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<document id="510B0107E5D4CD4078ED0998D3A41FC9" ID-DOI="10.11646/zootaxa.3894.1.4" ID-GBIF-Dataset="1bff28d0-db79-4a0f-8cfa-fc6cd82ce2ae" ID-ISSN="1175-5326" ID-Zenodo-Dep="251800" ID-ZooBank="70B9B045-9C14-47B1-B0C7-8DDA09AFE02B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460996815897" checkinUser="plazi" docAuthor="Marusik, Yuri M. &amp; Omelko, Mikhail M." docDate="2014" docId="03BD87E1FFA8DB7CF2C03F281DCCFA17" docLanguage="en" docName="zt03894p032.pdf" docOrigin="Zootaxa 3894 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gnaphosa sticta Kulczynski 1908" docType="treatment" docVersion="12" masterDocId="FF84FF99FFAEDB7BF25739751F7BFF99" masterDocTitle="A survey of East Palaearctic Gnaphosidae (Araneae). 3. On new and poorly known Gnaphosa Latreille, 1804" updateTime="1698562972859" updateUser="plazi">
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<mods:title id="64D282690B0741AAB61F36BB06F0EB39">A survey of East Palaearctic Gnaphosidae (Araneae). 3. On new and poorly known Gnaphosa Latreille, 1804</mods:title>
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<mods:namePart id="77E651A76FE05F25BCDEA9C186D68E2F">Marusik, Yuri M.</mods:namePart>
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<mods:date id="3FA0A14603ED1C1BE9E14771A70E2383">2014</mods:date>
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<treatment id="03BD87E1FFA8DB7CF2C03F281DCCFA17" ID-DOI="http://doi.org/10.5281/zenodo.5612616" ID-GBIF-Taxon="119597507" ID-Zenodo-Dep="5612616" LSID="urn:lsid:plazi:treatment:03BD87E1FFA8DB7CF2C03F281DCCFA17" httpUri="http://treatment.plazi.org/id/03BD87E1FFA8DB7CF2C03F281DCCFA17" lastPageId="7" pageId="6">
<subSubSection id="C30E657CFFA8DB7DF2C03F281E0BF900" pageId="6" type="nomenclature">
<paragraph id="8BAB36F7FFA8DB7DF2C03F281D4DF9E1" blockId="6.[151,566,1629,1689]" box="[151,566,1629,1656]" pageId="6">
<heading id="D0E3819BFFA8DB7DF2C03F281D4DF9E1" bold="true" box="[151,566,1629,1656]" fontSize="11" level="1" pageId="6" reason="1">
<taxonomicName id="4C144D74FFA8DB7DF2C03F281D4DF9E1" ID-CoL="3GL5Z" authority="Kulczynski, 1908" authorityName="Kulczynski" authorityYear="1908" box="[151,566,1629,1656]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="6" phylum="Arthropoda" rank="species" species="sticta">
<emphasis id="B960EAE5FFA8DB7DF2C03F281D4DF9E1" bold="true" box="[151,566,1629,1656]" pageId="6">
<emphasis id="B960EAE5FFA8DB7DF2C03F281E22F9EE" bold="true" box="[151,345,1629,1655]" italics="true" pageId="6">Gnaphosa sticta</emphasis>
Kulczyński, 1908
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BAB36F7FFA8DB7DF2C03FF51E0BF900" blockId="6.[151,566,1629,1689]" box="[151,368,1664,1689]" pageId="6">
