487 lines
36 KiB
XML
487 lines
36 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.6122624" ID-GBIF-Dataset="f88be4b8-a0da-444f-b8aa-c02388ba37cb" ID-GBIF-Taxon="116891755" ID-Zenodo-Dep="6122624" checkinTime="1457009644493" checkinUser="donat" docAuthor="David Singer, Anush Kosakyan, Amandine Pillonel, Edward A. D. Mitchell & Enrique Lara" docDate="2015" docId="03E18795BE1DDC625B09F8EE6E2CA823" docLanguage="en" docName="1-s2.0-S0932473914000819-main.pdf.imf" docOrigin="European Journal of Protistology 51 (51)" docStyle="DocumentStyle{}" docTitle="Nebela gimlii Singer, Kosakyan, Pillonel, Mitchell & Lara, 2015, n.sp." docType="treatment" docVersion="12" lastPageId="4" lastPageNumber="83" masterDocId="FFD8FFEDBE1CDC665856FFF86A55AE47" masterDocTitle="Eight species in the Nebela collaris complex: Nebela gimlii (Arcellinida, Hyalospheniidae), a new species described from a Swiss raised bog" masterLastPageNumber="85" masterPageNumber="79" pageId="1" pageNumber="80" updateTime="1645102876571" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Eight species in the Nebela collaris complex: Nebela gimlii (Arcellinida, Hyalospheniidae), a new species described from a Swiss raised bog</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>David Singer</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Anush Kosakyan</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Amandine Pillonel</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Edward A. D. Mitchell</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Enrique Lara</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>European Journal of Protistology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>51</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>51</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>79</mods:start>
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<mods:end>85</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1016/j.ejop.2014.11.004</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6122624" ID-GBIF-Taxon="116891755" ID-Zenodo-Dep="6122624" LSID="urn:lsid:plazi:treatment:03E18795BE1DDC625B09F8EE6E2CA823" httpUri="http://treatment.plazi.org/id/03E18795BE1DDC625B09F8EE6E2CA823" lastPageId="4" lastPageNumber="83" pageId="1" pageNumber="80">
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<subSubSection box="[863,1248,1814,1837]" pageId="1" pageNumber="80" type="nomenclature">
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<paragraph box="[863,1248,1814,1837]" pageId="1" pageNumber="80">
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<taxonomicName box="[863,1007,1814,1837]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="1" pageNumber="80" phylum="Amoebozoa" rank="species" species="gimlii" status="n.sp.">
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<emphasis bold="true" box="[863,1007,1814,1837]" italics="true" pageId="1" pageNumber="80">Nebela gimlii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1014,1064,1814,1837]" pageId="1" pageNumber="80" rank="species">n.sp.</taxonomicNameLabel>
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Singer and Lara.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="2" lastPageNumber="81" pageId="1" pageNumber="80" type="description">
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<paragraph lastPageId="2" lastPageNumber="81" pageId="1" pageNumber="80">
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The test is wide pyriform or drop-shaped, laterally compressed, with a protruding narrow neck. Two lateral pores are present ca.
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<number box="[1014,1023,1909,1932]" pageId="1" pageNumber="80" value="1.0">1</number>
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/
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<number box="[1033,1048,1909,1932]" pageId="1" pageNumber="80" value="4.0">4</number>
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from the distance from the pseudostome to the fundus of the test (
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<figureCitation box="[1179,1243,1941,1964]" captionStart="Fig. 1" captionStartId="2.[114,149,1179,1200]" captionTargetBox="[143,1423,189,1147]" captionTargetId="figure@2.[143,1423,189,1147]" captionTargetPageId="2" captionText="Fig. 1. (A – D) Light microscopy images of Nebela gimlii test. (A) test corresponding to Nebela gimlii el 3 1 (GenBank: KP 083297) – note the presence of two lateral pores (arrows) ca. 1 / 4 from the distance from the pseudostome to the fundus of the test, magnification of 400 × (B) test corresponding to Nebela gimlii el 3 4 (GenBank: KP 083298), magnification of 400 ×, (C) example of Nebela gimlii test showing the scales, magnification of 400 ×, (D) example of Nebela gimlii with focus on the aperture, magnification of 400 ×." pageId="1" pageNumber="80">
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Fig.
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<number box="[1230,1243,1941,1964]" pageId="1" pageNumber="80" value="1.0">1</number>
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</figureCitation>
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<collectionCode box="[1243,1262,1941,1964]" pageId="1" pageNumber="80">A</collectionCode>
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and
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<collectionCode box="[1319,1339,1941,1964]" pageId="1" pageNumber="80">B</collectionCode>
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). A variable number of small pores can be seen on SEM images in apparently random positions from near the aperture to the fundus of the test (
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<figureCitation box="[250,317,1388,1411]" captionStart="Fig. 2" captionStartId="3.[135,170,1523,1544]" captionTargetBox="[165,1445,187,1490]" captionTargetId="figure@3.[165,1445,187,1491]" captionTargetPageId="3" captionText="Fig. 2. (A – E) SEM micrographs of three individuals of Nebela gimlii. (A, C and E) – illustrating the shell general shape and composition; and (B, D and F) – detail view of the aperture of the same individuals. Arrows show some pores on the shell." pageId="2" pageNumber="81">
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Fig.
