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<document id="4A7AC25055E0A07A2E5DB3B4B4FB1215" ID-DOI="10.11646/zootaxa.3948.1.2" ID-GBIF-Dataset="9e109059-3d75-4e2c-997e-6f4217d201d0" ID-ISSN="1175-5326" ID-Zenodo-Dep="244003" ID-ZooBank="9A0330AD-EEF7-4A80-8D80-1B6116BD856F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461042933937" checkinUser="plazi" docAuthor="Filippini, Valentina, Galante, Eduardo &amp; Micó, Estefanía" docDate="2015" docId="03E1981EB533FFE4FF59F96F044DEFE6" docLanguage="en" docName="zt03948p040.pdf" docOrigin="Zootaxa 3948 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anomala aglaos Filippini, Galante" docType="treatment" docVersion="7" masterDocId="FFD8E066B530FFE2FFCEFFD3014CEC5D" masterDocTitle="Description of six new species of Anomalini from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae)" updateTime="1698576078296" updateUser="plazi">
<mods:mods id="7335236D95BCCBB23B7384707D8CDDAD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3C7E4743F7A7EB1884A786706D1C48C0">
<mods:title id="D81274DC3AB7FABB32B4F77FF9A455CC">Description of six new species of Anomalini from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae)</mods:title>
</mods:titleInfo>
<mods:name id="0622F86FE771C06A1E36B4AA00FA493B" type="personal">
<mods:role id="034012B716CABA91F272E410D27DB43A">
<mods:roleTerm id="9115611C180EA319BBB2ACCB3DB9286B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DB4E54CD6E2C6E180F499333EC21C991">Filippini, Valentina</mods:namePart>
</mods:name>
<mods:name id="FD61312B7218F8A3F27DD5F83537C277" type="personal">
<mods:role id="191DF422253E301E44E476D96DC53ABC">
<mods:roleTerm id="41317D63644F56C6BA0770006BEBCB39">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F6044389B7C62E675D45F5319B3A972F">Galante, Eduardo</mods:namePart>
</mods:name>
<mods:name id="969CE8A074FF3E6B3252866C851C2076" type="personal">
<mods:role id="F26E09A6BFC21EBCC3CA07A7232FA55A">
<mods:roleTerm id="1C51B6E64182AD1C58B007B3AA6F56D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B9E81D5618D556FD18DEDE8EBF41FE6E">Micó, Estefanía</mods:namePart>
</mods:name>
<mods:typeOfResource id="4E470B5698CD2DF39FAD51932BC2BAAF">text</mods:typeOfResource>
<mods:relatedItem id="69A52572ED3D513D5B64FA6942E85253" type="host">
<mods:titleInfo id="3B58D20B9E18531B9F35C38B14613D6E">
<mods:title id="BE761D033967A4D87CD16BD77633ECF7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9CD216D92098C0C751E7287CB28E08F6">
<mods:date id="EF337CCD9311BD9674FF2B2A595855CF">2015</mods:date>
<mods:detail id="34FBC9F1954E32B894A2086E4E59B1A0" type="volume">
<mods:number id="389B2B4B4101D9CD2AAEB894AE1922A6">3948</mods:number>
</mods:detail>
<mods:detail id="198D33665FE62541DD4333768CCF400C" type="issue">
<mods:number id="5FAB93FEF052188CC5BE760DE3FB56AF">1</mods:number>
</mods:detail>
</mods:part>
</mods:relatedItem>
<mods:classification id="CF24969857AE0AD05D9516C43C7F8BB0">journal volume</mods:classification>
<mods:identifier id="AF6087359455A0BF401B6BC7C458E6D9" type="DOI">10.11646/zootaxa.3948.1.2</mods:identifier>
<mods:identifier id="5A99C685B5A036FC077AACD4884F9F34" type="GBIF-Dataset">9e109059-3d75-4e2c-997e-6f4217d201d0</mods:identifier>
<mods:identifier id="36F5C3273E032A694A6068547B3B3C64" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="AD61ABFAD883E818274419BCCDA91F88" type="Zenodo-Dep">244003</mods:identifier>
<mods:identifier id="4D92477AF80372CED927AD6EDA15CEC4" type="ZooBank">9A0330AD-EEF7-4A80-8D80-1B6116BD856F</mods:identifier>
</mods:mods>
<treatment id="03E1981EB533FFE4FF59F96F044DEFE6" ID-DOI="http://doi.org/10.5281/zenodo.6122698" ID-GBIF-Taxon="119609993" ID-Zenodo-Dep="6122698" LSID="urn:lsid:plazi:treatment:03E1981EB533FFE4FF59F96F044DEFE6" httpUri="http://treatment.plazi.org/id/03E1981EB533FFE4FF59F96F044DEFE6" lastPageId="6" pageId="3">
