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<document id="3D524FDCFB8A5BE9942C863BCD13BE1D" ID-DOI="10.5281/zenodo.181581" ID-GBIF-Dataset="23f0fe20-030a-4a95-ba70-9e8880f04709" ID-ISSN="1175-5326" ID-Zenodo-Dep="181581" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459835706146" checkinUser="plazi" docAuthor="Botero-Trujillo, Ricardo" docDate="2008" docId="03BD87E9FFD7FF8E4BB92E21FC8870C2" docLanguage="en" docName="zt01743p042.pdf" docOrigin="Zootaxa 1743" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Chactopsis carolinae Botero-Trujillo, 2008, sp. nov." docType="treatment" docVersion="10" lastPageNumber="41" masterDocId="FF84FF91FFD5FF894B2E2D2CF9147278" masterDocTitle="First record of the scorpion genus Chactopsis Kraepelin in Colombia, with the description of Chactopsis carolinae sp. nov. (Scorpiones: Chactidae)" masterLastPageNumber="42" masterPageNumber="34" pageNumber="36" updateTime="1698223609098" updateUser="plazi">
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<mods:title id="E945765ACA1F0000C3B060C418336580">First record of the scorpion genus Chactopsis Kraepelin in Colombia, with the description of Chactopsis carolinae sp. nov. (Scorpiones: Chactidae)</mods:title>
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<mods:roleTerm id="6FFBE68386B91F0607E40DF8E9A037AC">Author</mods:roleTerm>
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<mods:namePart id="23C2CC9865B0966A4DE30167D8248683">Botero-Trujillo, Ricardo</mods:namePart>
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<mods:part id="839869C24961F308C2C10C4399492672">
<mods:date id="43E9D65BC3829BC070E97B3174DEF92E">2008</mods:date>
<mods:detail id="EBE3F3BAA0BBA827EA5945BF1A9DEE05" type="volume">
<mods:number id="0FB5A72F25241E7D1B4A0A735095210D">1743</mods:number>
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<mods:identifier id="E64C11090F6007CDC14AAE238312FAF2" type="DOI">10.5281/zenodo.181581</mods:identifier>
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<treatment id="03BD87E9FFD7FF8E4BB92E21FC8870C2" ID-DOI="http://doi.org/10.5281/zenodo.5678592" ID-GBIF-Taxon="119358836" ID-Zenodo-Dep="5678592" LSID="urn:lsid:plazi:treatment:03BD87E9FFD7FF8E4BB92E21FC8870C2" httpUri="http://treatment.plazi.org/id/03BD87E9FFD7FF8E4BB92E21FC8870C2" lastPageId="7" lastPageNumber="41" pageId="2" pageNumber="36">
<subSubSection id="C30E6574FFD7FF8B4BB92E21F8BE7132" pageId="2" pageNumber="36" type="nomenclature">
<paragraph id="8BAB36FFFFD7FF8B4BB92E21F8ED715F" blockId="2.[151,505,781,842]" box="[151,505,781,807]" pageId="2" pageNumber="36">
<heading id="D0E38193FFD7FF8B4BB92E21F8ED715F" bold="true" box="[151,505,781,807]" fontSize="11" level="1" pageId="2" pageNumber="36" reason="1">
<emphasis id="B960EAEDFFD7FF8B4BB92E21F8ED715F" bold="true" box="[151,505,781,807]" pageId="2" pageNumber="36">
<taxonomicName id="4C144D7CFFD7FF8B4BB92E21F880715F" box="[151,404,781,807]" class="Arachnida" family="Chactidae" genus="Chactopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="36" phylum="Arthropoda" rank="species" species="carolinae" status="sp. nov.">
<emphasis id="B960EAEDFFD7FF8B4BB92E21F880715F" bold="true" box="[151,404,781,807]" italics="true" pageId="2" pageNumber="36">Chactopsis carolinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2535796FFD7FF8B4AB52E21F8ED715F" box="[411,505,781,807]" pageId="2" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAB36FFFFD7FF8B4BB92E1CF8BE7132" blockId="2.[151,505,781,842]" box="[151,426,816,842]" pageId="2" pageNumber="36">
(
<figureCitation id="132F2A7AFFD7FF8B4BB12E1CF8297132" box="[159,317,816,842]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 5" captionStart-2="FIGURES 6 14" captionStartId-0="1.[151,255,2009,2033]" captionStartId-1="4.[151,269,1716,1740]" captionStartId-2="6.[151,269,1742,1766]" captionTargetBox-0="[231,1364,466,1977]" captionTargetBox-1="[153,1432,188,1688]" captionTargetBox-2="[210,1376,230,1689]" captionTargetId-0="figure@1.[231,1364,466,1977]" captionTargetId-1="figure@4.[151,1435,187,1690]" captionTargetId-2="figure@6.[185,1405,212,1701]" captionTargetPageId-0="1" captionTargetPageId-1="4" captionTargetPageId-2="6" captionText-0="FIGURE 1. Known distribution of Chactopsis carolinae sp. nov." captionText-1="FIGURES 2 5. Chactopsis carolinae sp. nov. 2 3. Male holotype (IAvH-E 100759), dorsal and ventral views. 4 5. Female paratype (IAvH-E 100760), dorsal and ventral views. Scale bars equal 5 mm." captionText-2="FIGURES 6 14. Chactopsis carolinae sp. nov., male holotype (IAvH-E 100759). 6 13. Distribution of the trichobothria. 6. Chela, dorsal view. 7. Chela, external view. 8. Chela, ventral view. 9. Chela, internal view. 10. Femur, dorsointernal view. 11. Patella, dorsointernal view. 12. Patella, ventral view. 13. Patella, external view. 14. Anterior margin of the carapace (coloration details not included). Scale bars equal 1 mm." httpUri-0="https://zenodo.org/record/181582/files/figure.png" httpUri-1="https://zenodo.org/record/181583/files/figure.png" httpUri-2="https://zenodo.org/record/181584/files/figure.png" pageId="2" pageNumber="36">Figures 114</figureCitation>
,
<tableCitation id="C6960344FFD7FF8B4A672E1CF88A7132" box="[329,414,816,842]" captionStart="TABLE 1" captionStartId="3.[151,239,151,175]" captionTargetBox="[156,1363,248,1979]" captionTargetPageId="3" captionText="TABLE 1. Morphometric measurements (in mm) of male holotype and adult female paratype of Chactopsis carolinae sp. nov." httpUri="http://table.plazi.org/id/DF6B6677FFD6FF8A4BB92DBBF9F972AA" pageId="2" pageNumber="36" tableUuid="DF6B6677FFD6FF8A4BB92DBBF9F972AA">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30E6574FFD7FF8B4BB92E53F81476F2" pageId="2" pageNumber="36" type="materials_examined">
<paragraph id="8BAB36FFFFD7FF8B4BB92E53F81476F2" blockId="2.[151,1436,895,2019]" pageId="2" pageNumber="36">
<emphasis id="B960EAEDFFD7FF8B4BB92E53F87D71E1" bold="true" box="[151,361,895,921]" pageId="2" pageNumber="36">
<typeStatus id="54AF885DFFD7FF8B4BB92E53F9C671E1" box="[151,210,895,921]" pageId="2" pageNumber="36">Type</typeStatus>
material.—
</emphasis>
<collectingCountry id="F303766FFFD7FF8B4A452EACFB1871E2" box="[363,524,896,922]" name="Colombia" pageId="2" pageNumber="36">COLOMBIA</collectingCountry>
. Male
<typeStatus id="54AF885DFFD7FF8B49732EACFBD071E2" box="[605,708,896,922]" pageId="2" pageNumber="36" type="holotype">holotype</typeStatus>
(adult) from Vichada department, Cumaribo, Selva de Matavén,
<geoCoordinate id="EE205038FFD7FF8B4BFF2E84F86571BA" box="[209,369,936,962]" direction="north" orientation="latitude" pageId="2" pageNumber="36" precision="15" value="4.486944">04°2913 N</geoCoordinate>
, -
<geoCoordinate id="EE205038FFD7FF8B4AA92E84FB3871BA" box="[391,556,936,962]" direction="west" orientation="longitude" pageId="2" pageNumber="36" precision="15" value="-68.00611">68°0022 W</geoCoordinate>
,
<quantity id="4CEC9B1AFFD7FF8B49172E84FB6B71BA" box="[569,639,936,962]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="36" unit="m" value="260.0">260m</quantity>
, Bosque de Planicies Arenosas, Pitfall,
<date id="FFAA103FFFD7FF8B4F772E84FDC971BA" box="[1113,1245,936,962]" pageId="2" pageNumber="36" value="2007-03-31">31/03/2007</date>
<date id="FFAA103FFFD7FF8B4FC32E84FC6171BA" box="[1261,1397,936,962]" pageId="2" pageNumber="36" value="2007-04-02">02/04/2007</date>
, L. E. Franco (IAvH-E
<date id="FFAA103FFFD7FF8B4A522EFCF8CD7192" box="[380,473,976,1002]" pageId="2" pageNumber="36" value="59-10-07">100759</date>
). One female
<typeStatus id="54AF885DFFD7FF8B49552EFCFBF47192" box="[635,736,976,1002]" pageId="2" pageNumber="36" type="paratype">paratype</typeStatus>
(adult) from Vichada department, Cumaribo, Selva de Matavén,
<geoCoordinate id="EE205038FFD7FF8B4BF12ED4F86B766A" box="[223,383,1016,1042]" direction="north" orientation="latitude" pageId="2" pageNumber="36" precision="15" value="4.6091666">04°3633 N</geoCoordinate>
, -
<geoCoordinate id="EE205038FFD7FF8B4AB82ED4FB2E766A" box="[406,570,1016,1042]" direction="west" orientation="longitude" pageId="2" pageNumber="36" precision="15" value="-67.86444">67°5152 W</geoCoordinate>
,
<quantity id="4CEC9B1AFFD7FF8B49662ED4FB99766A" box="[584,653,1016,1042]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="36" unit="m" value="300.0">300m</quantity>
, Bosque de Cerro Rocoso, Winkler,
<date id="FFAA103FFFD7FF8B4F6A2ED4FDEC766A" box="[1092,1272,1016,1042]" pageId="2" pageNumber="36" value="2007-03-06" valueMax="2007-03-08" valueMin="2007-03-06">0608/03/2007</date>
, L. E. Franco (IAvH-E
<date id="FFAA103FFFD7FF8B4A28290CF8777642" box="[262,355,1056,1082]" pageId="2" pageNumber="36" value="60-10-07">100760</date>
). One female
<typeStatus id="54AF885DFFD7FF8B4923290CFB667642" box="[525,626,1056,1082]" pageId="2" pageNumber="36" type="paratype">paratype</typeStatus>
(juvenile) from Vichada department, Cumaribo, Selva de Matavén,
<geoCoordinate id="EE205038FFD7FF8B4BB92964F823761A" box="[151,311,1096,1122]" direction="north" orientation="latitude" pageId="2" pageNumber="36" precision="15" value="4.558889">04°3332 N</geoCoordinate>
, -
<geoCoordinate id="EE205038FFD7FF8B4A602964F8E4761A" box="[334,496,1096,1122]" direction="west" orientation="longitude" pageId="2" pageNumber="36" precision="15" value="-68.1975">68°1151 W</geoCoordinate>
,
<quantity id="4CEC9B1AFFD7FF8B4AD02964FB50761A" box="[510,580,1096,1122]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="36" unit="m" value="270.0">270m</quantity>
, Bosque de Tierra Firme, Pitfall,
<date id="FFAA103FFFD7FF8B48FB2964FD9E761A" box="[981,1162,1096,1122]" pageId="2" pageNumber="36" value="2007-03-22" valueMax="2007-03-24" valueMin="2007-03-22">2224/03/2007</date>
, L. E. Franco (IAvH-E
<date id="FFAA103FFFD7FF8B4BB9295CF9E076F2" box="[151,244,1136,1162]" pageId="2" pageNumber="36" value="61-10-07">100761</date>
).
</paragraph>
</subSubSection>
<subSubSection id="C30E6574FFD7FF8B4BE829BBF89B777A" pageId="2" pageNumber="36" type="etymology">
<paragraph id="8BAB36FFFFD7FF8B4BE829BBF89B777A" blockId="2.[151,1436,895,2019]" pageId="2" pageNumber="36">
<emphasis id="B960EAEDFFD7FF8B4BE829BBF86476C9" bold="true" box="[198,368,1175,1201]" pageId="2" pageNumber="36">Etymology.—</emphasis>
Patronym dedicated to the authors girlfriend, Biviana
<taxonomicName id="4C144D7CFFD7FF8B4F1C29B4FC6476CA" authority="Baron Mendoza" authorityName="Baron Mendoza" box="[1074,1392,1176,1202]" class="Insecta" family="Scarabaeidae" genus="Carolina" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="36" phylum="Arthropoda" rank="genus">Carolina Barón Mendoza</taxonomicName>
, in acknowledgement to her support, encouragement and inspiration. It celebrates both past and forthcoming experiences together.
</paragraph>
</subSubSection>
<subSubSection id="C30E6574FFD7FF8C4BE82823F85F71FA" lastPageId="5" lastPageNumber="39" pageId="2" pageNumber="36" type="diagnosis">
<paragraph id="8BAB36FFFFD7FF8B4BE82823FA95759B" blockId="2.[151,1436,895,2019]" pageId="2" pageNumber="36">
<emphasis id="B960EAEDFFD7FF8B4BE82823F8707751" bold="true" box="[198,356,1295,1321]" pageId="2" pageNumber="36">Diagnosis.—</emphasis>
<taxonomicName id="4C144D7CFFD7FF8B4A4D283CFB4F7751" box="[355,603,1296,1321]" class="Arachnida" family="Chactidae" genus="Chactopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="36" phylum="Arthropoda" rank="species" species="carolinae" status="sp. nov.">
<emphasis id="B960EAEDFFD7FF8B4A4D283CFB4F7751" box="[355,603,1296,1321]" italics="true" pageId="2" pageNumber="36">Chactopsis carolinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2535796FFD7FF8B494C283CFBA87752" box="[610,700,1296,1322]" pageId="2" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
is most similar to
<taxonomicName id="4C144D7CFFD7FF8B48BC283CFD097751" box="[914,1053,1296,1321]" class="Arachnida" family="Chactidae" genus="Chactopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="36" phylum="Arthropoda" rank="species" species="anduzei">
<emphasis id="B960EAEDFFD7FF8B48BC283CFD097751" box="[914,1053,1296,1321]" italics="true" pageId="2" pageNumber="36">Ch. anduzei</emphasis>
</taxonomicName>
, with which it shares the following combination of features that distinguishes both species from all other in the genus: the trichobothrium
<emphasis id="B960EAEDFFD7FF8B4E452814FC957729" box="[1387,1409,1336,1361]" italics="true" pageId="2" pageNumber="36">ib</emphasis>
is not located in a basal position on the fixed finger of pedipalp chela (
<figureCitation id="132F2A7AFFD7FF8B4881284CFAE27702" box="[943,1014,1376,1402]" captionStart="FIGURES 6 14" captionStartId="6.[151,269,1742,1766]" captionTargetBox="[210,1376,230,1689]" captionTargetId="figure@6.[185,1405,212,1701]" captionTargetPageId="6" captionText="FIGURES 6 14. Chactopsis carolinae sp. nov., male holotype (IAvH-E 100759). 6 13. Distribution of the trichobothria. 6. Chela, dorsal view. 7. Chela, external view. 8. Chela, ventral view. 9. Chela, internal view. 10. Femur, dorsointernal view. 11. Patella, dorsointernal view. 12. Patella, ventral view. 13. Patella, external view. 14. Anterior margin of the carapace (coloration details not included). Scale bars equal 1 mm." httpUri="https://zenodo.org/record/181584/files/figure.png" pageId="2" pageNumber="36">Fig. 9</figureCitation>
;
<bibRefCitation id="EF854B0EFFD7FF8B4F2D284CFC0D7702" author="Gonzalez-Sponga" box="[1027,1305,1376,1402]" pageId="2" pageNumber="41" refString="Gonzalez-Sponga, M. A. (1982) Tres nuevas especies venezolanas del genero Chactopsis (Scorpionida: Chactidae). Boletin de la Academia de Ciencias Fisicas, Matematicas y Naturales de Venezuela, 42 (129 - 130), 127 - 146." type="journal article" year="1982">González-Sponga 1982</bibRefCitation>
:
<figureCitation id="132F2A7AFFD7FF8B4E0A284CFC7E7702" box="[1316,1386,1376,1402]" captionStart="FIGURES 2 5" captionStartId="4.[151,269,1716,1740]" captionTargetBox="[153,1432,188,1688]" captionTargetId="figure@4.[151,1435,187,1690]" captionTargetPageId="4" captionText="FIGURES 2 5. Chactopsis carolinae sp. nov. 2 3. Male holotype (IAvH-E 100759), dorsal and ventral views. 4 5. Female paratype (IAvH-E 100760), dorsal and ventral views. Scale bars equal 5 mm." httpUri="https://zenodo.org/record/181583/files/figure.png" pageId="2" pageNumber="36">Fig. 2</figureCitation>
);
<emphasis id="B960EAEDFFD7FF8B4EAE284CFC8F7701" box="[1408,1435,1376,1401]" italics="true" pageId="2" pageNumber="36">eb</emphasis>
and
<emphasis id="B960EAEDFFD7FF8B4BE728A4F9E477D9" box="[201,240,1416,1441]" italics="true" pageId="2" pageNumber="36">esb</emphasis>
are located between
<emphasis id="B960EAEDFFD7FF8B4AC728A4FB1977D9" box="[489,525,1416,1441]" italics="true" pageId="2" pageNumber="36">Db</emphasis>
and
<emphasis id="B960EAEDFFD7FF8B496828A4FB7777D9" box="[582,611,1416,1441]" italics="true" pageId="2" pageNumber="36">Dt</emphasis>
(
<figureCitation id="132F2A7AFFD7FF8B495A28A4FBF277DA" box="[628,742,1416,1442]" captionStart="FIGURES 6 14" captionStartId="6.[151,269,1742,1766]" captionTargetBox="[210,1376,230,1689]" captionTargetId="figure@6.[185,1405,212,1701]" captionTargetPageId="6" captionText="FIGURES 6 14. Chactopsis carolinae sp. nov., male holotype (IAvH-E 100759). 6 13. Distribution of the trichobothria. 6. Chela, dorsal view. 7. Chela, external view. 8. Chela, ventral view. 9. Chela, internal view. 10. Femur, dorsointernal view. 11. Patella, dorsointernal view. 12. Patella, ventral view. 13. Patella, external view. 14. Anterior margin of the carapace (coloration details not included). Scale bars equal 1 mm." httpUri="https://zenodo.org/record/181584/files/figure.png" pageId="2" pageNumber="36">Figs. 67</figureCitation>
;
<bibRefCitation id="EF854B0EFFD7FF8B49DC28A4FD1D77DA" author="Gonzalez-Sponga" box="[754,1033,1416,1442]" pageId="2" pageNumber="41" refString="Gonzalez-Sponga, M. A. (1982) Tres nuevas especies venezolanas del genero Chactopsis (Scorpionida: Chactidae). Boletin de la Academia de Ciencias Fisicas, Matematicas y Naturales de Venezuela, 42 (129 - 130), 127 - 146." type="journal article" year="1982">González-Sponga 1982</bibRefCitation>
:
<figureCitation id="132F2A7AFFD7FF8B4F3B28A4FD9177DA" box="[1045,1157,1416,1442]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 5" captionStartId-0="1.[151,255,2009,2033]" captionStartId-1="4.[151,269,1716,1740]" captionTargetBox-0="[231,1364,466,1977]" captionTargetBox-1="[153,1432,188,1688]" captionTargetId-0="figure@1.[231,1364,466,1977]" captionTargetId-1="figure@4.[151,1435,187,1690]" captionTargetPageId-0="1" captionTargetPageId-1="4" captionText-0="FIGURE 1. Known distribution of Chactopsis carolinae sp. nov." captionText-1="FIGURES 2 5. Chactopsis carolinae sp. nov. 2 3. Male holotype (IAvH-E 100759), dorsal and ventral views. 4 5. Female paratype (IAvH-E 100760), dorsal and ventral views. Scale bars equal 5 mm." httpUri-0="https://zenodo.org/record/181582/files/figure.png" httpUri-1="https://zenodo.org/record/181583/files/figure.png" pageId="2" pageNumber="36">Figs. 12</figureCitation>
); pedipalp patella bears 23 trichobothria on the external surface with
<emphasis id="B960EAEDFFD7FF8B4983289CFBC877B6" box="[685,732,1456,1486]" italics="true" pageId="2" pageNumber="36">esb5</emphasis>
absent (
<figureCitation id="132F2A7AFFD7FF8B486E289CFA8377B2" box="[832,919,1456,1482]" captionStart="FIGURES 6 14" captionStartId="6.[151,269,1742,1766]" captionTargetBox="[210,1376,230,1689]" captionTargetId="figure@6.[185,1405,212,1701]" captionTargetPageId="6" captionText="FIGURES 6 14. Chactopsis carolinae sp. nov., male holotype (IAvH-E 100759). 6 13. Distribution of the trichobothria. 6. Chela, dorsal view. 7. Chela, external view. 8. Chela, ventral view. 9. Chela, internal view. 10. Femur, dorsointernal view. 11. Patella, dorsointernal view. 12. Patella, ventral view. 13. Patella, external view. 14. Anterior margin of the carapace (coloration details not included). Scale bars equal 1 mm." httpUri="https://zenodo.org/record/181584/files/figure.png" pageId="2" pageNumber="36">Fig. 13</figureCitation>
;
<bibRefCitation id="EF854B0EFFD7FF8B488B289CFDA877B2" author="Gonzalez-Sponga" box="[933,1212,1456,1482]" pageId="2" pageNumber="41" refString="Gonzalez-Sponga, M. A. (1982) Tres nuevas especies venezolanas del genero Chactopsis (Scorpionida: Chactidae). Boletin de la Academia de Ciencias Fisicas, Matematicas y Naturales de Venezuela, 42 (129 - 130), 127 - 146." type="journal article" year="1982">González-Sponga 1982</bibRefCitation>
:
<figureCitation id="132F2A7AFFD7FF8B4FE6289CFC1B77B2" box="[1224,1295,1456,1482]" captionStart="FIGURES 2 5" captionStartId="4.[151,269,1716,1740]" captionTargetBox="[153,1432,188,1688]" captionTargetId="figure@4.[151,1435,187,1690]" captionTargetPageId="4" captionText="FIGURES 2 5. Chactopsis carolinae sp. nov. 2 3. Male holotype (IAvH-E 100759), dorsal and ventral views. 4 5. Female paratype (IAvH-E 100760), dorsal and ventral views. Scale bars equal 5 mm." httpUri="https://zenodo.org/record/181583/files/figure.png" pageId="2" pageNumber="36">Fig. 3</figureCitation>
), and seven ventral trichobothria of which
<date id="FFAA103FFFD7FF8B492B28F7FB797781" box="[517,621,1498,1529]" pageId="2" pageNumber="36">
<emphasis id="B960EAEDFFD7FF8B492B28F7FB0E7781" box="[517,538,1499,1529]" italics="true" pageId="2" pageNumber="36">v6</emphasis>
and
<emphasis id="B960EAEDFFD7FF8B497628F7FB797781" box="[600,621,1499,1529]" italics="true" pageId="2" pageNumber="36">v7</emphasis>
</date>
are not arranged linearly with
<date id="FFAA103FFFD7FF8B48CD28F7FD037781" box="[995,1047,1498,1529]" pageId="2" pageNumber="36">
<emphasis id="B960EAEDFFD7FF8B48CD28F7FD037781" box="[995,1047,1498,1529]" italics="true" pageId="2" pageNumber="36">
v1-v
<quantity id="4CEC9B1AFFD7FF8B4F2128C7FD037781" box="[1039,1047,1515,1529]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="2" pageNumber="36" unit="in" value="5.0">5</quantity>
</emphasis>
</date>
in the ventroexternal border but located near to the median longitudinal line of the segment (
<figureCitation id="132F2A7AFFD7FF8B48772B2AFAA47458" box="[857,944,1542,1568]" captionStart="FIGURES 6 14" captionStartId="6.[151,269,1742,1766]" captionTargetBox="[210,1376,230,1689]" captionTargetId="figure@6.[185,1405,212,1701]" captionTargetPageId="6" captionText="FIGURES 6 14. Chactopsis carolinae sp. nov., male holotype (IAvH-E 100759). 6 13. Distribution of the trichobothria. 6. Chela, dorsal view. 7. Chela, external view. 8. Chela, ventral view. 9. Chela, internal view. 10. Femur, dorsointernal view. 11. Patella, dorsointernal view. 12. Patella, ventral view. 13. Patella, external view. 14. Anterior margin of the carapace (coloration details not included). Scale bars equal 1 mm." httpUri="https://zenodo.org/record/181584/files/figure.png" pageId="2" pageNumber="36">Fig. 12</figureCitation>
;
<bibRefCitation id="EF854B0EFFD7FF8B48922B2AFDC77458" author="Gonzalez-Sponga" box="[956,1235,1542,1568]" pageId="2" pageNumber="41" refString="Gonzalez-Sponga, M. A. (1982) Tres nuevas especies venezolanas del genero Chactopsis (Scorpionida: Chactidae). Boletin de la Academia de Ciencias Fisicas, Matematicas y Naturales de Venezuela, 42 (129 - 130), 127 - 146." type="journal article" year="1982">González-Sponga 1982</bibRefCitation>
:
<figureCitation id="132F2A7AFFD7FF8B4FF02B2AFC327458" box="[1246,1318,1542,1568]" captionStart="FIGURES 2 5" captionStartId="4.[151,269,1716,1740]" captionTargetBox="[153,1432,188,1688]" captionTargetId="figure@4.[151,1435,187,1690]" captionTargetPageId="4" captionText="FIGURES 2 5. Chactopsis carolinae sp. nov. 2 3. Male holotype (IAvH-E 100759), dorsal and ventral views. 4 5. Female paratype (IAvH-E 100760), dorsal and ventral views. Scale bars equal 5 mm." httpUri="https://zenodo.org/record/181583/files/figure.png" pageId="2" pageNumber="36">Fig. 4</figureCitation>
). Furthermore,
<emphasis id="B960EAEDFFD7FF8B4BC92B02F9EC743F" box="[231,248,1582,1607]" italics="true" pageId="2" pageNumber="36">it</emphasis>
trichobothrium is located between
<emphasis id="B960EAEDFFD7FF8B49812B02FBDE743F" box="[687,714,1582,1607]" italics="true" pageId="2" pageNumber="36">eb</emphasis>
and
<emphasis id="B960EAEDFFD7FF8B48202B02FA21743F" box="[782,821,1582,1607]" italics="true" pageId="2" pageNumber="36">esb</emphasis>
, and
<emphasis id="B960EAEDFFD7FF8B48512B02FA82743F" box="[895,918,1582,1607]" italics="true" pageId="2" pageNumber="36">ib</emphasis>
is considerably basal to
<emphasis id="B960EAEDFFD7FF8B4FFF2B02FDF9743F" box="[1233,1261,1582,1607]" italics="true" pageId="2" pageNumber="36">eb</emphasis>
(
<figureCitation id="132F2A7AFFD7FF8B4E2C2B02FC617430" box="[1282,1397,1582,1608]" captionStart="FIGURES 6 14" captionStartId="6.[151,269,1742,1766]" captionTargetBox="[210,1376,230,1689]" captionTargetId="figure@6.[185,1405,212,1701]" captionTargetPageId="6" captionText="FIGURES 6 14. Chactopsis carolinae sp. nov., male holotype (IAvH-E 100759). 6 13. Distribution of the trichobothria. 6. Chela, dorsal view. 7. Chela, external view. 8. Chela, ventral view. 9. Chela, internal view. 10. Femur, dorsointernal view. 11. Patella, dorsointernal view. 12. Patella, ventral view. 13. Patella, external view. 14. Anterior margin of the carapace (coloration details not included). Scale bars equal 1 mm." httpUri="https://zenodo.org/record/181584/files/figure.png" pageId="2" pageNumber="36">Figs. 67</figureCitation>
)— although
<bibRefCitation id="EF854B0EFFD7FF8B4A242B7AFB617408" author="Gonzalez-Sponga" box="[266,629,1622,1648]" pageId="2" pageNumber="41" refString="Gonzalez-Sponga, M. A. (1982) Tres nuevas especies venezolanas del genero Chactopsis (Scorpionida: Chactidae). Boletin de la Academia de Ciencias Fisicas, Matematicas y Naturales de Venezuela, 42 (129 - 130), 127 - 146." type="journal article" year="1982">González-Sponga (1982: 128)</bibRefCitation>
erroneously noted that in
<emphasis id="B960EAEDFFD7FF8B48EF2B7AFD607417" box="[961,1140,1622,1647]" italics="true" pageId="2" pageNumber="36">
<taxonomicName id="4C144D7CFFD7FF8B48EF2B7AFD457417" box="[961,1105,1622,1647]" class="Arachnida" family="Chactidae" genus="Chactopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="36" phylum="Arthropoda" rank="species" species="anduzei">Ch. anduzei</taxonomicName>
ib
</emphasis>
and
<emphasis id="B960EAEDFFD7FF8B4F962B7AFDDD7417" box="[1208,1225,1622,1647]" italics="true" pageId="2" pageNumber="36">it</emphasis>
trichobothria are located between
<emphasis id="B960EAEDFFD7FF8B4A752B52F86374EF" box="[347,375,1662,1687]" italics="true" pageId="2" pageNumber="36">eb</emphasis>
and
<emphasis id="B960EAEDFFD7FF8B4A802B52F8C174EF" box="[430,469,1662,1687]" italics="true" pageId="2" pageNumber="36">esb</emphasis>
(see
<bibRefCitation id="EF854B0EFFD7FF8B493C2B52FA3C74E0" author="Gonzalez-Sponga" box="[530,808,1662,1688]" pageId="2" pageNumber="41" refString="Gonzalez-Sponga, M. A. (1982) Tres nuevas especies venezolanas del genero Chactopsis (Scorpionida: Chactidae). Boletin de la Academia de Ciencias Fisicas, Matematicas y Naturales de Venezuela, 42 (129 - 130), 127 - 146." type="journal article" year="1982">González-Sponga 1982</bibRefCitation>
:
<figureCitation id="132F2A7AFFD7FF8B481D2B52FA6D74E0" box="[819,889,1662,1688]" captionStart="FIGURES 2 5" captionStartId="4.[151,269,1716,1740]" captionTargetBox="[153,1432,188,1688]" captionTargetId="figure@4.[151,1435,187,1690]" captionTargetPageId="4" captionText="FIGURES 2 5. Chactopsis carolinae sp. nov. 2 3. Male holotype (IAvH-E 100759), dorsal and ventral views. 4 5. Female paratype (IAvH-E 100760), dorsal and ventral views. Scale bars equal 5 mm." httpUri="https://zenodo.org/record/181583/files/figure.png" pageId="2" pageNumber="36">Fig. 2</figureCitation>
). The new species can be readily distinguished from
<taxonomicName id="4C144D7CFFD7FF8B4BF62B8AF87174C7" box="[216,357,1702,1727]" class="Arachnida" family="Chactidae" genus="Chactopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="36" phylum="Arthropoda" rank="species" species="anduzei">
<emphasis id="B960EAEDFFD7FF8B4BF62B8AF87174C7" box="[216,357,1702,1727]" italics="true" pageId="2" pageNumber="36">Ch. anduzei</emphasis>
</taxonomicName>
since in the former the trichobothrium
<emphasis id="B960EAEDFFD7FF8B48152B8AFA4C74C7" box="[827,856,1702,1727]" italics="true" pageId="2" pageNumber="36">db</emphasis>
is basal in relation to
<emphasis id="B960EAEDFFD7FF8B4F4C2B8AFD9074BC" box="[1122,1156,1702,1732]" italics="true" pageId="2" pageNumber="36">Et4</emphasis>
(
<figureCitation id="132F2A7AFFD7FF8B4FBB2B8AFDCA74B8" box="[1173,1246,1702,1728]" captionStart="FIGURES 6 14" captionStartId="6.[151,269,1742,1766]" captionTargetBox="[210,1376,230,1689]" captionTargetId="figure@6.[185,1405,212,1701]" captionTargetPageId="6" captionText="FIGURES 6 14. Chactopsis carolinae sp. nov., male holotype (IAvH-E 100759). 6 13. Distribution of the trichobothria. 6. Chela, dorsal view. 7. Chela, external view. 8. Chela, ventral view. 9. Chela, internal view. 10. Femur, dorsointernal view. 11. Patella, dorsointernal view. 12. Patella, ventral view. 13. Patella, external view. 14. Anterior margin of the carapace (coloration details not included). Scale bars equal 1 mm." httpUri="https://zenodo.org/record/181584/files/figure.png" pageId="2" pageNumber="36">Fig. 7</figureCitation>
), patellar
<emphasis id="B960EAEDFFD7FF8B4E7B2B8AFC9474BC" box="[1365,1408,1702,1732]" italics="true" pageId="2" pageNumber="36">em2</emphasis>
is dorsal to
<emphasis id="B960EAEDFFD7FF8B4A2B2BFDF8397497" box="[261,301,1745,1775]" italics="true" pageId="2" pageNumber="36">est5</emphasis>
(
<figureCitation id="132F2A7AFFD7FF8B4A102BFDF8807493" box="[318,404,1745,1771]" captionStart="FIGURES 6 14" captionStartId="6.[151,269,1742,1766]" captionTargetBox="[210,1376,230,1689]" captionTargetId="figure@6.[185,1405,212,1701]" captionTargetPageId="6" captionText="FIGURES 6 14. Chactopsis carolinae sp. nov., male holotype (IAvH-E 100759). 6 13. Distribution of the trichobothria. 6. Chela, dorsal view. 7. Chela, external view. 8. Chela, ventral view. 9. Chela, internal view. 10. Femur, dorsointernal view. 11. Patella, dorsointernal view. 12. Patella, ventral view. 13. Patella, external view. 14. Anterior margin of the carapace (coloration details not included). Scale bars equal 1 mm." httpUri="https://zenodo.org/record/181584/files/figure.png" pageId="2" pageNumber="36">Fig. 13</figureCitation>
), sternite VII lacks any vestige of paramedian carinae, and metasomal segment V bears well-defined dorsolateral carinae. In contrast, in
<taxonomicName id="4C144D7CFFD7FF8B49F82BD0FA77756D" box="[726,867,1788,1813]" class="Arachnida" family="Chactidae" genus="Chactopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="36" phylum="Arthropoda" rank="species" species="anduzei">
<emphasis id="B960EAEDFFD7FF8B49F82BD0FA77756D" box="[726,867,1788,1813]" italics="true" pageId="2" pageNumber="36">Ch. anduzei</emphasis>
</taxonomicName>
the trichobothrium
<emphasis id="B960EAEDFFD7FF8B4F7E2BD0FD79756D" box="[1104,1133,1788,1813]" italics="true" pageId="2" pageNumber="36">db</emphasis>
is distal in relation to
<emphasis id="B960EAEDFFD7FF8B4E542BD0FC887562" box="[1402,1436,1788,1818]" italics="true" pageId="2" pageNumber="36">Et4</emphasis>
(
<bibRefCitation id="EF854B0EFFD7FF8B4BB12A0AF8AC7538" author="Gonzalez-Sponga" box="[159,440,1830,1856]" pageId="2" pageNumber="41" refString="Gonzalez-Sponga, M. A. (1982) Tres nuevas especies venezolanas del genero Chactopsis (Scorpionida: Chactidae). Boletin de la Academia de Ciencias Fisicas, Matematicas y Naturales de Venezuela, 42 (129 - 130), 127 - 146." type="journal article" year="1982">González-Sponga 1982</bibRefCitation>
:
<figureCitation id="132F2A7AFFD7FF8B4AED2A0AFB1D7538" box="[451,521,1830,1856]" captionStart="FIGURE 1" captionStartId="1.[151,255,2009,2033]" captionTargetBox="[231,1364,466,1977]" captionTargetId="figure@1.[231,1364,466,1977]" captionTargetPageId="1" captionText="FIGURE 1. Known distribution of Chactopsis carolinae sp. nov." httpUri="https://zenodo.org/record/181582/files/figure.png" pageId="2" pageNumber="36">Fig. 1</figureCitation>
;
<bibRefCitation id="EF854B0EFFD7FF8B49362A0AFA4E7538" author="Lourenco" box="[536,858,1830,1856]" pageId="2" pageNumber="42" refString="Lourenco, W. R. &amp; Francke, O. F. (1986) A new species of Chactopsis from Brazil (Scorpiones, Chactidae). Amazoniana, 9 (4), 549 - 558." type="journal article" year="1986">Lourenço &amp; Francke, 1986</bibRefCitation>
), patellar
<emphasis id="B960EAEDFFD7FF8B48FE2A0BFAEE753D" box="[976,1018,1831,1861]" italics="true" pageId="2" pageNumber="36">em2</emphasis>
is ventral to
<emphasis id="B960EAEDFFD7FF8B4FBB2A0BFDA9753D" box="[1173,1213,1831,1861]" italics="true" pageId="2" pageNumber="36">est5</emphasis>
(
<bibRefCitation id="EF854B0EFFD7FF8B4FE42A0AF83C7513" author="Gonzalez-Sponga" pageId="2" pageNumber="41" refString="Gonzalez-Sponga, M. A. (1982) Tres nuevas especies venezolanas del genero Chactopsis (Scorpionida: Chactidae). Boletin de la Academia de Ciencias Fisicas, Matematicas y Naturales de Venezuela, 42 (129 - 130), 127 - 146." type="journal article" year="1982">
González-Sponga 1982:
<figureCitation id="132F2A7AFFD7FF8B4BCF2A7DF83C7513" box="[225,296,1873,1899]" captionStart="FIGURES 2 5" captionStartId="4.[151,269,1716,1740]" captionTargetBox="[153,1432,188,1688]" captionTargetId="figure@4.[151,1435,187,1690]" captionTargetPageId="4" captionText="FIGURES 2 5. Chactopsis carolinae sp. nov. 2 3. Male holotype (IAvH-E 100759), dorsal and ventral views. 4 5. Female paratype (IAvH-E 100760), dorsal and ventral views. Scale bars equal 5 mm." httpUri="https://zenodo.org/record/181583/files/figure.png" pageId="2" pageNumber="36">Fig. 3</figureCitation>
</bibRefCitation>
), sternite VII presents paramedian carinae formed by vestigial granules, and metasomal segment V does not have well-defined dorsolateral carinae (
<bibRefCitation id="EF854B0EFFD7FF8B49C72A55FD1575EB" author="Gonzalez-Sponga" box="[745,1025,1913,1939]" pageId="2" pageNumber="41" refString="Gonzalez-Sponga, M. A. (1982) Tres nuevas especies venezolanas del genero Chactopsis (Scorpionida: Chactidae). Boletin de la Academia de Ciencias Fisicas, Matematicas y Naturales de Venezuela, 42 (129 - 130), 127 - 146." type="journal article" year="1982">González-Sponga 1982</bibRefCitation>
). In addition, the coloration pattern is different, especially on the tergites where a broad longitudinal yellowish band is present in the new species (
<figureCitation id="132F2A7AFFD7FF8B4BB12AE5F81B759B" box="[159,271,1993,2019]" captionStart="FIGURES 2 5" captionStartId="4.[151,269,1716,1740]" captionTargetBox="[153,1432,188,1688]" captionTargetId="figure@4.[151,1435,187,1690]" captionTargetPageId="4" captionText="FIGURES 2 5. Chactopsis carolinae sp. nov. 2 3. Male holotype (IAvH-E 100759), dorsal and ventral views. 4 5. Female paratype (IAvH-E 100760), dorsal and ventral views. Scale bars equal 5 mm." httpUri="https://zenodo.org/record/181583/files/figure.png" pageId="2" pageNumber="36">Figs. 2, 4</figureCitation>
) and absent in
<taxonomicName id="4C144D7CFFD7FF8B4AEF2AE5FA69759B" authority="Gonzalez-Sponga 1982" authorityName="Gonzalez-Sponga" authorityYear="1982" box="[449,893,1993,2019]" class="Arachnida" family="Chactidae" genus="Chactopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="36" phylum="Arthropoda" rank="species" species="anduzei">
<emphasis id="B960EAEDFFD7FF8B4AEF2AE5FB59759A" box="[449,589,1993,2018]" italics="true" pageId="2" pageNumber="36">Ch. anduzei</emphasis>
(
<bibRefCitation id="EF854B0EFFD7FF8B49722AE5FA60759B" author="Gonzalez-Sponga" box="[604,884,1993,2019]" pageId="2" pageNumber="41" refString="Gonzalez-Sponga, M. A. (1982) Tres nuevas especies venezolanas del genero Chactopsis (Scorpionida: Chactidae). Boletin de la Academia de Ciencias Fisicas, Matematicas y Naturales de Venezuela, 42 (129 - 130), 127 - 146." type="journal article" year="1982">González-Sponga 1982</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<caption id="DF6B6677FFD6FF8A4BB92DBBF9F972AA" ID-Table-UUID="DF6B6677FFD6FF8A4BB92DBBF9F972AA" httpUri="http://table.plazi.org/id/DF6B6677FFD6FF8A4BB92DBBF9F972AA" pageId="3" pageNumber="37" targetBox="[156,1363,248,1979]" targetIsTable="true" targetPageId="3">
<paragraph id="8BAB36FFFFD6FF8A4BB92DBBF9F972AA" blockId="3.[151,1435,151,210]" pageId="3" pageNumber="37">
<emphasis id="B960EAEDFFD6FF8A4BB92DBBF81E72D7" bold="true" box="[151,266,151,175]" pageId="3" pageNumber="37">TABLE 1.</emphasis>
Morphometric measurements (in mm) of male holotype and adult female paratype of
<taxonomicName id="4C144D7CFFD6FF8A4F962DB4FC8F72D7" box="[1208,1435,152,175]" class="Arachnida" family="Chactidae" genus="Chactopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="37" phylum="Arthropoda" rank="species" species="carolinae" status="sp. nov.">
<emphasis id="B960EAEDFFD6FF8A4F962DB4FC8F72D7" box="[1208,1435,152,175]" italics="true" pageId="3" pageNumber="37">Chactopsis carolinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2535796FFD6FF8A4BB92D96F9F972AA" box="[151,237,186,210]" pageId="3" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BAB36FFFFD6FF8A487D2DD4FC1E75C3" pageId="3" pageNumber="37">
<table id="F914C45FFFD600764BB22DD4FC4775C3" box="[156,1363,248,1979]" gridcols="3" gridrows="20" pageId="3" pageNumber="37">
<tr id="352434BDFFD600764BB22DD4FC47734A" box="[156,1363,248,306]" gridrow="0" pageId="3" pageNumber="37" rowspan-0="1">
<th id="76F55DC1FFD60076486C2DD4FAEA734A" box="[834,1022,248,306]" gridcol="1" gridrow="0" pageId="3" pageNumber="37">Male holotype (IAvH-E 100759)</th>
<th id="76F55DC1FFD600764FB92DD4FC47734A" box="[1175,1363,248,306]" gridcol="2" gridrow="0" pageId="3" pageNumber="37">Female paratype (IAvH-E 100760)</th>
</tr>
<tr id="352434BDFFD600764BB22C65FC4773FA" box="[156,1363,329,386]" gridrow="1" pageId="3" pageNumber="37">
<th id="76F55DC1FFD600764BB22C65FBAE73FA" box="[156,698,329,386]" gridcol="0" gridrow="1" pageId="3" pageNumber="37">Total body L (including telson)</th>
<td id="76F55DC1FFD60076486C2C65FAEA73FA" box="[834,1022,329,386]" gridcol="1" gridrow="1" pageId="3" pageNumber="37">18.99</td>
<td id="76F55DC1FFD600764FB92C65FC4773FA" box="[1175,1363,329,386]" gridcol="2" gridrow="1" pageId="3" pageNumber="37">22.67</td>
</tr>
<tr id="352434BDFFD600764BB22CB5FC4773A5" box="[156,1363,409,477]" gridrow="2" pageId="3" pageNumber="37">
<th id="76F55DC1FFD600764BB22CB5FBAE73A5" box="[156,698,409,477]" gridcol="0" gridrow="2" pageId="3" pageNumber="37">Carapace Length Anterior W</th>
<td id="76F55DC1FFD60076486C2CB5FAEA73A5" box="[834,1022,409,477]" gridcol="1" gridrow="2" pageId="3" pageNumber="37">2.64 1.66</td>
<td id="76F55DC1FFD600764FB92CB5FC4773A5" box="[1175,1363,409,477]" gridcol="2" gridrow="2" pageId="3" pageNumber="37">3.02 2.11</td>
</tr>
<tr id="352434BDFFD600764BB22CD8FC477040" box="[156,1363,500,568]" gridrow="3" pageId="3" pageNumber="37">
<th id="76F55DC1FFD600764BB22CD8FBAE7040" box="[156,698,500,568]" gridcol="0" gridrow="3" pageId="3" pageNumber="37">Posterior W Ocular diameter</th>
<td id="76F55DC1FFD60076486C2CD8FAEA7040" box="[834,1022,500,568]" gridcol="1" gridrow="3" pageId="3" pageNumber="37">2.52 0.12</td>
<td id="76F55DC1FFD600764FB92CD8FC477040" box="[1175,1363,500,568]" gridcol="2" gridrow="3" pageId="3" pageNumber="37">3.24 0.18</td>
</tr>
<tr id="352434BDFFD600764BB22F63FC4770EB" box="[156,1363,591,659]" gridrow="4" pageId="3" pageNumber="37">
<th id="76F55DC1FFD600764BB22F63FBAE70EB" box="[156,698,591,659]" gridcol="0" gridrow="4" pageId="3" pageNumber="37">Interocular distance Mesosoma Total L</th>
<td id="76F55DC1FFD60076486C2F63FAEA70EB" box="[834,1022,591,659]" gridcol="1" gridrow="4" pageId="3" pageNumber="37">0.14 5.08</td>
<td id="76F55DC1FFD600764FB92F63FC4770EB" box="[1175,1363,591,659]" gridcol="2" gridrow="4" pageId="3" pageNumber="37">0.16 6.42</td>
</tr>
<tr id="352434BDFFD600764BB22F86FC477096" box="[156,1363,682,750]" gridrow="5" pageId="3" pageNumber="37">
<th id="76F55DC1FFD600764BB22F86FBAE7096" box="[156,698,682,750]" gridcol="0" gridrow="5" pageId="3" pageNumber="37">Metasoma Total L (including telson) Segment I L</th>
<td id="76F55DC1FFD60076486C2F86FAEA7096" box="[834,1022,682,750]" gridcol="1" gridrow="5" pageId="3" pageNumber="37">11.27 0.99</td>
<td id="76F55DC1FFD600764FB92F86FC477096" box="[1175,1363,682,750]" gridcol="2" gridrow="5" pageId="3" pageNumber="37">13.23 1.23</td>
</tr>
<tr id="352434BDFFD600764BB22E29FC477131" box="[156,1363,773,841]" gridrow="6" pageId="3" pageNumber="37">
<th id="76F55DC1FFD600764BB22E29FBAE7131" box="[156,698,773,841]" gridcol="0" gridrow="6" pageId="3" pageNumber="37">Segment I W Segment I D</th>
<td id="76F55DC1FFD60076486C2E29FAEA7131" box="[834,1022,773,841]" gridcol="1" gridrow="6" pageId="3" pageNumber="37">1.56 1.21</td>
<td id="76F55DC1FFD600764FB92E29FC477131" box="[1175,1363,773,841]" gridcol="2" gridrow="6" pageId="3" pageNumber="37">2.04 1.39</td>
</tr>
<tr id="352434BDFFD600764BB22E4CFC4771DC" box="[156,1363,864,932]" gridrow="7" pageId="3" pageNumber="37">
<th id="76F55DC1FFD600764BB22E4CFBAE71DC" box="[156,698,864,932]" gridcol="0" gridrow="7" pageId="3" pageNumber="37">Segment II L Segment II W</th>
<td id="76F55DC1FFD60076486C2E4CFAEA71DC" box="[834,1022,864,932]" gridcol="1" gridrow="7" pageId="3" pageNumber="37">1.14 1.50</td>
<td id="76F55DC1FFD600764FB92E4CFC4771DC" box="[1175,1363,864,932]" gridcol="2" gridrow="7" pageId="3" pageNumber="37">1.36 1.95</td>
</tr>
<tr id="352434BDFFD600764BB22E97FC477187" box="[156,1363,955,1023]" gridrow="8" pageId="3" pageNumber="37">
<th id="76F55DC1FFD600764BB22E97FBAE7187" box="[156,698,955,1023]" gridcol="0" gridrow="8" pageId="3" pageNumber="37">Segment II D Segment III L</th>
<td id="76F55DC1FFD60076486C2E97FAEA7187" box="[834,1022,955,1023]" gridcol="1" gridrow="8" pageId="3" pageNumber="37">1.11 1.20</td>
<td id="76F55DC1FFD600764FB92E97FC477187" box="[1175,1363,955,1023]" gridcol="2" gridrow="8" pageId="3" pageNumber="37">1.44 1.44</td>
</tr>
<tr id="352434BDFFD600764BB2293AFC477655" box="[156,1363,1046,1069]" gridrow="9" pageId="3" pageNumber="37">
<th id="76F55DC1FFD600764BB2293AFBAE7655" box="[156,698,1046,1069]" gridcol="0" gridrow="9" pageId="3" pageNumber="37">Segment III W</th>
<td id="76F55DC1FFD60076486C293AFAEA7655" box="[834,1022,1046,1069]" gridcol="1" gridrow="9" pageId="3" pageNumber="37">1.60</td>
<td id="76F55DC1FFD600764FB9293AFC477655" box="[1175,1363,1046,1069]" gridcol="2" gridrow="9" pageId="3" pageNumber="37">1.95</td>
</tr>
<tr id="352434BDFFD600764BB22968FC4776CD" box="[156,1363,1092,1205]" gridrow="10" pageId="3" pageNumber="37">
<th id="76F55DC1FFD600764BB22968FBAE76CD" box="[156,698,1092,1205]" gridcol="0" gridrow="10" pageId="3" pageNumber="37">Segment III D Segment IV L Segment IV W</th>
<td id="76F55DC1FFD60076486C2968FAEA76CD" box="[834,1022,1092,1205]" gridcol="1" gridrow="10" pageId="3" pageNumber="37">1.21 1.63 1.60</td>
<td id="76F55DC1FFD600764FB92968FC4776CD" box="[1175,1363,1092,1205]" gridcol="2" gridrow="10" pageId="3" pageNumber="37">1.62 1.85 1.99</td>
</tr>
<tr id="352434BDFFD600764BB229E0FC47769B" box="[156,1363,1228,1251]" gridrow="11" pageId="3" pageNumber="37">
<th id="76F55DC1FFD600764BB229E0FBAE769B" box="[156,698,1228,1251]" gridcol="0" gridrow="11" pageId="3" pageNumber="37">Segment IV D</th>
<td id="76F55DC1FFD60076486C29E0FAEA769B" box="[834,1022,1228,1251]" gridcol="1" gridrow="11" pageId="3" pageNumber="37">1.18</td>
<td id="76F55DC1FFD600764FB929E0FC47769B" box="[1175,1363,1228,1251]" gridcol="2" gridrow="11" pageId="3" pageNumber="37">1.81</td>
</tr>
<tr id="352434BDFFD600764BB229D6FC477713" box="[156,1363,1274,1387]" gridrow="12" pageId="3" pageNumber="37">
<th id="76F55DC1FFD600764BB229D6FBAE7713" box="[156,698,1274,1387]" gridcol="0" gridrow="12" pageId="3" pageNumber="37">Segment V L Segment V W Segment V D</th>
<td id="76F55DC1FFD60076486C29D6FAEA7713" box="[834,1022,1274,1387]" gridcol="1" gridrow="12" pageId="3" pageNumber="37">3.03 1.60 1.31</td>
<td id="76F55DC1FFD600764FB929D6FC477713" box="[1175,1363,1274,1387]" gridcol="2" gridrow="12" pageId="3" pageNumber="37">3.57 2.01 1.74</td>
</tr>
<tr id="352434BDFFD600764BB228AEFC4777BE" box="[156,1363,1410,1478]" gridrow="13" pageId="3" pageNumber="37">
<th id="76F55DC1FFD600764BB228AEFBAE77BE" box="[156,698,1410,1478]" gridcol="0" gridrow="13" pageId="3" pageNumber="37">Telson L Vesicle W</th>
<td id="76F55DC1FFD60076486C28AEFAEA77BE" box="[834,1022,1410,1478]" gridcol="1" gridrow="13" pageId="3" pageNumber="37">3.28 1.24</td>
<td id="76F55DC1FFD600764FB928AEFC4777BE" box="[1175,1363,1410,1478]" gridcol="2" gridrow="13" pageId="3" pageNumber="37">3.78 1.52</td>
</tr>
<tr id="352434BDFFD600764BB228F1FC47778C" box="[156,1363,1501,1524]" gridrow="14" pageId="3" pageNumber="37">
<th id="76F55DC1FFD600764BB228F1FBAE778C" box="[156,698,1501,1524]" gridcol="0" gridrow="14" pageId="3" pageNumber="37">Vesicle D</th>
<td id="76F55DC1FFD60076486C28F1FAEA778C" box="[834,1022,1501,1524]" gridcol="1" gridrow="14" pageId="3" pageNumber="37">0.94</td>
<td id="76F55DC1FFD600764FB928F1FC47778C" box="[1175,1363,1501,1524]" gridcol="2" gridrow="14" pageId="3" pageNumber="37">1.19</td>
</tr>
<tr id="352434BDFFD600764BB22B27FC477404" box="[156,1363,1547,1660]" gridrow="15" pageId="3" pageNumber="37">
<th id="76F55DC1FFD600764BB22B27FBAE7404" box="[156,698,1547,1660]" gridcol="0" gridrow="15" pageId="3" pageNumber="37">Pedipalps Total L Femur L Femur W</th>
<td id="76F55DC1FFD60076486C2B27FAEA7404" box="[834,1022,1547,1660]" gridcol="1" gridrow="15" pageId="3" pageNumber="37">8.55 2.10 0.74</td>
<td id="76F55DC1FFD600764FB92B27FC477404" box="[1175,1363,1547,1660]" gridcol="2" gridrow="15" pageId="3" pageNumber="37">10.49 2.41 0.83</td>
</tr>
<tr id="352434BDFFD600764BB22BBFFC4774D2" box="[156,1363,1683,1706]" gridrow="16" pageId="3" pageNumber="37">
<th id="76F55DC1FFD600764BB22BBFFBAE74D2" box="[156,698,1683,1706]" gridcol="0" gridrow="16" pageId="3" pageNumber="37">Patella L</th>
<td id="76F55DC1FFD60076486C2BBFFAEA74D2" box="[834,1022,1683,1706]" gridcol="1" gridrow="16" pageId="3" pageNumber="37">2.81</td>
<td id="76F55DC1FFD600764FB92BBFFC4774D2" box="[1175,1363,1683,1706]" gridcol="2" gridrow="16" pageId="3" pageNumber="37">3.22</td>
</tr>
<tr id="352434BDFFD600764BB22BEDFC47757D" box="[156,1363,1729,1797]" gridrow="17" pageId="3" pageNumber="37">
<th id="76F55DC1FFD600764BB22BEDFBAE757D" box="[156,698,1729,1797]" gridcol="0" gridrow="17" pageId="3" pageNumber="37">Patella W Chela L</th>
<td id="76F55DC1FFD60076486C2BEDFAEA757D" box="[834,1022,1729,1797]" gridcol="1" gridrow="17" pageId="3" pageNumber="37">0.99 3.64</td>
<td id="76F55DC1FFD600764FB92BEDFC47757D" box="[1175,1363,1729,1797]" gridcol="2" gridrow="17" pageId="3" pageNumber="37">1.24 4.86</td>
</tr>
<tr id="352434BDFFD600764BB22A30FC4775F5" box="[156,1363,1820,1933]" gridrow="18" pageId="3" pageNumber="37">
<th id="76F55DC1FFD600764BB22A30FBAE75F5" box="[156,698,1820,1933]" gridcol="0" gridrow="18" pageId="3" pageNumber="37">Chela W Chela D Movable finger L</th>
<td id="76F55DC1FFD60076486C2A30FAEA75F5" box="[834,1022,1820,1933]" gridcol="1" gridrow="18" pageId="3" pageNumber="37">0.90 0.96 2.25</td>
<td id="76F55DC1FFD600764FB92A30FC4775F5" box="[1175,1363,1820,1933]" gridcol="2" gridrow="18" pageId="3" pageNumber="37">1.18 1.19 3.23</td>
</tr>
<tr id="352434BDFFD600764BB22A88FC4775C3" box="[156,1363,1956,1979]" gridrow="19" pageId="3" pageNumber="37">
<th id="76F55DC1FFD600764BB22A88FBAE75C3" box="[156,698,1956,1979]" gridcol="0" gridrow="19" pageId="3" pageNumber="37">Palm L</th>
<td id="76F55DC1FFD60076486C2A88FAEA75C3" box="[834,1022,1956,1979]" gridcol="1" gridrow="19" pageId="3" pageNumber="37">1.53</td>
<td id="76F55DC1FFD600764FB92A88FC4775C3" box="[1175,1363,1956,1979]" gridcol="2" gridrow="19" pageId="3" pageNumber="37">1.78</td>
</tr>
</table>
</paragraph>
<caption id="DF6B6677FFD1FF8D4BB92B98FD0F748B" httpUri="https://zenodo.org/record/181583/files/figure.png" pageId="4" pageNumber="38" targetBox="[153,1432,188,1688]" targetPageId="4">
<paragraph id="8BAB36FFFFD1FF8D4BB92B98FD0F748B" blockId="4.[151,1436,1716,1779]" pageId="4" pageNumber="38">
<emphasis id="B960EAEDFFD1FF8D4BB92B98F85274B4" bold="true" box="[151,326,1716,1740]" pageId="4" pageNumber="38">FIGURES 25.</emphasis>
<taxonomicName id="4C144D7CFFD1FF8D4A612B98FB2174B3" box="[335,565,1716,1739]" class="Arachnida" family="Chactidae" genus="Chactopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="38" phylum="Arthropoda" rank="species" species="carolinae" status="sp. nov.">
<emphasis id="B960EAEDFFD1FF8D4A612B98FB2174B3" box="[335,565,1716,1739]" italics="true" pageId="4" pageNumber="38">Chactopsis carolinae</emphasis>
</taxonomicName>
<emphasis id="B960EAEDFFD1FF8D49102B98FBDB74B4" bold="true" box="[574,719,1716,1740]" pageId="4" pageNumber="38">
<taxonomicNameLabel id="A2535796FFD1FF8D49102B98FB8374B4" box="[574,663,1716,1740]" pageId="4" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
23.
</emphasis>
Male holotype (IAvH-E 100759), dorsal and ventral views.
<emphasis id="B960EAEDFFD1FF8D4E432B98FC8874B4" bold="true" box="[1389,1436,1716,1740]" pageId="4" pageNumber="38">45.</emphasis>
Female paratype (IAvH-E 100760), dorsal and ventral views. Scale bars equal 5mm.
</paragraph>
</caption>
<paragraph id="8BAB36FFFFD1FF8D4BE82A06FA32753C" blockId="4.[151,1437,1834,2020]" box="[198,806,1834,1860]" pageId="4" pageNumber="38">
Description of the male
<typeStatus id="54AF885DFFD1FF8D4ACF2A06FB5C753C" box="[481,584,1834,1860]" pageId="4" pageNumber="38" type="holotype">holotype</typeStatus>
(IAvH-E
<date id="FFAA103FFFD1FF8D49922A06FA0D753C" box="[700,793,1834,1860]" pageId="4" pageNumber="38" value="59-10-07">100759</date>
).
</paragraph>
<paragraph id="8BAB36FFFFD1FF8C4BE82A7EF85F71FA" blockId="4.[151,1437,1834,2020]" lastBlockId="5.[151,1437,152,1983]" lastPageId="5" lastPageNumber="39" pageId="4" pageNumber="38">
<emphasis id="B960EAEDFFD1FF8D4BE82A7EF8797514" bold="true" box="[198,365,1874,1900]" italics="true" pageId="4" pageNumber="38">Coloration.—</emphasis>
Carapace predominantly brown with some lateral yellowish regions and a broad median longitudinal yellow area crossing from the anterior to the posterior border; median and lateral ocular tubercles black; a brown area is present from the median ocular tubercle to the anterior margin of the carapace, this is more conspicuous in the posterior region and is formed by inconspicuous spots anteriorly. Chelicerae predominantly yellow; coxa with some light-brown regions in dorsal view; tibia dorsally and externally with broad brown longitudinal reticulations and others thinner located between these, a rounded brown area is present just under the base of the fixed finger, ventral and internal surfaces with inconspicuous reticulations; fixed finger yellow, teeth reddish; movable finger light-brown, teeth reddish. Coxosternal region almost completely yellow, except for coxae II that bear few brownish spots in the inner half of the segment, and the sternum that is brownish anteriorly. Genital opercula, pectinal basal piece, pectines and sternites IIIIV completely yellow; sternites VVI with inconspicuous brown spots laterally; sternite VII with dark-brown regions on the position of lateral carinae. Tergites predominantly brown; a broad median longitudinal yellowish band is present in all tergites (
<figureCitation id="132F2A7AFFD0FF8C4BD32CF4F864738A" box="[253,368,472,498]" captionStart="FIGURES 2 5" captionStartId="4.[151,269,1716,1740]" captionTargetBox="[153,1432,188,1688]" captionTargetId="figure@4.[151,1435,187,1690]" captionTargetPageId="4" captionText="FIGURES 2 5. Chactopsis carolinae sp. nov. 2 3. Male holotype (IAvH-E 100759), dorsal and ventral views. 4 5. Female paratype (IAvH-E 100760), dorsal and ventral views. Scale bars equal 5 mm." httpUri="https://zenodo.org/record/181583/files/figure.png" pageId="5" pageNumber="39">Figs. 2, 4</figureCitation>
); each side of tergites IIVI with two transverse yellow lines converging near to the longitudinal line, resembling an arrow; lateral margins of all tergites yellow; tergite VII with brown mottling laterally. Metasoma with abundant dark-brown mottling in lateral and ventral surfaces; dorsal region of all segments completely yellow, except for segment V that bears lateral brown-spotted areas posteriorly. Telson yellow in dorsal surface, with few brown spots basally; lateral and ventral surfaces predominantly brown, with two ventromedian and two lateral yellow bands; aculeus reddish and spotless. Pedipalps predominantly light-brown; femur with brown regions in dorsointernal, dorsal, and ventroexternal surfaces, others yellow; patella with yellow areas in dorsal, external and internal surfaces, predominantly yellow ventrally; hand with longitudinal brown bands and few reticulations in all surfaces, except ventrally where almost completely yellow; fixed and movable fingers yellow. Legs with variegated pigmentation, except for coxae and trochanters that are yellow.
</paragraph>
</subSubSection>
<subSubSection id="C30E6574FFD0FF8E4BE82EA3FA8072A2" lastPageId="7" lastPageNumber="41" pageId="5" pageNumber="39" type="description">
<paragraph id="8BAB36FFFFD0FF8C4BE82EA3FD3D75C7" blockId="5.[151,1437,152,1983]" pageId="5" pageNumber="39">
<emphasis id="B960EAEDFFD0FF8C4BE82EA3F8E871D1" box="[198,508,911,937]" italics="true" pageId="5" pageNumber="39">
<emphasis id="B960EAEDFFD0FF8C4BE82EA3F86A71D1" bold="true" box="[198,382,911,937]" italics="true" pageId="5" pageNumber="39">Morphology.—</emphasis>
Carapace:
</emphasis>
subtriangular, densely covered with rounded granules throughout; anterior margin moderately emarginated and with a median tip directed anteriorly (
<figureCitation id="132F2A7AFFD0FF8C4F362E99FD6671B7" box="[1048,1138,949,975]" captionStart="FIGURES 6 14" captionStartId="6.[151,269,1742,1766]" captionTargetBox="[210,1376,230,1689]" captionTargetId="figure@6.[185,1405,212,1701]" captionTargetPageId="6" captionText="FIGURES 6 14. Chactopsis carolinae sp. nov., male holotype (IAvH-E 100759). 6 13. Distribution of the trichobothria. 6. Chela, dorsal view. 7. Chela, external view. 8. Chela, ventral view. 9. Chela, internal view. 10. Femur, dorsointernal view. 11. Patella, dorsointernal view. 12. Patella, ventral view. 13. Patella, external view. 14. Anterior margin of the carapace (coloration details not included). Scale bars equal 1 mm." httpUri="https://zenodo.org/record/181584/files/figure.png" pageId="5" pageNumber="39">Fig. 14</figureCitation>
); median ocular tubercle low, distinctly anterior to the centre of the carapace; lateral ocular tubercles each with thee ocelli, the posterior most is reduced (
<figureCitation id="132F2A7AFFD0FF8C4A4C292CF8AF7662" box="[354,443,1024,1050]" captionStart="FIGURES 6 14" captionStartId="6.[151,269,1742,1766]" captionTargetBox="[210,1376,230,1689]" captionTargetId="figure@6.[185,1405,212,1701]" captionTargetPageId="6" captionText="FIGURES 6 14. Chactopsis carolinae sp. nov., male holotype (IAvH-E 100759). 6 13. Distribution of the trichobothria. 6. Chela, dorsal view. 7. Chela, external view. 8. Chela, ventral view. 9. Chela, internal view. 10. Femur, dorsointernal view. 11. Patella, dorsointernal view. 12. Patella, ventral view. 13. Patella, external view. 14. Anterior margin of the carapace (coloration details not included). Scale bars equal 1 mm." httpUri="https://zenodo.org/record/181584/files/figure.png" pageId="5" pageNumber="39">Fig. 14</figureCitation>
).
<emphasis id="B960EAEDFFD0FF8C4AFF292CFB487661" box="[465,604,1024,1049]" italics="true" pageId="5" pageNumber="39">Chelicerae:</emphasis>
with abundant setae on internal and ventral surfaces; cheliceral dentition characteristic of the family
<taxonomicName id="4C144D7CFFD0FF8C49322909FA5B7647" authority="Vachon 1963" authorityName="Vachon" authorityYear="1963" box="[540,847,1061,1087]" class="Arachnida" family="Chactidae" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="39" phylum="Arthropoda" rank="family">
Chactidae (
<bibRefCitation id="EF854B0EFFD0FF8C498D2909FA527647" author="Vachon" box="[675,838,1061,1087]" pageId="5" pageNumber="42" refString="Vachon, M. (1963) De l'utilite, en systematique, d'une nomenclature des dents des cheliceres chez les scorpions. Bulletin du Museum national de Histoire naturelle (Paris) (2 e ser.), 35 (2), 161 - 166." type="journal article" year="1963">Vachon 1963</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="B960EAEDFFD0FF8C48732909FD587646" box="[861,1100,1061,1086]" italics="true" pageId="5" pageNumber="39">Coxosternal region:</emphasis>
sternum pentagonal, with a median longitudinal furrow; coxae sparsely punctulate, with few marginal weak tubercles and few setae; coxapophyses III with dense pilosity anteriorly.
<emphasis id="B960EAEDFFD0FF8C4983295CFD0376F1" box="[685,1047,1136,1161]" italics="true" pageId="5" pageNumber="39">Genital opercula and pectines:</emphasis>
genital opercula divided longitudinally; pectinal basal piece wider than long, scarcely punctulate; pectines long, surpassing the lateral margins of sternite III and with 11:11 teeth.
<emphasis id="B960EAEDFFD0FF8C49632997FBA876AC" box="[589,700,1211,1236]" italics="true" pageId="5" pageNumber="39">Sternites:</emphasis>
IIIVII smooth, densely punctulate; sternite VII with sparse weak granules on the lateral margins, without any vestige of paramedian carinae but with two slight elevations on the position of lateral carinae; spiracles small and oval.
<emphasis id="B960EAEDFFD0FF8C48682829FABD7766" box="[838,937,1285,1310]" italics="true" pageId="5" pageNumber="39">Tergites:</emphasis>
with similar granulation to that of the carapace; tergites IIIVI with vestigial axial and dorsolateral carinae represented by one or two granules on the posterior borders; tergite VII tetracarinate (paired dorsolateral and lateral carinae, incomplete and with strong but non-spinoid granules), a median elevation is present in the position of the axial carina, where some greater granules are located in unaligned position.
<emphasis id="B960EAEDFFD0FF8C49B628B7FA0877CC" box="[664,796,1435,1460]" italics="true" pageId="5" pageNumber="39">Metasoma:</emphasis>
with abundant long setae; segments IIV with sparse and weak granulation throughout; segment V with dense granulation; all carinae vestigial, slightly elevated except for dorsal and dorsolateral carinae that are stronger; segment I with eight complete carinae (paired ventrolateral, intermedian, dorsolateral and dorsal carinae; ventral carinae absent); segment II with ten (ventral carinae incomplete, reduced to a median elevation; intermedian carinae only represented posteriorly by a big tubercle; others complete); segment III with ten (intermedian carinae as in segment II; others complete); segments IVV with eight (intermedian carinae absent; others complete); in segment V dorsolateral carinae are only present on the anterior and median thirds of the segment, and ventral carinae are diffuse, difficult to distinguish from the remaining granules of the tegument. Telson densely punctulate throughout and with welldeveloped granules in lateral and ventral surfaces; aculeus short, thick basally.
<emphasis id="B960EAEDFFD0FF8C4F692BC7FDD2757C" box="[1095,1222,1771,1796]" italics="true" pageId="5" pageNumber="39">Pedipalps:</emphasis>
femur tricarinate, with sparse rounded granules in dorsal and internal surfaces; patella tricarinate, with few low granules dorsally; chela acarinate, smooth; all pedipalp segments densely punctulate, except dorsal surfaces of femur and patella. Trichobothriotaxy (
<figureCitation id="132F2A7AFFD0FF8C4AF92A76FB4D750C" box="[471,601,1882,1908]" captionStart="FIGURES 6 14" captionStartId="6.[151,269,1742,1766]" captionTargetBox="[210,1376,230,1689]" captionTargetId="figure@6.[185,1405,212,1701]" captionTargetPageId="6" captionText="FIGURES 6 14. Chactopsis carolinae sp. nov., male holotype (IAvH-E 100759). 6 13. Distribution of the trichobothria. 6. Chela, dorsal view. 7. Chela, external view. 8. Chela, ventral view. 9. Chela, internal view. 10. Femur, dorsointernal view. 11. Patella, dorsointernal view. 12. Patella, ventral view. 13. Patella, external view. 14. Anterior margin of the carapace (coloration details not included). Scale bars equal 1 mm." httpUri="https://zenodo.org/record/181584/files/figure.png" pageId="5" pageNumber="39">Figs. 613</figureCitation>
):
<typeStatus id="54AF885DFFD0FF8C49432A76FBB3750C" box="[621,679,1882,1908]" pageId="5" pageNumber="39">Type</typeStatus>
C (
<bibRefCitation id="EF854B0EFFD0FF8C49FE2A76FA7A750C" author="Vachon" box="[720,878,1882,1908]" pageId="5" pageNumber="42" refString="Vachon, M. (1973) Etude des caracteres utilises pour classer les familles et les genres de scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les scorpions. Bulletin du Museum national de Histoire naturelle (Paris) (3 e ser.), 104, 857 - 958." type="journal article" year="1973">Vachon 1973</bibRefCitation>
) with 62 trichobothria; femur with three trichobothria; patella with 33 of which 23 are external; chela with 26.
<emphasis id="B960EAEDFFD0FF8C48842AACFAFF75E1" box="[938,1003,1920,1945]" italics="true" pageId="5" pageNumber="39">Legs:</emphasis>
tibia, basitarsus and telotarsus with abundant setae not arranged in rows on external, internal and ventral surfaces.
</paragraph>
<caption id="DF6B6677FFD3FF8F4BB92BE2FA5B753D" httpUri="https://zenodo.org/record/181584/files/figure.png" pageId="6" pageNumber="40" targetBox="[210,1376,230,1689]" targetPageId="6">
<paragraph id="8BAB36FFFFD3FF8F4BB92BE2FA5B753D" blockId="6.[151,1436,1742,1861]" pageId="6" pageNumber="40">
<emphasis id="B960EAEDFFD3FF8F4BB92BE2F844749E" bold="true" box="[151,336,1742,1766]" pageId="6" pageNumber="40">FIGURES 614.</emphasis>
<taxonomicName id="4C144D7CFFD3FF8F4A792BE2FB2E749D" box="[343,570,1742,1765]" class="Arachnida" family="Chactidae" genus="Chactopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="40" phylum="Arthropoda" rank="species" species="carolinae" status="sp. nov.">
<emphasis id="B960EAEDFFD3FF8F4A792BE2FB2E749D" box="[343,570,1742,1765]" italics="true" pageId="6" pageNumber="40">Chactopsis carolinae</emphasis>
</taxonomicName>
<emphasis id="B960EAEDFFD3FF8F496F2BE2FB83749E" bold="true" box="[577,663,1742,1766]" pageId="6" pageNumber="40">
<taxonomicNameLabel id="A2535796FFD3FF8F496F2BE2FB83749E" box="[577,663,1742,1766]" pageId="6" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype (IAvH-E 100759).
<emphasis id="B960EAEDFFD3FF8F4F202BE2FD5E749E" bold="true" box="[1038,1098,1742,1766]" pageId="6" pageNumber="40">613.</emphasis>
Distribution of the trichobothria.
<emphasis id="B960EAEDFFD3FF8F4BEF2BC2F9C1757E" bold="true" box="[193,213,1774,1798]" pageId="6" pageNumber="40">6.</emphasis>
Chela, dorsal view.
<emphasis id="B960EAEDFFD3FF8F4A9E2BC2F8D0757E" bold="true" box="[432,452,1774,1798]" pageId="6" pageNumber="40">7.</emphasis>
Chela, external view.
<emphasis id="B960EAEDFFD3FF8F499A2BC2FBDD757E" bold="true" box="[692,713,1774,1798]" pageId="6" pageNumber="40">8.</emphasis>
Chela, ventral view.
<emphasis id="B960EAEDFFD3FF8F48822BC2FAD5757E" bold="true" box="[940,961,1774,1798]" pageId="6" pageNumber="40">9.</emphasis>
Chela, internal view.
<emphasis id="B960EAEDFFD3FF8F4F822BC2FDD9757E" bold="true" box="[1196,1229,1774,1798]" pageId="6" pageNumber="40">10.</emphasis>
Femur, dorsointernal view.
<emphasis id="B960EAEDFFD3FF8F4BD12A22F80A755E" bold="true" box="[255,286,1806,1830]" pageId="6" pageNumber="40">11.</emphasis>
Patella, dorsointernal view.
<emphasis id="B960EAEDFFD3FF8F497C2A22FB67755E" bold="true" box="[594,627,1806,1830]" pageId="6" pageNumber="40">12.</emphasis>
Patella, ventral view.
<emphasis id="B960EAEDFFD3FF8F484D2A22FA90755E" bold="true" box="[867,900,1806,1830]" pageId="6" pageNumber="40">13.</emphasis>
Patella, external view.
<emphasis id="B960EAEDFFD3FF8F4FAE2A22FDB5755E" bold="true" box="[1152,1185,1806,1830]" pageId="6" pageNumber="40">14.</emphasis>
Anterior margin of the carapace (coloration details not included). Scale bars equal 1mm.
</paragraph>
</caption>
<paragraph id="8BAB36FFFFD3FF8F4BE82A5AFABF75E8" blockId="6.[151,1436,1910,2016]" box="[198,939,1910,1936]" pageId="6" pageNumber="40">
Comparisons with the adult female
<typeStatus id="54AF885DFFD3FF8F49462A5AFBD975E8" box="[616,717,1910,1936]" pageId="6" pageNumber="40" type="paratype">paratype</typeStatus>
(IAvH-E
<date id="FFAA103FFFD3FF8F486F2A5AFA8A75E8" box="[833,926,1910,1936]" pageId="6" pageNumber="40" value="60-10-07">100760</date>
).
</paragraph>
<paragraph id="8BAB36FFFFD3FF8E4BE82AB2FA8072A2" blockId="6.[151,1436,1910,2016]" lastBlockId="7.[151,1437,151,698]" lastPageId="7" lastPageNumber="41" pageId="6" pageNumber="40">
The following features differ from the
<typeStatus id="54AF885DFFD3FF8F49882AB2FA0575C0" box="[678,785,1950,1976]" pageId="6" pageNumber="40" type="holotype">holotype</typeStatus>
:
<emphasis id="B960EAEDFFD3FF8F480E2AB2FAD375C0" bold="true" box="[800,967,1950,1976]" italics="true" pageId="6" pageNumber="40">Coloration.—</emphasis>
coxae II, sternites VVI, and pedipalp patella completely yellow; pedipalp femur predominantly yellow, except for two distal brown areas, one dorsal and the other ventral.
<emphasis id="B960EAEDFFD2FF8E4AEB2DBBFBDB72C9" box="[453,719,151,177]" italics="true" pageId="7" pageNumber="41">
<emphasis id="B960EAEDFFD2FF8E4AEB2DBBFB6972C9" bold="true" box="[453,637,151,177]" italics="true" pageId="7" pageNumber="41">Morphology.—</emphasis>
Ven t er:
</emphasis>
pectines with 8:8 teeth (
<emphasis id="B960EAEDFFD2FF8E48D62DB4FD2572C9" box="[1016,1073,152,177]" italics="true" pageId="7" pageNumber="41">idem</emphasis>
in the juvenile female).
<emphasis id="B960EAEDFFD2FF8E4E782DB4F9CB72A1" italics="true" pageId="7" pageNumber="41">Metasoma:</emphasis>
segment I bears vestigial, slightly elevated ventral carinae.
</paragraph>
</subSubSection>
<subSubSection id="C30E6574FFD2FF8E4BE82DCBFC8870C2" pageId="7" pageNumber="41" type="distribution">
<paragraph id="8BAB36FFFFD2FF8E4BE82DCBFA4A7302" blockId="7.[151,1437,151,698]" pageId="7" pageNumber="41">
<emphasis id="B960EAEDFFD2FF8E4BE82DCBFB517379" bold="true" box="[198,581,231,257]" pageId="7" pageNumber="41">Distribution and landscape.—</emphasis>
This species is known only from the
<typeStatus id="54AF885DFFD2FF8E4F3C2DC4FD50737A" box="[1042,1092,232,258]" pageId="7" pageNumber="41">type</typeStatus>
locality: Selva de Matavén, Cumaribo, Vichada department (
<figureCitation id="132F2A7AFFD2FF8E49372C3CFB4B7352" box="[537,607,272,298]" captionStart="FIGURE 1" captionStartId="1.[151,255,2009,2033]" captionTargetBox="[231,1364,466,1977]" captionTargetId="figure@1.[231,1364,466,1977]" captionTargetPageId="1" captionText="FIGURE 1. Known distribution of Chactopsis carolinae sp. nov." httpUri="https://zenodo.org/record/181582/files/figure.png" pageId="7" pageNumber="41">Fig. 1</figureCitation>
). This territory is conformed of alluvial valleys and rocky high plains (
<bibRefCitation id="EF854B0EFFD2FF8E4BB12C14F83B732A" author="IGAC" box="[159,303,312,338]" pageId="7" pageNumber="41" refString="IGAC (1996) Diccionario Geografico de Colombia. Third edition. Instituto Geografico Agustin Codazzi, Colombia, pp. 621 - 1244." type="book chapter" year="1996">IGAC 1996</bibRefCitation>
). The area presents semiarid zones, original forests, savannas both natural and produced through human activities, and relicts of felled forests (
<bibRefCitation id="EF854B0EFFD2FF8E49802C4CFA457302" author="Molano" box="[686,849,352,378]" pageId="7" pageNumber="42" refString="Molano, J. (1998) Biogeografia de la Orinoquia Colombiana. In: Dominguez, C. (Ed.). Colombia Orinoco. Fondo para la Proteccion del Medio Ambiente &quot; Jose Celestino Mutis &quot;, FEN, Colombia, pp. 69 - 101." type="book chapter" year="1998">Molano 1998</bibRefCitation>
).
</paragraph>
<paragraph id="8BAB36FFFFD2FF8E4BE82CA4FC8870C2" blockId="7.[151,1437,151,698]" pageId="7" pageNumber="41">
Even though the known distribution of
<taxonomicName id="4C144D7CFFD2FF8E49BD2CA4FA2773D9" box="[659,819,392,417]" class="Arachnida" family="Chactidae" genus="Chactopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="carolinae" status="sp. nov.">
<emphasis id="B960EAEDFFD2FF8E49BD2CA4FA2773D9" box="[659,819,392,417]" italics="true" pageId="7" pageNumber="41">Ch. carolinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2535796FFD2FF8E48142CA4FA8173DA" box="[826,917,392,418]" pageId="7" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
is relatively close to the
<typeStatus id="54AF885DFFD2FF8E4F972CA4FDF873DA" box="[1209,1260,392,418]" pageId="7" pageNumber="41">type</typeStatus>
locality of
<taxonomicName id="4C144D7CFFD2FF8E4E5C2CA4F9E673B1" class="Arachnida" family="Chactidae" genus="Chactopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="41" phylum="Arthropoda" rank="species" species="anduzei">
<emphasis id="B960EAEDFFD2FF8E4E5C2CA4F9E673B1" italics="true" pageId="7" pageNumber="41">Ch. anduzei</emphasis>
</taxonomicName>
(Caño Pava, between El Danto and Gavilán, Atures department,
<collectingCountry id="F303766FFFD2FF8E48D22C9CFD6E73B2" box="[1020,1146,432,458]" name="Venezuela" pageId="7" pageNumber="41">Venezuela</collectingCountry>
), the former inhabits the Negro-Branco Moist Forests ecoregion in
<collectingCountry id="F303766FFFD2FF8E49A82CF4FBE9738A" box="[646,765,472,498]" name="Colombia" pageId="7" pageNumber="41">Colombia</collectingCountry>
, whereas the second occurs in Venezuela—at least on the basis of González-Spongas (1996: 112) map—in the Guayanan Highlands Moist Forests (National Geographic Society 2001). The Negro-Branco Moist Forests ecoregion occurs along the Negro River basin, along the southwestern edge of the Guayana Shield in eastern
<collectingCountry id="F303766FFFD2FF8E48042F7CFAB57012" box="[810,929,592,618]" name="Colombia" pageId="7" pageNumber="41">Colombia</collectingCountry>
, northwestern
<collectingCountry id="F303766FFFD2FF8E4F602F7CFD8C7012" box="[1102,1176,592,618]" name="Brazil" pageId="7" pageNumber="41">Brazil</collectingCountry>
, and most of southern
<collectingCountry id="F303766FFFD2FF8E4BB92F54F80770EA" box="[151,275,632,658]" name="Venezuela" pageId="7" pageNumber="41">Venezuela</collectingCountry>
. It is strongly influenced by numerous rivers and streams that form the Negro River and Branco River watersheds, and is known to be a region of high species endemism (National Geographic Society 2001).
</paragraph>
</subSubSection>
</treatment>
</document>