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<number box="[304,317,1388,1411]" pageId="2" pageNumber="81" value="2.0">2</number>
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</figureCitation>
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<collectionCode box="[317,338,1388,1411]" pageId="2" pageNumber="81">A</collectionCode>
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,
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<collectionCode box="[355,375,1388,1411]" pageId="2" pageNumber="81">B</collectionCode>
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,
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<collectionCode box="[391,407,1388,1411]" pageId="2" pageNumber="81">E</collectionCode>
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and
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<collectionCode box="[470,487,1388,1411]" pageId="2" pageNumber="81">F</collectionCode>
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). The test is colourless or slightly brownish, composed of small particles (likely obtained from preys, e.g. euglyphid testate amoebae), which are covered by a thin layer of organic cement. The aperture is oval in frontal view, slightly curved in lateral view, surrounded by a thin organic collar characteristic of family
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<taxonomicName authority="Kosakyan et al., 2012" authorityName="Kosakyan et al." authorityYear="2012" box="[114,547,1579,1602]" class="Lobosa" genus="Hyalospheniidae" higherTaxonomySource="GBIF" kingdom="Protozoa" order="Arcellinida" pageId="2" pageNumber="81" phylum="Protozoa" rank="subSpecies" species="et" subGenus="Kosakyan" subSpecies="al">
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Hyalospheniidae (Kosakyan et al.,
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<number box="[483,536,1579,1602]" pageId="2" pageNumber="81" value="2012.0">2012</number>
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)
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</taxonomicName>
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. Dimensions based on
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<number box="[149,176,1611,1634]" pageId="2" pageNumber="81" value="14.0">14</number>
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individuals: length:
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<number box="[399,505,1611,1634]" pageId="2" pageNumber="81" value="72.65" valueMax="77.6" valueMin="67.7">67.7–77.6</number>
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μm (mean =
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<number box="[647,693,1611,1634]" pageId="2" pageNumber="81" value="72.8">72.8</number>
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μm), breadth:
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<number box="[212,331,1643,1666]" pageId="2" pageNumber="81" value="55.67" valueMax="61.64" valueMin="49.7">49.7–61.64</number>
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μm (mean =
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<number box="[476,522,1643,1666]" pageId="2" pageNumber="81" value="53.9">53.9</number>
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μm), width of aperture:
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<number box="[169,255,1675,1698]" pageId="2" pageNumber="81" value="19.25" valueMax="21.0" valueMin="17.5">17.5–21</number>
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μm (mean =
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<number box="[395,422,1675,1698]" pageId="2" pageNumber="81" value="19.0">19</number>
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μm).
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</paragraph>
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<caption pageId="2" pageNumber="81" targetBox="[143,1423,189,1147]" targetPageId="2">
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<paragraph pageId="2" pageNumber="81">
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<emphasis bold="true" box="[114,177,1179,1200]" pageId="2" pageNumber="81">Fig. 1.</emphasis>
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(A–D) Light microscopy images of
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<taxonomicName box="[535,666,1179,1200]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="2" pageNumber="81" phylum="Amoebozoa" rank="species" species="gimlii">
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<emphasis bold="true" box="[535,666,1179,1200]" italics="true" pageId="2" pageNumber="81">Nebela gimlii</emphasis>
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</taxonomicName>
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test. (A) test corresponding to
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<taxonomicName box="[966,1099,1179,1200]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="2" pageNumber="81" phylum="Amoebozoa" rank="species" species="gimlii">Nebela gimlii</taxonomicName>
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el3 1 (GenBank: KP083297) – note the presence of two lateral pores (arrows) ca. 1/4 from the distance from the pseudostome to the fundus of the test, magnification of 400× (B) test corresponding to
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<taxonomicName box="[356,490,1238,1259]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="2" pageNumber="81" phylum="Amoebozoa" rank="species" species="gimlii">Nebela gimlii</taxonomicName>
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el3 4 (GenBank: KP083298), magnification of 400×, (C) example of
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<taxonomicName box="[1164,1293,1238,1259]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="2" pageNumber="81" phylum="Amoebozoa" rank="species" species="gimlii">
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<emphasis bold="true" box="[1164,1293,1238,1259]" italics="true" pageId="2" pageNumber="81">Nebela gimlii</emphasis>
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</taxonomicName>
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test showing the scales, magnification of 400×, (D) example of
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<taxonomicName box="[564,694,1267,1288]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="2" pageNumber="81" phylum="Amoebozoa" rank="species" species="gimlii">
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<emphasis bold="true" box="[564,694,1267,1288]" italics="true" pageId="2" pageNumber="81">Nebela gimlii</emphasis>
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</taxonomicName>
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with focus on the aperture, magnification of 400×.
