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<document id="DF95656DF43EA4B65F5CDF801B0300EF" ID-DOI="10.5281/zenodo.3674835" ID-GBIF-Dataset="dde4db22-fb4f-4e91-b752-106ccadf9f90" ID-ISSN="1942-1354" ID-Zenodo-Dep="3674835" ID-ZooBank="D96A51F7-FA29-4CE5-A5CC-06BBDA2FEA37" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582111322982" checkinUser="felipe" docAuthor="Rázuri-Gonzales, Ernesto &amp; Armitage, Brian J." docDate="2019" docId="03F4364B126FFFABA287F223FBE1797C" docLanguage="en" docName="InsectaMundi.0710.1-13.pdf.imf" docOrigin="Insecta Mundi 710 (710)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Smicridea (Smicridea) dividua Razuri-Gonzales and Armitage 2019, sp. n." docType="treatment" docVersion="8" lastPageNumber="7" masterDocId="FFCD4E331268FFA3A247F04EFFA37848" masterDocTitle="The Trichoptera of Panama. XI. Three new species of caddisflies in the genus Smicridea McLachlan (Trichoptera: Hydropsychidae) from Omar Torrijos and Santa Fe National Parks" masterLastPageNumber="13" masterPageNumber="1" pageNumber="6" updateTime="1698719366723" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="723B026D8EF5C61B87F21A64637732CA">The Trichoptera of Panama. XI. Three new species of caddisflies in the genus Smicridea McLachlan (Trichoptera: Hydropsychidae) from Omar Torrijos and Santa Fe National Parks</mods:title>
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<mods:namePart id="AF51F9BDC1BFCB2AA7ED78D1D6164E8F">Rázuri-Gonzales, Ernesto</mods:namePart>
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<mods:namePart id="BB42833E3BD14F035F820B18B5D95355">Armitage, Brian J.</mods:namePart>
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<mods:title id="CA958656DCC58E3198879327571D9B35">Insecta Mundi</mods:title>
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<mods:date id="DB03F147D9CDD42E8648268DD315F494">2019</mods:date>
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<treatment id="03F4364B126FFFABA287F223FBE1797C" ID-DOI="http://doi.org/10.5281/zenodo.3680634" ID-GBIF-Taxon="162173221" ID-Zenodo-Dep="3680634" LSID="urn:lsid:plazi:treatment:03F4364B126FFFABA287F223FBE1797C" httpUri="http://treatment.plazi.org/id/03F4364B126FFFABA287F223FBE1797C" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6">
<subSubSection id="C347D4D6126FFFA4A287F223FBCE7ACE" box="[192,1133,621,646]" pageId="7" pageNumber="6" type="nomenclature">
<paragraph id="8BE2875D126FFFA4A287F223FBCE7ACE" blockId="7.[192,1133,621,679]" box="[192,1133,621,646]" pageId="7" pageNumber="6">
<heading id="D0AA3031126FFFA4A287F223FBCE7ACE" bold="true" box="[192,1133,621,646]" fontSize="10" level="2" pageId="7" pageNumber="6" reason="3">
<emphasis id="B9295B4F126FFFA4A287F223FBCE7ACE" bold="true" box="[192,1133,621,646]" pageId="7" pageNumber="6">
<taxonomicName id="4C5DFCDE126FFFA4A287F223FBBA7ACE" authority="Razuri-Gonzales and Armitage" authorityName="Razuri-Gonzales and Armitage" authorityYear="2019" box="[192,1049,621,646]" class="Insecta" family="Hydropsychidae" genus="Smicridea" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="dividua" status="sp. nov." subGenus="Smicridea">
<emphasis id="B9295B4F126FFFA4A287F223FEEE7ACD" bold="true" box="[192,333,621,645]" italics="true" pageId="7" pageNumber="6">Smicridea</emphasis>
(
<emphasis id="B9295B4F126FFFA4A318F223FE4F7ACD" bold="true" box="[351,492,621,645]" italics="true" pageId="7" pageNumber="6">Smicridea</emphasis>
)
<emphasis id="B9295B4F126FFFA4A3B9F223FDCA7ACD" bold="true" box="[510,617,621,645]" italics="true" pageId="7" pageNumber="6">dividua</emphasis>
Rázuri-Gonzales and Armitage