<subSubSection id="C3527A83B533FFE1FF59F96F0195EAAA" pageId="3" type="nomenclature">
<paragraph id="8BF72908B533FFE1FF59F96F0200EA8B" blockId="3.[151,844,1724,1783]" box="[151,844,1724,1750]" pageId="3">
<heading id="D0BF9E64B533FFE1FF59F96F0200EA8B" bold="true" box="[151,844,1724,1750]" fontSize="11" level="1" pageId="3" reason="1">
<emphasis id="B93CF51AB533FFE1FF59F96F0200EA8B" bold="true" box="[151,844,1724,1750]" pageId="3">
<taxonomicName id="4C48528BB533FFE1FF59F96F030EEA8B" authority="Filippini, Galante" authorityName="Filippini, Galante" box="[151,578,1724,1750]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="aglaos">
<emphasis id="B93CF51AB533FFE1FF59F96F0017EA8B" bold="true" box="[151,347,1724,1750]" italics="true" pageId="3">Anomala aglaos</emphasis>
<collectingCountry id="F35F6998B533FFE1FEACF96F0083EA8B" box="[354,463,1724,1750]" name="Philippines" pageId="3">Filippini</collectingCountry>
, Galante
</taxonomicName>
, &amp; Micó, new species
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF72908B533FFE1FF59F90D0195EAAA" blockId="3.[151,844,1724,1783]" box="[151,217,1758,1783]" pageId="3">
<figureCitation id="1373358DB533FFE1FF59F90D0195EAAA" box="[151,217,1758,1783]" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1977,2000]" captionTargetBox="[186,1400,705,1952]" captionTargetId="figure@4.[179,1408,697,1957]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Habitus. 1: Anomala aglaos (holotype). 2: A. estrella (holotype). 3: A. inbio (holotype). 4: A. pincelada (holotype). 5: Callistethus ruteloides (holotype). 6: C. yalizo (holotype). Scale = 5 mm." httpUri="https://zenodo.org/record/244004/files/figure.png" pageId="3">Fig. 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3527A83B533FFE4FF59F8F5020FEDFC" lastPageId="6" pageId="3" type="materials_examined">
<paragraph id="8BF72908B533FFE1FF59F8F50306EB3E" blockId="3.[151,1437,1830,2036]" pageId="3">
<emphasis id="B93CF51AB533FFE1FF59F8F500CBEB62" bold="true" box="[151,391,1830,1855]" pageId="3">Material examined.</emphasis>
<typeStatus id="54F397AAB533FFE1FE5BF8F5034EEB62" box="[405,514,1830,1855]" pageId="3" type="holotype">Holotype</typeStatus>
: ♂ &quot;La Montura, Prov. San José,
<collectingCountry id="F35F6998B533FFE1FC68F8F40564EB62" box="[934,1064,1830,1855]" name="Costa Rica" pageId="3">Costa Rica</collectingCountry>
<date id="FFF60FC8B533FFE1FBFBF8F405F8EB62" box="[1077,1204,1831,1855]" pageId="3" value="2007-11-07">07/11/2007</date>
Leg. M. Moraga /
<collectionCode id="ED59B1CDB533FFE1FF59F89801A8EB39" box="[151,228,1867,1892]" pageId="3">CEUA</collectionCode>
00106184&quot; deposited in
<taxonomicName id="4C48528BB533FFE1FE3BF898030AEB3E" box="[501,582,1867,1891]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="3" phylum="Arthropoda" rank="species" species="inbio">
<collectionCode id="ED59B1CDB533FFE1FE3BF898030AEB3E" box="[501,582,1867,1891]" pageId="3">INBIO</collectionCode>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF72908B533FFE6FF09F8A305A3ED40" blockId="3.[151,1437,1830,2036]" lastBlockId="4.[151,1437,151,644]" lastPageId="4" pageId="3">
<typeStatus id="54F397AAB533FFE1FF09F8A30078EBD5" box="[199,308,1904,1928]" pageId="3" type="paratype">Paratypes</typeStatus>
(6):
<specimenCount id="9D4EE281B533FFE1FEA4F8BC00D8EBDA" box="[362,404,1903,1927]" pageId="3" type="generic">1 ♂</specimenCount>
&quot;
<collectingCountry id="F35F6998B533FFE1FE6AF8BD0083EBDA" box="[420,463,1902,1927]" name="Iceland" pageId="3">Isla</collectingCountry>
Bonita, Prov. Alajuela,
<collectingCountry id="F35F6998B533FFE1FD13F8BC0216EBDA" box="[733,858,1902,1927]" name="Costa Rica" pageId="3">Costa Rica</collectingCountry>
.