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</paragraph>
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</caption>
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<paragraph pageId="2" pageNumber="81">
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Illustrations and morphological data of the new species are given in
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<figureCitation box="[206,278,1738,1761]" captionStart="Fig. 1" captionStartId="2.[114,149,1179,1200]" captionTargetBox="[143,1423,189,1147]" captionTargetId="figure@2.[143,1423,189,1147]" captionTargetPageId="2" captionText="Fig. 1. (A – D) Light microscopy images of Nebela gimlii test. (A) test corresponding to Nebela gimlii el 3 1 (GenBank: KP 083297) – note the presence of two lateral pores (arrows) ca. 1 / 4 from the distance from the pseudostome to the fundus of the test, magnification of 400 × (B) test corresponding to Nebela gimlii el 3 4 (GenBank: KP 083298), magnification of 400 ×, (C) example of Nebela gimlii test showing the scales, magnification of 400 ×, (D) example of Nebela gimlii with focus on the aperture, magnification of 400 ×." pageId="2" pageNumber="81">
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||
Figs.
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<number box="[265,278,1738,1761]" pageId="2" pageNumber="81" value="1.0">1</number>
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||
</figureCitation>
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||
and
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||
<figureCitation box="[330,343,1738,1761]" captionStart="Fig. 2" captionStartId="3.[135,170,1523,1544]" captionTargetBox="[165,1445,187,1490]" captionTargetId="figure@3.[165,1445,187,1491]" captionTargetPageId="3" captionText="Fig. 2. (A – E) SEM micrographs of three individuals of Nebela gimlii. (A, C and E) – illustrating the shell general shape and composition; and (B, D and F) – detail view of the aperture of the same individuals. Arrows show some pores on the shell." pageId="2" pageNumber="81">
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<number box="[330,343,1738,1761]" pageId="2" pageNumber="81" value="2.0">2</number>
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</figureCitation>
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and
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<tableCitation box="[395,483,1738,1761]" captionStart="Table 1" captionStartId="3.[135,191,1619,1640]" captionTargetBox="[135,1468,1703,1920]" captionText="Table 1. Biometrical characteristics of Nebela gimlii: M – median, SD – standard deviation, SE – standard error of the mean, CV – coefficient of variation, Min – minimum, Max – maximum, n – number of individuals examined (measurements in μm)." pageId="2" pageNumber="81">
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Tables
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<number box="[470,483,1738,1761]" pageId="2" pageNumber="81" value="1.0">1</number>
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||
</tableCitation>
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||
and
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<tableCitation box="[535,548,1738,1761]" captionStart="Table 2" captionStartId="4.[114,170,1611,1632]" captionTargetBox="[114,749,1721,1912]" captionText="Table 2. Comparison of morphological characters between Nebela" pageId="2" pageNumber="81">
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<number box="[535,548,1738,1761]" pageId="2" pageNumber="81" value="2.0">2</number>
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</tableCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="81" type="biology_ecology">
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<paragraph box="[114,213,1812,1838]" pageId="2" pageNumber="81">
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||
<emphasis bold="true" box="[114,213,1812,1838]" pageId="2" pageNumber="81">
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<heading bold="true" box="[114,213,1812,1838]" fontSize="11" level="2" pageId="2" pageNumber="81" reason="0">Ecology</heading>
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</emphasis>
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</paragraph>
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<paragraph pageId="2" pageNumber="81">
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<taxonomicName box="[140,285,1878,1901]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="2" pageNumber="81" phylum="Amoebozoa" rank="species" species="gimlii">
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<emphasis bold="true" box="[140,285,1878,1901]" italics="true" pageId="2" pageNumber="81">Nebela gimlii</emphasis>
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</taxonomicName>
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was found exclusively in the relatively dry
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<emphasis bold="true" box="[114,226,1910,1933]" italics="true" pageId="2" pageNumber="81">
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<taxonomicName box="[114,226,1910,1933]" class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" higherTaxonomySource="CoL" kingdom="Plantae" order="Sphagnales" pageId="2" pageNumber="81" phylum="Bryophyta" rank="genus">Sphagnum</taxonomicName>
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</emphasis>
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mosses in the forested part of a peat bog (influenced by lateral drainage) and the drained peatland margin a few meters from the base of old peat extraction walls. These habitats are characterised by low pH, moderate moisture and low nutrient content.
