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<document id="64DF92057E1F4F1191F4A3E9CC1EBC88" ID-CLB-Dataset="72867" ID-DOI="10.1080/00222933.2022.2046887" ID-GBIF-Dataset="93716e8b-7637-4345-8eaf-ecb3984baa19" ID-ISSN="1464-5262" ID-Zenodo-Dep="6530252" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1652081793889" checkinUser="plazi" docAuthor="Valois, Marcely C., Silva, Fernando A. B. &amp; Vaz-de-Mello, Fernando Z." docDate="2022" docId="03F487FF47714D159A63FDDCFBB1FC25" docLanguage="en" docName="JNATHIST.56.1-4.119-147.pdf" docOrigin="Journal of Natural History 56 (1 - 4)" docSource="http://dx.doi.org/10.1080/00222933.2022.2046887" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Dichotomius (Selenocopris) danieli Kohlmann and Solis 1997" docType="treatment" docVersion="7" lastPageNumber="132" masterDocId="FFCDFF87477D4D189B42FF9DFF91FFE2" masterDocTitle="A taxonomic revision of the globulus species group of Dichotomius Hope, 1838 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="147" masterPageNumber="119" pageNumber="131" updateTime="1699329890315" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F8022A0F561661C5B02976AE68E38997">A taxonomic revision of the globulus species group of Dichotomius Hope, 1838 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="14EC6F52E6D492F4E8531F133C0D0F7B">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<taxonomicName id="4C5D4D6A47714D149A63FDDCFB93FDB9" ID-CoL="35NBH" authority="Kohlmann and Solis, 1997" authorityName="Kohlmann and Solis" authorityYear="1997" box="[289,1026,577,603]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="danieli" subGenus="Selenocopris">
<emphasis id="B929EAFB47714D149A63FDDCFD58FDB9" bold="true" box="[289,713,577,603]" italics="true" pageId="12" pageNumber="131">Dichotomius (Selenocopris) danieli</emphasis>
<bibRefCitation id="EFCC4B1847714D149993FDDCFB93FDB9" author="Kohlmann B &amp; Solis A" box="[721,1026,577,603]" pageId="12" pageNumber="131" pagination="343 - 382" refId="ref14937" refString="Kohlmann B, Solis A. 1997. El genero Dichotomius en Costa Rica. Giorn. Ital. Entomol. 8: 343 - 382." type="journal article" year="1997">Kohlmann and Solís, 1997</bibRefCitation>
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<paragraph id="8BE236E947714D14996AFD1DFD6AFD78" blockId="12.[552,763,640,666]" box="[552,763,640,666]" pageId="12" pageNumber="131">
(
<figureCitation id="13662A6C47714D149972FD1DFD0DFD78" box="[560,668,640,666]" captionStart="Figure 2" captionStartId="5.[160,225,1505,1528]" captionTargetBox="[169,1149,142,1472]" captionTargetId="figure-3@5.[163,1153,142,1473]" captionTargetPageId="5" captionText="Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels." figureDoi="http://doi.org/10.5281/zenodo.6530256" httpUri="https://zenodo.org/record/6530256/files/figure.png" pageId="12" pageNumber="131">Figures 2</figureCitation>
(a–f), 6))
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<treatmentCitation id="0AFC10F847714D149B97FD28FC9DFD2C" author="Kohlmann B &amp; Solis A" box="[213,780,693,718]" page="345" pageId="12" pageNumber="131" year="1997">
<taxonomicName id="4C5D4D6A47714D149B97FD28FC9DFD2C" ID-CoL="35NBH" authority="Kohlmann and Solis 1997: 345" authorityName="Kohlmann and Solis" authorityPageNumber="345" authorityYear="1997" box="[213,780,693,718]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="danieli">
<emphasis id="B929EAFB47714D149B97FD28FE3EFD2C" box="[213,431,693,718]" italics="true" pageId="12" pageNumber="131">Dichotomius danieli</emphasis>
<bibRefCitation id="EFCC4B1847714D149AF4FD28FC9DFD2C" author="Kohlmann B &amp; Solis A" box="[438,780,693,718]" pageId="12" pageNumber="131" pagination="343 - 382" refId="ref14937" refString="Kohlmann B, Solis A. 1997. El genero Dichotomius en Costa Rica. Giorn. Ital. Entomol. 8: 343 - 382." type="journal article" year="1997">Kohlmann and Solís 1997: 345</bibRefCitation>