<figureCitation id="132F2A72FFA8DB7DF2C03FF51E6DF900" box="[151,278,1664,1689]" captionStart="FIGURES 7 15" captionStartId="3.[151,264,1913,1936]" captionTargetBox="[150,1434,471,1889]" captionTargetId="figure@3.[149,1434,471,1889]" captionTargetPageId="3" captionText="FIGURES 7 15. Epigynes of Gnaphosa koponeni sp. n. (7 9), G. wiehlei (10 12) and G. sticta (13 15, from Finland). 7, 10, 13 intact, ventral; 8, 11, 14 macerated, ventral; 9, 12, 15 macerated, dorsal. Scale = 0.2 mm. Abbreviations: Fb— bulge of fovea, Id— insemination duct, Lm— lateral margin, Mp— pocket of lateral margin, Pf— posterior furrow, Ps— pocket of scape, Re— receptacle, Rg— gland of receptacle." httpUri="https://zenodo.org/record/251802/files/figure.png" pageId="6">Figs 1315</figureCitation>
,
<figureCitation id="132F2A72FFA8DB7DF3753FF41E10F900" box="[290,363,1665,1689]" captionStart="FIGURES 27 30" captionStartId="7.[151,264,1052,1075]" captionTargetBox="[181,1436,200,1028]" captionTargetId="figure@7.[151,1436,195,1029]" captionTargetPageId="7" captionText="FIGURES 27 30. Copulatory organs of Gnaphosa sticta. 27 28 male palp, ventral and retrolateral, Finland; 29 30 epigyne, ventral and dorsal, Anadyr. Scale = 0.2 mm. Abbreviation: Ma— median apophysis." httpUri="https://zenodo.org/record/251805/files/figure.png" pageId="6">2730</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657CFFA8DB7DF2C03FBD1C3AF885" pageId="6" type="reference_group">
<paragraph id="8BAB36F7FFA8DB7DF2C03FBD1D2CF964" blockId="6.[151,1436,1736,1820]" pageId="6">
<treatmentCitationGroup id="AB0411D9FFA8DB7DF2C03FBD1D2CF964" pageId="6">
<taxonomicName id="4C144D74FFA8DB7DF2C03FBD1DC0F947" ID-CoL="3GL5Z" authority="Ovtsharenko &amp; Marusik 1988: 208" authorityName="Ovtsharenko &amp; Marusik" authorityPageNumber="208" authorityYear="1988" box="[151,699,1736,1758]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="6" phylum="Arthropoda" rank="species" species="sticta">
<emphasis id="B960EAE5FFA8DB7DF2C03FBD1E45F947" box="[151,318,1736,1758]" italics="true" pageId="6">Gnaphosa sticta</emphasis>
:
<treatmentCitation id="0AB510E6FFA8DB7DF31A3FBD1DC0F947" author="Ovtsharenko" box="[333,699,1736,1758]" page="208" pageId="6" year="1988">
<bibRefCitation id="EF854B06FFA8DB7DF31A3FBD1DFDF947" author="Ovtsharenko" box="[333,646,1736,1758]" pageId="6" refString="Ovtsharenko, V. I. &amp; Marusik, Y. M. (1988) Spiders of the family Gnaphosidae (Aranei) of the North-East of the USSR (Magadan Area). Entomologicheskoe Obozrenie, 61 (4), 204 - 217. [In Russian]" type="journal article" year="1988">Ovtsharenko &amp; Marusik 1988</bibRefCitation>
: 208
</treatmentCitation>
</taxonomicName>
, figs 1617, 20 (♂♀);
<treatmentCitation id="0AB510E6FFA8DB7DF1FC3FBD1BB4F947" author="Ovtsharenko" box="[939,1231,1736,1758]" page="48" pageId="6" year="1992">
<bibRefCitation id="EF854B06FFA8DB7DF1FC3FBD1BD3F947" author="Ovtsharenko" box="[939,1192,1736,1758]" pageId="6" refString="Ovtsharenko, V. I., Platnick, N. I. &amp; Song, D. X. (1992) A review of the North Asian ground spiders of the genus Gnaphosa (Araneae, Gnaphosidae). Bulletin of the American Museum of Natural History, 212, 1 - 88." type="book chapter" year="1992">Ovtsharenko et al. 1992</bibRefCitation>
: 48
</treatmentCitation>
, figs 163168 (♂♀);
<treatmentCitation id="0AB510E6FFA8DB7DF2903F921EEFF964" author="Almquist" box="[199,404,1767,1789]" page="404" pageId="6" year="2006">
<bibRefCitation id="EF854B06FFA8DB7DF2903F921E19F964" author="Almquist" box="[199,354,1767,1789]" pageId="6" refString="Almquist, S. (2006) Swedish Araneae, part 2 - families Dictynidae to Salticidae. Insect Systematics and Evolution Supplements, 63, 285 - 601." type="journal article" year="2006">Almquist 2006</bibRefCitation>
: 404
</treatmentCitation>
, figs 350ae (♂♀).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAB36F7FFA8DB7DF2C03E721C3AF885" blockId="6.[151,1436,1736,1820]" box="[151,833,1798,1820]" pageId="6">
For a complete synonym listing and references see
<bibRefCitation id="EF854B06FFA8DB7DF0CA3E731C46F885" author="Platnick" box="[669,829,1798,1820]" pageId="6" refString="Platnick, N. I. (2014) The World Spider Catalog, Version 14.5 American Museum of Natural History. Available from: http: // research. amnh. org / entomology / spiders / catalog / index. html (Accessed 28 May 2014)" type="journal article" year="2014">Platnick (2014)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657CFFA8DB7DF2C03E3D1DF0F830" pageId="6" type="materials_examined">
<paragraph id="8BAB36F7FFA8DB7DF2C03E3D1DF0F830" blockId="6.[151,1437,1864,1997]" pageId="6">
<emphasis id="B960EAE5FFA8DB7DF2C03E3D1E07F8F8" bold="true" box="[151,380,1864,1889]" pageId="6">Material examined</emphasis>
.
<collectingCountry id="F3037667FFA8DB7DF3D93E3C1D76F8F8" box="[398,525,1865,1889]" name="Finland" pageId="6">FINLAND</collectingCountry>
:
<specimenCount id="9D12FD7EFFA8DB7DF04C3E3C1D44F8F8" box="[539,575,1865,1889]" pageId="6" type="generic">1♂</specimenCount>
, 2♀ (
<collectionCode id="ED05AE32FFA8DB7DF0D33E3C1DA2F8F8" box="[644,729,1865,1889]" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="6">ZMUT</collectionCode>
), Utsjoki, Kevo, indoors, 5.09.1973-6.06.1974 (S. Koponen).
<collectingCountry id="F3037667FFA8DB7DF2C03E181F84F81C" box="[151,255,1901,1925]" name="Russia" pageId="6">RUSSIA</collectingCountry>
:
<emphasis id="B960EAE5FFA8DB7DF35D3E191E07F81C" bold="true" box="[266,380,1900,1925]" italics="true" pageId="6">Chukotka</emphasis>
: 1♀ (
<collectionCode id="ED05AE32FFA8DB7DF3E83E181D67F81C" box="[447,540,1901,1925]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="6">ZMMU</collectionCode>
),
<quantity id="4CEC9B12FFA8DB7DF0673E181D02F81C" box="[560,633,1900,1925]" metricMagnitude="4" metricUnit="m" metricValue="7.4" pageId="6" unit="km" value="74.0">74 km</quantity>
W of Anadyr City, Anadyr River near Omochi River mouth,
<quantity id="4CEC9B12FFA8DB7DF71A3E191AECF81D" box="[1357,1431,1900,1925]" metricMagnitude="2" metricUnit="m" metricValue="6.13" pageId="6" unit="m" value="613.0">613 m</quantity>
,
<geoCoordinate id="EE205030FFA8DB7DF2C03EE51F91F830" box="[151,234,1936,1961]" direction="north" orientation="latitude" pageId="6" precision="555" value="64.82">64.82N</geoCoordinate>
<geoCoordinate id="EE205030FFA8DB7DF2A63EE51E1BF830" box="[241,352,1936,1961]" direction="east" orientation="longitude" pageId="6" precision="55" value="175.967">175.967E</geoCoordinate>
, 7-8.2013 (O. Khrulyova).
</paragraph>
</subSubSection>
<subSubSection id="C30E657CFFA8DB7DF2903EC11CD2F854" box="[199,937,1972,1997]" pageId="6" type="description">
<paragraph id="8BAB36F7FFA8DB7DF2903EC11CD2F854" blockId="6.[151,1437,1864,1997]" box="[199,937,1972,1997]" pageId="6">
<emphasis id="B960EAE5FFA8DB7DF2903EC11E2AF854" bold="true" box="[199,337,1972,1997]" pageId="6">Description</emphasis>
. The species has been described by
<bibRefCitation id="EF854B06FFA8DB7DF0B03EC11CDFF854" author="Almquist" box="[743,932,1972,1997]" pageId="6" refString="Almquist, S. (2006) Swedish Araneae, part 2 - families Dictynidae to Salticidae. Insect Systematics and Evolution Supplements, 63, 285 - 601." type="journal article" year="2006">Almquist (2006)</bibRefCitation>
.
</paragraph>
</subSubSection>
<caption id="DF6B667FFFA9DB7CF2C03D691C94FBCB" httpUri="https://zenodo.org/record/251805/files/figure.png" pageId="7" targetBox="[181,1436,200,1028]" targetPageId="7">
<paragraph id="8BAB36F7FFA9DB7CF2C03D691C94FBCB" blockId="7.[151,1436,1052,1106]" pageId="7">
<emphasis id="B960EAE5FFA9DB7CF2C03D691E2EFBAB" bold="true" box="[151,341,1052,1075]" pageId="7">FIGURES 2730.</emphasis>
Copulatory organs of
<taxonomicName id="4C144D74FFA9DB7CF06A3D681D9FFBAA" box="[573,740,1053,1075]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="7" phylum="Arthropoda" rank="species" species="sticta">
<emphasis id="B960EAE5FFA9DB7CF06A3D681D9FFBAA" box="[573,740,1053,1075]" italics="true" pageId="7">Gnaphosa sticta</emphasis>
</taxonomicName>
. 2728 male palp, ventral and retrolateral, Finland; 2930 epigyne, ventral and dorsal, Anadyr. Scale = 0.2 mm. Abbreviation:
<emphasis id="B960EAE5FFA9DB7CF0AE3D481C4AFBCB" box="[761,817,1085,1106]" italics="true" pageId="7">Ma—</emphasis>
median apophysis.
</paragraph>
</caption>
<subSubSection id="C30E657CFFA9DB7CF2903D0C1E36FAB8" pageId="7" type="discussion">
<paragraph id="8BAB36F7FFA9DB7CF2903D0C1E36FAB8" blockId="7.[151,1436,1145,1422]" pageId="7">
<emphasis id="B960EAE5FFA9DB7CF2903D0C1E33FB0B" bold="true" box="[199,328,1145,1170]" pageId="7">Comments</emphasis>
. Here we provide figures of this species to show the similarity to females of
<taxonomicName id="4C144D74FFA9DB7CF69B3D0F1A2FFB0B" box="[1228,1364,1145,1170]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="7" phylum="Arthropoda" rank="species" species="koponeni" status="sp. nov.">
<emphasis id="B960EAE5FFA9DB7CF69B3D0F1A2FFB0B" box="[1228,1364,1145,1170]" italics="true" pageId="7">G. koponeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253579EFFA9DB7CF70A3D0F1AE7FB08" box="[1373,1436,1146,1169]" pageId="7" rank="species">sp. n.</taxonomicNameLabel>
Figures of epigynes provided by
<bibRefCitation id="EF854B06FFA9DB7CF0453DE81C42FB2F" box="[530,825,1181,1206]" pageId="7" refString="Ovtsharenko, V. I., Platnick, N. I. &amp; Song, D. X. (1992) A review of the North Asian ground spiders of the genus Gnaphosa (Araneae, Gnaphosidae). Bulletin of the American Museum of Natural History, 212, 1 - 88." type="book chapter">Ovtsharenko et al. (1992)</bibRefCitation>
,
<bibRefCitation id="EF854B06FFA9DB7CF1103DE81B78FB2F" author="Almquist" box="[839,1027,1181,1206]" pageId="7" refString="Almquist, S. (2006) Swedish Araneae, part 2 - families Dictynidae to Salticidae. Insect Systematics and Evolution Supplements, 63, 285 - 601." type="journal article" year="2006">Almquist (2006)</bibRefCitation>
and this study (
<figureCitation id="132F2A72FFA9DB7CF6EB3DE81A44FB2F" box="[1212,1343,1181,1206]" captionStart="FIGURES 7 15" captionStartId="3.[151,264,1913,1936]" captionTargetBox="[150,1434,471,1889]" captionTargetId="figure@3.[149,1434,471,1889]" captionTargetPageId="3" captionText="FIGURES 7 15. Epigynes of Gnaphosa koponeni sp. n. (7 9), G. wiehlei (10 12) and G. sticta (13 15, from Finland). 7, 10, 13 intact, ventral; 8, 11, 14 macerated, ventral; 9, 12, 15 macerated, dorsal. Scale = 0.2 mm. Abbreviations: Fb— bulge of fovea, Id— insemination duct, Lm— lateral margin, Mp— pocket of lateral margin, Pf— posterior furrow, Ps— pocket of scape, Re— receptacle, Rg— gland of receptacle." httpUri="https://zenodo.org/record/251802/files/figure.png" pageId="7">Figs 1315</figureCitation>
,
<figureCitation id="132F2A72FFA9DB7CF71A3DEB1AEFFB2F" box="[1357,1428,1182,1206]" captionStart="FIGURES 27 30" captionStartId="7.[151,264,1052,1075]" captionTargetBox="[181,1436,200,1028]" captionTargetId="figure@7.[151,1436,195,1029]" captionTargetPageId="7" captionText="FIGURES 27 30. Copulatory organs of Gnaphosa sticta. 27 28 male palp, ventral and retrolateral, Finland; 29 30 epigyne, ventral and dorsal, Anadyr. Scale = 0.2 mm. Abbreviation: Ma— median apophysis." httpUri="https://zenodo.org/record/251805/files/figure.png" pageId="7">2930</figureCitation>
) differ slightly in shape of lateral pockets, size and direction of glands and shape of receptacles, but all illustrated specimens have the same shaped insemination (copulatory) ducts (
<emphasis id="B960EAE5FFA9DB7CF1D33D901CE0FB67" box="[900,923,1253,1278]" italics="true" pageId="7">Id</emphasis>
). Males of
<taxonomicName id="4C144D74FFA9DB7CF6493D931B01FB64" box="[1054,1146,1254,1277]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="7" phylum="Arthropoda" rank="species" species="sticta">
<emphasis id="B960EAE5FFA9DB7CF6493D931B01FB64" box="[1054,1146,1254,1277]" italics="true" pageId="7">G. sticta</emphasis>
</taxonomicName>
have a very large median apophysis (
<emphasis id="B960EAE5FFA9DB7CF3403C7F1E47FAB8" box="[279,316,1290,1313]" italics="true" pageId="7">Ma</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C30E657CFFA9DB7CF2903C581DCCFA17" pageId="7" type="distribution">
<paragraph id="8BAB36F7FFA9DB7CF2903C581DCCFA17" blockId="7.[151,1436,1145,1422]" pageId="7">
<emphasis id="B960EAE5FFA9DB7CF2903C581E22FADF" bold="true" box="[199,345,1325,1350]" pageId="7">Distribution</emphasis>
. The present records from Chukotka coincide with the previous easternmost known locality of the species, which was in the upper reaches of Bolshaya Osinovaya River. This species is known from
<collectingCountry id="F3037667FFA9DB7CF7733C271A05FAF3" box="[1316,1406,1362,1386]" name="Norway" pageId="7">Norway</collectingCountry>
to East Chukotka and south to Tuva and Hokkaido.
</paragraph>
</subSubSection>
</treatment>
</document>