</taxonomicName>
,
<taxonomicNameLabel id="A21AE634126FFFA4A661F220FBCE7ACE" box="[1062,1133,622,646]" pageId="7" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C347D4D6126FFFA4A287F2C0FEAE7AEF" box="[192,269,654,679]" pageId="7" pageNumber="6" type="description">
<paragraph id="8BE2875D126FFFA4A287F2C0FEAE7AEF" blockId="7.[192,1133,621,679]" box="[192,269,654,679]" pageId="7" pageNumber="6">
<figureCitation id="13669BD8126FFFA4A287F2C0FEAB7AEF" box="[192,264,654,679]" captionStart="Figure 3" captionStartId="13.[192,275,1846,1868]" captionTargetBox="[307,1333,221,1809]" captionTargetId="figure@13.[297,1334,212,1812]" captionTargetPageId="13" captionText="Figure 3. Smicridea (Smicridea) dividua Rázuri-Gonzales and Armitage, sp. n., male genitalia. A) Lateral view. B) Dorsal view. C) Left inferior appendage, ventral view. D) Phallic apparatus, lateral view (inset: posterodorsal margin projection, dorsal view). E) Phallic apparatus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3674841" httpUri="https://zenodo.org/record/3674841/files/figure.png" pageId="7" pageNumber="6">Fig. 3</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C347D4D6126FFFA4A287F2F1FABC7BD2" pageId="7" pageNumber="6" type="diagnosis">
<paragraph id="8BE2875D126FFFA4A287F2F1FABC7BD2" blockId="7.[192,1440,703,922]" pageId="7" pageNumber="6">
<emphasis id="B9295B4F126FFFA4A287F2F1FEEC7A9F" bold="true" box="[192,335,703,727]" pageId="7" pageNumber="6">Diagnosis.</emphasis>
<taxonomicName id="4C5DFCDE126FFFA4A311F2F1FDCD7A9F" authority="Rázuri-Gonzales &amp; Armitage, 2019" authorityName="Rázuri-Gonzales &amp; Armitage" authorityYear="2019" box="[342,622,703,727]" class="Insecta" family="Hydropsychidae" genus="Smicridea" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="dividua" status="sp. nov." subGenus="Smicridea">
<emphasis id="B9295B4F126FFFA4A311F2F1FE777A9F" box="[342,468,703,727]" italics="true" pageId="7" pageNumber="6">Smicridea</emphasis>
(
<emphasis id="B9295B4F126FFFA4A3A2F2F1FE5D7A9E" box="[485,510,703,726]" italics="true" pageId="7" pageNumber="6">S.</emphasis>
)
<emphasis id="B9295B4F126FFFA4A048F2F1FDCD7A9F" box="[527,622,703,727]" italics="true" pageId="7" pageNumber="6">dividua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21AE634126FFFA4A031F2F1FD1B7A9E" box="[630,696,703,726]" pageId="7" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
is most similar to
<taxonomicName id="4C5DFCDE126FFFA4A1E7F2F1FB6D7A9F" authority="Flint, 1974" authorityName="Flint" authorityYear="1974" box="[928,1230,703,727]" class="Insecta" family="Hydropsychidae" genus="Smicridea" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="cholta" subGenus="Smicridea">
<emphasis id="B9295B4F126FFFA4A1E7F2F1FC1A7A9E" box="[928,953,703,726]" italics="true" pageId="7" pageNumber="6">S.</emphasis>
(
<emphasis id="B9295B4F126FFFA4A18DF2F1FC407A9E" box="[970,995,703,726]" italics="true" pageId="7" pageNumber="6">S.</emphasis>
)
<emphasis id="B9295B4F126FFFA4A1B3F2F1FB9C7A9F" box="[1012,1087,703,727]" italics="true" pageId="7" pageNumber="6">cholta</emphasis>
Flint, 1974
</taxonomicName>
from
<collectingCountry id="F34AC7CD126FFFA4A750F2F1FA037A9F" box="[1303,1440,703,727]" name="Guatemala" pageId="7" pageNumber="6">Guatemala</collectingCountry>
and
<collectingCountry id="F34AC7CD126FFFA4A2B0F291FED47ABF" box="[247,375,735,759]" name="Nicaragua" pageId="7" pageNumber="6">Nicaragua</collectingCountry>
based on the shape of tergum X and the phallic apparatus in ventral view. However,
<taxonomicName id="4C5DFCDE126FFFA4A287F34EFE757B50" authorityName="Rázuri-Gonzales &amp; Armitage" authorityYear="2019" box="[192,470,768,792]" class="Insecta" family="Hydropsychidae" genus="Smicridea" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="dividua" subGenus="Smicridea">
<emphasis id="B9295B4F126FFFA4A287F34EFE9D7B50" box="[192,318,768,792]" italics="true" pageId="7" pageNumber="6">Smicridea</emphasis>
(
<emphasis id="B9295B4F126FFFA4A309F34EFEC47B5F" box="[334,359,768,791]" italics="true" pageId="7" pageNumber="6">S.</emphasis>
)
<emphasis id="B9295B4F126FFFA4A330F34EFE757B50" box="[375,470,768,792]" italics="true" pageId="7" pageNumber="6">dividua</emphasis>
</taxonomicName>
is easily separated from
<taxonomicName id="4C5DFCDE126FFFA4A14FF34EFC067B50" authorityName="Flint" authorityYear="1974" box="[776,933,768,792]" class="Insecta" family="Hydropsychidae" genus="Smicridea" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="cholta" subGenus="Smicridea">
<emphasis id="B9295B4F126FFFA4A14FF34EFC827B5F" box="[776,801,768,791]" italics="true" pageId="7" pageNumber="6">S.</emphasis>
(
<emphasis id="B9295B4F126FFFA4A176F34EFCE87B5F" box="[817,843,768,791]" italics="true" pageId="7" pageNumber="6">S.</emphasis>
)
<emphasis id="B9295B4F126FFFA4A11DF34EFC067B50" box="[858,933,768,792]" italics="true" pageId="7" pageNumber="6">cholta</emphasis>
</taxonomicName>
due to the bilobed posteroventral process of the phallobase, which is broad and entire in
<taxonomicName id="4C5DFCDE126FFFA4A0BCF36FFC347B70" authorityName="Flint" authorityYear="1974" box="[763,919,800,825]" class="Insecta" family="Hydropsychidae" genus="Smicridea" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="cholta" subGenus="Smicridea">
<emphasis id="B9295B4F126FFFA4A0BCF36FFCB77B70" box="[763,788,801,824]" italics="true" pageId="7" pageNumber="6">S.</emphasis>
(
<emphasis id="B9295B4F126FFFA4A164F36FFC9E7B70" box="[803,829,801,824]" italics="true" pageId="7" pageNumber="6">S.</emphasis>
)
<emphasis id="B9295B4F126FFFA4A10BF36EFC347B70" box="[844,919,800,824]" italics="true" pageId="7" pageNumber="6">cholta</emphasis>
</taxonomicName>
, and the posterodorsal margin of the phallobase, which is produced into a slender lobe in the new species, but is undulate and bearing a few short spines on the lateral surface in
<taxonomicName id="4C5DFCDE126FFFA4A004F32CFD477B31" authorityName="Flint" authorityYear="1974" box="[579,740,865,890]" class="Insecta" family="Hydropsychidae" genus="Smicridea" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="cholta" subGenus="Smicridea">
<emphasis id="B9295B4F126FFFA4A004F32CFDFE7B31" box="[579,605,866,889]" italics="true" pageId="7" pageNumber="6">S.</emphasis>
(
<emphasis id="B9295B4F126FFFA4A02AF32CFD257B31" box="[621,646,866,889]" italics="true" pageId="7" pageNumber="6">S.</emphasis>
)
<emphasis id="B9295B4F126FFFA4A0D1F32FFD477B31" box="[662,740,865,889]" italics="true" pageId="7" pageNumber="6">cholta</emphasis>
</taxonomicName>
<emphasis id="B9295B4F126FFFA4A0A3F32FFD4B7B31" box="[740,744,865,889]" italics="true" pageId="7" pageNumber="6">.</emphasis>
Furthermore,
<emphasis id="B9295B4F126FFFA4A1D9F32CFC147B31" box="[926,951,866,889]" italics="true" pageId="7" pageNumber="6">S.</emphasis>
<taxonomicName id="4C5DFCDE126FFFA4A1F9F32CFBF37B31" authorityName="Rázuri-Gonzales &amp; Armitage" authorityYear="2019" box="[958,1104,865,890]" class="Insecta" family="Hydropsychidae" genus="Smicridea" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="dividua" subGenus="Smicridea">
(
<emphasis id="B9295B4F126FFFA4A180F32CFC7A7B31" box="[967,985,866,889]" italics="true" pageId="7" pageNumber="6">S</emphasis>
.)
<emphasis id="B9295B4F126FFFA4A1B6F32FFBF37B31" box="[1009,1104,865,889]" italics="true" pageId="7" pageNumber="6">dividua</emphasis>
</taxonomicName>
has a single sinuous spine, which shape is unique in this group whereas
<taxonomicName id="4C5DFCDE126FFFA4A0AEF3CCFC247BD2" authorityName="Flint" authorityYear="1974" box="[745,903,898,922]" class="Insecta" family="Hydropsychidae" genus="Smicridea" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="cholta" subGenus="Smicridea">
<emphasis id="B9295B4F126FFFA4A0AEF3CCFCA07BD1" box="[745,771,898,921]" italics="true" pageId="7" pageNumber="6">S.</emphasis>
(
<emphasis id="B9295B4F126FFFA4A154F3CCFC8F7BD1" box="[787,812,898,921]" italics="true" pageId="7" pageNumber="6">S.</emphasis>
)
<emphasis id="B9295B4F126FFFA4A17BF3CCFC247BD2" box="[828,903,898,922]" italics="true" pageId="7" pageNumber="6">cholta</emphasis>
</taxonomicName>
has a pair of long slender spines.
</paragraph>
</subSubSection>
<subSubSection id="C347D4D6126FFFA4A287F3FCFD787F6F" pageId="7" pageNumber="6" type="description">
<paragraph id="8BE2875D126FFFA4A287F3FCFA037CC6" blockId="7.[192,1440,946,1783]" pageId="7" pageNumber="6">
<emphasis id="B9295B4F126FFFA4A287F3FCFEC97B82" bold="true" box="[192,362,946,970]" pageId="7" pageNumber="6">Description.</emphasis>
<emphasis id="B9295B4F126FFFA4A336F3FCFDA17B82" box="[369,514,946,970]" italics="true" pageId="7" pageNumber="6">Adult male.</emphasis>
Forewing length average
<quantity id="4CA52AB8126FFFA4A104F3FDFC3C7B82" box="[835,927,947,971]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="7" pageNumber="6" unit="mm" value="4.3">4.3 mm</quantity>
(
<emphasis id="B9295B4F126FFFA4A1F7F3FDFC637B82" box="[944,960,947,970]" italics="true" pageId="7" pageNumber="6">n</emphasis>
= 7). Body color light brown (specimen preserved in alcohol). Head and thorax light brown with brown setae, dorsally thorax with dark brown marks anterad. Antennae and legs yellowish brown, antennae with short brown setae, legs with long brown setae. Maxillary palps 5-segmented; maxillary palps with long brown setae and a few thick long setae anteroventrally on segment II. Forewing membrane light brown, with brown and dark brown setae, with whitish transversal band in the nygmal area membrane, mostly denuded. Wing venation typical for the subgenus. Abdominal segments VI and VIII glands oval, larger than containing segment.
</paragraph>
<paragraph id="8BE2875D126FFFA4A2AFF4D8FAB77EBF" blockId="7.[192,1440,946,1783]" pageId="7" pageNumber="6">
<emphasis id="B9295B4F126FFFA4A2AFF4D8FE3C7CE6" box="[232,415,1174,1198]" italics="true" pageId="7" pageNumber="6">Male genitalia.</emphasis>
Segment IX in lateral view (
<figureCitation id="13669BD8126FFFA4A0BCF4D8FCF07CE6" box="[763,851,1174,1198]" captionStart="Figure 3" captionStartId="13.[192,275,1846,1868]" captionTargetBox="[307,1333,221,1809]" captionTargetId="figure@13.[297,1334,212,1812]" captionTargetPageId="13" captionText="Figure 3. Smicridea (Smicridea) dividua Rázuri-Gonzales and Armitage, sp. n., male genitalia. A) Lateral view. B) Dorsal view. C) Left inferior appendage, ventral view. D) Phallic apparatus, lateral view (inset: posterodorsal margin projection, dorsal view). E) Phallic apparatus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3674841" httpUri="https://zenodo.org/record/3674841/files/figure.png" pageId="7" pageNumber="6">Fig. 3A</figureCitation>
) much longer dorsally than ventrally, anterolateral margin slightly concave, anterodorsal corner not produced, forming nearly straight angle; anterior portion of dorsal margin long, slightly concave, mesal portion of dorsal margin convex, bearing small microsetae, closer to tergum X than to anterolateral margin of segment IX, posterior portion membranous; ventrolateral margin oblique, sinuous; ventral margin short, truncate; in dorsal view (
<figureCitation id="13669BD8126FFFA4A692F556FA8E7D78" box="[1237,1325,1304,1328]" captionStart="Figure 3" captionStartId="13.[192,275,1846,1868]" captionTargetBox="[307,1333,221,1809]" captionTargetId="figure@13.[297,1334,212,1812]" captionTargetPageId="13" captionText="Figure 3. Smicridea (Smicridea) dividua Rázuri-Gonzales and Armitage, sp. n., male genitalia. A) Lateral view. B) Dorsal view. C) Left inferior appendage, ventral view. D) Phallic apparatus, lateral view (inset: posterodorsal margin projection, dorsal view). E) Phallic apparatus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3674841" httpUri="https://zenodo.org/record/3674841/files/figure.png" pageId="7" pageNumber="6">Fig. 3B</figureCitation>
) anterior margin strongly concave, lateral margin sinuous, inflated mesally, mesally bearing microsetae. Tergum X in lateral view (
<figureCitation id="13669BD8126FFFA4A3DEF517FE517D39" box="[409,498,1369,1393]" captionStart="Figure 3" captionStartId="13.[192,275,1846,1868]" captionTargetBox="[307,1333,221,1809]" captionTargetId="figure@13.[297,1334,212,1812]" captionTargetPageId="13" captionText="Figure 3. Smicridea (Smicridea) dividua Rázuri-Gonzales and Armitage, sp. n., male genitalia. A) Lateral view. B) Dorsal view. C) Left inferior appendage, ventral view. D) Phallic apparatus, lateral view (inset: posterodorsal margin projection, dorsal view). E) Phallic apparatus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3674841" httpUri="https://zenodo.org/record/3674841/files/figure.png" pageId="7" pageNumber="6">Fig. 3A</figureCitation>
) narrow, setose, much shorter than segment IX, apex truncate, posterodorsal corner produced dorsad; in dorsal view (
<figureCitation id="13669BD8126FFFA4A0F8F537FCBF7DDA" box="[703,796,1401,1426]" captionStart="Figure 3" captionStartId="13.[192,275,1846,1868]" captionTargetBox="[307,1333,221,1809]" captionTargetId="figure@13.[297,1334,212,1812]" captionTargetPageId="13" captionText="Figure 3. Smicridea (Smicridea) dividua Rázuri-Gonzales and Armitage, sp. n., male genitalia. A) Lateral view. B) Dorsal view. C) Left inferior appendage, ventral view. D) Phallic apparatus, lateral view (inset: posterodorsal margin projection, dorsal view). E) Phallic apparatus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3674841" httpUri="https://zenodo.org/record/3674841/files/figure.png" pageId="7" pageNumber="6">Fig. 3B</figureCitation>
) narrow, with rounded apices, separated by mesal notch, less than half its length, membranous mesally. Inferior appendage (
<figureCitation id="13669BD8126FFFA4A608F5D4FB3D7DFA" box="[1103,1182,1434,1458]" captionStart="Figure 3" captionStartId="13.[192,275,1846,1868]" captionTargetBox="[307,1333,221,1809]" captionTargetId="figure@13.[297,1334,212,1812]" captionTargetPageId="13" captionText="Figure 3. Smicridea (Smicridea) dividua Rázuri-Gonzales and Armitage, sp. n., male genitalia. A) Lateral view. B) Dorsal view. C) Left inferior appendage, ventral view. D) Phallic apparatus, lateral view (inset: posterodorsal margin projection, dorsal view). E) Phallic apparatus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3674841" httpUri="https://zenodo.org/record/3674841/files/figure.png" pageId="7" pageNumber="6">Figs. 3</figureCitation>
A-C) two-segmented; first segment not extending beyond posterior margin of tergum X, covered with setae, inflated apically; second segment digitate, approximately 1/2 as long as first segment, with short setae on margins, in dorsal view apex acute, slightly inflated on distal half. Phallic apparatus (
<figureCitation id="13669BD8126FFFA4A60DF5B5FB397E5C" box="[1098,1178,1531,1556]" captionStart="Figure 3" captionStartId="13.[192,275,1846,1868]" captionTargetBox="[307,1333,221,1809]" captionTargetId="figure@13.[297,1334,212,1812]" captionTargetPageId="13" captionText="Figure 3. Smicridea (Smicridea) dividua Rázuri-Gonzales and Armitage, sp. n., male genitalia. A) Lateral view. B) Dorsal view. C) Left inferior appendage, ventral view. D) Phallic apparatus, lateral view (inset: posterodorsal margin projection, dorsal view). E) Phallic apparatus, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3674841" httpUri="https://zenodo.org/record/3674841/files/figure.png" pageId="7" pageNumber="6">Figs. 3</figureCitation>
E-D) with phallobase enlarged, open ventrally; apical section open, dorsolateral corner of phallobase produced into a slender lobe; posterodorsal margin of phallobase mesally produced, acute; posteroventral margin produced into two slender digitate lobes, the ventral one with subacute apex, directed laterad and the dorsal one with bilobed apex, directed dorsolaterad, without spines or processes on lateral surface; in ventral view, posteroventral process deeply notched, notch basally rounded and undulate, each lobe slender, bilobed apically, directed laterad; endothecal membranes in lateral view with a long, sinuous, apically acute spine, in ventral view, bilobed, broad basally and slender apically, deeply notched mesally.
</paragraph>
<paragraph id="8BE2875D126FFFA4A287F741FD787F6F" blockId="7.[192,731,1807,1831]" box="[192,731,1807,1831]" pageId="7" pageNumber="6">
<emphasis id="B9295B4F126FFFA4A287F741FDF77F6F" bold="true" box="[192,596,1807,1831]" pageId="7" pageNumber="6">Female and immature stages.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C347D4D6126FFFA4A287F70EFD177F10" box="[192,692,1856,1880]" pageId="7" pageNumber="6" type="distribution">
<paragraph id="8BE2875D126FFFA4A287F70EFD177F10" blockId="7.[192,692,1856,1880]" box="[192,692,1856,1880]" pageId="7" pageNumber="6">
<emphasis id="B9295B4F126FFFA4A287F70EFED07F10" bold="true" box="[192,371,1856,1880]" pageId="7" pageNumber="6">Distribution.</emphasis>
<collectingCountry id="F34AC7CD126FFFA4A33DF70EFE437F1F" box="[378,480,1856,1879]" name="Panama" pageId="7" pageNumber="6">Panama</collectingCountry>
(
<collectingRegion id="499949BF126FFFA4A3A9F70EFD0B7F10" box="[494,680,1856,1880]" country="Panama" name="Cocle" pageId="7" pageNumber="6">Coclé Province</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C347D4D6126FFFA4A287F73EFA9A7FC0" box="[192,1337,1904,1928]" pageId="7" pageNumber="6" type="etymology">
<paragraph id="8BE2875D126FFFA4A287F73EFA9A7FC0" blockId="7.[192,1337,1904,1928]" box="[192,1337,1904,1928]" pageId="7" pageNumber="6">
<emphasis id="B9295B4F126FFFA4A287F73EFEF87FC0" bold="true" box="[192,347,1904,1928]" pageId="7" pageNumber="6">Etymology.</emphasis>
From the Latin word
<emphasis id="B9295B4F126FFFA4A02DF73EFD757FC0" box="[618,726,1904,1928]" italics="true" pageId="7" pageNumber="6">dividuus</emphasis>
, referring to the unique divided endothecal spine.
</paragraph>
</subSubSection>
<subSubSection id="C347D4D61260FFABA287F097FBE1797C" pageId="8" pageNumber="7" type="materials_examined">
<paragraph id="8BE2875D1260FFABA287F097FBE1797C" blockId="8.[192,1440,217,308]" pageId="8" pageNumber="7">
<emphasis id="B9295B4F1260FFABA287F097FE6B78B9" bold="true" box="[192,456,217,241]" pageId="8" pageNumber="7">Material examined.</emphasis>
<materialsCitation id="3B358D001260FFABA38AF097FE9C797C" ID-GBIF-Occurrence="2569342516" collectingDate="2017-03-22" collectingDateMax="2017-03-26" collectingDateMin="2017-03-22" collectionCode="MALE, T, COZEM" collectorName="E. Alvarez &amp; E. Perez &amp; Rios" country="Panama" elevation="586" latitude="8.66168" location="Quebrada La Yayas" longLatPrecision="5" longitude="-80.5952" municipality="Omar Torrijos National Park" pageId="8" pageNumber="7" specimenCount="1" stateProvince="Cocle" typeStatus="holotype">
<emphasis id="B9295B4F1260FFABA38AF097FC1A78B9" bold="true" box="[461,953,217,241]" pageId="8" pageNumber="7">
<typeStatus id="54E639FF1260FFABA38AF097FDD278B9" box="[461,625,217,241]" pageId="8" pageNumber="7" type="holotype">HOLOTYPE</typeStatus>
<collectionCode id="ED4C1F981260FFABA030F097FD6D78B9" box="[631,718,217,241]" pageId="8" pageNumber="7">MALE</collectionCode>
.
<collectingCountry id="F34AC7CD1260FFABA09EF097FCF878B9" box="[729,859,217,241]" name="Panama" pageId="8" pageNumber="7">PANAMA</collectingCountry>
:
<collectingRegion id="499949BF1260FFABA121F097FC1078B9" box="[870,947,217,241]" country="Panama" name="Cocle" pageId="8" pageNumber="7">Coclé</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6B861D271260FFABA1F9F094FABE78B9" box="[958,1309,217,242]" pageId="8" pageNumber="7">Omar Torrijos National Park</collectingMunicipality>
,
<location id="8E82D1861260FFABA760F097FE97795B" LSID="urn:lsid:plazi:treatment:03F4364B126FFFABA287F223FBE1797C:8E82D1861260FFABA760F097FE97795B" country="Panama" latitude="8.66168" longLatPrecision="5" longitude="-80.5952" municipality="Omar Torrijos National Park" name="Quebrada La Yayas" pageId="8" pageNumber="7" stateProvince="Cocle">Quebrada La Yayas</location>
,
<geoCoordinate id="EE69E19A1260FFABA378F0B5FE61795B" box="[319,450,251,275]" degrees="8.66168" direction="north" orientation="latitude" pageId="8" pageNumber="7" precision="1" value="8.66168">8.66168°N</geoCoordinate>
,
<geoCoordinate id="EE69E19A1260FFABA38BF0B5FDF7795B" box="[460,596,251,275]" degrees="80.5952" direction="west" orientation="longitude" pageId="8" pageNumber="7" precision="5" value="-80.5952">80.5952°W</geoCoordinate>
,
<quantity id="4CA52AB81260FFABA019F0B5FD08795A" box="[606,683,251,275]" metricMagnitude="2" metricUnit="m" metricValue="5.86" pageId="8" pageNumber="7" unit="m" value="586.0">
<elevation id="0070606E1260FFABA019F0B5FD08795A" box="[606,683,251,275]" metricMagnitude="2" metricUnit="m" metricValue="5.86" pageId="8" pageNumber="7" unit="m" value="586.0">586 m</elevation>
</quantity>
,
<date id="FFE3A19D1260FFABA0F1F0B5FCC6795B" box="[694,869,251,275]" pageId="8" pageNumber="7" value="2017-03-22" valueMax="2017-03-26" valueMin="2017-03-22">
<collectingDate id="EFA758751260FFABA0F1F0B5FCC6795B" box="[694,869,251,275]" pageId="8" pageNumber="7" value="2017-03-22" valueMax="2017-03-26" valueMin="2017-03-22">2226.iii.2017</collectingDate>
</date>
,
<collectorName id="26A8E28B1260FFABA137F0B5FC52795A" box="[880,1009,247,274]" pageId="8" pageNumber="7">E. Álvarez</collectorName>
,
<collectorName id="26A8E28B1260FFABA1BBF0B5FBC6795B" box="[1020,1125,251,275]" pageId="8" pageNumber="7">E. Pérez</collectorName>
, and
<collectionCode id="ED4C1F981260FFABA6E4F0B5FB1B795A" box="[1187,1208,251,274]" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="7">T</collectionCode>
.
<collectorName id="26A8E28B1260FFABA684F0B5FB58795B" box="[1219,1275,251,275]" pageId="8" pageNumber="7">Ríos</collectorName>
,
<collectingMethod id="521CFF4A1260FFABA742F0B5FA03795A" box="[1285,1440,251,275]" pageId="8" pageNumber="7">Malaise trap</collectingMethod>
(
<emphasis id="B9295B4F1260FFABA28EF155FE97797B" bold="true" box="[201,308,283,307]" pageId="8" pageNumber="7">
<collectionCode id="ED4C1F981260FFABA28EF155FE97797B" box="[201,308,283,307]" pageId="8" pageNumber="7">COZEM</collectionCode>
</emphasis>
)
</materialsCitation>
.
<materialsCitation id="3B358D001260FFABA30CF155FBE1797C" ID-GBIF-Occurrence="2569342515" box="[331,1090,283,308]" pageId="8" pageNumber="7" typeStatus="paratype">
<emphasis id="B9295B4F1260FFABA30CF155FDA6797B" bold="true" box="[331,517,283,307]" pageId="8" pageNumber="7">
<typeStatus id="54E639FF1260FFABA30CF155FDA3797B" box="[331,512,283,307]" pageId="8" pageNumber="7" type="paratype">PARATYPES</typeStatus>
.
</emphasis>
Same data as
<typeStatus id="54E639FF1260FFABA0FEF152FC87797C" box="[697,804,284,308]" pageId="8" pageNumber="7" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D5B4CD41260FFABA168F152FCF1797C" box="[815,850,284,308]" pageId="8" pageNumber="7" type="male">6♂</specimenCount>
(
<emphasis id="B9295B4F1260FFABA125F155FB91797B" bold="true" box="[866,1074,283,307]" pageId="8" pageNumber="7">
<collectionCode id="ED4C1F981260FFABA125F155FC6C797B" box="[866,975,283,307]" pageId="8" pageNumber="7">COZEM</collectionCode>
,
<collectionCode id="ED4C1F981260FFABA19BF155FB91797B" box="[988,1074,283,307]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="8" pageNumber="7" type="University or college">UMSP</collectionCode>
</emphasis>
).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>