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, 362;
<treatmentCitation id="0AFC10F847714D14980EFD28FC6EFD2C" author="Krajcik M" box="[844,1023,693,718]" page="53" pageId="12" pageNumber="131" year="2006">
<bibRefCitation id="EFCC4B1847714D14980EFD28FC6EFD2C" author="Krajcik M" box="[844,1023,693,718]" pageId="12" pageNumber="131" pagination="1 - 189" refId="ref14964" refString="Krajcik M. 2006. Checklist of Scarabaeoidea of the World. 1. Scarabaeinae. Animma. X. Supplement 3: 1 - 189." type="journal article" year="2006">Krajcik 2006: 53</bibRefCitation>
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;
<treatmentCitation id="0AFC10F847714D149F4BFD28FBFBFD2C" author="Krajcik M" box="[1033,1130,693,718]" page="91" pageId="12" pageNumber="131" year="2012">
<bibRefCitation id="EFCC4B1847714D149F4BFD28FBFBFD2C" author="Krajcik M" box="[1033,1130,693,718]" pageId="12" pageNumber="131" pagination="278" refId="ref14991" refString="Krajcik M. 2012. Checklist of the World Scarabaeoidea. Anima. X Supplement 5: 278." type="journal article" year="2012">2012: 91</bibRefCitation>
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;
<treatmentCitation id="0AFC10F847714D149A6BFD4AFDF3FD12" author="Solis A &amp; Kohlmann B" box="[297,610,727,752]" page="6" pageId="12" pageNumber="131" year="2012">
<bibRefCitation id="EFCC4B1847714D149A6BFD4AFDF3FD12" author="Solis A &amp; Kohlmann B" box="[297,610,727,752]" pageId="12" pageNumber="131" pagination="1 - 32" refId="ref15591" refString="Solis A, Kohlmann B. 2012. Checklist and distribution atlas of the Scarabaeinae (Coleoptera: Scarabaeidae) of Costa Rica. Zootaxa. 3482 (1): 1 - 32." type="journal article" year="2012">Solís and Kohlmann 2012: 6</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AFC10F847714D14992CFD4AFC69FD12" author="Vaz-de-Mello FZ &amp; Nunes RV" box="[622,1016,727,752]" page="299" pageId="12" pageNumber="131" year="2016">
<bibRefCitation id="EFCC4B1847714D14992CFD4AFC69FD12" author="Vaz-de-Mello FZ &amp; Nunes RV" box="[622,1016,727,752]" pageId="12" pageNumber="131" pagination="294 - 300" refId="ref15886" refString="Vaz-de-Mello FZ, Nunes RV. 2016. New species, redescription and taxonomic notes in the Dichotomius (Selenocopris) batesi (Harold) species-group (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa. 4139 (2): 294 - 300." type="journal article" year="2016">Vaz-de-Mello and Nunes 2016: 299</bibRefCitation>
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.
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</paragraph>
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<subSubSection id="C347656247714D149BE2FC83FDAAFB92" pageId="12" pageNumber="131" type="diagnosis">
<paragraph id="8BE236E947714D149BE2FC83FE85FCDA" blockId="12.[160,1156,798,1136]" box="[160,276,798,824]" pageId="12" pageNumber="131">
<emphasis id="B929EAFB47714D149BE2FC83FE85FCDA" bold="true" box="[160,276,798,824]" italics="true" pageId="12" pageNumber="131">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BE236E947714D149BE2FCDFFDAAFB92" blockId="12.[160,1156,798,1136]" pageId="12" pageNumber="131">
<emphasis id="B929EAFB47714D149BE2FCDFFD91FCB9" box="[160,512,834,859]" italics="true" pageId="12" pageNumber="131">
<taxonomicName id="4C5D4D6A47714D149BE2FCDFFEECFCB9" authorityName="Kohlmann and Solis" authorityYear="1997" box="[160,381,834,859]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="danieli">Dichotomius danieli</taxonomicName>
,
<taxonomicName id="4C5D4D6A47714D149AC9FCDFFD91FCB9" authorityName="Kohlmann and Solis" authorityYear="1997" box="[395,512,834,859]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="rodrigoi">D. rodrigoi</taxonomicName>
</emphasis>
and
<taxonomicName id="4C5D4D6A47714D14997DFCDFFD16FCB9" authorityName="Kohlmann and Solis" authorityYear="1997" box="[575,647,834,859]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="favi">
