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<document id="6284E69EE2DDFB708305AA7CF4B04672" ID-DOI="10.11646/phytotaxa.434.1.5" ID-ISSN="1179-3163" ID-Zenodo-Dep="13875195" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1727819162264" checkinUser="felipe" docAuthor="Al-Shehbaz, Ihsan A." docDate="2020" docId="03E187FFEA5451526FF5F9272E50FC40" docLanguage="en" docName="phytotaxa.434.1.5.pdf" docOrigin="Phytotaxa 434 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.434.1.5" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Heliophila pseudoeximia Al-Shehbaz 2020, sp. nov." docType="treatment" docVersion="2" lastPageNumber="81" masterDocId="FFD8FF87EA5A51426F7DFFA12F19FFC9" masterDocTitle="Ten new species in the southern African genus Heliophila (Brassicaceae; Cruciferae)" masterLastPageNumber="88" masterPageNumber="65" pageNumber="79" updateTime="1727822826932" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9E17F49AC7B05637CB61B7CF194CAC83">Ten new species in the southern African genus Heliophila (Brassicaceae; Cruciferae)</mods:title>
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<mods:namePart id="22AEA3453D374F45DCEA3275BD782419">Al-Shehbaz, Ihsan A.</mods:namePart>
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<mods:title id="A2131C09EF0ECD8173898533D178A72E">Phytotaxa</mods:title>
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<mods:date id="93E1BB7615E4820BA413B3F57A7FDAFE">2020</mods:date>
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<mods:number id="F6679591B9D79C2EDBCA68A2DE1C88B6">2020-02-24</mods:number>
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<mods:identifier id="6B080B08D386E80EE6D9E2BD26C8FA30" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="C83E7D9A809778533BA6AFB1CB79B3E9" type="Zenodo-Dep">13875195</mods:identifier>
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<treatment id="03E187FFEA5451526FF5F9272E50FC40" ID-DOI="http://doi.org/10.5281/zenodo.13875223" ID-Zenodo-Dep="13875223" LSID="urn:lsid:plazi:treatment:03E187FFEA5451526FF5F9272E50FC40" httpUri="http://treatment.plazi.org/id/03E187FFEA5451526FF5F9272E50FC40" lastPageId="16" lastPageNumber="81" pageId="14" pageNumber="79">
<subSubSection id="C3526562EA54514C6FF5F9272D81F968" box="[136,664,1670,1697]" pageId="14" pageNumber="79" type="nomenclature">
<paragraph id="8BF736E9EA54514C6FF5F9272D81F968" blockId="14.[136,664,1670,1697]" box="[136,664,1670,1697]" pageId="14" pageNumber="79">
<heading id="D0BF8185EA54514C6FF5F9272D81F968" box="[136,664,1670,1697]" fontSize="11" level="2" pageId="14" pageNumber="79" reason="3">
<taxonomicName id="4C484D6AEA54514C6FF5F9272D2EF969" ID-CoL="C7848" authority="Al-Shehbaz" authorityName="Al-Shehbaz" authorityYear="2020" box="[136,567,1670,1697]" class="Magnoliopsida" family="Brassicaceae" genus="Heliophila" kingdom="Plantae" order="Brassicales" pageId="14" pageNumber="79" phylum="Tracheophyta" rank="species" species="pseudoeximia" status="sp. nov.">
<emphasis id="B93CEAFBEA54514C6FF5F9272EB0F968" bold="true" box="[136,425,1670,1697]" italics="true" pageId="14" pageNumber="79">Heliophila pseudoeximia</emphasis>
Al-Shehbaz
</taxonomicName>
,
<taxonomicNameLabel id="A20F5780EA54514C6D3FF9262D81F968" box="[578,664,1671,1697]" pageId="14" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3526562EA54514C6FF5F9732CCCF8E6" pageId="14" pageNumber="79" type="diagnosis">
<paragraph id="8BF736E9EA54514C6FF5F9732CCCF8E6" blockId="14.[136,1451,1746,1912]" pageId="14" pageNumber="79">
Diagnosis:—
<taxonomicName id="4C484D6AEA54514C6E7AF9732EE3F92E" authorityName="Al-Shehbaz" authorityYear="2020" box="[263,506,1746,1767]" class="Magnoliopsida" family="Brassicaceae" genus="Heliophila" kingdom="Plantae" order="Brassicales" pageId="14" pageNumber="79" phylum="Tracheophyta" rank="species" species="pseudoeximia">
<emphasis id="B93CEAFBEA54514C6E7AF9732EE3F92E" box="[263,506,1746,1767]" italics="true" pageId="14" pageNumber="79">Heliophila pseudoeximia</emphasis>
</taxonomicName>
is easily distinguished from the other glabrous perennial species of
<taxonomicName id="4C484D6AEA54514C6BDCF9732A11F92E" authorityName="Linnaeus" authorityYear="1763" box="[1185,1288,1746,1767]" class="Magnoliopsida" family="Brassicaceae" genus="Heliophila" kingdom="Plantae" order="Brassicales" pageId="14" pageNumber="79" phylum="Tracheophyta" rank="genus">
<emphasis id="B93CEAFBEA54514C6BDCF9732A11F92E" box="[1185,1288,1746,1767]" italics="true" pageId="14" pageNumber="79">Heliophila</emphasis>
</taxonomicName>
with pinnatisect leaves and muricate-papillate lower stems by having paired glands at the pedicels and petioles bases, unappendaged lateral filaments and petal bases, narrowly oblong and non-moniliform fruit, and 68-ovuled ovary.
</paragraph>
</subSubSection>
<subSubSection id="C3526562EA54514C6FF5F89F2A60F8BE" pageId="14" pageNumber="79" type="materials_examined">
<paragraph id="8BF736E9EA54514C6FF5F89F2A60F8BE" blockId="14.[136,1451,1746,1912]" pageId="14" pageNumber="79">
<emphasis id="B93CEAFBEA54514C6FF5F89F2FA3F89A" bold="true" box="[136,186,1854,1875]" pageId="14" pageNumber="79">Type</emphasis>
:—
<materialsCitation id="3B203CB4EA54514C6FABF89F2A6CF8BE" collectingDate="1977-08-30" collectionCode="N, K" collectorName="Oliver &amp; Tolken &amp; Venter" country="South Africa" county="Richtersveld" elevation="1060" location="Lelieshoek" municipality="Rosyntjiesberg" pageId="14" pageNumber="79" specimenCount="2" stateProvince="Northern Cape" typeStatus="holotype">
<collectingCountry id="F35F7679EA54514C6FABF89F2E93F89A" box="[214,394,1854,1875]" name="South Africa" pageId="14" pageNumber="79">SOUTH AFRICA</collectingCountry>
. [
<collectingRegion id="498CF80BEA54514C6EE7F89F2D37F89A" box="[410,558,1854,1876]" country="South Africa" name="Northern Cape" pageId="14" pageNumber="79">Northern Cape</collectingRegion>
, Namakwa],
<collectingCounty id="62964E65EA54514C6DCDF89F2C35F89D" box="[688,812,1854,1876]" pageId="14" pageNumber="79">Richtersveld</collectingCounty>
,
<collectingMunicipality id="6B93AC93EA54514C6C4AF89F2CD0F89D" box="[823,969,1854,1876]" pageId="14" pageNumber="79">Rosyntjiesberg</collectingMunicipality>
, neck
<collectionCode id="ED59AE2CEA54514C6B75F89F2B00F89A" box="[1032,1049,1854,1875]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="14" pageNumber="79" type="Herbarium">N</collectionCode>
of
<location id="8E976032EA54514C6B41F89F2BB1F89D" LSID="urn:lsid:plazi:treatment:03E187FFEA5451526FF5F9272E50FC40:8E976032EA54514C6B41F89F2BB1F89D" box="[1084,1192,1854,1876]" country="South Africa" county="Richtersveld" municipality="Rosyntjiesberg" name="Lelieshoek" pageId="14" pageNumber="79" stateProvince="Northern Cape">Lelieshoek</location>
, southern slope to the east with large quartz outcrops,
<date id="FFF61029EA54514C6EB9F8C32D51F8BE" box="[452,584,1890,1911]" pageId="14" pageNumber="79" value="1977-08-30">
<collectingDate id="EFB2E9C1EA54514C6EB9F8C32D51F8BE" box="[452,584,1890,1911]" pageId="14" pageNumber="79" value="1977-08-30">30 Aug. 1977</collectingDate>
</date>
,
<quantity id="4CB09B0CEA54514C6D2FF8C32D84F8BE" box="[594,669,1890,1912]" metricMagnitude="3" metricUnit="m" metricValue="1.06" pageId="14" pageNumber="79" unit="m" value="1060.0">
<elevation id="0065D1DAEA54514C6D2FF8C32D84F8BE" box="[594,669,1890,1912]" metricMagnitude="3" metricUnit="m" metricValue="1.06" pageId="14" pageNumber="79" unit="m" value="1060.0">1060 m</elevation>
</quantity>
,
<emphasis id="B93CEAFBEA54514C6DDAF8C32CACF8BE" box="[679,949,1890,1911]" italics="true" pageId="14" pageNumber="79">
<collectorName id="26BD533FEA54514C6DDAF8C32DFDF8BE" box="[679,740,1890,1911]" pageId="14" pageNumber="79">Oliver</collectorName>
,
<collectorName id="26BD533FEA54514C6D93F8C32C36F8BE" box="[750,815,1890,1911]" pageId="14" pageNumber="79">Tölken</collectorName>
&amp;
<collectorName id="26BD533FEA54514C6C33F8C32C92F8BE" box="[846,907,1890,1911]" pageId="14" pageNumber="79">Venter</collectorName>
301
</emphasis>
(
<typeStatus id="54F3884BEA54514C6CBCF8C32B00F8B1" box="[961,1049,1890,1912]" pageId="14" pageNumber="79" type="holotype">holotype</typeStatus>
, PRE-650231;
<typeStatus id="54F3884BEA54514C6BD1F8C32BEDF8B1" box="[1196,1268,1890,1912]" pageId="14" pageNumber="79" type="isotype">isotype</typeStatus>
,
<collectionCode id="ED59AE2CEA54514C6B80F8C32A08F8BE" box="[1277,1297,1890,1911]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="14" pageNumber="79" type="Herbarium">K</collectionCode>
).
<figureCitation id="13732A6CEA54514C6A5FF8C32A6CF8BE" box="[1314,1397,1890,1912]" captionStart="FIGURE 8" captionStartId="15.[136,229,1953,1975]" captionTargetBox="[204,1383,190,1925]" captionTargetId="figure-18@15.[204,1383,190,1928]" captionTargetPageId="15" captionText="FIGURE 8. Heliophila pseudoeximia. A. Plant. B. Sepal. C. Petal. D. Stamen. E. Fruiting pedicel and young fruit. Drawn by Al-Shehbaz from holotype (PRE-6502510)." figureDoi="http://doi.org/10.5281/zenodo.13875217" httpUri="https://zenodo.org/record/13875217/files/figure.png" pageId="14" pageNumber="79">Figure 8</figureCitation>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3526562EA5451526FF5F8072D54FD8D" lastPageId="16" lastPageNumber="81" pageId="14" pageNumber="79" type="description">
<paragraph id="8BF736E9EA5451526FF5F8072E92FDE9" blockId="14.[136,1452,1958,2093]" lastBlockId="16.[136,1452,158,905]" lastPageId="16" lastPageNumber="81" pageId="14" pageNumber="79">
<emphasis id="B93CEAFBEA54514C6FF5F8072E05F809" bold="true" box="[136,284,1958,1984]" pageId="14" pageNumber="79">Description:</emphasis>
—Plants perennial herbs, with woody caudex, not glaucous. Trichomes absent. Stems herbaceous,
<quantity id="4CB09B0CEA54514C6A0EF8072FB3F82C" metricMagnitude="-1" metricUnit="m" metricValue="2.35" metricValueMax="4.2" metricValueMin="0.5" pageId="14" pageNumber="79" unit="cm" value="23.5" valueMax="42.0" valueMin="5.0">542 cm</quantity>
tall, terete, erect to ascending, simple or branched at base, glabrous. Lowermost cauline leaves not fleshy,
<quantity id="4CB09B0CEA54514C6A25F86B2AB2F82C" box="[1368,1451,1994,2021]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="9.0" metricValueMin="2.0" pageId="14" pageNumber="79" unit="cm" value="5.5" valueMax="9.0" valueMin="2.0">29 cm</quantity>
long; petiole
<quantity id="4CB09B0CEA54514C6E66F84F2E77F7C0" box="[283,366,2030,2057]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="14" pageNumber="79" unit="cm" value="2.5" valueMax="4.0" valueMin="1.0">14 cm</quantity>
long; blade pinnatisect, 59-lobed, segments linear,
<quantity id="4CB09B0CEA54514C6CC7F84F2B2EF7C0" box="[954,1079,2030,2057]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.7" metricValueMin="0.8" pageId="14" pageNumber="79" unit="cm" value="1.75" valueMax="2.7" valueMin="0.8">0.82.7 cm</quantity>
×
<quantity id="4CB09B0CEA54514C6B28F84F2BADF7C0" box="[1109,1204,2030,2057]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="14" pageNumber="79" unit="mm" value="3.0" valueMax="5.0" valueMin="1.0">15 mm</quantity>
; upper cauline similar to lowermost ones, narrower upwards, base not auriculate, all with a pair of stipule-like glands at node, glabrous, base not decurrent or articulate. Racemes terminal, lax, 1731-flowered, elongated in fruit, not secund; rachis straight, glabrous; fruiting pedicels with a pair of basal bract-like glands, not articulate at base, glabrous, slender, divaricate, arcuate-descending, curved, persistent, lowermost
<quantity id="4CB09B0CEA4A51526DBFFF472C32FEC8" box="[706,811,230,257]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.7" metricValueMin="1.0" pageId="16" pageNumber="81" unit="cm" value="1.35" valueMax="1.7" valueMin="1.0">11.7 cm</quantity>
long; buds oblong. Sepals ascending, oblong,
<quantity id="4CB09B0CEA4A51526A44FF472AB5FEC8" box="[1337,1452,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.5" metricValueMin="4.0" pageId="16" pageNumber="81" unit="mm" value="4.75" valueMax="5.5" valueMin="4.0">45.5 mm</quantity>
long, glabrous, caducous, membranous margined, not cucullate at apex, not saccate at base. Petals white, broadly obovate, 912 ×
<quantity id="4CB09B0CEA4A51526E30FE8F2EB7FE80" box="[333,430,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="16" pageNumber="81" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">56 mm</quantity>
, ascending, rounded at apex, unappendaged, not papillate; claw absent. Stamens slightly tetradynamous; filaments
<quantity id="4CB09B0CEA4A51526ED6FEF32D1EFEA4" box="[427,519,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="16" pageNumber="81" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
long, glabrous at base, unappendaged; anthers oblong-linear,
<quantity id="4CB09B0CEA4A51526BC6FEF32A5BFEA4" box="[1211,1346,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="16" pageNumber="81" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.82.2 mm</quantity>
long, not apiculate at apex. Nectar glands lateral, median glands absent. Ovary glabrous; ovules 68 per ovary. Fruit (immature) silique, capsular, narrowly oblong, ca.
<quantity id="4CB09B0CEA4A51526D3FFE3B2D89FE7C" box="[578,656,410,437]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="16" pageNumber="81" unit="cm" value="1.7">1.7 cm</quantity>
×
<quantity id="4CB09B0CEA4A51526DCFFE3B2DEFFE7C" box="[690,758,410,437]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="16" pageNumber="81" unit="mm" value="3.0">3 mm</quantity>
, latiseptate, glabrous, not moniliform, straight along replum, divaricate to descending; valves papery, midvein obscure, lateral veins obscure, margin not thickened; gynophore ca.
<quantity id="4CB09B0CEA4A51526FF5FE432FF9FE34" box="[136,224,482,509]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="81" unit="mm" value="0.5">0.5 mm</quantity>
long; style in fruit
<quantity id="4CB09B0CEA4A51526EBCFE432D39FE34" box="[449,544,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="16" pageNumber="81" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
long, cylindrical, slender, glabrous; stigma entire. Seeds (immature) uniseriate, suborbicular, flattened.
</paragraph>
<caption id="DF376661EA55514D6FF5F8002EAFF812" ID-DOI="http://doi.org/10.5281/zenodo.13875217" ID-Zenodo-Dep="13875217" httpUri="https://zenodo.org/record/13875217/files/figure.png" pageId="15" pageNumber="80" startId="15.[136,229,1953,1975]" targetBox="[204,1383,190,1925]" targetPageId="15" targetType="figure">
<paragraph id="8BF736E9EA55514D6FF5F8002EAFF812" blockId="15.[136,1451,1953,2011]" pageId="15" pageNumber="80">
<emphasis id="B93CEAFBEA55514D6FF5F8002FE4F87F" bold="true" box="[136,253,1953,1975]" pageId="15" pageNumber="80">FIGURE 8.</emphasis>
<emphasis id="B93CEAFBEA55514D6E7EF8002EE2F87F" box="[259,507,1953,1974]" italics="true" pageId="15" pageNumber="80">
<taxonomicName id="4C484D6AEA55514D6E7EF8002EEFF87F" authorityName="Al-Shehbaz" authorityYear="2020" box="[259,502,1953,1974]" class="Magnoliopsida" family="Brassicaceae" genus="Heliophila" kingdom="Plantae" order="Brassicales" pageId="15" pageNumber="80" phylum="Tracheophyta" rank="species" species="pseudoeximia">Heliophila pseudoeximia</taxonomicName>
.
</emphasis>
A. Plant. B. Sepal. C. Petal. D. Stamen. E. Fruiting pedicel and young fruit. Drawn by Al-Shehbaz from holotype (PRE-6502510).
</paragraph>
</caption>
<paragraph id="8BF736E9EA4A51526FC0FD8B2D54FD8D" blockId="16.[136,1452,158,905]" box="[189,589,554,580]" pageId="16" pageNumber="81">
<emphasis id="B93CEAFBEA4A51526FC0FD8B2E5AFD8D" bold="true" box="[189,323,554,580]" pageId="16" pageNumber="81">Phenology:</emphasis>
—Flowering in August.
</paragraph>
</subSubSection>
<subSubSection id="C3526562EA4A51526FC0FDEF2BC1FDA1" box="[189,1240,590,617]" pageId="16" pageNumber="81" type="distribution">
<paragraph id="8BF736E9EA4A51526FC0FDEF2BC1FDA1" blockId="16.[136,1452,158,905]" box="[189,1240,590,617]" pageId="16" pageNumber="81">
<emphasis id="B93CEAFBEA4A51526FC0FDEF2E40FDA1" bold="true" box="[189,345,590,616]" pageId="16" pageNumber="81">Distribution:</emphasis>
—Known only from the
<typeStatus id="54F3884BEA4A51526D13FDEE2D87FDA0" box="[622,670,591,617]" pageId="16" pageNumber="81">type</typeStatus>
locality in Namakwa district of the
<collectingRegion id="498CF80BEA4A51526B4FFDEF2BC1FDA1" box="[1074,1240,590,616]" country="South Africa" name="Northern Cape" pageId="16" pageNumber="81">Northern Cape</collectingRegion>
</paragraph>
</subSubSection>
<subSubSection id="C3526562EA4A51526FC0FDD32C40FD79" pageId="16" pageNumber="81" type="etymology">
<paragraph id="8BF736E9EA4A51526FC0FDD32A01FD45" blockId="16.[136,1452,158,905]" box="[189,1304,626,653]" pageId="16" pageNumber="81">
<emphasis id="B93CEAFBEA4A51526FC0FDD32E51FD45" bold="true" box="[189,328,626,652]" pageId="16" pageNumber="81">Etymology:</emphasis>
—The name reflects the similarity to the related
<taxonomicName id="4C484D6AEA4A51526C15FDD32A0DFD45" authority="Marais (1966: 103)" authorityName="Marais" authorityPageNumber="103" authorityYear="1966" box="[872,1300,626,652]" class="Magnoliopsida" family="Brassicaceae" genus="Heliophila" kingdom="Plantae" order="Brassicales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="species" species="eximia">
<emphasis id="B93CEAFBEA4A51526C15FDD32B2BFD45" box="[872,1074,626,652]" italics="true" pageId="16" pageNumber="81">Heliophila eximia</emphasis>
<bibRefCitation id="EFD94B18EA4A51526B45FDD32A0DFD45" author="Marais, W." box="[1080,1300,626,652]" pageId="16" pageNumber="81" pagination="97 - 112" refId="ref14026" refString="Marais, W. (1966) Notes on South African Cruciferae. Bothalia 9: 97 - 112. https: // doi. org / 10.4102 / abc. v 9 i 1.1577" type="journal article" year="1966">Marais (1966: 103)</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF736E9EA4A51526FC0FD372C40FD79" blockId="16.[136,1452,158,905]" box="[189,857,662,689]" pageId="16" pageNumber="81">
<emphasis id="B93CEAFBEA4A51526FC0FD372E3AFD79" bold="true" box="[189,291,662,688]" pageId="16" pageNumber="81">Habitat:</emphasis>
—Well-drained stony clay loam, karoo vegetation.
</paragraph>
</subSubSection>
<subSubSection id="C3526562EA4A51526FC0FD1B2E50FC40" pageId="16" pageNumber="81" type="discussion">
<paragraph id="8BF736E9EA4A51526FC0FD1B2E50FC40" blockId="16.[136,1452,158,905]" pageId="16" pageNumber="81">
<emphasis id="B93CEAFBEA4A51526FC0FD1B2E5CFD1D" bold="true" box="[189,325,698,724]" pageId="16" pageNumber="81">Discussion:</emphasis>
<taxonomicName id="4C484D6AEA4A51526E1CFD1B2D65FD1D" authorityName="Al-Shehbaz" authorityYear="2020" box="[353,636,698,724]" class="Magnoliopsida" family="Brassicaceae" genus="Heliophila" kingdom="Plantae" order="Brassicales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="species" species="pseudoeximia">
<emphasis id="B93CEAFBEA4A51526E1CFD1B2D65FD1D" box="[353,636,698,724]" italics="true" pageId="16" pageNumber="81">Heliophila pseudoeximia</emphasis>
</taxonomicName>
is most closely related to
<taxonomicName id="4C484D6AEA4A51526CD0FD1A2B04FD1D" authorityName="Marais" authorityYear="1966" box="[941,1053,699,724]" class="Magnoliopsida" family="Brassicaceae" genus="Heliophila" kingdom="Plantae" order="Brassicales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="species" species="eximia">
<emphasis id="B93CEAFBEA4A51526CD0FD1A2B04FD1D" box="[941,1053,699,724]" italics="true" pageId="16" pageNumber="81">H. eximia</emphasis>
</taxonomicName>
, which it resembles in fruit shape, slender pedicels, presence of gland pairs at bases of petioles and pedicels, and unappendaged lateral filaments and petals. It differs from the latter by being a perennial herb (vs. subshrubs) with basally non-decurrent (vs. decurrent), and pinnatisect, 59-lobed leaves with linear segments (vs. undivided or apically palmate and 35(7)-lobed and obovate or subflabellate to spatulate leaves), membranous- (vs. non-membranous)-margined sepals, and 57 (vs. 814) ovules per ovary.
</paragraph>
</subSubSection>
</treatment>
</document>