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</paragraph>
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</subSubSection>
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<subSubSection box="[815,977,1471,1497]" pageId="2" pageNumber="81" type="description">
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||
<paragraph box="[815,977,1471,1497]" pageId="2" pageNumber="81">
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||
<heading bold="true" box="[815,977,1471,1497]" fontSize="11" level="2" pageId="2" pageNumber="81" reason="0">
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<emphasis bold="true" box="[815,977,1471,1497]" pageId="2" pageNumber="81">Hapantotype</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="81" type="materials_examined">
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<paragraph pageId="2" pageNumber="81">
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<materialsCitation ID-GBIF-Occurrence="1233222613" collectedFrom="Sphagnum" country="Switzerland" latitude="47.00423" location="Le Cachot, Jura Mountains" longitude="6.664675" pageId="2" pageNumber="81" specimenCode-GBK-0="KP083297" specimenCode-GBK-1="KP083298" specimenCount="1" stateProvince="Neuchatel">
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The tests were collected from a
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||
<collectedFrom box="[1219,1331,1537,1560]" pageId="2" pageNumber="81">
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||
<emphasis bold="true" box="[1219,1331,1537,1560]" italics="true" pageId="2" pageNumber="81">
|
||
<taxonomicName box="[1219,1331,1537,1560]" class="Sphagnopsida" family="Sphagnaceae" genus="Sphagnum" higherTaxonomySource="CoL" kingdom="Plantae" order="Sphagnales" pageId="2" pageNumber="81" phylum="Bryophyta" rank="genus">Sphagnum</taxonomicName>
|
||
</emphasis>
|
||
</collectedFrom>
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||
sample in a peatland in
|
||
<location LSID="urn:lsid:plazi:treatment:03E18795BE1DDC625B09F8EE6E2CA823:8E976058BE1EDC645B8CF9D96F4CA87F" box="[986,1305,1569,1592]" country="Switzerland" name="Le Cachot, Jura Mountains" pageId="2" pageNumber="81" stateProvince="Neuchatel">Le Cachot, Jura Mountains</location>
|
||
,
|
||
<collectingCountry box="[1326,1453,1569,1592]" name="Switzerland" pageId="2" pageNumber="81" value="Switzerland">Switzerland</collectingCountry>
|
||
(
|
||
<geoCoordinate box="[823,988,1595,1624]" pageId="2" pageNumber="81">
|
||
<number box="[823,851,1601,1624]" pageId="2" pageNumber="81" value="47.0">47</number>
|
||
º00'
|
||
<number box="[895,955,1601,1624]" pageId="2" pageNumber="81" value="15.23">15.23</number>
|
||
''N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[999,1148,1595,1624]" pageId="2" pageNumber="81">
|
||
<number box="[999,1012,1601,1624]" pageId="2" pageNumber="81" value="6.0">6</number>
|
||
º
|
||
<number box="[1023,1050,1601,1624]" pageId="2" pageNumber="81" value="39.0">39</number>
|
||
'
|
||
<number box="[1057,1117,1601,1624]" pageId="2" pageNumber="81" value="52.83">52.83</number>
|
||
''E
|
||
</geoCoordinate>
|
||
). One SEM stub with several specimens is deposited at the Natural History Museum of Neuchâtel (Ref. Nr.: UniNe-EM-
|
||
<number box="[1175,1186,1664,1687]" pageId="2" pageNumber="81" value="5.0">5</number>
|
||
). COI gene sequences of
|
||
<taxonomicName box="[815,957,1697,1720]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="2" pageNumber="81" phylum="Amoebozoa" rank="species" species="gimlii">
|
||
<emphasis bold="true" box="[815,957,1697,1720]" italics="true" pageId="2" pageNumber="81">Nebela gimlii</emphasis>
|
||
</taxonomicName>
|
||
el
|
||
<number box="[982,994,1696,1719]" pageId="2" pageNumber="81" value="3.0">3</number>
|
||
<number box="[1004,1017,1696,1719]" pageId="2" pageNumber="81" value="1.0">1</number>
|
||
(
|
||
<number box="[1029,1070,1696,1719]" pageId="2" pageNumber="81" value="499.0">499</number>
|
||
bp) and
|
||
<taxonomicName box="[1158,1300,1697,1720]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="2" pageNumber="81" phylum="Amoebozoa" rank="species" species="gimlii">
|
||
<emphasis bold="true" box="[1158,1300,1697,1720]" italics="true" pageId="2" pageNumber="81">Nebela gimlii</emphasis>
|
||
</taxonomicName>
|
||
el
|
||
<number box="[1325,1337,1696,1719]" pageId="2" pageNumber="81" value="3.0">3</number>
|
||
<number box="[1346,1359,1696,1719]" pageId="2" pageNumber="81" value="4.0">4</number>
|
||
(
|
||
<number box="[1372,1413,1696,1719]" pageId="2" pageNumber="81" value="499.0">499</number>
|
||
bp) were deposited in GenBank (
|
||
<taxonomicName box="[1119,1268,1728,1751]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="2" pageNumber="81" phylum="Amoebozoa" rank="species" species="gimlii">Nebela gimlii</taxonomicName>
|
||
<specimenCode box="[1278,1453,1728,1751]" pageId="2" pageNumber="81">
|
||
el
|
||
<number box="[1299,1311,1728,1751]" pageId="2" pageNumber="81" value="3.0">3</number>
|
||
<number box="[1320,1333,1728,1751]" pageId="2" pageNumber="81" value="1.0">1</number>
|
||
KP083297
|
||
</specimenCode>
|
||
and, respectively,
|
||
<taxonomicName box="[1006,1153,1760,1783]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="2" pageNumber="81" phylum="Amoebozoa" rank="species" species="gimlii">Nebela gimlii</taxonomicName>
|
||
el
|
||
<number box="[1184,1196,1760,1783]" pageId="2" pageNumber="81" value="3.0">3</number>
|
||
<number box="[1205,1218,1760,1783]" pageId="2" pageNumber="81" value="4.0">4</number>
|
||
KP083298)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="4" lastPageNumber="83" pageId="2" pageNumber="81" type="etymology">
|
||
<paragraph box="[815,950,1843,1869]" pageId="2" pageNumber="81">
|
||
<emphasis bold="true" box="[815,950,1843,1869]" pageId="2" pageNumber="81">
|
||
<heading bold="true" box="[815,950,1843,1869]" fontSize="11" level="2" pageId="2" pageNumber="81" reason="0">Etymology</heading>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph lastPageId="4" lastPageNumber="83" pageId="2" pageNumber="81">
|
||
The name of this species refers to the name of Gimli, one of the dwarfs in J.R.R. Tolkien’s masterpiece “The Lord of the Rings”, because of its small size (the smallest known member of the
|
||
<emphasis bold="true" box="[275,438,1421,1444]" italics="true" pageId="4" pageNumber="83">
|
||
<taxonomicName box="[275,438,1421,1444]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="4" pageNumber="83" phylum="Amoebozoa" rank="species" species="collaris">Nebela collaris</taxonomicName>
|
||
</emphasis>
|
||
complex) and stout shape. In addition, it has been found abundantly in a forest, and Gimli was unique among his kind to have been travelling in the woods.
|
||
</paragraph>
|
||
<caption pageId="3" pageNumber="82" targetBox="[165,1445,187,1490]" targetPageId="3">
|
||
<paragraph pageId="3" pageNumber="82">
|
||
<emphasis bold="true" box="[135,199,1523,1544]" pageId="3" pageNumber="82">Fig. 2.</emphasis>
|
||
(A–E) SEM micrographs of three individuals of
|
||
<emphasis bold="true" box="[679,810,1523,1544]" italics="true" pageId="3" pageNumber="82">
|
||
<taxonomicName box="[679,810,1523,1544]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="3" pageNumber="82" phylum="Amoebozoa" rank="species" species="gimlii">Nebela gimlii</taxonomicName>
|
||
</emphasis>
|
||
. (A, C and E) – illustrating the shell general shape and composition; and (B, D and F) – detail view of the aperture of the same individuals. Arrows show some pores on the shell.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="3" pageNumber="82" targetBox="[135,1468,1703,1920]" targetIsTable="true" targetPageId="3">
|
||
<paragraph pageId="3" pageNumber="82">
|
||
<emphasis bold="true" box="[135,215,1619,1640]" pageId="3" pageNumber="82">Table 1.</emphasis>
|
||
Biometrical characteristics of
|
||
<taxonomicName box="[512,641,1620,1641]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="3" pageNumber="82" phylum="Amoebozoa" rank="species" species="gimlii">
|
||
<emphasis bold="true" box="[512,641,1620,1641]" italics="true" pageId="3" pageNumber="82">Nebela gimlii</emphasis>
|
||
</taxonomicName>
|
||
: M – median, SD – standard deviation, SE – standard error of the mean, CV –coefficient of variation, Min – minimum, Max – maximum,
|
||
<emphasis bold="true" box="[603,615,1649,1670]" italics="true" pageId="3" pageNumber="82">n</emphasis>
|
||
– number of individuals examined (measurements in μm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="3" pageNumber="82">
|
||
<table box="[135,1468,1703,1920]" pageId="3" pageNumber="82">
|
||
<tr box="[135,1468,1703,1724]" pageId="3" pageNumber="82">
|
||
<td box="[135,237,1703,1724]" pageId="3" pageNumber="82">Characters</td>
|
||
<td box="[437,491,1703,1724]" pageId="3" pageNumber="82">Mean</td>
|
||
<td box="[584,605,1703,1724]" pageId="3" pageNumber="82">M</td>
|
||
<td box="[731,762,1703,1724]" pageId="3" pageNumber="82">SD</td>
|
||
<td box="[867,895,1703,1724]" pageId="3" pageNumber="82">SE</td>
|
||
<td box="[1002,1035,1703,1724]" pageId="3" pageNumber="82">CV</td>
|
||
<td box="[1149,1189,1703,1724]" pageId="3" pageNumber="82">Min</td>
|
||
<td box="[1297,1341,1703,1724]" pageId="3" pageNumber="82">Max</td>
|
||
<td box="[1444,1456,1703,1724]" pageId="3" pageNumber="82">
|
||
<emphasis bold="true" box="[1444,1456,1703,1724]" italics="true" pageId="3" pageNumber="82">n</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr box="[135,1468,1752,1773]" pageId="3" pageNumber="82">
|
||
<td box="[135,203,1752,1773]" pageId="3" pageNumber="82">Length</td>
|
||
<td box="[437,479,1752,1773]" pageId="3" pageNumber="82">72.8</td>
|
||
<td box="[584,638,1752,1773]" pageId="3" pageNumber="82">73.23</td>
|
||
<td box="[731,773,1752,1773]" pageId="3" pageNumber="82">2.92</td>
|
||
<td box="[867,909,1752,1773]" pageId="3" pageNumber="82">0.78</td>
|
||
<td box="[1014,1026,1752,1773]" pageId="3" pageNumber="82">4</td>
|
||
<td box="[1149,1203,1752,1773]" pageId="3" pageNumber="82">67.71</td>
|
||
<td box="[1297,1351,1752,1773]" pageId="3" pageNumber="82">77.61</td>
|
||
<td box="[1444,1468,1752,1773]" pageId="3" pageNumber="82">14</td>
|
||
</tr>
|
||
<tr box="[135,1468,1781,1802]" pageId="3" pageNumber="82">
|
||
<td box="[135,211,1781,1802]" pageId="3" pageNumber="82">Breadth</td>
|
||
<td box="[437,479,1781,1802]" pageId="3" pageNumber="82">53.9</td>
|
||
<td box="[584,638,1781,1802]" pageId="3" pageNumber="82">53.12</td>
|
||
<td box="[731,773,1781,1802]" pageId="3" pageNumber="82">3.67</td>
|
||
<td box="[867,909,1781,1802]" pageId="3" pageNumber="82">0.98</td>
|
||
<td box="[1014,1056,1781,1802]" pageId="3" pageNumber="82">6.81</td>
|
||
<td box="[1149,1203,1781,1802]" pageId="3" pageNumber="82">49.68</td>
|
||
<td box="[1297,1351,1781,1802]" pageId="3" pageNumber="82">61.64</td>
|
||
<td box="[1444,1468,1781,1802]" pageId="3" pageNumber="82">14</td>
|
||
</tr>
|
||
<tr box="[135,1468,1811,1832]" pageId="3" pageNumber="82">
|
||
<td box="[135,193,1811,1832]" pageId="3" pageNumber="82">Depth</td>
|
||
<td box="[437,479,1811,1832]" pageId="3" pageNumber="82">31.4</td>
|
||
<td box="[584,626,1811,1832]" pageId="3" pageNumber="82">30.7</td>
|
||
<td box="[731,761,1811,1832]" pageId="3" pageNumber="82">1.4</td>
|
||
<td box="[867,909,1811,1832]" pageId="3" pageNumber="82">0.37</td>
|
||
<td box="[1014,1056,1811,1832]" pageId="3" pageNumber="82">4.45</td>
|
||
<td box="[1149,1191,1811,1832]" pageId="3" pageNumber="82">30.5</td>
|
||
<td box="[1297,1321,1811,1832]" pageId="3" pageNumber="82">35</td>
|
||
<td box="[1444,1468,1811,1832]" pageId="3" pageNumber="82">14</td>
|
||
</tr>
|
||
<tr box="[135,1468,1840,1861]" pageId="3" pageNumber="82">
|
||
<td box="[135,280,1840,1861]" pageId="3" pageNumber="82">Aperture width</td>
|
||
<td box="[437,461,1840,1861]" pageId="3" pageNumber="82">19</td>
|
||
<td box="[584,638,1840,1861]" pageId="3" pageNumber="82">18.71</td>
|
||
<td box="[731,773,1840,1861]" pageId="3" pageNumber="82">1.03</td>
|
||
<td box="[867,909,1840,1861]" pageId="3" pageNumber="82">0.28</td>
|
||
<td box="[1014,1056,1840,1861]" pageId="3" pageNumber="82">5.45</td>
|
||
<td box="[1149,1191,1840,1861]" pageId="3" pageNumber="82">17.5</td>
|
||
<td box="[1297,1351,1840,1861]" pageId="3" pageNumber="82">21.06</td>
|
||
<td box="[1444,1468,1840,1861]" pageId="3" pageNumber="82">14</td>
|
||
</tr>
|
||
<tr box="[135,1468,1869,1920]" pageId="3" pageNumber="82">
|
||
<td box="[135,344,1869,1920]" pageId="3" pageNumber="82">Ratio (L/B) Biovolume (104 μm3)</td>
|
||
<td box="[449,491,1869,1919]" pageId="3" pageNumber="82">1.4 8.25</td>
|
||
<td box="[596,638,1869,1919]" pageId="3" pageNumber="82">1.35 7.82</td>
|
||
<td box="[731,773,1869,1919]" pageId="3" pageNumber="82">0.07 1.08</td>
|
||
<td box="[867,909,1869,1919]" pageId="3" pageNumber="82">0.02 0.28</td>
|
||
<td box="[1002,1056,1869,1919]" pageId="3" pageNumber="82">5.45 13.12</td>
|
||
<td box="[1161,1203,1869,1919]" pageId="3" pageNumber="82">1.22 6.94</td>
|
||
<td box="[1297,1351,1869,1919]" pageId="3" pageNumber="82">1.5 10.47</td>
|
||
<td box="[1444,1468,1869,1919]" pageId="3" pageNumber="82">14 14</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption pageId="4" pageNumber="83" targetBox="[143,1423,187,1212]" targetPageId="4">
|
||
<paragraph pageId="4" pageNumber="83">
|
||
<emphasis bold="true" box="[114,177,1243,1264]" pageId="4" pageNumber="83">Fig. 3.</emphasis>
|
||
Bayesian Markov Chain Monte Carlo consensus tree of 31
|
||
<emphasis bold="true" box="[760,907,1244,1265]" italics="true" pageId="4" pageNumber="83">
|
||
<taxonomicName box="[760,907,1244,1265]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="4" pageNumber="83" phylum="Amoebozoa" rank="species" species="collaris">Nebela collaris</taxonomicName>
|
||
</emphasis>
|
||
s.I. testate amoeba. Based on mitochondrial cytochrome oxidase, subunit I (COI) gene.
|
||
<taxonomicName box="[422,555,1273,1294]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="4" pageNumber="83" phylum="Amoebozoa" rank="species" species="gimlii">
|
||
<emphasis bold="true" box="[422,555,1273,1294]" italics="true" pageId="4" pageNumber="83">Nebela gimlii</emphasis>
|
||
</taxonomicName>
|
||
is denoted in bold. The numbers along the branches represent, respectively, the bootstraps obtained by maximum likelihood method and the posterior probabilities as calculated with Bayesian analyses. Only values above 50/0.50 are shown. The tree was rooted with outgroup
|
||
<emphasis bold="true" box="[523,705,1332,1353]" italics="true" pageId="4" pageNumber="83">
|
||
<taxonomicName authority="Loeblich & Tappan, 1961" authorityName="Loeblich & Tappan" authorityYear="1961" box="[523,705,1332,1353]" class="Lobosa" family="Nebelidae" genus="Certesella" higherTaxonomySource="GBIF" kingdom="Protista" order="Arcellinida" pageId="4" pageNumber="83" phylum="Sarcomastigophora" rank="species" species="martiali">Certesella martiali</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis bold="true" box="[751,907,1332,1353]" italics="true" pageId="4" pageNumber="83">
|
||
<taxonomicName box="[751,907,1332,1353]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="4" pageNumber="83" phylum="Amoebozoa" rank="species" species="tubulosa">Nebela tubulosa</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption box="[114,752,1611,1633]" pageId="4" pageNumber="83" targetBox="[114,749,1721,1912]" targetIsTable="true" targetPageId="4">
|
||
<paragraph box="[114,752,1611,1633]" pageId="4" pageNumber="83">
|
||
<emphasis bold="true" box="[114,193,1611,1632]" pageId="4" pageNumber="83">Table 2.</emphasis>
|
||
Comparison of morphological characters between
|
||
<taxonomicName box="[684,752,1612,1633]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="4" pageNumber="83" phylum="Amoebozoa" rank="genus">Nebela</taxonomicName>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph box="[114,751,1641,1662]" pageId="4" pageNumber="83">
|
||
<emphasis bold="true" box="[114,170,1641,1662]" italics="true" pageId="4" pageNumber="83">
|
||
<taxonomicName box="[114,170,1641,1662]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="4" pageNumber="83" phylum="Amoebozoa" rank="species" species="gimlii">gimlii</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis bold="true" box="[212,305,1641,1662]" italics="true" pageId="4" pageNumber="83">
|
||
<taxonomicName box="[212,305,1641,1662]" class="Lobosa" family="Nebelidae" genus="Nebela" genus.bestMatchDistance="81" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="4" pageNumber="83" phylum="Amoebozoa" rank="species" species="guttata">N. guttata</taxonomicName>
|
||
</emphasis>
|
||
. Min – minimum, Max – maximum,
|
||
<emphasis bold="true" box="[647,659,1641,1662]" italics="true" pageId="4" pageNumber="83">n</emphasis>
|
||
– number
|
||
</paragraph>
|
||
<paragraph box="[114,574,1670,1692]" pageId="4" pageNumber="83">of individuals examined (measurements in μm).</paragraph>
|
||
<paragraph pageId="4" pageNumber="83">
|
||
<table box="[114,749,1721,1912]" pageId="4" pageNumber="83">
|
||
<tr box="[114,749,1721,1746]" pageId="4" pageNumber="83">
|
||
<td box="[114,216,1725,1746]" pageId="4" pageNumber="83">Characters</td>
|
||
<td box="[314,444,1725,1746]" pageId="4" pageNumber="83">
|
||
<taxonomicName box="[314,444,1725,1746]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="4" pageNumber="83" phylum="Amoebozoa" rank="species" species="gimlii">
|
||
<emphasis bold="true" box="[314,444,1725,1746]" italics="true" pageId="4" pageNumber="83">Nebela gimlii</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td box="[576,725,1721,1746]" pageId="4" pageNumber="83">
|
||
<emphasis bold="true" box="[576,718,1725,1746]" italics="true" pageId="4" pageNumber="83">
|
||
<taxonomicName box="[576,718,1725,1746]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="4" pageNumber="83" phylum="Amoebozoa" rank="species" species="guttata">Nebela guttata</taxonomicName>
|
||
</emphasis>
|
||
*
|
||
</td>
|
||
</tr>
|
||
<tr box="[114,749,1754,1775]" pageId="4" pageNumber="83">
|
||
<td box="[314,500,1754,1775]" colspan="2" colspanLeft="1" pageId="4" pageNumber="83">