<emphasis id="B929EAFB47714D14997DFCDFFD16FCB9" box="[575,647,834,859]" italics="true" pageId="12" pageNumber="131">D. favi</emphasis>
</taxonomicName>
are the only known species of the
<taxonomicName id="4C5D4D6A47714D149F65FCDFFB12FCB9" baseAuthorityName="Felsche" baseAuthorityYear="1901" box="[1063,1155,834,859]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="globulus">
<emphasis id="B929EAFB47714D149F65FCDFFB12FCB9" box="[1063,1155,834,859]" italics="true" pageId="12" pageNumber="131">globulus</emphasis>
</taxonomicName>
group recorded in
<collectingCountry id="F34A767947714D149A3CFCF8FE66FC9C" box="[382,503,869,894]" name="Costa Rica" pageId="12" pageNumber="131">Costa Rica</collectingCountry>
. As pre-apical teeth are absent on the dorsal surface of paramera,
<taxonomicName id="4C5D4D6A47714D149A5AFC1AFEEEFC42" authorityName="Kohlmann and Solis" authorityYear="1997" box="[280,383,903,928]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="danieli">
<emphasis id="B929EAFB47714D149A5AFC1AFEEEFC42" box="[280,383,903,928]" italics="true" pageId="12" pageNumber="131">D. danieli</emphasis>
</taxonomicName>
is suggested to be more closely related to
<taxonomicName id="4C5D4D6A47714D149826FC1AFC3BFC42" authorityName="Kohlmann and Solis" authorityYear="1997" box="[868,938,903,928]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="favi">
<emphasis id="B929EAFB47714D149826FC1AFC3BFC42" box="[868,938,903,928]" italics="true" pageId="12" pageNumber="131">D. favi</emphasis>
</taxonomicName>
than to
<taxonomicName id="4C5D4D6A47714D149F49FC1AFBEFFC42" authorityName="Kohlmann and Solis" authorityYear="1997" box="[1035,1150,903,928]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="rodrigoi">
<emphasis id="B929EAFB47714D149F49FC1AFBEFFC42" box="[1035,1150,903,928]" italics="true" pageId="12" pageNumber="131">D. rodrigoi</emphasis>
</taxonomicName>
. The following characters can be used to distinguish
<taxonomicName id="4C5D4D6A47714D14984AFC37FCE2FC21" authorityName="Kohlmann and Solis" authorityYear="1997" box="[776,883,938,963]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="danieli">
<emphasis id="B929EAFB47714D14984AFC37FCE2FC21" box="[776,883,938,963]" italics="true" pageId="12" pageNumber="131">D. danieli</emphasis>
</taxonomicName>
from
<taxonomicName id="4C5D4D6A47714D1498FFFC37FB96FC21" authorityName="Kohlmann and Solis" authorityYear="1997" box="[957,1031,938,963]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="favi">
<emphasis id="B929EAFB47714D1498FFFC37FB96FC21" box="[957,1031,938,963]" italics="true" pageId="12" pageNumber="131">D. favi</emphasis>
</taxonomicName>
and other species in the
<taxonomicName id="4C5D4D6A47714D149A0BFC50FE34FC04" baseAuthorityName="Felsche" baseAuthorityYear="1901" box="[329,421,973,998]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="131" phylum="Arthropoda" rank="species" species="globulus">
<emphasis id="B929EAFB47714D149A0BFC50FE34FC04" box="[329,421,973,998]" italics="true" pageId="12" pageNumber="131">globulus</emphasis>
</taxonomicName>
group: surface of head covered with dense ocellate punctures; pronotal punctures equally spaced from each other; anteromedian region of pronotum with a feeble central impression; elytral striae wide, crenulate, punctures almost as wide as striae; elytral interstriae opaque with indistinct smooth dots; paramera rectangular, apex rounded in dorsal view (
<figureCitation id="13662A6C47714D149AF1FBCAFD81FB92" box="[435,528,1111,1136]" captionStart="Figure 2" captionStartId="5.[160,225,1505,1528]" captionTargetBox="[169,1149,142,1472]" captionTargetId="figure-3@5.[163,1153,142,1473]" captionTargetPageId="5" captionText="Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels." figureDoi="http://doi.org/10.5281/zenodo.6530256" httpUri="https://zenodo.org/record/6530256/files/figure.png" pageId="12" pageNumber="131">Figure 2</figureCitation>
(d)).
</paragraph>
</subSubSection>
<subSubSection id="C347656247714D159BE2FB03FD67FD17" lastPageId="13" lastPageNumber="132" pageId="12" pageNumber="131" type="description">
<paragraph id="8BE236E947714D149BE2FB03FE03FB5A" blockId="12.[160,1156,1182,1728]" box="[160,402,1182,1208]" pageId="12" pageNumber="131">
<emphasis id="B929EAFB47714D149BE2FB03FE03FB5A" bold="true" box="[160,402,1182,1208]" italics="true" pageId="12" pageNumber="131">Redescription (male)</emphasis>
</paragraph>
<paragraph id="8BE236E947714D159BE2FB5FFD67FD17" blockId="12.[160,1156,1182,1728]" lastBlockId="13.[160,1156,142,757]" lastPageId="13" lastPageNumber="132" pageId="12" pageNumber="131">
<emphasis id="B929EAFB47714D149BE2FB5FFEA1FB39" bold="true" box="[160,304,1218,1243]" pageId="12" pageNumber="131">Body length</emphasis>
:
<quantity id="4CA59B0C47714D149A7FFB5FFE03FB39" box="[317,402,1218,1243]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="12" pageNumber="131" unit="mm" value="9.5">9.5 mm</quantity>
. Dorsum black, barely shiny (
<figureCitation id="13662A6C47714D149997FB5FFCBEFB39" box="[725,815,1218,1243]" captionStart="Figure 2" captionStartId="5.[160,225,1505,1528]" captionTargetBox="[169,1149,142,1472]" captionTargetId="figure-3@5.[163,1153,142,1473]" captionTargetPageId="5" captionText="Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels." figureDoi="http://doi.org/10.5281/zenodo.6530256" httpUri="https://zenodo.org/record/6530256/files/figure.png" pageId="12" pageNumber="131">Figure 2</figureCitation>
(a)).
<emphasis id="B929EAFB47714D14981CFB5FFC0DFB39" bold="true" box="[862,924,1218,1243]" pageId="12" pageNumber="131">Head</emphasis>
: clypeal edge bidentate, teeth acute, separated by a deep V-shaped emargination. Clypeogenal junction rounded. Surface with different-sized ocellate punctures; punctures forming transverse rows at anterior clypeal region. Clypeogenal suture present. Cephalic process produced into a small central tubercle (
<figureCitation id="13662A6C47714D149AB7FAD0FDC3FA84" box="[501,594,1357,1382]" captionStart="Figure 2" captionStartId="5.[160,225,1505,1528]" captionTargetBox="[169,1149,142,1472]" captionTargetId="figure-3@5.[163,1153,142,1473]" captionTargetPageId="5" captionText="Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels." figureDoi="http://doi.org/10.5281/zenodo.6530256" httpUri="https://zenodo.org/record/6530256/files/figure.png" pageId="12" pageNumber="131">Figure 2</figureCitation>
(a)). Antennal club with light brown tumescence.
<emphasis id="B929EAFB47714D149BE2FAF2FF63FA6A" bold="true" box="[160,242,1391,1416]" pageId="12" pageNumber="131">Thorax</emphasis>
: pronotum approximately twice wider than long. Pronotal disc convex, anteromedian region with a shallow central excavation; surface with ocellate punctures as dense as those on anterior angles (
<figureCitation id="13662A6C47714D149A83FA28FD8AFA2C" box="[449,539,1461,1486]" captionStart="Figure 2" captionStartId="5.[160,225,1505,1528]" captionTargetBox="[169,1149,142,1472]" captionTargetId="figure-3@5.[163,1153,142,1473]" captionTargetPageId="5" captionText="Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels." figureDoi="http://doi.org/10.5281/zenodo.6530256" httpUri="https://zenodo.org/record/6530256/files/figure.png" pageId="12" pageNumber="131">Figure 2</figureCitation>
(a)). Surface of hypomera with sparse, coarse setiferous punctures; setae short, and usually absent in some punctures. Prosternum filled with ocellate punctures. Mesoventrite filled with ocellate punctures usually bearing short, orange setae. Metaventrite surface with ocellate punctures, except on disc; punctures on anterior region usually bearing short setae. Meso-metaventral suture distinct. Mid-longitudinal sulcus feebly marked posteriorly. Lateral lobes of metaventrite with ocellate punctures larger than those on anterior region.
<emphasis id="B929EAFB47714D14997EF918FD11F97C" bold="true" box="[572,640,1669,1694]" pageId="12" pageNumber="131">Elytra</emphasis>
: slightly convex in lateral view. Humeral callus distinct. Striae thin, with well-defined punctures spaced by their diameter. Interstriae I–VII slight convex, barely shiny, with shagreened microsculpture; medial region with distinct smooth dots.
<emphasis id="B929EAFB47704D159A79FF2CFEE0FF28" bold="true" box="[315,369,177,202]" pageId="13" pageNumber="132">Legs</emphasis>
: protibiae with three distinct teeth externally. Protibial spur curved at apical fifth. Metatibial spur deeply emarginate apically. Ventral surface of the profemora covered with coarse setiferous punctures along its entire extension, setae short. Ventral surface of the mesofemora with ocellate punctures at apical half. Ventral surface of the metafemora with ocellate punctures concentrated on apical half; basal half with sparse, scattered punctures. Meso and metafemora bearing a thin longitudinal groove along the entire extension of posterior margin, wider on apical half.
<emphasis id="B929EAFB47704D159853FE1CFC14FE78" bold="true" box="[785,901,385,410]" pageId="13" pageNumber="132">Abdomen</emphasis>
: ventrites covered with ocellate punctures laterally, punctures medially disposed in a single row along anterior edge. Pygidium twice wider than long, disc strongly convex apically. Surface of pygidium with ocellate punctures concentrated along anterior half, and spaced by half their diameter. Apex of the pygidium margined.
<emphasis id="B929EAFB47704D159952FD91FDE1FDC7" bold="true" box="[528,624,524,549]" pageId="13" pageNumber="132">Tegmen</emphasis>
: in dorsal view, paramera rectangular, one-third longer than its width medially, apex truncate; inner edges of paramera curved outward along its whole length (
<figureCitation id="13662A6C47704D159AE0FDCCFE6DFD88" box="[418,508,593,618]" captionStart="Figure 2" captionStartId="5.[160,225,1505,1528]" captionTargetBox="[169,1149,142,1472]" captionTargetId="figure-3@5.[163,1153,142,1473]" captionTargetPageId="5" captionText="Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels." figureDoi="http://doi.org/10.5281/zenodo.6530256" httpUri="https://zenodo.org/record/6530256/files/figure.png" pageId="13" pageNumber="132">Figure 2</figureCitation>
(d)). Dorsal surface of paramera smooth (
<figureCitation id="13662A6C47704D1598F5FDCCFB80FD88" box="[951,1041,593,618]" captionStart="Figure 2" captionStartId="5.[160,225,1505,1528]" captionTargetBox="[169,1149,142,1472]" captionTargetId="figure-3@5.[163,1153,142,1473]" captionTargetPageId="5" captionText="Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels." figureDoi="http://doi.org/10.5281/zenodo.6530256" httpUri="https://zenodo.org/record/6530256/files/figure.png" pageId="13" pageNumber="132">Figure 2</figureCitation>
(d)). Apical half noticeably narrower than base in lateral view (
<figureCitation id="13662A6C47704D1599FCFDE9FC86FD6F" box="[702,791,628,653]" captionStart="Figure 2" captionStartId="5.[160,225,1505,1528]" captionTargetBox="[169,1149,142,1472]" captionTargetId="figure-3@5.[163,1153,142,1473]" captionTargetPageId="5" captionText="Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels." figureDoi="http://doi.org/10.5281/zenodo.6530256" httpUri="https://zenodo.org/record/6530256/files/figure.png" pageId="13" pageNumber="132">Figure 2</figureCitation>
(f)).Ventral sclerotised membrane between paramera thinner and curved at basal half; rounded laterally (
<figureCitation id="13662A6C47704D1598E0FD0BFC6AFD4D" box="[930,1019,662,687]" captionStart="Figure 2" captionStartId="5.[160,225,1505,1528]" captionTargetBox="[169,1149,142,1472]" captionTargetId="figure-3@5.[163,1153,142,1473]" captionTargetPageId="5" captionText="Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels." figureDoi="http://doi.org/10.5281/zenodo.6530256" httpUri="https://zenodo.org/record/6530256/files/figure.png" pageId="13" pageNumber="132">Figure 2</figureCitation>
(e)).
<emphasis id="B929EAFB47704D159F6AFD0BFBECFD4D" bold="true" box="[1064,1149,662,687]" pageId="13" pageNumber="132">Female</emphasis>
: differs from male by clypeal teeth acute and separated by a deep V-shaped emargination, and sixth abdominal ventrite 3 times longer than fifth.
</paragraph>
</subSubSection>
<subSubSection id="C347656247704D159BE2FCBFFBB1FC25" pageId="13" pageNumber="132" type="materials_examined">
<paragraph id="8BE236E947704D159BE2FCBFFE2EFCDE" blockId="13.[160,1156,802,967]" box="[160,447,802,828]" pageId="13" pageNumber="132">
<emphasis id="B929EAFB47704D159BE2FCBFFE2EFCDE" bold="true" box="[160,447,802,828]" italics="true" pageId="13" pageNumber="132">Type material examined</emphasis>
</paragraph>
<paragraph id="8BE236E947704D159BE2FCDBFBB1FC25" blockId="13.[160,1156,802,967]" pageId="13" pageNumber="132">
<typeStatus id="54E6884B47704D159BE2FCDBFE96FCBD" box="[160,263,838,863]" pageId="13" pageNumber="132" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C5D4D6A47704D159A69FCDBFD95FCBD" authorityName="Kohlmann and Solis" authorityYear="1997" box="[299,516,838,863]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="132" phylum="Arthropoda" rank="species" species="danieli">
<emphasis id="B929EAFB47704D159A69FCDBFD95FCBD" box="[299,516,838,863]" italics="true" pageId="13" pageNumber="132">Dichotomius danieli</emphasis>
</taxonomicName>
<specimenCount id="9D5BFD6047704D159948FCDBFD8FFCBF" box="[522,542,838,861]" pageId="13" pageNumber="132" type="male">♂</specimenCount>
[
<collectionCode id="ED4CAE2C47704D159968FCDBFDE6FCBD" box="[554,631,838,863]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="13" pageNumber="132">MNCR</collectionCode>
]: ‘Est. Pitilla,
<quantity id="4CA59B0C47704D159840FCDBFCABFCBD" box="[770,826,838,863]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="132" unit="km" value="9.0">9 km</quantity>
S. Santa Cecilia, Prov./Guana,
<collectingCountry id="F34A767947704D159BE2FCF5FEBDFC63" box="[160,300,872,897]" name="Costa Rica" pageId="13" pageNumber="132">COSTA RICA</collectingCountry>
,
<specimenCount id="9D5BFD6047704D159A75FCF5FE14FC63" box="[311,389,872,897]" pageId="13" pageNumber="132" type="generic">
<quantity id="4CA59B0C47704D159A75FCF5FE10FC63" box="[311,385,872,897]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="132" unit="m" value="700.0">700 m</quantity>
.
</specimenCount>
<date id="FFE3102947704D159ACFFCF5FE6DFC63" box="[397,508,872,897]" pageId="13" pageNumber="132" value="1994-08">Ago 1994</date>
/C. Moranga, L N 330200_380200 # 3198’ // ‘
<taxonomicName id="4C5D4D6A47704D1598B0FCF5FF7AFC46" authorityName="Kohlmann and Solis" authorityYear="1997" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="132" phylum="Arthropoda" rank="species" species="danieli">Dichotomius danieli</taxonomicName>
/Kohlmann and Solís’/‘
<collectingCountry id="F34A767947704D159A9CFC16FDFAFC46" box="[478,619,907,932]" name="Costa Rica" pageId="13" pageNumber="132">COSTA RICA</collectingCountry>
INBIO/[barcode] CRI002/029748’ // ‘HOLOTYPO/
<taxonomicName id="4C5D4D6A47704D159BE2FC33FE10FC25" authorityName="Kohlmann and Solis" authorityYear="1997" box="[160,385,942,967]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="132" phylum="Arthropoda" rank="species" species="danieli">Dichotomius danieli</taxonomicName>
/Kohlmann y Solís’. Examined via photographs (
<figureCitation id="13662A6C47704D1598D9FC33FC66FC25" box="[923,1015,942,967]" captionStart="Figure 2" captionStartId="5.[160,225,1505,1528]" captionTargetBox="[169,1149,142,1472]" captionTargetId="figure-3@5.[163,1153,142,1473]" captionTargetPageId="5" captionText="Figure 2. Holotypes. Dichotomius danieli, male: (a,b) Dorsal and ventral views; (c) labels; (d–f) tegmen in dorsal, ventral and lateral views; D. favi, male: (g,h) dorsal and ventral views; (i) labels; (j–l) tegmen in dorsal, ventral and lateral views. D. rodrigoi, male: (m,n) dorsal and ventral views; (o) labels; (p–r) tegmen in dorsal, ventral view and lateral views. Lectotype D. globulus, male: (s,t) dorsal and ventral views; (u) labels." figureDoi="http://doi.org/10.5281/zenodo.6530256" httpUri="https://zenodo.org/record/6530256/files/figure.png" pageId="13" pageNumber="132">Figure 2</figureCitation>
(c)).
</paragraph>
</subSubSection>
</treatment>
</document>
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