<date id="FFF60FC8B533FFE1FCABF8BD02AFEBDA" box="[869,995,1902,1927]" pageId="3" value="2007-06-11">11/06/2007</date>
Leg. M. Moraga /
<collectionCode id="ED59B1CDB533FFE1FB76F8BD044BEBDA" box="[1208,1287,1902,1927]" pageId="3">CEUA</collectionCode>
00106183&quot;;
<specimenCount id="9D4EE281B533FFE1FA40F8BC01E0EBF6" pageId="3" type="generic">1 ♂</specimenCount>
&quot;
<collectingCountry id="F35F6998B533FFE1FF73F840002AEBF6" box="[189,358,1939,1963]" name="Costa Rica" pageId="3">COSTA RICA</collectingCountry>
Prov. San José. P.N. Braulio Carrillo. Sitio La Montura.
<quantity id="4CB084EDB533FFE1FC38F840050AEBF6" box="[1014,1094,1939,1963]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="3" unit="m" value="1100.0">1100m</quantity>
.
<date id="FFF60FC8B533FFE1FB9DF84005BDEBF6" box="[1107,1265,1939,1963]" pageId="3" value="2007-06-13">13 JUN 2007</date>
. A. García, M. Moraga, M.A. Zumbado. Tp. Luz 1. L_N_232500_539125 #92217 /
<collectionCode id="ED59B1CDB533FFE1FC53F86502A0EB92" box="[925,1004,1974,1999]" pageId="3">CEUA</collectionCode>
00106185&quot;; 1 ♀ &quot;
<collectingCountry id="F35F6998B533FFE1FB63F8640414EB92" box="[1197,1368,1975,1999]" name="Costa Rica" pageId="3">COSTA RICA</collectingCountry>
. Prov. Alajuela. Guatuso. P.N. Volcán Tenorio. Punto 1: Catarata Río Buenavista.
<specimenCount id="9D4EE281B533FFE1FC27F808057DEBAE" box="[1001,1073,2011,2035]" pageId="3" type="generic">
<quantity id="4CB084EDB533FFE1FC27F8080560EBAE" box="[1001,1068,2011,2035]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" unit="m" value="800.0">800m</quantity>
.
</specimenCount>
<date id="FFF60FC8B533FFE1FBF6F8080588EBAE" box="[1080,1220,2011,2036]" pageId="3" value="2008-06-05">5 JUN 2008</date>
. J. A. Azofeifa. Tp de Luz 2. L_N_298380_427850 #93849 /
<collectionCode id="ED59B1CDB534FFE6FD4BFF440398ECED" box="[645,724,151,176]" pageId="4">CEUA</collectionCode>
00106186&quot;; 1 ♀ &quot;
<collectingCountry id="F35F6998B534FFE6FC51FF4B0505ECED" box="[927,1097,152,176]" name="Costa Rica" pageId="4">COSTA RICA</collectingCountry>
Prov. San José. P.N. Braulio Carrillo. Sitio La Montura.
<quantity id="4CB084EDB534FFE6FE05FF6F0357EC89" box="[459,539,188,212]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="4" unit="m" value="1100.0">1100m</quantity>
.
<date id="FFF60FC8B534FFE6FDE8FF6F03F0EC89" box="[550,700,188,212]" pageId="4" value="2007-07-12">12 JUL 2007</date>
. A. García, M. Moraga. Tp. Luz 2. L_N_232500_539125 #92245 /
<collectionCode id="ED59B1CDB534FFE6FF66FF0C01BBECA5" box="[168,247,223,248]" pageId="4">CEUA</collectionCode>
00106187&quot;; 1 ♀ &quot;La Montura Prov. San José
<collectingCountry id="F35F6998B534FFE6FCC8FF3302CEECA5" box="[774,898,223,248]" name="Costa Rica" pageId="4">Costa Rica</collectingCountry>
<date id="FFF60FC8B534FFE6FC42FF0C0540ECA5" box="[908,1036,223,248]" pageId="4" value="2007-06-13">13/06/2007</date>
Det. D. Briceño&quot; INBIOCRI; 1 ♀ &quot;
<collectingCountry id="F35F6998B534FFE6FF6EFED70187ED40" box="[160,203,260,285]" name="Iceland" pageId="4">Isla</collectingCountry>
Bonita, Prov. Alajuela,
<collectingCountry id="F35F6998B534FFE6FE16FED7031AED40" box="[472,598,260,285]" name="Costa Rica" pageId="4">Costa Rica</collectingCountry>
.
<date id="FFF60FC8B534FFE6FDAEFED7039DED40" box="[608,721,260,285]" pageId="4" value="2007-11-06">6/11/2007</date>
Leg. A. García, M. Moraga /
<collectionCode id="ED59B1CDB534FFE6FBEEFED70523ED40" box="[1056,1135,260,285]" pageId="4">CEUA</collectionCode>
00106189&quot;.
</paragraph>
<paragraph id="8BF72908B534FFE6FF09FEF4053BED62" blockId="4.[151,1437,151,644]" box="[199,1143,295,320]" pageId="4">
<emphasis id="B93CF51AB534FFE6FF09FEF40344ED1D" bold="true" box="[199,520,295,320]" pageId="4">
<typeStatus id="54F397AAB534FFE6FF09FEF40078ED1D" box="[199,308,295,320]" pageId="4" type="holotype">Holotype</typeStatus>
male description.