|
||
(Min/Max) (
|
||
<emphasis bold="true" box="[434,446,1754,1775]" italics="true" pageId="4" pageNumber="83">n</emphasis>
|
||
= 14)
|
||
</td>
|
||
<td box="[576,749,1754,1775]" pageId="4" pageNumber="83">
|
||
(Min/Max) (
|
||
<emphasis bold="true" box="[696,708,1754,1775]" italics="true" pageId="4" pageNumber="83">n</emphasis>
|
||
= 4)
|
||
</td>
|
||
</tr>
|
||
<tr box="[114,749,1803,1824]" pageId="4" pageNumber="83">
|
||
<td box="[114,182,1803,1824]" pageId="4" pageNumber="83">Length</td>
|
||
<td box="[314,369,1803,1824]" pageId="4" pageNumber="83">68/78</td>
|
||
<td box="[576,630,1803,1824]" pageId="4" pageNumber="83">80/89</td>
|
||
</tr>
|
||
<tr box="[114,749,1832,1853]" pageId="4" pageNumber="83">
|
||
<td box="[114,190,1832,1853]" pageId="4" pageNumber="83">Breadth</td>
|
||
<td box="[314,369,1832,1853]" pageId="4" pageNumber="83">50/62</td>
|
||
<td box="[576,631,1832,1853]" pageId="4" pageNumber="83">53/63</td>
|
||
</tr>
|
||
<tr box="[114,749,1861,1882]" pageId="4" pageNumber="83">
|
||
<td box="[114,259,1861,1882]" pageId="4" pageNumber="83">Aperture width</td>
|
||
<td box="[314,386,1861,1882]" pageId="4" pageNumber="83">17.5/21</td>
|
||
<td box="[576,630,1861,1882]" pageId="4" pageNumber="83">20/21</td>
|
||
</tr>
|
||
<tr box="[114,749,1891,1912]" pageId="4" pageNumber="83">
|
||
<td box="[114,224,1891,1912]" pageId="4" pageNumber="83">Ratio (L/B)</td>
|
||
<td box="[326,393,1891,1912]" pageId="4" pageNumber="83">1.2/1.4</td>
|
||
<td box="[588,654,1891,1912]" pageId="4" pageNumber="83">1.4/1.5</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph box="[135,435,1936,1959]" pageId="4" pageNumber="83">
|
||
* Data from Kosakyan et al. (
|
||
<number box="[379,423,1940,1959]" pageId="4" pageNumber="83" value="2013.0">2013</number>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="83" type="discussion">
|
||
<paragraph box="[815,873,1388,1414]" pageId="4" pageNumber="83">
|
||
<emphasis bold="true" box="[815,873,1388,1414]" pageId="4" pageNumber="83">
|
||
<heading bold="true" box="[815,873,1388,1414]" fontSize="11" level="2" pageId="4" pageNumber="83" reason="0">Note</heading>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph pageId="4" pageNumber="83">
|
||
<emphasis bold="true" box="[842,989,1454,1477]" italics="true" pageId="4" pageNumber="83">
|
||
<taxonomicName box="[842,989,1454,1477]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="4" pageNumber="83" phylum="Amoebozoa" rank="species" species="gimlii">Nebela gimlii</taxonomicName>
|
||
</emphasis>
|
||
can be confused with
|
||
<emphasis bold="true" box="[1243,1403,1454,1477]" italics="true" pageId="4" pageNumber="83">
|
||
<taxonomicName box="[1243,1403,1454,1477]" class="Lobosa" family="Nebelidae" genus="Nebela" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="4" pageNumber="83" phylum="Amoebozoa" rank="species" species="guttata">Nebela guttata</taxonomicName>
|
||
</emphasis>
|
||
, but clearly differs in the wider shape of the shell and the smaller dimensions of the test (
|
||
<tableCitation box="[1078,1159,1518,1541]" captionStart="Table 2" captionStartId="4.[114,170,1611,1632]" captionTargetBox="[114,749,1721,1912]" captionText="Table 2. Comparison of morphological characters between Nebela" pageId="4" pageNumber="83">
|
||
Table
|
||
<number box="[1146,1159,1518,1541]" pageId="4" pageNumber="83" value="2.0">2</number>
|
||
</tableCitation>
|
||
). Both species are closely related and differ by
|
||
<number box="[1039,1072,1549,1572]" pageId="4" pageNumber="83" value="3.7">3.7</number>
|
||
% on the considered COI fragment. The two sequences of
|
||
<taxonomicName box="[1056,1149,1582,1605]" class="Lobosa" family="Nebelidae" genus="Nebela" genus.bestMatchDistance="24" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Protozoa" order="Arcellinida" pageId="4" pageNumber="83" phylum="Amoebozoa" rank="species" species="gimlii">
|
||
<emphasis bold="true" box="[1056,1149,1582,1605]" italics="true" pageId="4" pageNumber="83">N. gimlii</emphasis>
|
||
</taxonomicName>
|
||
obtained from two different cells share
|
||
<number box="[933,980,1613,1636]" pageId="4" pageNumber="83" value="99.8">99.8</number>
|
||
% of similarity.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |