pageId="21" pageNumber="28" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D5BB9E7A23A305DFCF80255FC76FA8E" box="[876,912,1288,1311]" pageId="21" pageNumber="28" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED4CEAABA23A305DFC0A0255FC33FA8E" box="[926,981,1288,1311]" pageId="21" pageNumber="28">SEM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34721E5A23A305DFD310263FB40FAC4" box="[677,1190,1342,1365]" pageId="21" pageNumber="28" type="etymology">
<paragraph id="8BE2726EA23A305DFD310263FB40FAC4" blockId="21.[677,1190,1342,1365]" box="[677,1190,1342,1365]" pageId="21" pageNumber="28">ETYMOLOGY. — Honours Petar Beron, the collector.</paragraph>
</subSubSection>
<subSubSection id="C34721E5A23A305DFD31022EFCDAFA65" pageId="21" pageNumber="28" type="diagnosis">
<paragraph id="8BE2726EA23A305DFD31022EFCDAFA65" blockId="21.[675,1210,1395,1524]" pageId="21" pageNumber="28">DIAGNOSIS. — Differs by the crests just below and above the ozopore-bearing crests being divided into three, not two, transverse rows of tubercles, coupled with an evident mediodorsal tubercle present in front of the caudal edge of the epiproct.</paragraph>
</subSubSection>
<subSubSection id="C34721E5A23A305FFD310176FED5F994" lastPageId="23" lastPageNumber="30" pageId="21" pageNumber="28" type="description">
<paragraph id="8BE2726EA23A305DFD310176FCD3F9D2" blockId="21.[676,1209,1579,1700]" box="[677,821,1579,1605]" pageId="21" pageNumber="28">DESCRIPTION</paragraph>
<paragraph id="8BE2726EA23A305EFD310117FEA9F994" blockId="21.[676,1209,1579,1700]" lastBlockId="22.[113,646,1484,1700]" lastPageId="22" lastPageNumber="29" pageId="21" pageNumber="28">
Length of adults 11-17 (
<specimenCount id="9D5BB9E7A23A305DFC2B0110FC33F9F2" box="[959,981,1613,1635]" pageId="21" pageNumber="28" type="male">♂</specimenCount>
) or
<quantity id="4CA5DF8BA23A305DFB9D0117FB64F9F5" box="[1033,1154,1610,1636]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.4" metricValueMin="1.2" pageId="21" pageNumber="28" unit="mm" value="18.0" valueMax="24.0" valueMin="12.0">12-24 mm</quantity>
(
<specimenCount id="9D5BB9E7A23A305DFB010110FB4EF9F2" box="[1173,1192,1613,1635]" pageId="21" pageNumber="28" type="female">♀</specimenCount>
), width 0.8-1.0 (
<specimenCount id="9D5BB9E7A23A305DFCDC0130FCB8F912" box="[840,862,1645,1667]" pageId="21" pageNumber="28" type="male">♂</specimenCount>
) or
<quantity id="4CA5DF8BA23A305DFC1E0137FBEDF915" box="[906,1035,1642,1668]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.3" metricValueMin="0.8" pageId="21" pageNumber="28" unit="mm" value="1.05" valueMax="1.3" valueMin="0.8">0.8-1.3 mm</quantity>
(
<specimenCount id="9D5BB9E7A23A305DFB8E0130FBCBF912" box="[1050,1069,1645,1667]" pageId="21" pageNumber="28" type="female">♀</specimenCount>
). Coloration nearly entirely light yellowish, only ocellaria brown, as well as sometimes a vague lateral line level to ozoporiferous crests.
</paragraph>
<caption id="DF2222E6A239305EFFE5021CFEF2FAEE" ID-DOI="http://doi.org/10.5281/zenodo.5392115" ID-Zenodo-Dep="5392115" httpUri="https://zenodo.org/record/5392115/files/figure.png" pageId="22" pageNumber="29" startId="22.[113,124,1345,1361]" targetBox="[121,1201,215,1301]" targetPageId="22">
<paragraph id="8BE2726EA239305EFFE5021CFEF2FAEE" blockId="22.[113,1210,1344,1407]" pageId="22" pageNumber="29">
FIG. 17. —
<taxonomicName id="4C5D09EDA239305EFF5F021DFE89FAC0" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[203,367,1344,1361]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="29" phylum="Arthropoda" rank="species" species="bedosae" status="sp. nov.">
<emphasis id="B929AE7CA239305EFF5F021DFE89FAC0" box="[203,367,1344,1361]" italics="true" pageId="22" pageNumber="29">Glyphiulus bedosae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A1307A239305EFEE0021CFE42FAC0" box="[372,420,1345,1361]" pageId="22" pageNumber="29" rank="species">n. sp.</taxonomicNameLabel>
, ♀ paratype:
<emphasis id="B929AE7CA239305EFD86021DFDF9FAC0" bold="true" box="[530,543,1344,1361]" pageId="22" pageNumber="29">A</emphasis>
, anterior part of body, lateral view;
<emphasis id="B929AE7CA239305EFCD7021CFCB6FAC0" bold="true" box="[835,848,1345,1361]" pageId="22" pageNumber="29">B</emphasis>
, same, dorsal view;
<emphasis id="B929AE7CA239305EFC6F021DFBEFFAC0" bold="true" box="[1019,1033,1344,1361]" pageId="22" pageNumber="29">C</emphasis>
, middle part of body, lateral view;
<emphasis id="B929AE7CA239305EFF480205FF0CFAF9" bold="true" box="[220,234,1368,1384]" pageId="22" pageNumber="29">D</emphasis>
, same, dorsal view;
<emphasis id="B929AE7CA239305EFE0F0205FE41FAF9" bold="true" box="[411,423,1368,1384]" pageId="22" pageNumber="29">E</emphasis>
, caudal part of body, lateral view;
<emphasis id="B929AE7CA239305EFD440205FD3DFAF9" bold="true" box="[720,731,1368,1384]" pageId="22" pageNumber="29">F</emphasis>
, telson, ventral view. Scale bars: A, C, 0.5 mm; B, D, E, 0.2 mm; F, 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BE2726EA239305EFF1C0156FF0DF935" blockId="22.[113,646,1484,1700]" pageId="22" pageNumber="29">
Adults with 28-40p+5-2a+T (
<specimenCount id="9D5BB9E7A239305EFE5A0153FE02F9B5" box="[462,484,1550,1572]" pageId="22" pageNumber="29" type="male">♂</specimenCount>
) or 28-53p+5- 1a+T (
<specimenCount id="9D5BB9E7A239305EFF560173FF33F9D5" box="[194,213,1582,1604]" pageId="22" pageNumber="29" type="female">♀</specimenCount>
); largest juvenile
<specimenCount id="9D5BB9E7A239305EFE3F0173FE27F9D5" box="[427,449,1582,1604]" pageId="22" pageNumber="29" type="male">♂</specimenCount>
with 27p+5a+T, largest juvenile
<specimenCount id="9D5BB9E7A239305EFEB90110FEA7F9F2" box="[301,321,1613,1635]" pageId="22" pageNumber="29" type="female">♀</specimenCount>
with 27p+4a+T. Length of
<typeStatus id="54E6CCCCA239305EFFE50137FF31F915" box="[113,215,1642,1668]" pageId="22" pageNumber="29" type="holotype">holotype</typeStatus>
about
<quantity id="4CA5DF8BA239305EFEBA0137FE61F915" box="[302,391,1642,1668]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="22" pageNumber="29" unit="mm" value="17.0">17 mm</quantity>
, width 1.0 mm, with 40p+2a+T.
</paragraph>
<paragraph id="8BE2726EA239305EFD2F0291FBB9F935" blockId="22.[675,1211,1484,1700]" pageId="22" pageNumber="29">
Carinotaxy of collum typical (
<figureCitation id="13666EEBA239305EFC680291FB9DFA77" box="[1020,1147,1484,1510]" captionStart="FIG" captionStartId="20.[113,124,1397,1413]" captionTargetBox="[122,1200,215,1354]" captionTargetId="figure-44@20.[391,662,783,974]" captionTargetPageId="20" captionText="FIG. 15. — Glyphiulus beroni n.sp., ♀ paratype:A, anterior part of body, lateral view; B, same, dorsofrontal view;C, middle part of body, dorsal view; D, same, lateral view; E, caudal part of body, dorsal view; F, same, ventral view. Scale bars: A-E, 0.5 mm; F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5392111" httpUri="https://zenodo.org/record/5392111/files/figure.png" pageId="22" pageNumber="29">Fig. 15A, B</figureCitation>
), that of trunk segments generally typical but, starting from segment 7 until 6 or 7 penultimate segments, crests just below as well as between ozopores divided into three, not two, transverse rows of tubercles (
<figureCitation id="13666EEBA239305EFD3A0137FCD2F915" box="[686,820,1642,1668]" captionStart="FIG" captionStartId="20.[113,124,1397,1413]" captionTargetBox="[122,1200,215,1354]" captionTargetId="figure-44@20.[391,662,783,974]" captionTargetPageId="20" captionText="FIG. 15. — Glyphiulus beroni n.sp., ♀ paratype:A, anterior part of body, lateral view; B, same, dorsofrontal view;C, middle part of body, dorsal view; D, same, lateral view; E, caudal part of body, dorsal view; F, same, ventral view. Scale bars: A-E, 0.5 mm; F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5392111" httpUri="https://zenodo.org/record/5392111/files/figure.png" pageId="22" pageNumber="29">Fig. 15C, D</figureCitation>
). An evident middorsal tubercle just above caudal edge of epiproct (
<figureCitation id="13666EEBA239305EFC6601D7FBB5F935" box="[1010,1107,1674,1700]" captionStart="FIG" captionStartId="20.[113,124,1397,1413]" captionTargetBox="[122,1200,215,1354]" captionTargetId="figure-44@20.[391,662,783,974]" captionTargetPageId="20" captionText="FIG. 15. — Glyphiulus beroni n.sp., ♀ paratype:A, anterior part of body, lateral view; B, same, dorsofrontal view;C, middle part of body, dorsal view; D, same, lateral view; E, caudal part of body, dorsal view; F, same, ventral view. Scale bars: A-E, 0.5 mm; F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5392111" httpUri="https://zenodo.org/record/5392111/files/figure.png" pageId="22" pageNumber="29">Fig. 15E</figureCitation>
).
</paragraph>
<caption id="DF2222E6A238305FFFE503F8FF01FB72" ID-DOI="http://doi.org/10.5281/zenodo.5392117" ID-Zenodo-Dep="5392117" httpUri="https://zenodo.org/record/5392117/files/figure.png" pageId="23" pageNumber="30" startId="23.[113,124,1189,1205]" targetBox="[110,1225,182,1181]" targetPageId="23">
<paragraph id="8BE2726EA238305FFFE503F8FF01FB72" blockId="23.[112,1210,1188,1251]" pageId="23" pageNumber="30">
FIG. 18. —
<taxonomicName id="4C5D09EDA238305FFF5C03F9FE8FFB24" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[200,361,1188,1205]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="30" phylum="Arthropoda" rank="species" species="bedosae" status="sp. nov.">
<emphasis id="B929AE7CA238305FFF5C03F9FE8FFB24" box="[200,361,1188,1205]" italics="true" pageId="23" pageNumber="30">Glyphiulus bedosae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A1307A238305FFEF903F8FE7AFB24" box="[365,412,1189,1205]" pageId="23" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
, ♂ paratype:
<emphasis id="B929AE7CA238305FFD9C03F9FDF3FB24" bold="true" box="[520,533,1188,1205]" pageId="23" pageNumber="30">A</emphasis>
, antenna;
<emphasis id="B929AE7CA238305FFDFE03F8FD91FB24" bold="true" box="[618,631,1189,1205]" pageId="23" pageNumber="30">B</emphasis>
, gnathochilarium, caudal view;
<emphasis id="B929AE7CA238305FFCEC03F9FC60FB24" bold="true" box="[888,902,1188,1205]" pageId="23" pageNumber="30">C</emphasis>
, legs 1, caudal view;
<emphasis id="B929AE7CA238305FFBA603F8FBA6FB24" bold="true" box="[1074,1088,1189,1205]" pageId="23" pageNumber="30">D</emphasis>
, legs 2, caudal view;
<emphasis id="B929AE7CA238305FFF3603E1FF48FB5D" bold="true" box="[162,174,1212,1228]" pageId="23" pageNumber="30">E</emphasis>
, leg 3;
<emphasis id="B929AE7CA238305FFF7B03E1FF1CFB5D" bold="true" box="[239,250,1212,1228]" pageId="23" pageNumber="30">F</emphasis>
, midbody leg;
<emphasis id="B929AE7CA238305FFEED03E6FE61FB5D" bold="true" box="[377,391,1211,1228]" pageId="23" pageNumber="30">G</emphasis>
, anterior gonopods, caudal view;
<emphasis id="B929AE7CA238305FFD3303E1FD53FB5D" bold="true" box="[679,693,1212,1228]" pageId="23" pageNumber="30">H</emphasis>
, posterior gonopod, subcaudal view. Scale bar: A-F, 0.2 mm; G, H, 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BE2726EA238305FFF1C0253FED5F994" blockId="23.[113,649,1294,1541]" pageId="23" pageNumber="30">
Antennae (
<figureCitation id="13666EEBA238305FFF6F0253FE9FFAB9" box="[251,377,1294,1320]" captionStart="FIG" captionStartId="20.[113,124,1397,1413]" captionTargetBox="[122,1200,215,1354]" captionTargetId="figure-44@20.[391,662,783,974]" captionTargetPageId="20" captionText="FIG. 15. — Glyphiulus beroni n.sp., ♀ paratype:A, anterior part of body, lateral view; B, same, dorsofrontal view;C, middle part of body, dorsal view; D, same, lateral view; E, caudal part of body, dorsal view; F, same, ventral view. Scale bars: A-E, 0.5 mm; F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5392111" httpUri="https://zenodo.org/record/5392111/files/figure.png" pageId="23" pageNumber="30">Figs 15A, B</figureCitation>
;
<figureCitation id="13666EEBA238305FFE100253FE52FAB9" box="[388,436,1294,1320]" captionStart="FIG" captionStartId="21.[113,124,1170,1186]" captionTargetBox="[112,1211,214,1130]" captionTargetId="figure-14@21.[112,1211,214,1130]" captionTargetPageId="21" captionText="FIG. 16. — Glyphiulus beroni n. sp., ♂ paratype: A, antenna; B, legs 1, caudal view; C, legs 2, caudal view; D, anterior gonopods, caudal view; E, posterior gonopods, caudal view; F, same, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5392113" httpUri="https://zenodo.org/record/5392113/files/figure.png" pageId="23" pageNumber="30">16A</figureCitation>
), legs (
<figureCitation id="13666EEBA238305FFE6F0253FD9EFAB9" box="[507,632,1294,1320]" captionStart="FIG" captionStartId="21.[113,124,1170,1186]" captionTargetBox="[112,1211,214,1130]" captionTargetId="figure-14@21.[112,1211,214,1130]" captionTargetPageId="21" captionText="FIG. 16. — Glyphiulus beroni n. sp., ♂ paratype: A, antenna; B, legs 1, caudal view; C, legs 2, caudal view; D, anterior gonopods, caudal view; E, posterior gonopods, caudal view; F, same, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5392113" httpUri="https://zenodo.org/record/5392113/files/figure.png" pageId="23" pageNumber="30">Fig. 16B, C</figureCitation>
), claws, posterior gonopods (
<figureCitation id="13666EEBA238305FFE060270FDE8FAD6" box="[402,526,1325,1351]" captionStart="FIG" captionStartId="21.[113,124,1170,1186]" captionTargetBox="[112,1211,214,1130]" captionTargetId="figure-14@21.[112,1211,214,1130]" captionTargetPageId="21" captionText="FIG. 16. — Glyphiulus beroni n. sp., ♂ paratype: A, antenna; B, legs 1, caudal view; C, legs 2, caudal view; D, anterior gonopods, caudal view; E, posterior gonopods, caudal view; F, same, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5392113" httpUri="https://zenodo.org/record/5392113/files/figure.png" pageId="23" pageNumber="30">Fig. 16E, F</figureCitation>
), etc., as in
<taxonomicName id="4C5D09EDA238305FFFE60210FF19FAF6" baseAuthorityName="Gervais" baseAuthorityYear="1847" box="[114,255,1357,1383]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="30" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B929AE7CA238305FFFE60210FF19FAF6" box="[114,255,1357,1383]" italics="true" pageId="23" pageNumber="30">G. granulatus</emphasis>
</taxonomicName>
, but body size (often much larger), segment shape (slightly compressed laterally) and anterior gonopods (
<figureCitation id="13666EEBA238305FFEDF02D1FE56FA37" box="[331,432,1420,1446]" captionStart="FIG" captionStartId="21.[113,124,1170,1186]" captionTargetBox="[112,1211,214,1130]" captionTargetId="figure-14@21.[112,1211,214,1130]" captionTargetPageId="21" captionText="FIG. 16. — Glyphiulus beroni n. sp., ♂ paratype: A, antenna; B, legs 1, caudal view; C, legs 2, caudal view; D, anterior gonopods, caudal view; E, posterior gonopods, caudal view; F, same, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5392113" httpUri="https://zenodo.org/record/5392113/files/figure.png" pageId="23" pageNumber="30">Fig. 16D</figureCitation>
) more like those of
<taxonomicName id="4C5D09EDA238305FFFE702F1FED4FA57" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[115,306,1452,1478]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="30" phylum="Arthropoda" rank="species" species="semigranulatus" status="sp. nov.">
<emphasis id="B929AE7CA238305FFFE702F1FED4FA57" box="[115,306,1452,1478]" italics="true" pageId="23" pageNumber="30">G. semigranulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A1307A238305FFEAD02F1FE91FA57" box="[313,375,1452,1478]" pageId="23" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
Adults with 5-13 ocelli. Male legs 1 with tubercular leg vestiges near base of prongs (
<figureCitation id="13666EEBA238305FFF5302B6FEC0F994" box="[199,294,1515,1541]" captionStart="FIG" captionStartId="21.[113,124,1170,1186]" captionTargetBox="[112,1211,214,1130]" captionTargetId="figure-14@21.[112,1211,214,1130]" captionTargetPageId="21" captionText="FIG. 16. — Glyphiulus beroni n. sp., ♂ paratype: A, antenna; B, legs 1, caudal view; C, legs 2, caudal view; D, anterior gonopods, caudal view; E, posterior gonopods, caudal view; F, same, frontal view. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5392113" httpUri="https://zenodo.org/record/5392113/files/figure.png" pageId="23" pageNumber="30">Fig. 16B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34721E5A238305FFFE50176FCCAFAD6" pageId="23" pageNumber="30" type="discussion">
<paragraph id="8BE2726EA238305FFFE50176FF33F9D2" blockId="23.[111,646,1579,1700]" box="[113,213,1579,1605]" pageId="23" pageNumber="30">REMARKS</paragraph>
<paragraph id="8BE2726EA238305FFFE50117FCCAFAD6" blockId="23.[111,646,1579,1700]" lastBlockId="23.[679,1211,1294,1351]" pageId="23" pageNumber="30">In the absence of any clearly troglomorphic traits, except perhaps for its nearly completely unpigmented body and the somewhat reduced ocellaria, this species can hardly be considered as more than troglophilic.</paragraph>
</subSubSection>
</treatment>
</document>
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