</emphasis>
Body shape elongate. Length
<quantity id="4CB084EDB534FFE6FC93FEF40288ED62" box="[861,964,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" pageId="4" unit="mm" value="17.5">17.5 mm</quantity>
. Width
<quantity id="4CB084EDB534FFE6FBD4FEFB053FED62" box="[1050,1139,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="4" unit="mm" value="9.7">9.7 mm</quantity>
.
</paragraph>
<paragraph id="8BF72908B534FFE6FF09FE9F0334EDF1" blockId="4.[151,1437,151,644]" pageId="4">Color. Head, pronotum, and scutellum metallic green. Elytra metallic green with branched ochre lines along apical portion of costae and coppery luster. Pygidium bronze. Legs metallic green with coppery luster. Venter dark metallic green, abdominal sternites copper.</paragraph>
<paragraph id="8BF72908B534FFE6FF09FE6405FAEE45" blockId="4.[151,1437,151,644]" pageId="4">Clypeus trapezoidal, surface densely punctate, convex. Clypeus anterior margin slightly sinuate, anteriorly thick. Clypeus ratio width/length 2.5. Frons moderately punctate, flat. Ocular canthum long, thin, with rounded apex. Interocular ratio (interocular width/width of eye): 3.0. Antenna: ratio funiculus/club 0.7.</paragraph>
<paragraph id="8BF72908B534FFE6FF09FDF60047EED9" blockId="4.[151,1437,151,644]" pageId="4">Pronotum subtrapezoidal, ratio width/length 1.9. Lateral margins sinuate. Anterior angles acute, sharp; posterior angles obtuse, blunt. Basal margin sinuate, subapical bead obliterated in the middle. Surface with fine punctures.</paragraph>
<caption id="DF377980B534FFE6FF59F86A0540EBB2" httpUri="https://zenodo.org/record/244004/files/figure.png" pageId="4" targetBox="[186,1400,705,1952]" targetPageId="4">
<paragraph id="8BF72908B534FFE6FF59F86A0540EBB2" blockId="4.[151,1436,1977,2031]" pageId="4">
<emphasis id="B93CF51AB534FFE6FF59F86A0072EB92" bold="true" box="[151,318,1977,2000]" pageId="4">FIGURES 1–6.</emphasis>
Habitus. 1:
<taxonomicName id="4C48528BB534FFE6FE08F869033CEB92" box="[454,624,1978,1999]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="aglaos">
<emphasis id="B93CF51AB534FFE6FE08F869033CEB92" box="[454,624,1978,1999]" italics="true" pageId="4">Anomala aglaos</emphasis>
</taxonomicName>
(holotype). 2:
<taxonomicName id="4C48528BB534FFE6FCDDF8680232EB92" box="[787,894,1978,2000]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="estrella">
<emphasis id="B93CF51AB534FFE6FCDDF8680232EB92" box="[787,894,1978,2000]" italics="true" pageId="4">A. estrella</emphasis>
</taxonomicName>
(holotype). 3:
<taxonomicName id="4C48528BB534FFE6FBEFF8680539EB92" box="[1057,1141,1978,2000]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="inbio">
<emphasis id="B93CF51AB534FFE6FBEFF8680539EB92" box="[1057,1141,1978,2000]" italics="true" pageId="4">A. inbio</emphasis>
</taxonomicName>
(holotype). 4:
<taxonomicName id="4C48528BB534FFE6FAD6F86804D0EB92" box="[1304,1436,1978,2000]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="pincelada">
<emphasis id="B93CF51AB534FFE6FAD6F86804D0EB92" box="[1304,1436,1978,2000]" italics="true" pageId="4">A. pincelada</emphasis>
</taxonomicName>
(holotype). 5:
<taxonomicName id="4C48528BB534FFE6FEE9F80A0347EBB3" box="[295,523,2009,2030]" class="Insecta" family="Rutelidae" genus="Callistethus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="ruteloides">
<emphasis id="B93CF51AB534FFE6FEE9F80A0347EBB3" box="[295,523,2009,2030]" italics="true" pageId="4">Callistethus ruteloides</emphasis>
</taxonomicName>
(holotype). 6:
<taxonomicName id="4C48528BB534FFE6FD6CF80A03B0EBB3" box="[674,764,2009,2030]" class="Insecta" family="Rutelidae" genus="Callistethus" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="yalizo">
<emphasis id="B93CF51AB534FFE6FD6CF80A03B0EBB3" box="[674,764,2009,2030]" italics="true" pageId="4">C. yalizo</emphasis>
</taxonomicName>
(holotype). Scale = 5 mm.
</paragraph>
</caption>
<paragraph id="8BF72908B535FFE7FF59FA0400C3EADB" blockId="5.[151,1437,1495,1670]" pageId="5">
<emphasis id="B93CF51AB535FFE7FF59FA04006DE9B1" bold="true" box="[151,289,1495,1517]" pageId="5">Figures 7–12</emphasis>
. Shape of protibia in 7:
<taxonomicName id="4C48528BB535FFE7FDDDFA0403F4E9B1" box="[531,696,1495,1516]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="aglaos">
<emphasis id="B93CF51AB535FFE7FDDDFA0403F4E9B1" box="[531,696,1495,1516]" italics="true" pageId="5">Anomala aglaos</emphasis>
</taxonomicName>
(male:
<typeStatus id="54F397AAB535FFE7FCCAFA04022CE9B0" box="[772,864,1495,1517]" pageId="5" type="holotype">holotype</typeStatus>
; female:
<typeStatus id="54F397AAB535FFE7FC75FA0B0559E9B0" box="[955,1045,1496,1517]" pageId="5" type="paratype">paratype</typeStatus>
,
<collectingCountry id="F35F6998B535FFE7FBD1FA04050FE9B0" box="[1055,1091,1495,1517]" name="Iceland" pageId="5">Isla</collectingCountry>
Bonita,
<collectingCountry id="F35F6998B535FFE7FB57FA040440E9B0" box="[1177,1292,1495,1517]" name="Costa Rica" pageId="5">Costa Rica</collectingCountry>
). 8:
<taxonomicName id="4C48528BB535FFE7FAFBFA0B04D0E9B1" box="[1333,1436,1495,1517]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="estrella">
<emphasis id="B93CF51AB535FFE7FAFBFA0B04D0E9B1" box="[1333,1436,1495,1517]" italics="true" pageId="5">A. estrella</emphasis>
</taxonomicName>
(male:
<typeStatus id="54F397AAB535FFE7FF2DFA250073EA51" box="[227,319,1526,1548]" pageId="5" type="holotype">holotype</typeStatus>
; female:
<typeStatus id="54F397AAB535FFE7FE68FA24034CEA51" box="[422,512,1527,1548]" pageId="5" type="paratype">paratype</typeStatus>
, Los Almendros,
<collectingCountry id="F35F6998B535FFE7FD0DFA250270EA51" box="[707,828,1526,1548]" name="Costa Rica" pageId="5">Costa Rica</collectingCountry>
). 9:
<taxonomicName id="4C48528BB535FFE7FCBFFA25028AEA56" box="[881,966,1526,1547]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="inbio">
<emphasis id="B93CF51AB535FFE7FCBFFA25028AEA56" box="[881,966,1526,1547]" italics="true" pageId="5">A. inbio</emphasis>
</taxonomicName>
(male:
<typeStatus id="54F397AAB535FFE7FBD0FA250536EA51" box="[1054,1146,1526,1548]" pageId="5" type="holotype">holotype</typeStatus>
; female:
<typeStatus id="54F397AAB535FFE7FB2FFA240477EA51" box="[1249,1339,1527,1548]" pageId="5" type="paratype">paratype</typeStatus>
, Reserva Biológica Hitoy Cerere,
<collectingCountry id="F35F6998B535FFE7FE55F9C7035EEA77" box="[411,530,1556,1578]" name="Costa Rica" pageId="5">Costa Rica</collectingCountry>
). 10:
<taxonomicName id="4C48528BB535FFE7FD9FF9C60399EA74" box="[593,725,1556,1578]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="pincelada">
<emphasis id="B93CF51AB535FFE7FD9FF9C60399EA74" box="[593,725,1556,1578]" italics="true" pageId="5">A. pincelada</emphasis>
</taxonomicName>
(male:
<typeStatus id="54F397AAB535FFE7FCE4F9C602C8EA77" box="[810,900,1557,1578]" pageId="5" type="paratype">paratype</typeStatus>
, Parque Nacional Santa Rosa,
<collectingCountry id="F35F6998B535FFE7FB02F9C7040EEA77" box="[1228,1346,1556,1578]" name="Costa Rica" pageId="5">Costa Rica</collectingCountry>
; female:
<typeStatus id="54F397AAB535FFE7FF59F9E701BDEA14" box="[151,241,1588,1609]" pageId="5" type="paratype">paratype</typeStatus>
, Tierras Morenas,
<collectingCountry id="F35F6998B535FFE7FE61F9E0036FEA14" box="[431,547,1587,1609]" name="Costa Rica" pageId="5">Costa Rica</collectingCountry>
). 11:
<taxonomicName id="4C48528BB535FFE7FD95F9E0020CEA15" box="[603,832,1587,1608]" class="Insecta" family="Rutelidae" genus="Callistethus" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="ruteloides">
<emphasis id="B93CF51AB535FFE7FD95F9E0020CEA15" box="[603,832,1587,1608]" italics="true" pageId="5">Callistethus ruteloides</emphasis>
</taxonomicName>
(male:
<typeStatus id="54F397AAB535FFE7FC41F9E002A7EA14" box="[911,1003,1587,1609]" pageId="5" type="holotype">holotype</typeStatus>
; female:
<typeStatus id="54F397AAB535FFE7FB87F9E705EFEA14" box="[1097,1187,1588,1609]" pageId="5" type="paratype">paratype</typeStatus>
, El Copal, Cartago). 12: Shape of protibia of
<taxonomicName id="4C48528BB535FFE7FEA6F9810368EA3A" box="[360,548,1618,1639]" class="Insecta" family="Rutelidae" genus="Callistethus" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="yalizo">
<emphasis id="B93CF51AB535FFE7FEA6F9810368EA3A" box="[360,548,1618,1639]" italics="true" pageId="5">Callistethus yalizo</emphasis>
</taxonomicName>
(male:
<typeStatus id="54F397AAB535FFE7FDBCF9800380EA35" box="[626,716,1619,1640]" pageId="5" type="paratype">paratype</typeStatus>
, Esperanza del Guarco, Cartago; female:
<typeStatus id="54F397AAB535FFE7FBBFF9800587EA35" box="[1137,1227,1619,1640]" pageId="5" type="paratype">paratype</typeStatus>
, Quebrada Segunda, Cartago). Scale =
<quantity id="4CB084EDB535FFE7FE83F9A300C7EADB" box="[333,395,1648,1670]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="5" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
<paragraph id="8BF72908B535FFE7FF09F91F058EEAB8" blockId="5.[151,1437,1740,2020]" box="[199,1218,1740,1765]" pageId="5">Scutellum. Shape subtriangular, with rounded sides and blunt apex. Ratio width/length 1.4.</paragraph>
<paragraph id="8BF72908B535FFE7FF09F92300FCEB70" blockId="5.[151,1437,1740,2020]" pageId="5">Elytra with costae defined by irregular rows of sparse punctures. Interstices with sparse, shallow punctures; surface slightly irregular.</paragraph>
<paragraph id="8BF72908B535FFE7FF09F8EB0534EB0C" blockId="5.[151,1437,1740,2020]" box="[199,1144,1848,1873]" pageId="5">Pygidium sparsely punctate with long setae at apical margin. Ratio width/length 1.5.</paragraph>
<paragraph id="8BF72908B535FFE7FF09F88E0552EBE2" blockId="5.[151,1437,1740,2020]" pageId="5">Space between the mesocoxae narrow, flat. Mesometasternal suture well defined, at base of mesocoxae. Abdominal sternites with 1–2 rows of setae (2–3 rows on 2nd sternite) and 6–8 rows of punctures per sternite. Last sternite punctate, with subapical bead thin and sinuate, apical membrane narrow.</paragraph>
<paragraph id="8BF72908B535FFE4FF09F818053FECA5" blockId="5.[151,1437,1740,2020]" lastBlockId="6.[151,1437,151,955]" lastPageId="6" pageId="5">Protibia (Fig. 7) bidentate; apical tooth long and curved; basal tooth at same level as internal apex of protibia, triangular in shape, acute. Metatibia slender, fusiform. Ratio length/width 3.2. First external carina obliterated. Surface punctate above second external carina and rugose below. Protarsal claws: internal claw bifurcate, with upper branch of same length and 2/3 the width of the lower one, inferior margin sinuate.</paragraph>
<paragraph id="8BF72908B536FFE4FF59FED7020FEDFC" blockId="6.[151,1437,151,955]" pageId="6">
Aedeagus (
<figureCitation id="1373358DB536FFE4FEDAFED7002BED41" box="[276,359,260,285]" captionStart="FIGURES 13 – 18" captionStartId="6.[151,264,1885,1908]" captionTargetBox="[153,1429,1020,1837]" captionTargetId="figure@6.[151,1436,1010,1864]" captionTargetPageId="6" captionText="FIGURES 13 – 18. Shape of aedeagus, lateral view (top), dorsal view (bottom left), ventral view (bottom right). 13: Anomala aglaos (paratype, Isla Bonita, Costa Rica). 14: A. estrella (holotype). 15: A. inbio (holotype). 16: A. pincelada (holotype). 17: Callistethus ruteloides (holotype). 18: C. yalizo (Cerro Chompipe, Heredia). Scale = 1 mm." httpUri="https://zenodo.org/record/244005/files/figure.png" pageId="6">Fig. 13</figureCitation>
): parameres large and stout in a lateral view, but narrow dorsally. Ventral plate with curved apex in a lateral view. Endophallus (
<figureCitation id="1373358DB536FFE4FDC2FEF6032EED63" box="[524,610,293,318]" captionStart="FIGURES 19 – 22" captionStartId="7.[151,264,1709,1732]" captionTargetBox="[155,1421,200,1673]" captionTargetId="figure@7.[151,1436,193,1688]" captionTargetPageId="7" captionText="FIGURES 19 – 22. Endophallus of 19: Anomala aglaos, (a) particular in frontal view, (b) lateral view (paratype, Isla Bonita, Costa Rica). 20: A. estrella (paratype, Hacienda Tiquires, Costa Rica). 21: A. inbio (holotype). 22: A. pincelada (paratype, Cuajiniquil, Costa Rica). Scale = 1 mm." httpUri="https://zenodo.org/record/244006/files/figure.png" pageId="6">Fig. 19</figureCitation>
) with 2 long sacculi; 1 ending with a pair of thick, long, sclerotized projections with a long diverticle in between, and with a patch of sclerotized spines medially; the other sacculi with swollen base, where ejaculatory sclerite and duct opening are located, and a ridged plate of thick spines at apex. Median lobes asymmetric, 1 with a triangular fold near apex.
</paragraph>
</subSubSection>
<subSubSection id="C3527A83B536FFE4FF09FE79039DEE2A" pageId="6" type="description">
<paragraph id="8BF72908B536FFE4FF09FE7902DBEE57" blockId="6.[151,1437,151,955]" pageId="6">
<emphasis id="B93CF51AB536FFE4FF09FE790077ED9E" bold="true" box="[199,315,426,451]" pageId="6">Variation.</emphasis>
Ochre lines on elytra of variable length; elytra with green or copper luster. Female. Similar to male, protibia (Fig. 7) with longer and wider apical tooth; basal tooth above internal apex; internal protarsal claw narrower with curved inferior margin; last sternite with curved apex.
</paragraph>
<paragraph id="8BF72908B536FFE4FF09FDC5039DEE2A" blockId="6.[151,1437,151,955]" pageId="6">
Body length
<quantity id="4CB084EDB536FFE4FEAEFDC50345EE73" box="[352,521,534,559]" metricMagnitude="-2" metricUnit="m" metricValue="1.815" metricValueMax="1.88" metricValueMin="1.75" pageId="6" unit="mm" value="18.15" valueMax="18.8" valueMin="17.5">17.5–18.8 mm</quantity>
, body width
<quantity id="4CB084EDB536FFE4FD69FDC4020FEE73" box="[679,835,535,559]" metricMagnitude="-3" metricUnit="m" metricValue="9.95" metricValueMax="10.2" metricValueMin="9.7" pageId="6" unit="mm" value="9.95" valueMax="10.2" valueMin="9.7">9.7–10.2 mm</quantity>
. Clypeus w/l: 2.3–2.7. Interocular ratio (interocular width/width of eye): 3.0–3.4. Antenna: ratio funiculus/club 0.7–0.9. Pronotum w/l: 1.8–1.9. Scutellum w/l: 1.4– 1.6. Pygidium w/l: 1.5–1.6. Metatibia l/w: 2.9–3.5.
</paragraph>
</subSubSection>
<subSubSection id="C3527A83B536FFE4FF09FD51028DEF12" pageId="6" type="diagnosis">
<paragraph id="8BF72908B536FFE4FF09FD5104D8EEE2" blockId="6.[151,1437,151,955]" pageId="6">
<emphasis id="B93CF51AB536FFE4FF09FD51000DEEC6" bold="true" box="[199,321,642,667]" pageId="6">Diagnosis.</emphasis>
Size large, shape elongate, metallic green color with branched ochre lines on apical portion of elytra, elytral surface irregular, parameres large and stout, endophallus with a pair of long, sclerotized projections.
</paragraph>
<paragraph id="8BF72908B536FFE4FF09FD19028DEF12" blockId="6.[151,1437,151,955]" pageId="6">
This species is similar to
<taxonomicName id="4C48528BB536FFE4FE29FD1803B6EEBE" authority="Ohaus, 1933" authorityName="Ohaus" authorityYear="1933" box="[487,762,714,739]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="obovata">
<emphasis id="B93CF51AB536FFE4FE29FD18032DEEBE" box="[487,609,715,739]" italics="true" pageId="6">A. obovata</emphasis>
Ohaus, 1933
</taxonomicName>
, with the slightly bigger size, oblong shape, and dark green color; but
<taxonomicName id="4C48528BB536FFE4FEC1FD3C00C6EF5B" box="[271,394,750,774]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="obovata">
<emphasis id="B93CF51AB536FFE4FEC1FD3C00C6EF5B" box="[271,394,750,774]" italics="true" pageId="6">A. obovata</emphasis>
</taxonomicName>
lacks the ochre lines on elytra, its elytral surface is strongly irregular with large and deep coalescing punctures, and it has slender parameres (
<figureCitation id="1373358DB536FFE4FD3EFCC1020BEF76" box="[752,839,786,811]" captionStart="FIGURES 27 – 33. 27" captionStartId="14.[151,264,1539,1562]" captionTargetBox="[156,1433,641,1510]" captionTargetId="figure@14.[151,1436,630,1517]" captionTargetPageId="14" captionText="FIGURES 27 – 33. 27: Aedeagus of Anomala obovata, lateral view (Quebrada Kuisa, Costa Rica, INBIO). 28: Endophallus of A. obovata (Cerro Chompipe, Costa Rica, INBIO). 29: Aedeagus of A. jansoni, lateral view (Monte Rotondo, Costa Rica, syntype, MNHUB). 30: Aedeagus of A. antica, lateral view (Estación Palo Verde, Costa Rica, INBIO). 31: Aedeagus of A. luciae, lateral view (Santa Lucia, Antilles, lectotype, MNHN). 32: Aedeagus of A. quiche, lateral view (Estación Maritza, Costa Rica, INBIO). 33: Aedeagus of A. inconstans, (a) lateral view, (b) other side of parameres (Bogotá, Colombia, syntype, MLUH). Scale = 1 mm." httpUri="https://zenodo.org/record/244009/files/figure.png" pageId="6">Fig. 27</figureCitation>
), endophallus with a sacculus with a large patch of dense setae on basal portion, instead of sclerotized projections (
<figureCitation id="1373358DB536FFE4FC91FCE502F8EF12" box="[863,948,822,847]" captionStart="FIGURES 27 – 33. 27" captionStartId="14.[151,264,1539,1562]" captionTargetBox="[156,1433,641,1510]" captionTargetId="figure@14.[151,1436,630,1517]" captionTargetPageId="14" captionText="FIGURES 27 – 33. 27: Aedeagus of Anomala obovata, lateral view (Quebrada Kuisa, Costa Rica, INBIO). 28: Endophallus of A. obovata (Cerro Chompipe, Costa Rica, INBIO). 29: Aedeagus of A. jansoni, lateral view (Monte Rotondo, Costa Rica, syntype, MNHUB). 30: Aedeagus of A. antica, lateral view (Estación Palo Verde, Costa Rica, INBIO). 31: Aedeagus of A. luciae, lateral view (Santa Lucia, Antilles, lectotype, MNHN). 32: Aedeagus of A. quiche, lateral view (Estación Maritza, Costa Rica, INBIO). 33: Aedeagus of A. inconstans, (a) lateral view, (b) other side of parameres (Bogotá, Colombia, syntype, MLUH). Scale = 1 mm." httpUri="https://zenodo.org/record/244009/files/figure.png" pageId="6">Fig. 28</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3527A83B536FFE4FF09FC89035DEFCA" pageId="6" type="etymology">
<paragraph id="8BF72908B536FFE4FF09FC89035DEFCA" blockId="6.[151,1437,151,955]" pageId="6">
<emphasis id="B93CF51AB536FFE4FF09FC890002EF2E" bold="true" box="[199,334,858,883]" pageId="6">Etymology.</emphasis>
From the Greek adjective
<taxonomicName id="4C48528BB536FFE4FDB2FC88038BEF2E" box="[636,711,859,883]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="aglaos">
<emphasis id="B93CF51AB536FFE4FDB2FC88038BEF2E" box="[636,711,859,883]" italics="true" pageId="6">aglaós</emphasis>
</taxonomicName>
, splendid, beautiful, treated as a noun in apposition, for the nice and elegant aspect of this species.
</paragraph>
</subSubSection>
<subSubSection id="C3527A83B536FFE4FF09FC71044DEFE6" box="[199,1281,930,955]" pageId="6" type="distribution">
<paragraph id="8BF72908B536FFE4FF09FC71044DEFE6" blockId="6.[151,1437,151,955]" box="[199,1281,930,955]" pageId="6">
<emphasis id="B93CF51AB536FFE4FF09FC71002CEFE6" bold="true" box="[199,352,930,955]" pageId="6">Distribution.</emphasis>
Guanacaste and Central mountain ranges,
<collectingCountry id="F35F6998B536FFE4FC8EFC7002F1EFE6" box="[832,957,930,955]" name="Costa Rica" pageId="6">Costa Rica</collectingCountry>
, from
<quantity id="4CB084EDB536FFE4FBCBFC7005DDEFE7" box="[1029,1169,931,955]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="6" unit="m" value="950.0" valueMax="1100.0" valueMin="800.0">800–1100 m</quantity>
(
<figureCitation id="1373358DB536FFE4FB6FFC7105B9EFE6" box="[1185,1269,930,955]" captionStart="FIGURE 34 – 37" captionStartId="15.[151,250,1947,1969]" captionTargetBox="[158,1427,933,1891]" captionTargetId="figure@15.[151,1436,925,1898]" captionTargetPageId="15" captionText="FIGURE 34 – 37. Distribution of 34: Anomala aglaos and A. estrella. 35: A. inbio and A. pincelada. 36: Callistethus ruteloides. 37: C. yalizo." httpUri="https://zenodo.org/record/244010/files/figure.png" pageId="6">Fig. 34</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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