308 lines
35 KiB
XML
308 lines
35 KiB
XML
<document id="C6E28F585DDD5A6FF3CE9823418E1878" ID-DOI="10.11646/zootaxa.3710.1.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="10099429" ID-ZooBank="F10A5AE0-831F-4AAC-ABA1-890E34AAA2A7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629900861594" checkinUser="felipe" docAuthor="Lara, María Belén & Lukashevich, Elena D." docDate="2013" docId="03E1D82612288832FF4886A6B767FE78" docLanguage="en" docName="zt03710p092.pdf" docOrigin="Zootaxa 3710 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trihennigma Lara & Lukashevich 2013, gen. nov." docType="treatment" docVersion="3" lastPageNumber="89" masterDocId="FFD8A05E122B883AFFDF8031B676FFA8" masterDocTitle="The first Triassic dipteran (Insecta) from South America, with review of Hennigmatidae" masterLastPageNumber="92" masterPageNumber="81" pageNumber="84" updateTime="1699595441112" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="D132975798CCBD4AAB71D337E2DE8419" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1F40FD152D02F8E370AE53D20B4902A1">
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<mods:title id="E1A835479C808317F383817D2F02C9EA">The first Triassic dipteran (Insecta) from South America, with review of Hennigmatidae</mods:title>
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</mods:titleInfo>
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<mods:name id="69420F3C513C48FBA5F52A68CBE5037D" type="personal">
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<mods:role id="3A4E243255835299D84D2F79A3628199">
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<mods:roleTerm id="91FE7B31BB92D3380337ED6442391150">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="73E355BB9AC36315EB4C02B78F96CB43">Lara, María Belén</mods:namePart>
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<mods:affiliation id="C3038AE740D312B45C0C5823191FABC8">Area Entomología, Universidad Nacional del Nordeste and Area Paleontología, Centro de Ecología Aplicada del Litoral (CONICET), Casilla de Correo 128, 3400 Corrientes, Argentina. E-mail: lara. maria. belen @ live. com. ar</mods:affiliation>
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</mods:name>
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<mods:name id="E08D2FD4B61E2335524BFD7515FA4A76" type="personal">
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<mods:role id="C87F8D72FBD77255908482F3A6F3C076">
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<mods:roleTerm id="AAB08D1B7B9E140CC1B0F424AE41AC16">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8D0BD673F7F826E481837E34B587C768">Lukashevich, Elena D.</mods:namePart>
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<mods:affiliation id="99F0D28EED7FAB89DB2A1971BD17B6FF">Borissiak Paleontological Institue RAS, Moscow, Russia. E-mail: elukashevich @ hotmail. com</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="5C15A953BCE00B2011FE42977BA08D06">text</mods:typeOfResource>
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<mods:titleInfo id="DBF8AD549489E2C53623636830FD5E16">
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<mods:title id="095C9D4B7FAF59917436A0471AA27C75">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F0ADC776A490A981515A2FF4E5D58AFE">
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<mods:date id="41CC12503636A6E1FF122B11C229D5D7">2013</mods:date>
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<mods:detail id="F826773133C1D2E7643430FA86E97737" type="pubDate">
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<mods:number id="B4D65FA8F6EDD18256AABD266432D252">2013-09-11</mods:number>
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</mods:detail>
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<mods:detail id="114E12FFB4EEA04DCB382F363709A9F1" type="volume">
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<mods:number id="A3F8FC261BF63F988ADA3999BDDD684E">3710</mods:number>
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</mods:detail>
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<mods:detail id="5EA981C17006F1A08F9A3004672C94EE" type="issue">
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<mods:number id="64CA031089C2CF78C50CDC4338388496">1</mods:number>
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</mods:detail>
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<mods:extent id="759C4DEEE27436328ACB150A56030691" unit="page">
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<mods:start id="8CF48B2068787A4B01974FD70EFA31E1">81</mods:start>
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<mods:end id="6B552FA32FBD6272DDFBCC03E0127C1F">92</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="4C4AFAD43213261F3596C299D439F30E">journal article</mods:classification>
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<mods:identifier id="305B6A793B670FA07E743B96C950C938" type="DOI">10.11646/zootaxa.3710.1.6</mods:identifier>
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<mods:identifier id="C60E571ACE466C7CC210C69D9A2E6512" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="2BBF9C3A451F2A73D7E05A48E946C97D" type="Zenodo-Dep">10099429</mods:identifier>
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<mods:identifier id="7A51C947B5305A5B55940AA24BF87DA1" type="ZooBank">F10A5AE0-831F-4AAC-ABA1-890E34AAA2A7</mods:identifier>
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</mods:mods>
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<treatment id="03E1D82612288832FF4886A6B767FE78" LSID="urn:lsid:plazi:treatment:03E1D82612288832FF4886A6B767FE78" httpUri="http://treatment.plazi.org/id/03E1D82612288832FF4886A6B767FE78" lastPageId="8" lastPageNumber="89" pageId="3" pageNumber="84">
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<subSubSection id="C3523ABB12288839FF4886A6B7E8F918" box="[151,414,1687,1712]" pageId="3" pageNumber="84" type="nomenclature">
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<paragraph id="8BF7693012288839FF4886A6B7E8F918" blockId="3.[151,414,1687,1712]" box="[151,414,1687,1712]" pageId="3" pageNumber="84">
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<heading id="D0BFDE5C12288839FF4886A6B7E8F918" bold="true" box="[151,414,1687,1712]" fontSize="10" level="2" pageId="3" pageNumber="84" reason="2">
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<emphasis id="B93CB52212288839FF4886A6B7E8F918" bold="true" box="[151,414,1687,1712]" pageId="3" pageNumber="84">
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<taxonomicName id="4C4812B312288839FF4886A6B759F918" ID-CoL="89RN9" authority="Lara & Lukashevich, 2013" authorityName="Lara & Lukashevich" authorityYear="2013" box="[151,303,1687,1712]" class="Insecta" family="Hennigmatidae" genus="Trihennigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B93CB52212288839FF4886A6B759F918" bold="true" box="[151,303,1687,1712]" italics="true" pageId="3" pageNumber="84">Trihennigma</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20F085912288839FEEA86A6B7E8F918" box="[309,414,1687,1712]" pageId="3" pageNumber="84" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3523ABB12288839FF4886EEB4D7F95F" box="[151,673,1759,1784]" pageId="3" pageNumber="84" type="type_taxon">
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<paragraph id="8BF7693012288839FF4886EEB4D7F95F" blockId="3.[151,1436,1759,2008]" box="[151,673,1759,1784]" pageId="3" pageNumber="84">
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<emphasis id="B93CB52212288839FF4886EEB744F950" bold="true" box="[151,306,1759,1784]" inLineHeading="true" pageId="3" pageNumber="84" reason="1">
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<typeStatus id="54F3D79212288839FF4886EEB6A4F950" box="[151,210,1759,1784]" pageId="3" pageNumber="84">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C4812B312288839FEE686EEB436F95F" ID-CoL="8FJDK" authority="Lara & Lukashevich, 2013" authorityName="Lara & Lukashevich" authorityYear="2013" box="[313,576,1759,1784]" class="Insecta" family="Hennigmatidae" genus="Trihennigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="84" phylum="Arthropoda" rank="species" species="zavattierii" status="sp. nov.">
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<emphasis id="B93CB52212288839FEE686EEB436F95F" box="[313,576,1759,1784]" italics="true" pageId="3" pageNumber="84">Trihennigma zavattierii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20F085912288839FD9886EEB4D7F95F" box="[583,673,1759,1784]" pageId="3" pageNumber="84" rank="species">
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<emphasis id="B93CB52212288839FD9886EEB4ECF950" bold="true" box="[583,666,1759,1784]" pageId="3" pageNumber="84">sp. nov</emphasis>
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.
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</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3523ABB1228883DFF188732B468F956" lastPageId="7" lastPageNumber="88" pageId="3" pageNumber="84" type="diagnosis">
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<paragraph id="8BF7693012288839FF188732B219F870" blockId="3.[151,1436,1759,2008]" pageId="3" pageNumber="84">
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<emphasis id="B93CB52212288839FF188732B7A6F8B3" box="[199,464,1795,1820]" inLineHeading="true" pageId="3" pageNumber="84" reason="1">
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<emphasis id="B93CB52212288839FF188732B7BFF8B4" bold="true" box="[199,457,1795,1820]" pageId="3" pageNumber="84">Differential diagnosis</emphasis>
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.
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</emphasis>
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Wing rather broad, with vestigial pterostigma and without spots at crossveins; long strong Sc ending just before R
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<subScript id="17CC6B7512288839FDDE8707B46FF8EC" attach="left" box="[513,537,1846,1860]" fontSize="6" pageId="3" pageNumber="84">2+3</subScript>
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bifurcation and possibly distal to M
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<subScript id="17CC6B7512288839FC1B8707B5AAF8EC" attach="left" box="[964,988,1846,1860]" fontSize="6" pageId="3" pageNumber="84">1+2</subScript>
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bifurcation; postsectoral
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<emphasis id="B93CB52212288839FADE8719B35BF897" box="[1281,1325,1832,1855]" italics="true" pageId="3" pageNumber="84">sc-r</emphasis>
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probably nearly level with R
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<subScript id="17CC6B7512288839FEB4876CB705F8C3" attach="left" box="[363,371,1885,1899]" fontSize="6" pageId="3" pageNumber="84">5</subScript>
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origin; supernumerary crossveins
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<emphasis id="B93CB52212288839FD2B877EB56BF8CE" box="[756,797,1871,1894]" italics="true" pageId="3" pageNumber="84">r-rs</emphasis>
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,
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<emphasis id="B93CB52212288839FCF4877CB5D9F8CE" box="[811,943,1869,1894]" italics="true" pageId="3" pageNumber="84">r3-r4, r4-r5</emphasis>
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and
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<emphasis id="B93CB52212288839FC3B877CB25FF8CE" box="[996,1065,1869,1894]" italics="true" pageId="3" pageNumber="84">r5-m1</emphasis>
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; R
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<subScript id="17CC6B7512288839FB95876CB224F8C3" attach="left" box="[1098,1106,1885,1899]" fontSize="6" pageId="3" pageNumber="84">2</subScript>
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longer than R
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<subScript id="17CC6B7512288839FB2F876CB37EF8C3" attach="left" box="[1264,1288,1885,1899]" fontSize="6" pageId="3" pageNumber="84">2+3</subScript>
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, R
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<subScript id="17CC6B7512288839FAFA876CB35BF8C3" attach="left" box="[1317,1325,1885,1899]" fontSize="6" pageId="3" pageNumber="84">4</subScript>
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almost as long as half of wing, entering margin at wing tip;
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<emphasis id="B93CB52212288839FD218744B56CF824" box="[766,794,1909,1932]" italics="true" pageId="3" pageNumber="84">im</emphasis>
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not preserved,
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<emphasis id="B93CB52212288839FC078744B266F824" box="[984,1040,1909,1932]" italics="true" pageId="3" pageNumber="84">m-cu</emphasis>
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just before M
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<subScript id="17CC6B7512288839FB1F87B2B2AEF839" attach="left" box="[1216,1240,1923,1937]" fontSize="6" pageId="3" pageNumber="84">3+4</subScript>
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fork; M
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<subScript id="17CC6B7512288839FAE487B2B335F839" attach="left" box="[1339,1347,1923,1937]" fontSize="6" pageId="3" pageNumber="84">4</subScript>
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curved backwards before apex; CuA section before
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<emphasis id="B93CB52212288839FD5187ADB4B0F81B" box="[654,710,1948,1971]" italics="true" pageId="3" pageNumber="84">m-cu</emphasis>
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twice as long as section beyond it; anal loop broad, subequal to 1A beyond it, the latter terminating at deep alular incision, anal lobe possibly produced.
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</paragraph>
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<caption id="DF3739B8122F883EFF488764B4D9F822" ID-DOI="http://doi.org/10.5281/zenodo.10099437" ID-Zenodo-Dep="10099437" httpUri="https://zenodo.org/record/10099437/files/figure.png" pageId="4" pageNumber="85" startId="4.[151,249,1877,1899]" targetBox="[171,1415,193,1852]" targetPageId="4">
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<paragraph id="8BF76930122F883EFF488764B4D9F822" blockId="4.[151,1436,1877,1930]" pageId="4" pageNumber="85">
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<emphasis id="B93CB522122F883EFF488764B760F8C2" bold="true" box="[151,278,1877,1899]" pageId="4" pageNumber="85">FIGURE 3.</emphasis>
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A,
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<taxonomicName id="4C4812B3122F883EFE9D8764B50DF8C3" authority="(Shcherbakov, 1995)" baseAuthorityName="Shcherbakov" baseAuthorityYear="1995" box="[322,891,1877,1899]" class="Insecta" family="Hennigmatidae" genus="Metatrichopteridium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="cladistorum">
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<emphasis id="B93CB522122F883EFE9D8764B4E1F8C3" box="[322,663,1877,1899]" italics="true" pageId="4" pageNumber="85">Metatrichopteridium cladistorum</emphasis>
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(
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<bibRefCitation id="EFD914C1122F883EFD788764B504F8C3" author="Shcherbakov, D. & Lukashevich, E. D. & Blagoderov, V. A." box="[679,882,1877,1899]" pageId="4" pageNumber="85" pagination="75 - 115" refId="ref6207" refString="Shcherbakov, D. E, Lukashevich, E. D. & Blagoderov, V. A. (1995) Triassic Diptera and initial radiation of the order. International Journal of Dipterolological Research, 6, 75 - 115." type="journal article" year="1995">Shcherbakov, 1995</bibRefCitation>
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)
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</taxonomicName>
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: holotype PIN 3965/2789; B,
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<taxonomicName id="4C4812B3122F883EFB658764B7E2F822" authority="Lukashevich et al., 2006" authorityName="Lukashevich" authorityYear="2006" class="Insecta" family="Hennigmatidae" genus="Daohennigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="85" phylum="Arthropoda" rank="species" species="panops">
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<emphasis id="B93CB522122F883EFB658764B3EAF8C3" box="[1210,1436,1877,1899]" italics="true" pageId="4" pageNumber="85">Daohennigma panops</emphasis>
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<bibRefCitation id="EFD914C1122F883EFF488745B7E2F822" author="Lukashevich, E. D. & Huang, D. - Y. & Lin, Q. - B." box="[151,404,1908,1930]" pageId="4" pageNumber="85" pagination="127 - 143" refId="ref5720" refString="Lukashevich, E. D., Huang, D. - Y. & Lin, Q. - B. (2006) Rare families of lower Diptera (Hennigmatidae, Blephariceridae, Perissommatidae) from the Jurassic of China. Studia dipterologica, 13, 127 - 143." type="journal article" year="2006">
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Lukashevich
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<emphasis id="B93CB522122F883EFEC18744B73DF822" box="[286,331,1908,1930]" italics="true" pageId="4" pageNumber="85">et al</emphasis>
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., 2006
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</bibRefCitation>
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</taxonomicName>
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: holotype NIGPAS 133706.
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</paragraph>
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</caption>
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<caption id="DF3739B8122E883FFF48871EB4DCF8CB" ID-DOI="http://doi.org/10.5281/zenodo.10099439" ID-Zenodo-Dep="10099439" httpUri="https://zenodo.org/record/10099439/files/figure.png" pageId="5" pageNumber="86" startId="5.[151,249,1839,1861]" targetBox="[184,1422,213,1804]" targetPageId="5">
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<paragraph id="8BF76930122E883FFF48871EB4DCF8CB" blockId="5.[151,1437,1839,1891]" pageId="5" pageNumber="86">
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<emphasis id="B93CB522122E883FFF48871EB762F8EC" bold="true" box="[151,276,1839,1861]" pageId="5" pageNumber="86">FIGURE 4.</emphasis>
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<taxonomicName id="4C4812B3122E883FFEC3871EB47AF8ED" authority="Lara & Lukashevich, 2013" authorityName="Lara & Lukashevich" authorityYear="2013" box="[284,524,1839,1861]" class="Insecta" family="Hennigmatidae" genus="Trihennigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="86" phylum="Arthropoda" rank="species" species="zavattierii" status="gen. et sp. nov.">
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<emphasis id="B93CB522122E883FFEC3871EB47AF8ED" box="[284,524,1839,1861]" italics="true" pageId="5" pageNumber="86">Trihennigma zavattierii</emphasis>
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</taxonomicName>
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<emphasis id="B93CB522122E883FFDCB871EB4C8F8EC" bold="true" box="[532,702,1839,1860]" pageId="5" pageNumber="86">
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<taxonomicNameLabel id="A20F0859122E883FFDCB871EB4CFF8EC" box="[532,697,1839,1860]" pageId="5" pageNumber="86" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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:
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</emphasis>
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holotype MCNAM-PI 24607: A, photo, B, drawing, C, reconstruction. Scale bar 1.7 mm. Abbreviation: ai—alular incision.
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</paragraph>
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</caption>
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<caption id="DF3739B8122D883CFF4887BAB758F876" ID-DOI="http://doi.org/10.5281/zenodo.10099441" ID-Zenodo-Dep="10099441" httpUri="https://zenodo.org/record/10099441/files/figure.png" pageId="6" pageNumber="87" startId="6.[151,249,1931,1953]" targetBox="[201,1387,193,1893]" targetPageId="6">
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<paragraph id="8BF76930122D883CFF4887BAB758F876" blockId="6.[151,1436,1931,2014]" pageId="6" pageNumber="87">
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<emphasis id="B93CB522122D883CFF4887BAB765F808" bold="true" box="[151,275,1931,1953]" pageId="6" pageNumber="87">FIGURE 5.</emphasis>
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Basiala of
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<taxonomicName id="4C4812B3122D883CFE5787BAB451F809" authorityName="Shcherbakov" authorityYear="1995" box="[392,551,1931,1953]" class="Insecta" family="Hennigmatidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="87" phylum="Arthropoda" rank="subFamily" subFamily="Hennigmatinae">Hennigmatinae</taxonomicName>
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: A, B,
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<taxonomicName id="4C4812B3122D883CFDB087BAB529F809" authority="Lara & Lukashevich, 2013" authorityName="Lara & Lukashevich" authorityYear="2013" box="[623,863,1931,1953]" class="Insecta" family="Hennigmatidae" genus="Trihennigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="87" phylum="Arthropoda" rank="species" species="zavattierii" status="gen. et sp. nov.">
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<emphasis id="B93CB522122D883CFDB087BAB529F809" box="[623,863,1931,1953]" italics="true" pageId="6" pageNumber="87">Trihennigma zavattierii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20F0859122D883CFCB987BAB270F809" box="[870,1030,1931,1953]" pageId="6" pageNumber="87" rank="species">
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<emphasis id="B93CB522122D883CFCB987BAB276F808" bold="true" box="[870,1024,1931,1952]" pageId="6" pageNumber="87">gen. et sp. nov</emphasis>
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.
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</taxonomicNameLabel>
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: holotype MCNAM-PI 24607, part and inverted counterpart; C,
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<taxonomicName id="4C4812B3122D883CFE4C8798B5B0F817" authority="(Shcherbakov, 1995)" baseAuthorityName="Shcherbakov" baseAuthorityYear="1995" box="[403,966,1961,1983]" class="Insecta" family="Hennigmatidae" genus="Metatrichopteridium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="87" phylum="Arthropoda" rank="species" species="cladistorum">
|
||
<emphasis id="B93CB522122D883CFE4C8798B490F817" box="[403,742,1961,1983]" italics="true" pageId="6" pageNumber="87">Metatrichopteridium cladistorum</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD914C1122D883CFD2B8798B5C8F817" author="Shcherbakov, D. & Lukashevich, E. D. & Blagoderov, V. A." box="[756,958,1961,1983]" pageId="6" pageNumber="87" pagination="75 - 115" refId="ref6207" refString="Shcherbakov, D. E, Lukashevich, E. D. & Blagoderov, V. A. (1995) Triassic Diptera and initial radiation of the order. International Journal of Dipterolological Research, 6, 75 - 115." type="journal article" year="1995">Shcherbakov, 1995</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
: holotype PIN 3965/2789. Abbreviation: ai— alular incision.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3739B8122C883DFF48832DB552FCF9" ID-DOI="http://doi.org/10.5281/zenodo.10099443" ID-Zenodo-Dep="10099443" httpUri="https://zenodo.org/record/10099443/files/figure.png" pageId="7" pageNumber="88" startId="7.[151,249,796,818]" targetBox="[168,1436,193,775]" targetPageId="7">
|
||
<paragraph id="8BF76930122C883DFF48832DB552FCF9" blockId="7.[151,1435,796,849]" pageId="7" pageNumber="88">
|
||
<emphasis id="B93CB522122C883DFF48832DB765FC99" bold="true" box="[151,275,796,818]" pageId="7" pageNumber="88">FIGURE 6.</emphasis>
|
||
Proximal part of the wing of
|
||
<taxonomicName id="4C4812B3122C883DFD9E832DB207FC9A" authority="(Shcherbakov, 1995)" baseAuthorityName="Shcherbakov" baseAuthorityYear="1995" box="[577,1137,796,818]" class="Insecta" family="Hennigmatidae" genus="Metatrichopteridium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="88" phylum="Arthropoda" rank="species" species="cladistorum">
|
||
<emphasis id="B93CB522122C883DFD9E832DB5E5FC9A" box="[577,915,796,818]" italics="true" pageId="7" pageNumber="88">Metatrichopteridium cladistorum</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD914C1122C883DFC7E832DB21FFC9A" author="Shcherbakov, D. & Lukashevich, E. D. & Blagoderov, V. A." box="[929,1129,796,818]" pageId="7" pageNumber="88" pagination="75 - 115" refId="ref6207" refString="Shcherbakov, D. E, Lukashevich, E. D. & Blagoderov, V. A. (1995) Triassic Diptera and initial radiation of the order. International Journal of Dipterolological Research, 6, 75 - 115." type="journal article" year="1995">Shcherbakov, 1995</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
: holotype PIN 3965/2789: A, drawing; B, SEM image (SE). Abbreviation: ai—alular incision.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3739B8122C883DFF488694B4FFF913" ID-DOI="http://doi.org/10.5281/zenodo.10099445" ID-Zenodo-Dep="10099445" box="[151,649,1701,1723]" httpUri="https://zenodo.org/record/10099445/files/figure.png" pageId="7" pageNumber="88" startId="7.[151,249,1701,1723]" targetBox="[152,1425,901,1674]" targetPageId="7">
|
||
<paragraph id="8BF76930122C883DFF488694B4FFF913" blockId="7.[151,649,1701,1723]" box="[151,649,1701,1723]" pageId="7" pageNumber="88">
|
||
<emphasis id="B93CB522122C883DFF488694B764F912" bold="true" box="[151,274,1701,1723]" pageId="7" pageNumber="88">FIGURE 7.</emphasis>
|
||
Distribution map of
|
||
<taxonomicName id="4C4812B3122C883DFE388694B4F3F913" authorityName="Shcherbakov" authorityYear="1995" box="[487,645,1701,1723]" class="Insecta" family="Hennigmatidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="88" phylum="Arthropoda" rank="family">Hennigmatidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF76930122C883DFF1886D4B468F956" blockId="7.[151,1437,1765,2011]" box="[199,542,1765,1790]" pageId="7" pageNumber="88">
|
||
<emphasis id="B93CB522122C883DFF1886D4B7E5F955" box="[199,403,1765,1790]" inLineHeading="true" pageId="7" pageNumber="88" reason="1">
|
||
<emphasis id="B93CB522122C883DFF1886D4B7FAF956" bold="true" box="[199,396,1765,1790]" pageId="7" pageNumber="88">Species included</emphasis>
|
||
.
|
||
</emphasis>
|
||
Monobasic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3523ABB122C883DFF188738B5B9F889" box="[199,975,1801,1826]" pageId="7" pageNumber="88" type="etymology">
|
||
<paragraph id="8BF76930122C883DFF188738B5B9F889" blockId="7.[151,1437,1765,2011]" box="[199,975,1801,1826]" pageId="7" pageNumber="88">
|
||
<emphasis id="B93CB522122C883DFF188738B738F88A" bold="true" box="[199,334,1801,1826]" inLineHeading="true" pageId="7" pageNumber="88" reason="1">Etymology.</emphasis>
|
||
From the Triassic period and the genus name
|
||
<taxonomicName id="4C4812B3122C883DFC8C873BB5BEF889" authorityName="Shcherbakov" authorityYear="1995" box="[851,968,1802,1825]" class="Insecta" family="Hennigmatidae" genus="Hennigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93CB522122C883DFC8C873BB5BEF889" box="[851,968,1802,1825]" italics="true" pageId="7" pageNumber="88">Hennigma</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3523ABB122C8832FF18871CB767FE78" lastPageId="8" lastPageNumber="89" pageId="7" pageNumber="88" type="discussion">
|
||
<paragraph id="8BF76930122C8832FF18871CB2A0FF50" blockId="7.[151,1437,1765,2011]" lastBlockId="8.[151,1436,151,464]" lastPageId="8" lastPageNumber="89" pageId="7" pageNumber="88">
|
||
<emphasis id="B93CB522122C883DFF18871CB74DF8EE" bold="true" box="[199,315,1837,1862]" inLineHeading="true" pageId="7" pageNumber="88" reason="1">Remarks.</emphasis>
|
||
Combination of long R
|
||
<subScript id="17CC6B75122C883DFD92870DB413F8E2" attach="left" box="[589,613,1852,1866]" fontSize="6" pageId="7" pageNumber="88">2+3</subScript>
|
||
fork, curved M
|
||
<subScript id="17CC6B75122C883DFCC8870DB569F8E2" attach="left" box="[791,799,1852,1866]" fontSize="6" pageId="7" pageNumber="88">4</subScript>
|
||
and broad anal loop has been known within the family only in the monobasic superfamily
|
||
<taxonomicName id="4C4812B3122C883DFD898765B562F8C5" authorityName="Lukashevich" authorityYear="1995" box="[598,788,1876,1901]" class="Insecta" family="Hennigmatidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="88" phylum="Arthropoda" rank="subFamily" subFamily="Kuperwoodiinae">Kuperwoodiinae</taxonomicName>
|
||
. The new genus resembles
|
||
<taxonomicName id="4C4812B3122C883DFBA68765B314F8C5" authorityName="Handlirsch" authorityYear="1939" box="[1145,1378,1876,1901]" class="Insecta" family="Hennigmatidae" genus="Metatrichopteridium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93CB522122C883DFBA68765B314F8C5" box="[1145,1378,1876,1901]" italics="true" pageId="7" pageNumber="88">Metatrichopteridium</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4812B3122C883DFF488749B740F839" authorityName="Lukashevich, Huang et Lin" authorityYear="2006" box="[151,310,1912,1937]" class="Insecta" family="Hennigmatidae" genus="Daohennigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93CB522122C883DFF488749B740F839" box="[151,310,1912,1937]" italics="true" pageId="7" pageNumber="88">Daohennigma</emphasis>
|
||
</taxonomicName>
|
||
in the long Sc and extremely short M stem (the correspondingly long
|
||
<emphasis id="B93CB522122C883DFB9C8749B227F839" box="[1091,1105,1912,1937]" italics="true" pageId="7" pageNumber="88">d</emphasis>
|
||
cell is hypothesized), and the positions of R
|
||
<subScript id="17CC6B75122C883DFEE0879AB731F811" attach="left" box="[319,327,1963,1977]" fontSize="6" pageId="7" pageNumber="88">4</subScript>
|
||
(ending at wing tip) and
|
||
<emphasis id="B93CB522122C883DFD5C87ACB4CDF81C" box="[643,699,1949,1972]" italics="true" pageId="7" pageNumber="88">m-cu</emphasis>
|
||
(just before M
|
||
<subScript id="17CC6B75122C883DFCAC879AB5FCF811" attach="left" box="[883,906,1963,1977]" fontSize="6" pageId="7" pageNumber="88">3+4</subScript>
|
||
bifurcation). Such strong Sc, as well as 1A terminating at a deep alular incision, has been recorded in both species of
|
||
<taxonomicName id="4C4812B3122C883DFC0987F3B2C9F873" authorityName="Handlirsch" authorityYear="1939" box="[982,1215,1986,2011]" class="Insecta" family="Hennigmatidae" genus="Metatrichopteridium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="88" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93CB522122C883DFC0987F3B2C9F873" box="[982,1215,1986,2011]" italics="true" pageId="7" pageNumber="88">Metatrichopteridium</emphasis>
|
||
</taxonomicName>
|
||
(basiala is not seen on the single specimen of
|
||
<taxonomicName id="4C4812B312238832FE1880A6B413FF18" authorityName="Lukashevich, Huang et Lin" authorityYear="2006" box="[455,613,151,176]" class="Insecta" family="Hennigmatidae" genus="Daohennigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93CB52212238832FE1880A6B413FF18" box="[455,613,151,176]" italics="true" pageId="8" pageNumber="89">Daohennigma</emphasis>
|
||
</taxonomicName>
|
||
); a similar set of supernumerary crossveins is known in
|
||
<taxonomicName id="4C4812B312238832FB2980A6B3E2FF18" authorityName="Lukashevich, Huang et Lin" authorityYear="2006" box="[1270,1428,151,176]" class="Insecta" family="Hennigmatidae" genus="Daohennigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93CB52212238832FB2980A6B3E2FF18" box="[1270,1428,151,176]" italics="true" pageId="8" pageNumber="89">Daohennigma</emphasis>
|
||
</taxonomicName>
|
||
. The wing under description is distinguishable from
|
||
<taxonomicName id="4C4812B312238832FD38808DB5F0FF7D" authorityName="Lukashevich, Huang et Lin" authorityYear="2006" box="[743,902,188,213]" class="Insecta" family="Hennigmatidae" genus="Daohennigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93CB52212238832FD38808DB5F0FF7D" box="[743,902,188,213]" italics="true" pageId="8" pageNumber="89">Daohennigma</emphasis>
|
||
</taxonomicName>
|
||
by the more distal position of unaligned
|
||
<emphasis id="B93CB52212238832FABE808DB3EAFF7D" box="[1377,1436,188,213]" italics="true" pageId="8" pageNumber="89">r3-r4</emphasis>
|
||
and
|
||
<emphasis id="B93CB52212238832FF1980EEB774FF50" box="[198,258,223,248]" italics="true" pageId="8" pageNumber="89">r4-r5</emphasis>
|
||
, and the absence of spots on crossveins; in fact, these may be species-level differences.
|
||
</paragraph>
|
||
<paragraph id="8BF7693012238832FF188135B767FE78" blockId="8.[151,1436,151,464]" pageId="8" pageNumber="89">
|
||
We agree with
|
||
<bibRefCitation id="EFD914C112238832FEAC8135B453FEB5" author="Ansorge, J." box="[371,549,260,285]" pageId="8" pageNumber="89" pagination="97 - 102" refId="ref4602" refString="Ansorge, J. (2001) Lower Jurassic Hennigmatidae (Diptera) from Germany. Studia dipterologica, 8, 97 - 102." type="journal article" year="2001">Ansorge (2001)</bibRefCitation>
|
||
that the subcostal veinlets mentioned in the original description of
|
||
<taxonomicName id="4C4812B312238832FAF88134B3EDFEB4" authorityName="Shcherbakov" authorityYear="1995" box="[1319,1435,261,284]" class="Insecta" family="Hennigmatidae" genus="Hennigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93CB52212238832FAF88134B3EDFEB4" box="[1319,1435,261,284]" italics="true" pageId="8" pageNumber="89">Hennigma</emphasis>
|
||
</taxonomicName>
|
||
are artifacts of preservation and are absent in the
|
||
<typeStatus id="54F3D79212238832FD668116B56DFEE8" box="[697,795,295,320]" pageId="8" pageNumber="89" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4812B312238832FCE08118B59CFEE8" baseAuthorityName="Shcherbakov" baseAuthorityYear="1995" box="[831,1002,295,320]" class="Insecta" family="Hennigmatidae" genus="Metatrichopteridium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="cladistorum">
|
||
<emphasis id="B93CB52212238832FCE08118B59CFEE8" box="[831,1002,295,320]" italics="true" pageId="8" pageNumber="89">M. cladistorum</emphasis>
|
||
</taxonomicName>
|
||
, as well as in other hennigmatids (
|
||
<figureCitation id="137375B512238832FAB58116B6D3FECD" captionStart="FIGURE 1" captionStartId="2.[151,249,995,1017]" captionTargetBox="[158,1417,396,958]" captionTargetId="figure-804@2.[151,1436,378,970]" captionTargetPageId="2" captionText="FIGURE 1. Wing venation of Hennigmatidae (holotypes): A, Kuperwoodia benefica Lukashevich, 1995; B, Anemeca liya Shcherbakov, 1995; С, Metatrichopteridium cladistorum (Shcherbakov, 1995); D, Daohennigma panops Lukashevich et al., 2006; modified after Shcherbakov et al., 1995 (A–C) and Lukashevich et al., 2006 (D)." figureDoi="http://doi.org/10.5281/zenodo.10099431" httpUri="https://zenodo.org/record/10099431/files/figure.png" pageId="8" pageNumber="89">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="137375B512238832FF6B817DB6B2FECD" box="[180,196,332,357]" captionStart="FIGURE 5" captionStartId="6.[151,249,1931,1953]" captionTargetBox="[201,1387,193,1893]" captionTargetId="figure-52@6.[201,1387,193,1894]" captionTargetPageId="6" captionText="FIGURE 5. Basiala of Hennigmatinae: A, B, Trihennigma zavattierii gen. et sp. nov.: holotype MCNAM-PI 24607, part and inverted counterpart; C, Metatrichopteridium cladistorum (Shcherbakov, 1995): holotype PIN 3965/2789. Abbreviation: ai— alular incision." figureDoi="http://doi.org/10.5281/zenodo.10099441" httpUri="https://zenodo.org/record/10099441/files/figure.png" pageId="8" pageNumber="89">5</figureCitation>
|
||
,
|
||
<figureCitation id="137375B512238832FF0F817DB696FECD" box="[208,224,332,357]" captionStart="FIGURE 6" captionStartId="7.[151,249,796,818]" captionTargetBox="[168,1436,193,775]" captionTargetId="figure-153@7.[151,1436,193,775]" captionTargetPageId="7" captionText="FIGURE 6. Proximal part of the wing of Metatrichopteridium cladistorum (Shcherbakov, 1995): holotype PIN 3965/2789: A, drawing; B, SEM image (SE). Abbreviation: ai—alular incision." figureDoi="http://doi.org/10.5281/zenodo.10099443" httpUri="https://zenodo.org/record/10099443/files/figure.png" pageId="8" pageNumber="89">6</figureCitation>
|
||
). However, his redrawing of the wing base is incorrect (
|
||
<bibRefCitation id="EFD914C112238832FC84817DB589FECD" author="Ansorge, J." box="[859,1023,332,357]" pageId="8" pageNumber="89" pagination="97 - 102" refId="ref4602" refString="Ansorge, J. (2001) Lower Jurassic Hennigmatidae (Diptera) from Germany. Studia dipterologica, 8, 97 - 102." type="journal article" year="2001">Ansorge 2001</bibRefCitation>
|
||
:
|
||
<figureCitation id="137375B512238832FBD5817DB22AFECD" box="[1034,1116,332,357]" captionStart="FIGURE 3" captionStartId="4.[151,249,1877,1899]" captionTargetBox="[171,1415,193,1852]" captionTargetId="figure-38@4.[171,1415,193,1856]" captionTargetPageId="4" captionText="FIGURE 3. A, Metatrichopteridium cladistorum (Shcherbakov, 1995): holotype PIN 3965/2789; B, Daohennigma panops Lukashevich et al., 2006: holotype NIGPAS 133706." figureDoi="http://doi.org/10.5281/zenodo.10099437" httpUri="https://zenodo.org/record/10099437/files/figure.png" pageId="8" pageNumber="89">Fig. 3c</figureCitation>
|
||
):
|
||
<emphasis id="B93CB52212238832FBAD817CB2D2FECC" box="[1138,1188,333,356]" italics="true" pageId="8" pageNumber="89">cu-a</emphasis>
|
||
is certainly preserved as it was shown in
|
||
<bibRefCitation id="EFD914C112238832FEB4815EB4FFFE20" author="Shcherbakov, D. & Lukashevich, E. D. & Blagoderov, V. A." box="[363,649,367,392]" pageId="8" pageNumber="89" pagination="75 - 115" refId="ref6207" refString="Shcherbakov, D. E, Lukashevich, E. D. & Blagoderov, V. A. (1995) Triassic Diptera and initial radiation of the order. International Journal of Dipterolological Research, 6, 75 - 115." type="journal article" year="1995">
|
||
Shcherbakov
|
||
<emphasis id="B93CB52212238832FDDB8140B44AFE20" box="[516,572,367,392]" italics="true" pageId="8" pageNumber="89">et al.</emphasis>
|
||
(1995
|
||
</bibRefCitation>
|
||
: Fig. 33) and can be seen in
|
||
<taxonomicName id="4C4812B312238832FC18815EB220FE20" authority="Lara & Lukashevich, 2013" authorityName="Lara & Lukashevich" authorityYear="2013" box="[967,1110,367,392]" class="Insecta" family="Hennigmatidae" genus="Trihennigma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B93CB52212238832FC18815EB220FE20" box="[967,1110,367,392]" italics="true" pageId="8" pageNumber="89">Trihennigma</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93CB52212238832FB82815EB2B0FE20" bold="true" box="[1117,1222,367,392]" pageId="8" pageNumber="89">
|
||
<taxonomicNameLabel id="A20F085912238832FB82815EB2B0FE20" box="[1117,1222,367,392]" pageId="8" pageNumber="89" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; moreover, in both genera, 2A is not marginal and a small alula is visible in
|
||
<taxonomicName id="4C4812B312238832FCD281A4B5CEFE05" baseAuthorityName="Shcherbakov" baseAuthorityYear="1995" box="[781,952,404,429]" class="Insecta" family="Hennigmatidae" genus="Metatrichopteridium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="cladistorum">
|
||
<emphasis id="B93CB52212238832FCD281A4B5CEFE05" box="[781,952,404,429]" italics="true" pageId="8" pageNumber="89">M. cladistorum</emphasis>
|
||
</taxonomicName>
|
||
and is likely to be present in the new genus (
|
||
<figureCitation id="137375B512238832FF408186B691FE78" box="[159,231,439,464]" captionStart="FIGURE 5" captionStartId="6.[151,249,1931,1953]" captionTargetBox="[201,1387,193,1893]" captionTargetId="figure-52@6.[201,1387,193,1894]" captionTargetPageId="6" captionText="FIGURE 5. Basiala of Hennigmatinae: A, B, Trihennigma zavattierii gen. et sp. nov.: holotype MCNAM-PI 24607, part and inverted counterpart; C, Metatrichopteridium cladistorum (Shcherbakov, 1995): holotype PIN 3965/2789. Abbreviation: ai— alular incision." figureDoi="http://doi.org/10.5281/zenodo.10099441" httpUri="https://zenodo.org/record/10099441/files/figure.png" pageId="8" pageNumber="89">Figs 5</figureCitation>
|
||
,
|
||
<figureCitation id="137375B512238832FF2C8186B775FE78" box="[243,259,439,464]" captionStart="FIGURE 6" captionStartId="7.[151,249,796,818]" captionTargetBox="[168,1436,193,775]" captionTargetId="figure-153@7.[151,1436,193,775]" captionTargetPageId="7" captionText="FIGURE 6. Proximal part of the wing of Metatrichopteridium cladistorum (Shcherbakov, 1995): holotype PIN 3965/2789: A, drawing; B, SEM image (SE). Abbreviation: ai—alular incision." figureDoi="http://doi.org/10.5281/zenodo.10099443" httpUri="https://zenodo.org/record/10099443/files/figure.png" pageId="8" pageNumber="89">6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |