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<document id="F6AFDB479108C2E2232AB7C22CC3E578" ID-DOI="10.5281/zenodo.199529" ID-GBIF-Dataset="1652c47d-63ad-42b7-b1ac-3c4b0cef9fd2" ID-ISSN="1175-5326" ID-Zenodo-Dep="199529" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460339251317" checkinUser="plazi" docAuthor="Stiassny, Melanie L. J., Marchi, Giuseppe De &amp; Lamboj, Anton" docDate="2010" docId="03BD87B7FFE7222DE1E6F8B3BDA937E2" docLanguage="en" docName="zt02690p052.pdf" docOrigin="Zootaxa 2690" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Danakilia dinicolai Stiassny, Marchi &amp; Lamboj, 2010, new species" docType="treatment" docVersion="8" lastPageNumber="51" masterDocId="FF84FFCFFFE52225E171FFFEB8243172" masterDocTitle="A new species of Danakilia (Teleostei, Cichlidae) from Lake Abaeded in the Danakil Depression of Eritrea (East Africa)" masterLastPageNumber="52" masterPageNumber="43" pageNumber="45" updateTime="1698260734143" updateUser="plazi">
<mods:mods id="86DD25FDEF0B1DFA6EC3E859C3B81F6E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B1EF3B709B6E43A117261F29049D8548">
<mods:title id="5002D92FD844349F7E68C8D1B90482D3">A new species of Danakilia (Teleostei, Cichlidae) from Lake Abaeded in the Danakil Depression of Eritrea (East Africa)</mods:title>
</mods:titleInfo>
<mods:name id="7C4CB318F246DD03D358E2A0406A1180" type="personal">
<mods:role id="CB25E94E0641D36D56AADF04361FB855">
<mods:roleTerm id="2D997FC7B911BACD91A2321FD49F832B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="43B36B8F19D1B5E8F6E909A3DEB5B4C1">Stiassny, Melanie L. J.</mods:namePart>
</mods:name>
<mods:name id="27C879F7712E264303982768D5E62560" type="personal">
<mods:role id="EE60CE667401C264FBA2927AECF2F616">
<mods:roleTerm id="2FA7F37E2A9DE97CCA1ED686747E7ED0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F06484B6E17BFCE66333D5D6771E23C7">Marchi, Giuseppe De</mods:namePart>
</mods:name>
<mods:name id="29ED45BEA32F721C428EC65CD9590D0B" type="personal">
<mods:role id="019A15390ECF41F746B0CE6C996E912F">
<mods:roleTerm id="799F8899D9305474367BDA28B9E08254">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0667273EEAB68A49B3CBAEC5F911731A">Lamboj, Anton</mods:namePart>
</mods:name>
<mods:typeOfResource id="FCC22B22D6810FFE3B03F51BB77D772B">text</mods:typeOfResource>
<mods:relatedItem id="3E82659C5B47B85514D5F1375560D97C" type="host">
<mods:titleInfo id="E46B671FB9253645CCB9E0E5DFC6C06B">
<mods:title id="E58150F0EB2330C8D3CE87531C58BE4B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E51BC8403E4B5530751F4053E26EDB1D">
<mods:date id="62495A1E538DBEE5258911E13731BA77">2010</mods:date>
<mods:detail id="7C74402A54BF45A986B696E5D1D81EDF" type="volume">
<mods:number id="6245DE14088085C229EB4A5F843E320E">2690</mods:number>
</mods:detail>
<mods:extent id="493C80357BD5BFB43CA3AE728372A7F9" unit="page">
<mods:start id="7D7BDBD52BB1FF69CE330701B2F1087D">43</mods:start>
<mods:end id="B2BC91FBF3CF54E24AE6E3CAE4AE2A13">52</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="406E7DC48B8C4AE31E608237771A2DF8">journal article</mods:classification>
<mods:identifier id="5C6DE7A804AA322A2C26235AD9C51C8F" type="DOI">10.5281/zenodo.199529</mods:identifier>
<mods:identifier id="B8D9CF34931E4A20F2A3CEB7485D970A" type="GBIF-Dataset">1652c47d-63ad-42b7-b1ac-3c4b0cef9fd2</mods:identifier>
<mods:identifier id="22F95D09E0664A23BA0E9EE07B1BF187" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E238373D4FF750C625EFA10922D66F40" type="Zenodo-Dep">199529</mods:identifier>
</mods:mods>
<treatment id="03BD87B7FFE7222DE1E6F8B3BDA937E2" ID-DOI="http://doi.org/10.5281/zenodo.5624407" ID-GBIF-Taxon="119406035" ID-Zenodo-Dep="5624407" LSID="urn:lsid:plazi:treatment:03BD87B7FFE7222DE1E6F8B3BDA937E2" httpUri="http://treatment.plazi.org/id/03BD87B7FFE7222DE1E6F8B3BDA937E2" lastPageId="8" lastPageNumber="51" pageId="2" pageNumber="45">
<subSubSection id="C30E652AFFE72227E1E6F8B3B94E36F8" pageId="2" pageNumber="45" type="nomenclature">
<paragraph id="8BAB36A1FFE72227E1E6F8B3BA3B3615" blockId="2.[151,543,1869,1930]" box="[151,543,1869,1895]" pageId="2" pageNumber="45">
<heading id="D0E381CDFFE72227E1E6F8B3BA3B3615" bold="true" box="[151,543,1869,1895]" fontSize="11" level="1" pageId="2" pageNumber="45" reason="1">
<emphasis id="B960EAB3FFE72227E1E6F8B3BA3B3615" bold="true" box="[151,543,1869,1895]" pageId="2" pageNumber="45">
<taxonomicName id="4C144D22FFE72227E1E6F8B3B9A53615" ID-CoL="343RR" box="[151,385,1869,1895]" class="Actinopterygii" family="Cichlidae" genus="Danakilia" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="45" phylum="Chordata" rank="species" species="dinicolai" status="sp. nov.">
<emphasis id="B960EAB3FFE72227E1E6F8B3B9A53615" bold="true" box="[151,385,1869,1895]" italics="true" pageId="2" pageNumber="45">Danakilia dinicolai</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25357C8FFE72227E0FEF8B3BA3B3615" box="[399,543,1869,1895]" pageId="2" pageNumber="45" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAB36A1FFE72227E1E6F88EB94E36F8" blockId="2.[151,543,1869,1930]" box="[151,362,1904,1930]" pageId="2" pageNumber="45">
<figureCitation id="132F2A24FFE72227E1E6F88EB92C36F8" box="[151,264,1904,1930]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="3.[151,255,1398,1422]" captionStartId-1="5.[151,255,1914,1938]" captionStartId-2="6.[151,255,1486,1510]" captionTargetBox-0="[162,1432,883,1354]" captionTargetBox-1="[248,1362,206,1893]" captionTargetBox-2="[172,1414,216,1443]" captionTargetId-0="figure@3.[151,1436,850,1375]" captionTargetId-1="figure@5.[219,1378,194,1904]" captionTargetId-2="figure@6.[151,1436,194,1465]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIGURE 2. Danakilia dinicolai, holotype, MSNM 4888, male, 96.4 mm SL, Lake Abaeded, Eritrea." captionText-1="FIGURE 3. Lower pharyngeal jaw (dorsal, posterior and lateral views), left mandible and isolated tooth in (a) Danakilia dinicolai, (b) Danakilia franchettii, (c) Iranocichla hormuzensis. Elements drawn in corresponding scale for each species. Posterior neurocranium and anterior vertebral elements, arrows indicate inferior vertebral apophysis of (d) Iranocichla hormuzensis." captionText-2="FIGURE 4. Danakilia dinicolai (a) wild caught female and (b) wild caught male. Specimens not preserved or measured." httpUri-0="https://zenodo.org/record/199531/files/figure.png" httpUri-1="https://zenodo.org/record/199532/files/figure.png" httpUri-2="https://zenodo.org/record/199533/files/figure.png" pageId="2" pageNumber="45">Figs. 2–4</figureCitation>
;
<tableCitation id="C696031AFFE72227E062F88EB94E36F8" box="[275,362,1904,1930]" captionStart="TABLE 2" captionStartId="4.[151,239,151,175]" captionTargetBox="[159,1313,330,951]" captionTargetPageId="4" captionText="TABLE 2. Morphometric and meristic data for the holotype and 13 paratypes of Danakilia dinicolai." httpUri="http://table.plazi.org/id/DF6B6629FFE12221E1E6FF69BCEA31DD" pageId="2" pageNumber="45" tableUuid="DF6B6629FFE12221E1E6FF69BCEA31DD">Table 2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30E652AFFE72226E1E6F84BB8FD301E" lastPageId="3" lastPageNumber="46" pageId="2" pageNumber="45" type="materials_examined">
<paragraph id="8BAB36A1FFE72227E1E6F84BB9BB3686" blockId="2.[151,1436,1973,2036]" pageId="2" pageNumber="45">
<typeStatus id="54AF8803FFE72227E1E6F84BB92936BD" box="[151,269,1973,1999]" pageId="2" pageNumber="45" type="holotype">
<emphasis id="B960EAB3FFE72227E1E6F84BB92936BD" bold="true" box="[151,269,1973,1999]" pageId="2" pageNumber="45">Holotype</emphasis>
</typeStatus>
.
<collectionCode id="ED05AE64FFE72227E06CF84BB95C36BD" box="[285,376,1973,1999]" pageId="2" pageNumber="45">MSNM</collectionCode>
4888, male,
<quantity id="4CEC9B44FFE72227E368F84BBAA136BD" box="[537,645,1973,1999]" metricMagnitude="-2" metricUnit="m" metricValue="9.64" pageId="2" pageNumber="45" unit="mm" value="96.4">96.4 mm</quantity>
SL,
<collectingCountry id="F3037631FFE72227E3B0F84BBB3D36BD" box="[705,793,1973,1999]" name="Eritrea" pageId="2" pageNumber="45">Eritrea</collectingCountry>
: Lake Abaeded. 14˚ 35' 17&quot; N, 40˚ 05' 04&quot; E., G.
<collectingCountry id="F3037631FFE72227E408F84BBDB836BD" box="[1401,1436,1973,1999]" name="South Georgia and the South Sandwich Islands" pageId="2" pageNumber="45">De</collectingCountry>
Marchi,
<date id="FFAA1061FFE72227E18BF824B9BE3686" box="[250,410,2010,2036]" pageId="2" pageNumber="45" value="1998-01">January 1998</date>
.
</paragraph>
<paragraph id="8BAB36A1FFE62226E1B4FF69B8FD301E" blockId="3.[151,1437,151,812]" pageId="3" pageNumber="46">
<emphasis id="B960EAB3FFE62226E1B4FF69B96E31C3" bold="true" box="[197,330,151,177]" pageId="3" pageNumber="46">
<typeStatus id="54AF8803FFE62226E1B4FF69B96131C3" box="[197,325,151,177]" pageId="3" pageNumber="46" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED05AE64FFE62226E020FF66B98F31C0" box="[337,427,152,178]" pageId="3" pageNumber="46">MSNM</collectionCode>
4889, male,
<quantity id="4CEC9B44FFE62226E335FF66BA8931C0" box="[580,685,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="7.29" pageId="3" pageNumber="46" unit="mm" value="72.9">72.9 mm</quantity>
SL, same data as
<typeStatus id="54AF8803FFE62226E2F2FF66BBC931C0" box="[899,1005,152,178]" pageId="3" pageNumber="46" type="holotype">holotype</typeStatus>
.
<collectionCode id="ED05AE64FFE62226E289FF66BC7531C0" box="[1016,1105,152,178]" pageId="3" pageNumber="46">MSNM</collectionCode>
4890, female,
<quantity id="4CEC9B44FFE62226E473FF66BD4F31C0" box="[1282,1387,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="6.39" pageId="3" pageNumber="46" unit="mm" value="63.9">63.9 mm</quantity>
SL, same data as
<typeStatus id="54AF8803FFE62226E048FF43B98331A5" box="[313,423,189,215]" pageId="3" pageNumber="46" type="holotype">holotype</typeStatus>
.
<collectionCode id="ED05AE64FFE62226E0C2FF43BA2A31A5" box="[435,526,189,215]" pageId="3" pageNumber="46">MSNM</collectionCode>
4891, male,
<quantity id="4CEC9B44FFE62226E3DFFF43BB3E31A5" box="[686,794,189,215]" metricMagnitude="-2" metricUnit="m" metricValue="5.67" pageId="3" pageNumber="46" unit="mm" value="56.7">56.7 mm</quantity>
SL, same data as
<typeStatus id="54AF8803FFE62226E288FF43BC4331A5" box="[1017,1127,189,215]" pageId="3" pageNumber="46" type="holotype">holotype</typeStatus>
.
<collectionCode id="ED05AE64FFE62226E502FF43BCEA31A5" LSID="urn:lsid:biocol.org:col:34925" box="[1139,1230,189,215]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="46">AMNH</collectionCode>
248881, 5 ex.,
<specimenCount id="9D12FD28FFE62226E4FCFF43B8FA318E" pageId="3" pageNumber="46" type="male">3 males</specimenCount>
,
<specimenCount id="9D12FD28FFE62226E198FF1CB978318E" box="[233,348,226,252]" pageId="3" pageNumber="46" type="female">2 females</specimenCount>
,
<quantity id="4CEC9B44FFE62226E019FF1CBA36318E" box="[360,530,226,252]" metricMagnitude="-2" metricUnit="m" metricValue="7.64" metricValueMax="8.29" metricValueMin="6.99" pageId="3" pageNumber="46" unit="mm" value="76.4" valueMax="82.9" valueMin="69.9">69.9–82.9 mm</quantity>
SL,
<collectingCountry id="F3037631FFE62226E339FF1CBAB8318E" box="[584,668,226,252]" name="Eritrea" pageId="3" pageNumber="46">Eritrea</collectingCountry>
: Lake Abaeded, G.
<collectingCountry id="F3037631FFE62226E20FFF1CBB84318E" box="[894,928,226,252]" name="South Georgia and the South Sandwich Islands" pageId="3" pageNumber="46">De</collectingCountry>
Marchi,
<date id="FFAA1061FFE62226E578FF1CBCBE318E" box="[1033,1178,226,252]" pageId="3" pageNumber="46" value="2001-03">March 2001</date>
.
<collectionCode id="ED05AE64FFE62226E5D4FF1CBCDB318E" LSID="urn:lsid:biocol.org:col:34925" box="[1189,1279,226,252]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="46">AMNH</collectionCode>
251633 4 ex. (1C&amp;S),
<specimenCount id="9D12FD28FFE62226E074FEF6B97D3050" box="[261,345,264,290]" pageId="3" pageNumber="46" type="male">1 male</specimenCount>
,
<specimenCount id="9D12FD28FFE62226E017FEF6B9F93050" box="[358,477,264,290]" pageId="3" pageNumber="46" type="female">3 females</specimenCount>
,
<quantity id="4CEC9B44FFE62226E09BFEF6BABC3050" box="[490,664,264,290]" metricMagnitude="-2" metricUnit="m" metricValue="7.005" metricValueMax="7.65" metricValueMin="6.36" pageId="3" pageNumber="46" unit="mm" value="70.05" valueMax="76.5" valueMin="63.6">63.6–76.5 mm</quantity>
SL,
<collectingCountry id="F3037631FFE62226E3A2FEF6BB0D3050" box="[723,809,264,290]" name="Eritrea" pageId="3" pageNumber="46">Eritrea</collectingCountry>
: Lake Abaeded, G.
<collectingCountry id="F3037631FFE62226E565FEF6BC133050" box="[1044,1079,264,290]" name="South Georgia and the South Sandwich Islands" pageId="3" pageNumber="46">De</collectingCountry>
Marchi,
<date id="FFAA1061FFE62226E5D4FEF6BD1E3050" box="[1189,1338,264,290]" pageId="3" pageNumber="46" value="2001-03">March 2001</date>
.
<collectionCode id="ED05AE64FFE62226E437FEF6BDB83050" box="[1350,1436,264,290]" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="3" pageNumber="46">MSNG</collectionCode>
55210,
<specimenCount id="9D12FD28FFE62226E180FED3B9783035" box="[241,348,301,327]" pageId="3" pageNumber="46" type="female">1 female</specimenCount>
, 73.0 mm SL,
<collectingCountry id="F3037631FFE62226E37FFED3BA403035" box="[526,612,301,327]" name="Eritrea" pageId="3" pageNumber="46">Eritrea</collectingCountry>
: Lake Abaeded. 14˚ 35' 17&quot; N, 40˚ 05' 04&quot;, E., G.
<collectingCountry id="F3037631FFE62226E5CFFED3BCC53035" box="[1214,1249,301,327]" name="South Georgia and the South Sandwich Islands" pageId="3" pageNumber="46">De</collectingCountry>
Marchi,
<date id="FFAA1061FFE62226E43EFED3B8F1301E" pageId="3" pageNumber="46" value="2001-03">March 2001</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E652AFFE62226E1B7FE89BC6E3355" pageId="3" pageNumber="46" type="diagnosis">
<paragraph id="8BAB36A1FFE62226E1B7FE89BC6E3355" blockId="3.[151,1437,151,812]" pageId="3" pageNumber="46">
<emphasis id="B960EAB3FFE62226E1B7FE89B96230E3" bold="true" box="[198,326,375,401]" pageId="3" pageNumber="46">Diagnosis.</emphasis>
Differs from its only congener,
<taxonomicName id="4C144D22FFE62226E3B0FE86BB9C30E3" box="[705,952,376,401]" class="Actinopterygii" family="Cichlidae" genus="Danakilia" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="46" phylum="Chordata" rank="species" species="franchettii">
<emphasis id="B960EAB3FFE62226E3B0FE86BB9C30E3" box="[705,952,376,401]" italics="true" pageId="3" pageNumber="46">Danakilia franchettii</emphasis>
</taxonomicName>
, in the possession of longer pectoral fins (31.1–38.7% SL vs. 21.7–30.6% SL). Outer row teeth in both oral jaws composed of stout tricuspids, considerably more robust and fewer in number than those of
<taxonomicName id="4C144D22FFE62226E212FE3CBC2430A9" box="[867,1024,450,475]" class="Actinopterygii" family="Cichlidae" genus="Danakilia" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="46" phylum="Chordata" rank="species" species="franchettii">
<emphasis id="B960EAB3FFE62226E212FE3CBC2430A9" box="[867,1024,450,475]" italics="true" pageId="3" pageNumber="46">D. franchettii</emphasis>
</taxonomicName>
. Lower pharyngeal bone relatively massive, longer than wide, length 30–32% HL (vs. small, only slightly longer than wide, length 25–27% HL in
<taxonomicName id="4C144D22FFE62226E1C4FDF3B9773354" box="[181,339,525,550]" class="Actinopterygii" family="Cichlidae" genus="Danakilia" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="46" phylum="Chordata" rank="species" species="franchettii">
<emphasis id="B960EAB3FFE62226E1C4FDF3B9773354" box="[181,339,525,550]" italics="true" pageId="3" pageNumber="46">D. franchettii</emphasis>
</taxonomicName>
), with 40–48 teeth in posterior row (vs.
<quantity id="4CEC9B44FFE62226E25DFDF3BBB03355" box="[812,916,525,551]" metricMagnitude="-1" metricUnit="m" metricValue="7.874" metricValueMax="9.144" metricValueMin="6.604" pageId="3" pageNumber="46" unit="in" value="31.0" valueMax="36.0" valueMin="26.0">26–36 in</quantity>
<taxonomicName id="4C144D22FFE62226E2EAFDF3BC1D3354" box="[923,1081,525,550]" class="Actinopterygii" family="Cichlidae" genus="Danakilia" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="46" phylum="Chordata" rank="species" species="franchettii">
<emphasis id="B960EAB3FFE62226E2EAFDF3BC1D3354" box="[923,1081,525,550]" italics="true" pageId="3" pageNumber="46">D. franchettii</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C30E652AFFE6222DE1B7FDCCBD7C30C5" lastPageId="8" lastPageNumber="51" pageId="3" pageNumber="46" type="description">
<paragraph id="8BAB36A1FFE62226E1B7FDCCBC86325E" blockId="3.[151,1437,151,812]" pageId="3" pageNumber="46">
<emphasis id="B960EAB3FFE62226E1B7FDCCB97A333E" bold="true" box="[198,350,562,588]" pageId="3" pageNumber="46">Description.</emphasis>
Based on
<typeStatus id="54AF8803FFE62226E0ADFDCCBA67333E" box="[476,579,562,588]" pageId="3" pageNumber="46" type="holotype">holotype</typeStatus>
and 13
<typeStatus id="54AF8803FFE62226E3D3FDCCBB32333E" box="[674,790,562,588]" pageId="3" pageNumber="46" type="paratype">paratypes</typeStatus>
. Maximum size
<quantity id="4CEC9B44FFE62226E2ABFDCCBC67333E" box="[986,1091,562,588]" metricMagnitude="-2" metricUnit="m" metricValue="9.64" pageId="3" pageNumber="46" unit="mm" value="96.4">96.4 mm</quantity>
SL. Counts and proportional measurements are given in
<tableCitation id="C696031AFFE62226E090FDA6BA193300" box="[481,573,600,626]" captionStart="TABLE 2" captionStartId="4.[151,239,151,175]" captionTargetBox="[159,1313,330,951]" captionTargetPageId="4" captionText="TABLE 2. Morphometric and meristic data for the holotype and 13 paratypes of Danakilia dinicolai." httpUri="http://table.plazi.org/id/DF6B6629FFE12221E1E6FF69BCEA31DD" pageId="3" pageNumber="46" tableUuid="DF6B6629FFE12221E1E6FF69BCEA31DD">Table 2</tableCitation>
. Snout acute, mouth terminal, lips thickened but not fleshy. Head large, snout conical with straight profile, head smoothly convex behind nape to dorsal fin, except in largest males with a bulbous frontal gibbosity, particularly evident in life (
<figureCitation id="132F2A24FFE62226E20FFD5CBBF733CE" box="[894,979,674,700]" captionStart="FIGURE 2" captionStartId="3.[151,255,1398,1422]" captionTargetBox="[162,1432,883,1354]" captionTargetId="figure@3.[151,1436,850,1375]" captionTargetPageId="3" captionText="FIGURE 2. Danakilia dinicolai, holotype, MSNM 4888, male, 96.4 mm SL, Lake Abaeded, Eritrea." httpUri="https://zenodo.org/record/199531/files/figure.png" pageId="3" pageNumber="46">Figs. 2</figureCitation>
&amp;
<figureCitation id="132F2A24FFE62226E28AFD5CBC2833CE" box="[1019,1036,674,700]" captionStart="FIGURE 4" captionStartId="6.[151,255,1486,1510]" captionTargetBox="[172,1414,216,1443]" captionTargetId="figure@6.[151,1436,194,1465]" captionTargetPageId="6" captionText="FIGURE 4. Danakilia dinicolai (a) wild caught female and (b) wild caught male. Specimens not preserved or measured." httpUri="https://zenodo.org/record/199533/files/figure.png" pageId="3" pageNumber="46">4</figureCitation>
B). Greatest body depth between insertion of dorsal and pelvic fins. Caudal peduncle always longer than deep. Caudal fin truncate. Dorsal XIII–XIV 9–11. Anal
<date id="FFAA1061FFE62226E0EAFD13B9EA3275" box="[411,462,749,775]" pageId="3" pageNumber="46">III 7</date>
or 8. Pectoral fin long, reaching to anal fin origin or beyond. First pelvic ray longer in males than females but pelvic fin always reaching anus, and often beyond anal origin.
</paragraph>
<caption id="DF6B6629FFE62226E1E6FA88BCEA34FC" box="[151,1230,1398,1422]" httpUri="https://zenodo.org/record/199531/files/figure.png" pageId="3" pageNumber="46" targetBox="[162,1432,883,1354]" targetPageId="3">
<paragraph id="8BAB36A1FFE62226E1E6FA88BCEA34FC" blockId="3.[151,1230,1398,1422]" box="[151,1230,1398,1422]" pageId="3" pageNumber="46">
<emphasis id="B960EAB3FFE62226E1E6FA88B93D34FC" bold="true" box="[151,281,1398,1422]" pageId="3" pageNumber="46">FIGURE 2.</emphasis>
<taxonomicName id="4C144D22FFE62226E051FA89B9D434FC" box="[288,496,1399,1422]" class="Actinopterygii" family="Cichlidae" genus="Danakilia" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="46" phylum="Chordata" rank="species" species="dinicolai">
<emphasis id="B960EAB3FFE62226E051FA89B9D434FC" box="[288,496,1399,1422]" italics="true" pageId="3" pageNumber="46">Danakilia dinicolai</emphasis>
</taxonomicName>
, holotype, MSNM 4888, male, 96.4 mm SL, Lake Abaeded, Eritrea.
</paragraph>
</caption>
<paragraph id="8BAB36A1FFE62226E1B7FA3EBD3837AD" blockId="3.[151,1437,1472,2030]" pageId="3" pageNumber="46">
Jaws isognathous, both with 2–4 rows of tricuspid teeth. Outer row teeth in both jaws stout tricuspids with small, symmetrically rounded lateral cusps and larger medial cusp (
<figureCitation id="132F2A24FFE62226E2B3FA1BBC2D348D" box="[962,1033,1509,1535]" captionStart="FIGURE 3" captionStartId="5.[151,255,1914,1938]" captionTargetBox="[248,1362,206,1893]" captionTargetId="figure@5.[219,1378,194,1904]" captionTargetPageId="5" captionText="FIGURE 3. Lower pharyngeal jaw (dorsal, posterior and lateral views), left mandible and isolated tooth in (a) Danakilia dinicolai, (b) Danakilia franchettii, (c) Iranocichla hormuzensis. Elements drawn in corresponding scale for each species. Posterior neurocranium and anterior vertebral elements, arrows indicate inferior vertebral apophysis of (d) Iranocichla hormuzensis." httpUri="https://zenodo.org/record/199532/files/figure.png" pageId="3" pageNumber="46">Fig. 3</figureCitation>
A), considerably more robust and fewer in number than in
<taxonomicName id="4C144D22FFE62226E0C6F9F5BA713756" box="[439,597,1547,1572]" class="Actinopterygii" family="Cichlidae" genus="Danakilia" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="46" phylum="Chordata" rank="species" species="franchettii">
<emphasis id="B960EAB3FFE62226E0C6F9F5BA713756" box="[439,597,1547,1572]" italics="true" pageId="3" pageNumber="46">D. franchettii</emphasis>
</taxonomicName>
(
<figureCitation id="132F2A24FFE62226E314F9F4BA8F3756" box="[613,683,1546,1572]" captionStart="FIGURE 3" captionStartId="5.[151,255,1914,1938]" captionTargetBox="[248,1362,206,1893]" captionTargetId="figure@5.[219,1378,194,1904]" captionTargetPageId="5" captionText="FIGURE 3. Lower pharyngeal jaw (dorsal, posterior and lateral views), left mandible and isolated tooth in (a) Danakilia dinicolai, (b) Danakilia franchettii, (c) Iranocichla hormuzensis. Elements drawn in corresponding scale for each species. Posterior neurocranium and anterior vertebral elements, arrows indicate inferior vertebral apophysis of (d) Iranocichla hormuzensis." httpUri="https://zenodo.org/record/199532/files/figure.png" pageId="3" pageNumber="46">Fig. 3</figureCitation>
B). Lower pharyngeal bone relatively massive, longer than wide (
<figureCitation id="132F2A24FFE62226E1D1F9CEB8C83738" box="[160,236,1584,1610]" captionStart="FIGURE 3" captionStartId="5.[151,255,1914,1938]" captionTargetBox="[248,1362,206,1893]" captionTargetId="figure@5.[219,1378,194,1904]" captionTargetPageId="5" captionText="FIGURE 3. Lower pharyngeal jaw (dorsal, posterior and lateral views), left mandible and isolated tooth in (a) Danakilia dinicolai, (b) Danakilia franchettii, (c) Iranocichla hormuzensis. Elements drawn in corresponding scale for each species. Posterior neurocranium and anterior vertebral elements, arrows indicate inferior vertebral apophysis of (d) Iranocichla hormuzensis." httpUri="https://zenodo.org/record/199532/files/figure.png" pageId="3" pageNumber="46">Fig. 3</figureCitation>
A), length 30–32% HL (versus small, only slightly longer than wide, length 25–27% HL in
<taxonomicName id="4C144D22FFE62226E40FF9CEB93F371C" class="Actinopterygii" family="Cichlidae" genus="Danakilia" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="46" phylum="Chordata" rank="species" species="franchettii">
<emphasis id="B960EAB3FFE62226E40FF9CEB93F371C" italics="true" pageId="3" pageNumber="46">D. franchettii</emphasis>
</taxonomicName>
(
<figureCitation id="132F2A24FFE62226E05FF9ABB95E371D" box="[302,378,1621,1647]" captionStart="FIGURE 3" captionStartId="5.[151,255,1914,1938]" captionTargetBox="[248,1362,206,1893]" captionTargetId="figure@5.[219,1378,194,1904]" captionTargetPageId="5" captionText="FIGURE 3. Lower pharyngeal jaw (dorsal, posterior and lateral views), left mandible and isolated tooth in (a) Danakilia dinicolai, (b) Danakilia franchettii, (c) Iranocichla hormuzensis. Elements drawn in corresponding scale for each species. Posterior neurocranium and anterior vertebral elements, arrows indicate inferior vertebral apophysis of (d) Iranocichla hormuzensis." httpUri="https://zenodo.org/record/199532/files/figure.png" pageId="3" pageNumber="46">Fig. 3</figureCitation>
B). Pharyngeal teeth in anterior field of jaw, robust unicuspids, sparsely implanted, becoming increasingly elongate, recurved and spatulate posteriorly, densely implanted over posterior field of jaw, 40–48 teeth in posterior row (
<figureCitation id="132F2A24FFE62226E332F95EBAA837C8" box="[579,652,1696,1722]" captionStart="FIGURE 3" captionStartId="5.[151,255,1914,1938]" captionTargetBox="[248,1362,206,1893]" captionTargetId="figure@5.[219,1378,194,1904]" captionTargetPageId="5" captionText="FIGURE 3. Lower pharyngeal jaw (dorsal, posterior and lateral views), left mandible and isolated tooth in (a) Danakilia dinicolai, (b) Danakilia franchettii, (c) Iranocichla hormuzensis. Elements drawn in corresponding scale for each species. Posterior neurocranium and anterior vertebral elements, arrows indicate inferior vertebral apophysis of (d) Iranocichla hormuzensis." httpUri="https://zenodo.org/record/199532/files/figure.png" pageId="3" pageNumber="46">Fig. 3</figureCitation>
A). Pharyngeal teeth in
<taxonomicName id="4C144D22FFE62226E2C4F95EBC7F37CB" box="[949,1115,1696,1721]" class="Actinopterygii" family="Cichlidae" genus="Danakilia" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="46" phylum="Chordata" rank="species" species="franchettii">
<emphasis id="B960EAB3FFE62226E2C4F95EBC7F37CB" box="[949,1115,1696,1721]" italics="true" pageId="3" pageNumber="46">D. franchettii</emphasis>
</taxonomicName>
are similarly arrayed, but coarser and less densely implanted in posterior field of jaw, 26–36 teeth in posterior row (
<figureCitation id="132F2A24FFE62226E5C3F93BBCDC37AD" box="[1202,1272,1733,1759]" captionStart="FIGURE 3" captionStartId="5.[151,255,1914,1938]" captionTargetBox="[248,1362,206,1893]" captionTargetId="figure@5.[219,1378,194,1904]" captionTargetPageId="5" captionText="FIGURE 3. Lower pharyngeal jaw (dorsal, posterior and lateral views), left mandible and isolated tooth in (a) Danakilia dinicolai, (b) Danakilia franchettii, (c) Iranocichla hormuzensis. Elements drawn in corresponding scale for each species. Posterior neurocranium and anterior vertebral elements, arrows indicate inferior vertebral apophysis of (d) Iranocichla hormuzensis." httpUri="https://zenodo.org/record/199532/files/figure.png" pageId="3" pageNumber="46">Fig. 3</figureCitation>
B).
</paragraph>
<paragraph id="8BAB36A1FFE62226E1B4F914BBFD3658" blockId="3.[151,1437,1472,2030]" pageId="3" pageNumber="46">Eleven to 14 small, moderately tuberculate gill rakers along lower limb of first arch (including raker at angle), 2 or 3 pointed epibranchial rakers. Microbranchiospines absent.</paragraph>
<paragraph id="8BAB36A1FFE62226E1B7F8CBBD183698" blockId="3.[151,1437,1472,2030]" pageId="3" pageNumber="46">
Scales cycloid, 2 or 3 rows on cheek, 2–4 horizontal rows on opercle. Dark scaleless spot on outer edge of opercle. Chest and belly scales deeply embedded, markedly smaller than on flank, 4 or 5 scales between pectoral and pelvic fins. Upper lateral line separated from dorsal fin anteriorly by 3 or 4 scale rows, at 8th pored scale by 2 scale rows, at last pored scale by 1–1.5 scale rows.
<collectingCountry id="F3037631FFE62226E2B2F856BBD636B0" box="[963,1010,1960,1986]" name="India" pageId="3" pageNumber="46">End</collectingCountry>
of upper lateral line separated from lower lateral line by 1 or 2 scale rows. Basal 1/3 of caudal fin covered with scales; other fins unscaled.
</paragraph>
<caption id="DF6B6629FFE12221E1E6FF69BCEA31DD" ID-Table-UUID="DF6B6629FFE12221E1E6FF69BCEA31DD" box="[151,1230,151,175]" httpUri="http://table.plazi.org/id/DF6B6629FFE12221E1E6FF69BCEA31DD" pageId="4" pageNumber="47" targetBox="[159,1313,330,951]" targetIsTable="true" targetPageId="4">
<paragraph id="8BAB36A1FFE12221E1E6FF69BCEA31DD" blockId="4.[151,1313,151,268]" box="[151,1230,151,175]" pageId="4" pageNumber="47">
<emphasis id="B960EAB3FFE12221E1E6FF69B92C31DD" bold="true" box="[151,264,151,175]" pageId="4" pageNumber="47">TABLE 2.</emphasis>
Morphometric and meristic data for the holotype and 13 paratypes of
<taxonomicName id="4C144D22FFE12221E286FF66BCE331DD" box="[1015,1223,152,175]" class="Actinopterygii" family="Cichlidae" genus="Danakilia" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="47" phylum="Chordata" rank="species" species="dinicolai">
<emphasis id="B960EAB3FFE12221E286FF66BCE331DD" box="[1015,1223,152,175]" italics="true" pageId="4" pageNumber="47">Danakilia dinicolai</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BAB36A1FFE12221E303FF34BD1834C5" blockId="4.[151,1313,151,268]" lastBlockId="4.[159,1340,1440,1505]" pageId="4" pageNumber="47">
<typeStatus id="54AF8803FFE12221E303FF34BAEB3193" box="[626,719,202,225]" pageId="4" pageNumber="47" type="holotype">holotype</typeStatus>
mean SD Range Standard length 96.4 77.9 63.6–96.4 Caudal peduncle length % as of depth 120.2 118.7 8.3 102.8–128.8
</paragraph>
<paragraph id="8BAB36A1FFE12221E1EEFEB4BD0432C5" pageId="4" pageNumber="47">
<table id="F914C401FFE1DDDAE1EEFEB4BD0532C5" box="[159,1313,330,951]" gridcols="4" gridrows="15" pageId="4" pageNumber="47">
<tr id="352434E3FFE1DDDAE1EEFEB4BD053013" box="[159,1313,330,353]" gridrow="0" pageId="4" pageNumber="47" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76F55D9FFFE1DDDAE1EEFEB4B9843013" box="[159,416,330,353]" gridcol="0" gridrow="0" pageId="4" pageNumber="47">% SL</th>
</tr>
<tr id="352434E3FFE1DDDAE1EEFE8BBD0530FE" box="[159,1313,373,396]" gridrow="1" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFE8BB98430FE" box="[159,416,373,396]" gridcol="0" gridrow="1" pageId="4" pageNumber="47">Body depth</th>
<td id="76F55D9FFFE1DDDAE31EFE8BBAB930FE" box="[623,669,373,396]" gridcol="1" gridrow="1" pageId="4" pageNumber="47">41.0</td>
<td id="76F55D9FFFE1DDDAE214FE8BBBB730FE" box="[869,915,373,396]" gridcol="2" gridrow="1" pageId="4" pageNumber="47">39.4</td>
<td id="76F55D9FFFE1DDDAE55EFE8BBD0530FE" box="[1071,1313,373,396]" gridcol="3" gridrow="1" pageId="4" pageNumber="47">1.2 37.2–41.3</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEFE5EBD0530C5" box="[159,1313,416,439]" gridrow="2" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFE5EB98430C5" box="[159,416,416,439]" gridcol="0" gridrow="2" pageId="4" pageNumber="47">Head length</th>
<td id="76F55D9FFFE1DDDAE31EFE5EBAB930C5" box="[623,669,416,439]" gridcol="1" gridrow="2" pageId="4" pageNumber="47">39.2</td>
<td id="76F55D9FFFE1DDDAE214FE5EBBB730C5" box="[869,915,416,439]" gridcol="2" gridrow="2" pageId="4" pageNumber="47">38.9</td>
<td id="76F55D9FFFE1DDDAE55EFE5EBD0530C5" box="[1071,1313,416,439]" gridcol="3" gridrow="2" pageId="4" pageNumber="47">1.2 37.0–40.9</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEFE34BD053093" box="[159,1313,458,481]" gridrow="3" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFE34B9843093" box="[159,416,458,481]" gridcol="0" gridrow="3" pageId="4" pageNumber="47">Caudal peduncle length</th>
<td id="76F55D9FFFE1DDDAE31EFE34BAB93093" box="[623,669,458,481]" gridcol="1" gridrow="3" pageId="4" pageNumber="47">14.2</td>
<td id="76F55D9FFFE1DDDAE214FE34BBB73093" box="[869,915,458,481]" gridcol="2" gridrow="3" pageId="4" pageNumber="47">14.2</td>
<td id="76F55D9FFFE1DDDAE55EFE34BD053093" box="[1071,1313,458,481]" gridcol="3" gridrow="3" pageId="4" pageNumber="47">0.9 12.5–15.2</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEFE0BBD05337E" box="[159,1313,501,524]" gridrow="4" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFE0BB984337E" box="[159,416,501,524]" gridcol="0" gridrow="4" pageId="4" pageNumber="47">Caudal peduncle depth</th>
<td id="76F55D9FFFE1DDDAE31EFE0BBAB9337E" box="[623,669,501,524]" gridcol="1" gridrow="4" pageId="4" pageNumber="47">11.8</td>
<td id="76F55D9FFFE1DDDAE214FE0BBBB7337E" box="[869,915,501,524]" gridcol="2" gridrow="4" pageId="4" pageNumber="47">11.9</td>
<td id="76F55D9FFFE1DDDAE55EFE0BBD05337E" box="[1071,1313,501,524]" gridcol="3" gridrow="4" pageId="4" pageNumber="47">0.5 11.4–13.4</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEFDDEBD053345" box="[159,1313,544,567]" gridrow="5" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFDDEB9843345" box="[159,416,544,567]" gridcol="0" gridrow="5" pageId="4" pageNumber="47">Dorsal-fin base</th>
<td id="76F55D9FFFE1DDDAE31EFDDEBAB93345" box="[623,669,544,567]" gridcol="1" gridrow="5" pageId="4" pageNumber="47">49.2</td>
<td id="76F55D9FFFE1DDDAE214FDDEBBB73345" box="[869,915,544,567]" gridcol="2" gridrow="5" pageId="4" pageNumber="47">48.7</td>
<td id="76F55D9FFFE1DDDAE55EFDDEBD053345" box="[1071,1313,544,567]" gridcol="3" gridrow="5" pageId="4" pageNumber="47">1.3 46.8–51.0</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEFDB4BD053313" box="[159,1313,586,609]" gridrow="6" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFDB4B9843313" box="[159,416,586,609]" gridcol="0" gridrow="6" pageId="4" pageNumber="47">Anal-fin base</th>
<td id="76F55D9FFFE1DDDAE31EFDB4BAB93313" box="[623,669,586,609]" gridcol="1" gridrow="6" pageId="4" pageNumber="47">13.7</td>
<td id="76F55D9FFFE1DDDAE214FDB4BBB73313" box="[869,915,586,609]" gridcol="2" gridrow="6" pageId="4" pageNumber="47">13.5</td>
<td id="76F55D9FFFE1DDDAE55EFDB4BD053313" box="[1071,1313,586,609]" gridcol="3" gridrow="6" pageId="4" pageNumber="47">1.0 12.0–14.9</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEFD8BBD0533FE" box="[159,1313,629,652]" gridrow="7" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFD8BB98433FE" box="[159,416,629,652]" gridcol="0" gridrow="7" pageId="4" pageNumber="47">Predorsal distance</th>
<td id="76F55D9FFFE1DDDAE31EFD8BBAB933FE" box="[623,669,629,652]" gridcol="1" gridrow="7" pageId="4" pageNumber="47">39.2</td>
<td id="76F55D9FFFE1DDDAE214FD8BBBB733FE" box="[869,915,629,652]" gridcol="2" gridrow="7" pageId="4" pageNumber="47">40.0</td>
<td id="76F55D9FFFE1DDDAE55EFD8BBD0533FE" box="[1071,1313,629,652]" gridcol="3" gridrow="7" pageId="4" pageNumber="47">1.0 38.7–41.7</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEFD5EBD0533C5" box="[159,1313,672,695]" gridrow="8" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFD5EB98433C5" box="[159,416,672,695]" gridcol="0" gridrow="8" pageId="4" pageNumber="47">Preanal distance</th>
<td id="76F55D9FFFE1DDDAE31EFD5EBAB933C5" box="[623,669,672,695]" gridcol="1" gridrow="8" pageId="4" pageNumber="47">75.7</td>
<td id="76F55D9FFFE1DDDAE214FD5EBBB733C5" box="[869,915,672,695]" gridcol="2" gridrow="8" pageId="4" pageNumber="47">73.3</td>
<td id="76F55D9FFFE1DDDAE55EFD5EBD0533C5" box="[1071,1313,672,695]" gridcol="3" gridrow="8" pageId="4" pageNumber="47">2.6 69.5–76.3</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEFD34BD053393" box="[159,1313,714,737]" gridrow="9" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFD34B9843393" box="[159,416,714,737]" gridcol="0" gridrow="9" pageId="4" pageNumber="47">Prepectoral distance</th>
<td id="76F55D9FFFE1DDDAE31EFD34BAB93393" box="[623,669,714,737]" gridcol="1" gridrow="9" pageId="4" pageNumber="47">44.5</td>
<td id="76F55D9FFFE1DDDAE214FD34BBB73393" box="[869,915,714,737]" gridcol="2" gridrow="9" pageId="4" pageNumber="47">42.1</td>
<td id="76F55D9FFFE1DDDAE55EFD34BD053393" box="[1071,1313,714,737]" gridcol="3" gridrow="9" pageId="4" pageNumber="47">1.0 40.8–44.5</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEFD0BBD05327E" box="[159,1313,757,780]" gridrow="10" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFD0BB984327E" box="[159,416,757,780]" gridcol="0" gridrow="10" pageId="4" pageNumber="47">Prepelvic distance</th>
<td id="76F55D9FFFE1DDDAE31EFD0BBAB9327E" box="[623,669,757,780]" gridcol="1" gridrow="10" pageId="4" pageNumber="47">47.4</td>
<td id="76F55D9FFFE1DDDAE214FD0BBBB7327E" box="[869,915,757,780]" gridcol="2" gridrow="10" pageId="4" pageNumber="47">45.3</td>
<td id="76F55D9FFFE1DDDAE55EFD0BBD05327E" box="[1071,1313,757,780]" gridcol="3" gridrow="10" pageId="4" pageNumber="47">1.3 43.5–47.4</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEFCDEBD053245" box="[159,1313,800,823]" gridrow="11" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFCDEB9843245" box="[159,416,800,823]" gridcol="0" gridrow="11" pageId="4" pageNumber="47">Longest dorsal-fin ray</th>
<td id="76F55D9FFFE1DDDAE31EFCDEBAB93245" box="[623,669,800,823]" gridcol="1" gridrow="11" pageId="4" pageNumber="47">28.3</td>
<td id="76F55D9FFFE1DDDAE214FCDEBBB73245" box="[869,915,800,823]" gridcol="2" gridrow="11" pageId="4" pageNumber="47">17.1</td>
<td id="76F55D9FFFE1DDDAE55EFCDEBD053245" box="[1071,1313,800,823]" gridcol="3" gridrow="11" pageId="4" pageNumber="47">3.9 13.2–28.3</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEFCB4BD053213" box="[159,1313,842,865]" gridrow="12" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFCB4B9843213" box="[159,416,842,865]" gridcol="0" gridrow="12" pageId="4" pageNumber="47">Longest anal-fin ray</th>
<td id="76F55D9FFFE1DDDAE31EFCB4BAB93213" box="[623,669,842,865]" gridcol="1" gridrow="12" pageId="4" pageNumber="47">25.4</td>
<td id="76F55D9FFFE1DDDAE214FCB4BBB73213" box="[869,915,842,865]" gridcol="2" gridrow="12" pageId="4" pageNumber="47">16.1</td>
<td id="76F55D9FFFE1DDDAE55EFCB4BD053213" box="[1071,1313,842,865]" gridcol="3" gridrow="12" pageId="4" pageNumber="47">3.1 13.8–25.4</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEFC8BBD0532FE" box="[159,1313,885,908]" gridrow="13" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFC8BB98432FE" box="[159,416,885,908]" gridcol="0" gridrow="13" pageId="4" pageNumber="47">Longest pectoral-fin ray</th>
<td id="76F55D9FFFE1DDDAE31EFC8BBAB932FE" box="[623,669,885,908]" gridcol="1" gridrow="13" pageId="4" pageNumber="47">31.1</td>
<td id="76F55D9FFFE1DDDAE214FC8BBBB732FE" box="[869,915,885,908]" gridcol="2" gridrow="13" pageId="4" pageNumber="47">33.6</td>
<td id="76F55D9FFFE1DDDAE55EFC8BBD0532FE" box="[1071,1313,885,908]" gridcol="3" gridrow="13" pageId="4" pageNumber="47">2.5 31.1–38.7</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEFC5EBD0532C5" box="[159,1313,928,951]" gridrow="14" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFC5EB98432C5" box="[159,416,928,951]" gridcol="0" gridrow="14" pageId="4" pageNumber="47">Longest pelvic-fin ray</th>
<td id="76F55D9FFFE1DDDAE31EFC5EBAB932C5" box="[623,669,928,951]" gridcol="1" gridrow="14" pageId="4" pageNumber="47">32.6</td>
<td id="76F55D9FFFE1DDDAE214FC5EBBB732C5" box="[869,915,928,951]" gridcol="2" gridrow="14" pageId="4" pageNumber="47">24.0</td>
<td id="76F55D9FFFE1DDDAE55EFC5EBD0532C5" box="[1071,1313,928,951]" gridcol="3" gridrow="14" pageId="4" pageNumber="47">2.9 21.2–32.6</td>
</tr>
</table>
</paragraph>
<paragraph id="8BAB36A1FFE12221E1EEFC0BBD043413" pageId="4" pageNumber="47">
<table id="F914C401FFE1DDDAE1EEFC0BBD053413" box="[159,1313,1013,1377]" gridcols="4" gridrows="9" pageId="4" pageNumber="47">
<tr id="352434E3FFE1DDDAE1EEFC0BBD05357E" box="[159,1313,1013,1036]" gridrow="0" pageId="4" pageNumber="47" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76F55D9FFFE1DDDAE1EEFC0BB951357E" box="[159,373,1013,1036]" gridcol="0" gridrow="0" pageId="4" pageNumber="47">% HL</th>
</tr>
<tr id="352434E3FFE1DDDAE1EEFBDEBD053545" box="[159,1313,1056,1079]" gridrow="1" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFBDEB9513545" box="[159,373,1056,1079]" gridcol="0" gridrow="1" pageId="4" pageNumber="47">Head depth</th>
<td id="76F55D9FFFE1DDDAE31EFBDEBAB93545" box="[623,669,1056,1079]" gridcol="1" gridrow="1" pageId="4" pageNumber="47">96.0</td>
<td id="76F55D9FFFE1DDDAE214FBDEBBB73545" box="[869,915,1056,1079]" gridcol="2" gridrow="1" pageId="4" pageNumber="47">70.0</td>
<td id="76F55D9FFFE1DDDAE55EFBDEBD053545" box="[1071,1313,1056,1079]" gridcol="3" gridrow="1" pageId="4" pageNumber="47">8.8 61.7–96.0</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEFBB4BD053513" box="[159,1313,1098,1121]" gridrow="2" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFBB4B9513513" box="[159,373,1098,1121]" gridcol="0" gridrow="2" pageId="4" pageNumber="47">Snout length</th>
<td id="76F55D9FFFE1DDDAE31EFBB4BAB93513" box="[623,669,1098,1121]" gridcol="1" gridrow="2" pageId="4" pageNumber="47">42.2</td>
<td id="76F55D9FFFE1DDDAE214FBB4BBB73513" box="[869,915,1098,1121]" gridcol="2" gridrow="2" pageId="4" pageNumber="47">36.5</td>
<td id="76F55D9FFFE1DDDAE55EFBB4BD053513" box="[1071,1313,1098,1121]" gridcol="3" gridrow="2" pageId="4" pageNumber="47">2.5 31.7–42.2</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEFB8BBD0535FE" box="[159,1313,1141,1164]" gridrow="3" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFB8BB95135FE" box="[159,373,1141,1164]" gridcol="0" gridrow="3" pageId="4" pageNumber="47">Eye diameter</th>
<td id="76F55D9FFFE1DDDAE31EFB8BBAB935FE" box="[623,669,1141,1164]" gridcol="1" gridrow="3" pageId="4" pageNumber="47">19.2</td>
<td id="76F55D9FFFE1DDDAE214FB8BBBB735FE" box="[869,915,1141,1164]" gridcol="2" gridrow="3" pageId="4" pageNumber="47">24.1</td>
<td id="76F55D9FFFE1DDDAE55EFB8BBD0535FE" box="[1071,1313,1141,1164]" gridcol="3" gridrow="3" pageId="4" pageNumber="47">2.0 19.2–27.0</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEFB5EBD0535C5" box="[159,1313,1184,1207]" gridrow="4" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFB5EB95135C5" box="[159,373,1184,1207]" gridcol="0" gridrow="4" pageId="4" pageNumber="47">Postorbital distance</th>
<td id="76F55D9FFFE1DDDAE31EFB5EBAB935C5" box="[623,669,1184,1207]" gridcol="1" gridrow="4" pageId="4" pageNumber="47">38.6</td>
<td id="76F55D9FFFE1DDDAE214FB5EBBB735C5" box="[869,915,1184,1207]" gridcol="2" gridrow="4" pageId="4" pageNumber="47">39.4</td>
<td id="76F55D9FFFE1DDDAE55EFB5EBD0535C5" box="[1071,1313,1184,1207]" gridcol="3" gridrow="4" pageId="4" pageNumber="47">1.8 37.0–41.9</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEFB34BD053593" box="[159,1313,1226,1249]" gridrow="5" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFB34B9513593" box="[159,373,1226,1249]" gridcol="0" gridrow="5" pageId="4" pageNumber="47">Interorbital distance</th>
<td id="76F55D9FFFE1DDDAE31EFB34BAB93593" box="[623,669,1226,1249]" gridcol="1" gridrow="5" pageId="4" pageNumber="47">31.5</td>
<td id="76F55D9FFFE1DDDAE214FB34BBB73593" box="[869,915,1226,1249]" gridcol="2" gridrow="5" pageId="4" pageNumber="47">29.5</td>
<td id="76F55D9FFFE1DDDAE55EFB34BD053593" box="[1071,1313,1226,1249]" gridcol="3" gridrow="5" pageId="4" pageNumber="47">1.7 25.6–31.5</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEFB0BBD05347E" box="[159,1313,1269,1292]" gridrow="6" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFB0BB951347E" box="[159,373,1269,1292]" gridcol="0" gridrow="6" pageId="4" pageNumber="47">Cheek depth</th>
<td id="76F55D9FFFE1DDDAE31EFB0BBAB9347E" box="[623,669,1269,1292]" gridcol="1" gridrow="6" pageId="4" pageNumber="47">32.2</td>
<td id="76F55D9FFFE1DDDAE214FB0BBBB7347E" box="[869,915,1269,1292]" gridcol="2" gridrow="6" pageId="4" pageNumber="47">31.7</td>
<td id="76F55D9FFFE1DDDAE55EFB0BBD05347E" box="[1071,1313,1269,1292]" gridcol="3" gridrow="6" pageId="4" pageNumber="47">1.4 28.9–34.8</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEFADEBD053445" box="[159,1313,1312,1335]" gridrow="7" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFADEB9513445" box="[159,373,1312,1335]" gridcol="0" gridrow="7" pageId="4" pageNumber="47">Lower jaw length</th>
<td id="76F55D9FFFE1DDDAE31EFADEBAB93445" box="[623,669,1312,1335]" gridcol="1" gridrow="7" pageId="4" pageNumber="47">32.7</td>
<td id="76F55D9FFFE1DDDAE214FADEBBB73445" box="[869,915,1312,1335]" gridcol="2" gridrow="7" pageId="4" pageNumber="47">32.2</td>
<td id="76F55D9FFFE1DDDAE55EFADEBD053445" box="[1071,1313,1312,1335]" gridcol="3" gridrow="7" pageId="4" pageNumber="47">1.9 29.9–37.2</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEFAB4BD053413" box="[159,1313,1354,1377]" gridrow="8" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFAB4B9513413" box="[159,373,1354,1377]" gridcol="0" gridrow="8" pageId="4" pageNumber="47">Preorbital distance</th>
<td id="76F55D9FFFE1DDDAE31EFAB4BAB93413" box="[623,669,1354,1377]" gridcol="1" gridrow="8" pageId="4" pageNumber="47">25.9</td>
<td id="76F55D9FFFE1DDDAE214FAB4BBB73413" box="[869,915,1354,1377]" gridcol="2" gridrow="8" pageId="4" pageNumber="47">25.6</td>
<td id="76F55D9FFFE1DDDAE55EFAB4BD053413" box="[1071,1313,1354,1377]" gridcol="3" gridrow="8" pageId="4" pageNumber="47">1.5 23.2–27.6</td>
</tr>
</table>
</paragraph>
<paragraph id="8BAB36A1FFE12221E214FA34BB973493" blockId="4.[159,1340,1440,1505]" box="[869,947,1482,1505]" pageId="4" pageNumber="47">median</paragraph>
<paragraph id="8BAB36A1FFE12221E1EEFA0BBCDC3613" pageId="4" pageNumber="47">
<table id="F914C401FFE1DDDAE1EEFA0BBD183613" box="[159,1340,1525,1889]" gridcols="4" gridrows="9" pageId="4" pageNumber="47">
<tr id="352434E3FFE1DDDAE1EEFA0BBD18377E" box="[159,1340,1525,1548]" gridrow="0" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEFA0BB988377E" box="[159,428,1525,1548]" gridcol="0" gridrow="0" pageId="4" pageNumber="47">Upper lateral-line scales</th>
<th id="76F55D9FFFE1DDDAE31EFA0BBAAD377E" box="[623,649,1525,1548]" gridcol="1" gridrow="0" pageId="4" pageNumber="47">16</th>
<th id="76F55D9FFFE1DDDAE214FA0BBB5B377E" box="[869,895,1525,1548]" gridcol="2" gridrow="0" pageId="4" pageNumber="47">17</th>
<th id="76F55D9FFFE1DDDAE5C7FA0BBD18377E" box="[1206,1340,1525,1548]" gridcol="3" gridrow="0" pageId="4" pageNumber="47">16–18</th>
</tr>
<tr id="352434E3FFE1DDDAE1EEF9DEBD183745" box="[159,1340,1568,1591]" gridrow="1" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEF9DEB9883745" box="[159,428,1568,1591]" gridcol="0" gridrow="1" pageId="4" pageNumber="47">Lower lateral-line scales</th>
<td id="76F55D9FFFE1DDDAE31EF9DEBAAD3745" box="[623,649,1568,1591]" gridcol="1" gridrow="1" pageId="4" pageNumber="47">9</td>
<td id="76F55D9FFFE1DDDAE214F9DEBB5B3745" box="[869,895,1568,1591]" gridcol="2" gridrow="1" pageId="4" pageNumber="47">9</td>
<td id="76F55D9FFFE1DDDAE5C7F9DEBD183745" box="[1206,1340,1568,1591]" gridcol="3" gridrow="1" pageId="4" pageNumber="47">8–11</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEF9B4BD183713" box="[159,1340,1610,1633]" gridrow="2" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEF9B4B9883713" box="[159,428,1610,1633]" gridcol="0" gridrow="2" pageId="4" pageNumber="47">Total lateral-line scales</th>
<td id="76F55D9FFFE1DDDAE31EF9B4BAAD3713" box="[623,649,1610,1633]" gridcol="1" gridrow="2" pageId="4" pageNumber="47">27</td>
<td id="76F55D9FFFE1DDDAE214F9B4BB5B3713" box="[869,895,1610,1633]" gridcol="2" gridrow="2" pageId="4" pageNumber="47">27</td>
<td id="76F55D9FFFE1DDDAE5C7F9B4BD183713" box="[1206,1340,1610,1633]" gridcol="3" gridrow="2" pageId="4" pageNumber="47">27–29</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEF98BBD1837FE" box="[159,1340,1653,1676]" gridrow="3" pageId="4" pageNumber="47" rowspan-3="1">
<th id="76F55D9FFFE1DDDAE1EEF98BB98837FE" box="[159,428,1653,1676]" gridcol="0" gridrow="3" pageId="4" pageNumber="47">Circumpeduncular scales</th>
<td id="76F55D9FFFE1DDDAE31EF98BBAAD37FE" box="[623,649,1653,1676]" gridcol="1" gridrow="3" pageId="4" pageNumber="47">16</td>
<td id="76F55D9FFFE1DDDAE214F98BBB5B37FE" box="[869,895,1653,1676]" gridcol="2" gridrow="3" pageId="4" pageNumber="47">16</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEF95EBD1837C5" box="[159,1340,1696,1719]" gridrow="4" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEF95EB98837C5" box="[159,428,1696,1719]" gridcol="0" gridrow="4" pageId="4" pageNumber="47">Dorsal-fin spines</th>
<td id="76F55D9FFFE1DDDAE31EF95EBAAD37C5" box="[623,649,1696,1719]" gridcol="1" gridrow="4" pageId="4" pageNumber="47">14</td>
<td id="76F55D9FFFE1DDDAE214F95EBB5B37C5" box="[869,895,1696,1719]" gridcol="2" gridrow="4" pageId="4" pageNumber="47">14</td>
<td id="76F55D9FFFE1DDDAE5C7F95EBD1837C5" box="[1206,1340,1696,1719]" gridcol="3" gridrow="4" pageId="4" pageNumber="47">13–14</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEF934BD183793" box="[159,1340,1738,1761]" gridrow="5" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEF934B9883793" box="[159,428,1738,1761]" gridcol="0" gridrow="5" pageId="4" pageNumber="47">Dorsal-fin rays</th>
<td id="76F55D9FFFE1DDDAE31EF934BAAD3793" box="[623,649,1738,1761]" gridcol="1" gridrow="5" pageId="4" pageNumber="47">10</td>
<td id="76F55D9FFFE1DDDAE214F934BB5B3793" box="[869,895,1738,1761]" gridcol="2" gridrow="5" pageId="4" pageNumber="47">10</td>
<td id="76F55D9FFFE1DDDAE5C7F934BD183793" box="[1206,1340,1738,1761]" gridcol="3" gridrow="5" pageId="4" pageNumber="47">9–11</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEF90BBD18367E" box="[159,1340,1781,1804]" gridrow="6" pageId="4" pageNumber="47" rowspan-3="1">
<th id="76F55D9FFFE1DDDAE1EEF90BB988367E" box="[159,428,1781,1804]" gridcol="0" gridrow="6" pageId="4" pageNumber="47">Anal-fin spines</th>
<td id="76F55D9FFFE1DDDAE31EF90BBAAD367E" box="[623,649,1781,1804]" gridcol="1" gridrow="6" pageId="4" pageNumber="47">3</td>
<td id="76F55D9FFFE1DDDAE214F90BBB5B367E" box="[869,895,1781,1804]" gridcol="2" gridrow="6" pageId="4" pageNumber="47">3</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEF8DEBD183645" box="[159,1340,1824,1847]" gridrow="7" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEF8DEB9883645" box="[159,428,1824,1847]" gridcol="0" gridrow="7" pageId="4" pageNumber="47">Anal-fin rays</th>
<td id="76F55D9FFFE1DDDAE31EF8DEBAAD3645" box="[623,649,1824,1847]" gridcol="1" gridrow="7" pageId="4" pageNumber="47">7</td>
<td id="76F55D9FFFE1DDDAE214F8DEBB5B3645" box="[869,895,1824,1847]" gridcol="2" gridrow="7" pageId="4" pageNumber="47">8</td>
<td id="76F55D9FFFE1DDDAE5C7F8DEBD183645" box="[1206,1340,1824,1847]" gridcol="3" gridrow="7" pageId="4" pageNumber="47">7 or 8</td>
</tr>
<tr id="352434E3FFE1DDDAE1EEF8B4BD183613" box="[159,1340,1866,1889]" gridrow="8" pageId="4" pageNumber="47">
<th id="76F55D9FFFE1DDDAE1EEF8B4B9883613" box="[159,428,1866,1889]" gridcol="0" gridrow="8" pageId="4" pageNumber="47">Pectoral-fin rays</th>
<td id="76F55D9FFFE1DDDAE31EF8B4BAAD3613" box="[623,649,1866,1889]" gridcol="1" gridrow="8" pageId="4" pageNumber="47">13</td>
<td id="76F55D9FFFE1DDDAE214F8B4BB5B3613" box="[869,895,1866,1889]" gridcol="2" gridrow="8" pageId="4" pageNumber="47">12</td>
<td id="76F55D9FFFE1DDDAE5C7F8B4BD183613" box="[1206,1340,1866,1889]" gridcol="3" gridrow="8" pageId="4" pageNumber="47">11–13</td>
</tr>
</table>
</paragraph>
<paragraph id="8BAB36A1FFE12221E1EEF88BBCDD36C5" blockId="4.[159,1273,1909,1975]" pageId="4" pageNumber="47">
Gill rakers on lower limb of first arch
<date id="FFAA1061FFE12221E31EF88BBCD336FE" box="[623,1271,1909,1932]" pageId="4" pageNumber="47" value="1912-12-11" valueMax="1912-12-14" valueMin="1912-12-11">12 12 11–14</date>
Total gill rakers on first arch 14 14 13–17
</paragraph>
<caption id="DF6B6629FFE02220E1E6F884B9823683" httpUri="https://zenodo.org/record/199532/files/figure.png" pageId="5" pageNumber="48" targetBox="[248,1362,206,1893]" targetPageId="5">
<paragraph id="8BAB36A1FFE02220E1E6F884B9823683" blockId="5.[151,1437,1914,2033]" pageId="5" pageNumber="48">
<emphasis id="B960EAB3FFE02220E1E6F884B93D36E0" bold="true" box="[151,281,1914,1938]" pageId="5" pageNumber="48">FIGURE 3.</emphasis>
Lower pharyngeal jaw (dorsal, posterior and lateral views), left mandible and isolated tooth in (a)
<taxonomicName id="4C144D22FFE02220E440F884B8DF36C3" class="Actinopterygii" family="Cichlidae" genus="Danakilia" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="dinicolai">
<emphasis id="B960EAB3FFE02220E440F884B8DF36C3" italics="true" pageId="5" pageNumber="48">Danakilia dinicolai</emphasis>
</taxonomicName>
, (b)
<taxonomicName id="4C144D22FFE02220E042F864BA0436C3" box="[307,544,1946,1969]" class="Actinopterygii" family="Cichlidae" genus="Danakilia" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="franchettii">
<emphasis id="B960EAB3FFE02220E042F864BA0436C3" box="[307,544,1946,1969]" italics="true" pageId="5" pageNumber="48">Danakilia franchettii</emphasis>
</taxonomicName>
, (c)
<taxonomicName id="4C144D22FFE02220E326F864BB4936C3" box="[599,877,1946,1969]" class="Actinopterygii" family="Cichlidae" genus="Iranocichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="hormuzensis">
<emphasis id="B960EAB3FFE02220E326F864BB4936C3" box="[599,877,1946,1969]" italics="true" pageId="5" pageNumber="48">Iranocichla hormuzensis</emphasis>
</taxonomicName>
. Elements drawn in corresponding scale for each species. Posterior neurocranium and anterior vertebral elements, arrows indicate inferior vertebral apophysis of (d)
<taxonomicName id="4C144D22FFE02220E1E6F824B9BB3683" box="[151,415,2010,2033]" class="Actinopterygii" family="Cichlidae" genus="Iranocichla" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="hormuzensis">
<emphasis id="B960EAB3FFE02220E1E6F824B9BB3683" box="[151,415,2010,2033]" italics="true" pageId="5" pageNumber="48">Iranocichla hormuzensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF6B6629FFE32223E1E6FA30BDB93494" box="[151,1437,1486,1510]" httpUri="https://zenodo.org/record/199533/files/figure.png" pageId="6" pageNumber="49" targetBox="[172,1414,216,1443]" targetPageId="6">
<paragraph id="8BAB36A1FFE32223E1E6FA30BDB93494" blockId="6.[151,1437,1486,1510]" box="[151,1437,1486,1510]" pageId="6" pageNumber="49">
<emphasis id="B960EAB3FFE32223E1E6FA30B93D3494" bold="true" box="[151,281,1486,1510]" pageId="6" pageNumber="49">FIGURE 4.</emphasis>
<taxonomicName id="4C144D22FFE32223E06EFA31B9CB3494" box="[287,495,1487,1510]" class="Actinopterygii" family="Cichlidae" genus="Danakilia" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="49" phylum="Chordata" rank="species" species="dinicolai">
<emphasis id="B960EAB3FFE32223E06EFA31B9CB3494" box="[287,495,1487,1510]" italics="true" pageId="6" pageNumber="49">Danakilia dinicolai</emphasis>
</taxonomicName>
(a) wild caught female and (b) wild caught male. Specimens not preserved or measured.
</paragraph>
</caption>
<paragraph id="8BAB36A1FFE32223E1B7F9F0B9BF379B" blockId="6.[151,1437,1550,2031]" pageId="6" pageNumber="49">
<emphasis id="B960EAB3FFE32223E1B7F9F0B9D3375A" bold="true" box="[198,503,1550,1576]" pageId="6" pageNumber="49">Miscellaneous osteology.</emphasis>
Infraorbital series complete, with lachrymal (first infraorbital) and four tubular infraorbitals, lachrymal with five openings of laterosensory system. Total of 25 or 26 vertebrae, 11–13 precaudal and 12–14 caudal. Inferior vertebral apophysis (spondylophysial apophysis) supporting anterior end of gas bladder composed of elongate ventro-posteriorly directed process on 3rd vertebral centrum which abuts a stout ventro-posteriorly directed process on 4th centrum; similar in form and composition to that of
<taxonomicName id="4C144D22FFE32223E1E6F92EB904379B" box="[151,288,1744,1769]" class="Actinopterygii" family="Cichlidae" genus="Iranocichla" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="49" phylum="Chordata" rank="genus">
<emphasis id="B960EAB3FFE32223E1E6F92EB904379B" box="[151,288,1744,1769]" italics="true" pageId="6" pageNumber="49">Iranocichla</emphasis>
</taxonomicName>
(
<figureCitation id="132F2A24FFE32223E05EF931B950379B" box="[303,372,1743,1769]" captionStart="FIGURE 3" captionStartId="5.[151,255,1914,1938]" captionTargetBox="[248,1362,206,1893]" captionTargetId="figure@5.[219,1378,194,1904]" captionTargetPageId="5" captionText="FIGURE 3. Lower pharyngeal jaw (dorsal, posterior and lateral views), left mandible and isolated tooth in (a) Danakilia dinicolai, (b) Danakilia franchettii, (c) Iranocichla hormuzensis. Elements drawn in corresponding scale for each species. Posterior neurocranium and anterior vertebral elements, arrows indicate inferior vertebral apophysis of (d) Iranocichla hormuzensis." httpUri="https://zenodo.org/record/199532/files/figure.png" pageId="6" pageNumber="49">Fig. 3</figureCitation>
D).
</paragraph>
<paragraph id="8BAB36A1FFE32222E1B7F90ABA953050" blockId="6.[151,1437,1550,2031]" lastBlockId="7.[151,1436,152,290]" lastPageId="7" lastPageNumber="50" pageId="6" pageNumber="49">
<emphasis id="B960EAB3FFE32223E1B7F90AB98F367C" bold="true" box="[198,427,1780,1806]" pageId="6" pageNumber="49">Coloration in life.</emphasis>
Females and non-dominant males (
<figureCitation id="132F2A24FFE32223E211F90BBB8F367D" box="[864,939,1781,1807]" captionStart="FIGURE 4" captionStartId="6.[151,255,1486,1510]" captionTargetBox="[172,1414,216,1443]" captionTargetId="figure@6.[151,1436,194,1465]" captionTargetPageId="6" captionText="FIGURE 4. Danakilia dinicolai (a) wild caught female and (b) wild caught male. Specimens not preserved or measured." httpUri="https://zenodo.org/record/199533/files/figure.png" pageId="6" pageNumber="49">Fig. 4</figureCitation>
A). Opercule and body pale silvery-grey, snout and nape dark brownish-black. Dark scaleless spot on outer edge of opercle variously intense, but always present. Upper lip with bluish cast, lower lip grey. Throat and branchiostegal membranes pale silvergrey. Two dark longitudinal stripes sometimes visible on flanks, upper stripe at level of upper lateral line, passing to end of dorsal-fin base, lower stripe from edge of opercle to caudal base, not extending into caudal fin. Sometimes 5–7 dark vertical bars on body from dorsal-fin base to ventrum. Eye golden-yellow to brownish-black. Dominant males (
<figureCitation id="132F2A24FFE32223E35EF82BBA5C369D" box="[559,632,2005,2031]" captionStart="FIGURE 4" captionStartId="6.[151,255,1486,1510]" captionTargetBox="[172,1414,216,1443]" captionTargetId="figure@6.[151,1436,194,1465]" captionTargetPageId="6" captionText="FIGURE 4. Danakilia dinicolai (a) wild caught female and (b) wild caught male. Specimens not preserved or measured." httpUri="https://zenodo.org/record/199533/files/figure.png" pageId="6" pageNumber="49">Fig. 4</figureCitation>
B). Dorsal, caudal and anal fins pale blue, with numerous paler blue dots. White margin of dorsal fin flushed pale red-orange distally. Thin white margin on upper caudal lobe. Pelvics blue, paler distally. Body scales with iridescent, pale blue posterior margins, centrally grey to greyishblack. Scales on chest and belly silvery grey, with darker grey margins. Prominent frontal gibbosity blackishblue to greyish-blue. Snout and lips pale blue.
</paragraph>
<caption id="DF6B6629FFE22222E1E6F834BC703693" box="[151,1108,1994,2018]" httpUri="https://zenodo.org/record/199534/files/figure.png" pageId="7" pageNumber="50" targetBox="[193,1406,366,1959]" targetPageId="7">
<paragraph id="8BAB36A1FFE22222E1E6F834BC703693" blockId="7.[151,1108,1994,2018]" box="[151,1108,1994,2018]" pageId="7" pageNumber="50">
<emphasis id="B960EAB3FFE22222E1E6F834B93E3690" bold="true" box="[151,282,1994,2018]" pageId="7" pageNumber="50">FIGURE 5.</emphasis>
Geographic location and photograph of type locality, Lake Abaeded, Eritrea.
</paragraph>
</caption>
<paragraph id="8BAB36A1FFED222DE1B7FF69B9E4318E" blockId="8.[151,1437,151,1680]" pageId="8" pageNumber="51">
<emphasis id="B960EAB3FFED222DE1B7FF69BA3231C3" bold="true" box="[198,534,151,177]" pageId="8" pageNumber="51">Coloration in preservation</emphasis>
(
<figureCitation id="132F2A24FFED222DE359FF66BA5731C0" box="[552,627,152,178]" captionStart="FIGURE 2" captionStartId="3.[151,255,1398,1422]" captionTargetBox="[162,1432,883,1354]" captionTargetId="figure@3.[151,1436,850,1375]" captionTargetPageId="3" captionText="FIGURE 2. Danakilia dinicolai, holotype, MSNM 4888, male, 96.4 mm SL, Lake Abaeded, Eritrea." httpUri="https://zenodo.org/record/199531/files/figure.png" pageId="8" pageNumber="51">Fig. 2</figureCitation>
). Both sexes with head and body brown, darker dorsally, with dark lines or bars often visible as described in living specimens. Dorsal, anal and caudal fins heavily spotted in males, less so in females.
</paragraph>
<paragraph id="8BAB36A1FFED222DE1B7FEF9BDB830E0" blockId="8.[151,1437,151,1680]" pageId="8" pageNumber="51">
<emphasis id="B960EAB3FFED222DE1B7FEF9B9913053" bold="true" box="[198,437,263,289]" pageId="8" pageNumber="51">Feeding behaviour</emphasis>
. No field observations made, but gut morphology and contents suggests a diet primarily consisting of algae grazed from the surface of plants and rocks. Guts are highly coiled, and extend to 7.6 times SL, most are filled with floculent algae with some sand grains and vegetable debris. However, it is likely that this species is an opportunistic feeder, as individuals were caught on lines baited with grasshoppers.
</paragraph>
<paragraph id="8BAB36A1FFED222DE1B7FE63BD7C30C5" blockId="8.[151,1437,151,1680]" box="[198,1368,413,439]" pageId="8" pageNumber="51">
<emphasis id="B960EAB3FFED222DE1B7FE63B99830C5" bold="true" box="[198,444,413,439]" pageId="8" pageNumber="51">Breeding behaviour</emphasis>
. Unknown, but large males guarding pits over sand were observed in the field.
</paragraph>
</subSubSection>
<subSubSection id="C30E652AFFED222DE1B7FE3CBB7F3512" pageId="8" pageNumber="51" type="distribution">
<paragraph id="8BAB36A1FFED222DE1B7FE3CBB7F3512" blockId="8.[151,1437,151,1680]" pageId="8" pageNumber="51">
<emphasis id="B960EAB3FFED222DE1B7FE3CB9DE30AE" bold="true" box="[198,506,450,476]" pageId="8" pageNumber="51">Distribution and habitat</emphasis>
(
<figureCitation id="132F2A24FFED222DE37AFE3CBA7030AE" box="[523,596,450,476]" captionStart="FIGURE 5" captionStartId="7.[151,255,1994,2018]" captionTargetBox="[193,1406,366,1959]" captionTargetId="figure@7.[186,1410,343,2000]" captionTargetPageId="7" captionText="FIGURE 5. Geographic location and photograph of type locality, Lake Abaeded, Eritrea." httpUri="https://zenodo.org/record/199534/files/figure.png" pageId="8" pageNumber="51">Fig. 5</figureCitation>
). Lake Abaeded occupies a small oval depression (maximum width of
<quantity id="4CEC9B44FFED222DE1E6FE16B8C23370" box="[151,230,488,514]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="51" unit="m" value="450.0">450 m</quantity>
, maximum depth
<quantity id="4CEC9B44FFED222DE0CFFE16B9D93370" box="[446,509,488,514]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="51" unit="m" value="10.0">10 m</quantity>
) in a volcanic crater at the northwestern border of the Danakil Depression in
<collectingCountry id="F3037631FFED222DE1E6FDF3B8CE3355" box="[151,234,525,551]" name="Eritrea" pageId="8" pageNumber="51">Eritrea</collectingCountry>
,
<quantity id="4CEC9B44FFED222DE187FDF3B96B3355" box="[246,335,525,551]" metricMagnitude="5" metricUnit="m" metricValue="1.3199999999999998" pageId="8" pageNumber="51" unit="km" value="132.0">132 km</quantity>
southeast of the coastal town of Massawa. Lake surface is about
<quantity id="4CEC9B44FFED222DE526FDF3BCB73355" box="[1111,1171,525,551]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="51" unit="m" value="30.0">30 m</quantity>
below sea level. Sand from dunes along the eastern and southern rim is frequently blown into the lake. The lake is fed by many hot springs at surface level and underwater, and as there is no surface outlet evaporation in this hyper arid region (rainfall of ~
<quantity id="4CEC9B44FFED222DE041FD83B97933E5" box="[304,349,637,663]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="51" unit="m" value="5.0">5 m</quantity>
/year) probably fully compensates for the input from springs to maintain a stable lake level year round. In
<date id="FFAA1061FFED222DE030FD5CB9E533CE" box="[321,449,674,700]" pageId="8" pageNumber="51" value="1998-04">April 1998</date>
water temperature was 29°C but reached 45°C close to the hot springs. Total salinity was 14.5 gr/l. The most abundant ions were Na+ (111 EPM), Ca++ (99 EPM), Mg++ (18 EPM), Cl - (217 EPM) and SO4 - (41 EPM).
<collectingCountry id="F3037631FFED222DE0E5FD0DB990327F" box="[404,436,755,781]" name="Philippines" pageId="8" pageNumber="51">Ph</collectingCountry>
was 7.46 and EC was 23100. High concentrations of Na+, Ca++, Cl - and SO4 - are certainly the result of circulation of water heated at depth through layers of gypsum and halite that cover the bottom of the northern part of the Danakil Depression (
<bibRefCitation id="EF854B50FFED222DE220FCBFBC783229" author="Gasse" box="[849,1116,833,859]" pageId="8" pageNumber="51" refString="Gasse, F. &amp; Street, F. A. (1978). Late Quaternary lake level fluctuations and environments of the Northern Rift Valley and Afar Region (Ethiopia and Djibouti). Paleogeography, Paleoclimatology, Paleoecology, 24, 279 - 325." type="journal article" year="1978">Gasse &amp; Street 1978</bibRefCitation>
). The lakeshore is almost completely encircled by reeds (
<taxonomicName id="4C144D22FFED222DE378FC98BADB320D" box="[521,767,870,895]" class="Liliopsida" family="Poaceae" genus="Phragmites" kingdom="Plantae" order="Poales" pageId="8" pageNumber="51" phylum="Tracheophyta" rank="species" species="australis">
<emphasis id="B960EAB3FFED222DE378FC98BADB320D" box="[521,767,870,895]" italics="true" pageId="8" pageNumber="51">Phragmites australis</emphasis>
</taxonomicName>
) and stands of tamarisk (
<taxonomicName id="4C144D22FFED222DE55AFC98BCC8320D" box="[1067,1260,870,895]" class="Magnoliopsida" family="Tamaricaceae" genus="Tamarix" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="8" pageNumber="51" phylum="Magnoliophyta" rank="species" species="aphylla">
<emphasis id="B960EAB3FFED222DE55AFC98BCC8320D" box="[1067,1260,870,895]" italics="true" pageId="8" pageNumber="51">Tamarix aphylla</emphasis>
</taxonomicName>
), but is free of vegetation in the proximity of hot springs. A single non-piscine vertebrate inhabiting the lake is a frog, probably
<taxonomicName id="4C144D22FFED222DE07AFC4FBA3232B8" box="[267,534,945,970]" class="Amphibia" family="Ptychadenidae" genus="Ptychadena" kingdom="Animalia" order="Anura" pageId="8" pageNumber="51" phylum="Chordata" rank="species" species="anchietae">
<emphasis id="B960EAB3FFED222DE07AFC4FBA3232B8" box="[267,534,945,970]" italics="true" pageId="8" pageNumber="51">Ptychadena anchietae</emphasis>
</taxonomicName>
. Fishes are abundant in the lake and concentrate among the reeds, but are found also on sandy substrate near the shore, where temperature is not too high. The bottom of the sandy areas is covered with crater-like nests dug and defended by large colourful males. The species has been found so far only in Lake Abaeded, but no research has yet been conducted in the nearby Lake Badda, a variable surface reed-fringed lake only
<quantity id="4CEC9B44FFED222DE0D3FBB8B9FA3512" box="[418,478,1094,1120]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="51" unit="km" value="8.0">8 km</quantity>
north of Lake Abaeded (
<figureCitation id="132F2A24FFED222DE275FBB8BB6F3512" box="[772,843,1094,1120]" captionStart="FIGURE 5" captionStartId="7.[151,255,1994,2018]" captionTargetBox="[193,1406,366,1959]" captionTargetId="figure@7.[186,1410,343,2000]" captionTargetPageId="7" captionText="FIGURE 5. Geographic location and photograph of type locality, Lake Abaeded, Eritrea." httpUri="https://zenodo.org/record/199534/files/figure.png" pageId="8" pageNumber="51">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30E652AFFED222DE1B7FB95BB1435D9" pageId="8" pageNumber="51" type="etymology">
<paragraph id="8BAB36A1FFED222DE1B7FB95BB1435D9" blockId="8.[151,1437,151,1680]" pageId="8" pageNumber="51">
<emphasis id="B960EAB3FFED222DE1B7FB95B97735F7" bold="true" box="[198,339,1131,1157]" pageId="8" pageNumber="51">Etymology.</emphasis>
Named in memory of Ernesto
<collectingCountry id="F3037631FFED222DE3B0FB95BAFA35F7" box="[705,734,1131,1157]" name="Iceland" pageId="8" pageNumber="51">Di</collectingCountry>
Nicola, a member of the 2001 expedition to Lake Abaeded who died in a car accident while returning from the lake.
</paragraph>
</subSubSection>
<subSubSection id="C30E652AFFED222DE1B7FB4BBDA937E2" pageId="8" pageNumber="51" type="discussion">
<paragraph id="8BAB36A1FFED222DE1B7FB4BBDA937E2" blockId="8.[151,1437,151,1680]" pageId="8" pageNumber="51">
<emphasis id="B960EAB3FFED222DE1B7FB4BB97535BD" bold="true" box="[198,337,1205,1231]" pageId="8" pageNumber="51">Discussion.</emphasis>
The discovery of a second species of
<taxonomicName id="4C144D22FFED222DE27BFB48BBA435BD" box="[778,896,1206,1231]" class="Actinopterygii" family="Cichlidae" genus="Danakilia" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="51" phylum="Chordata" rank="genus">
<emphasis id="B960EAB3FFED222DE27BFB48BBA435BD" box="[778,896,1206,1231]" italics="true" pageId="8" pageNumber="51">Danakilia</emphasis>
</taxonomicName>
in a small lake currently isolated by some
<quantity id="4CEC9B44FFED222DE401FB48B8993587" metricMagnitude="5" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="51" unit="km" value="160.0">160 km</quantity>
of hyper-arid desert from its congener,
<taxonomicName id="4C144D22FFED222DE3E6FB25BBB43586" box="[663,912,1243,1268]" class="Actinopterygii" family="Cichlidae" genus="Danakilia" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="51" phylum="Chordata" rank="species" species="franchettii">
<emphasis id="B960EAB3FFED222DE3E6FB25BBB43586" box="[663,912,1243,1268]" italics="true" pageId="8" pageNumber="51">Danakilia franchettii</emphasis>
</taxonomicName>
, raises the question of the age and origin of these two isolated species. The geological history of the region indicates that the Danakil Depression formed as a result of tectonic movements related to the Afro-Arabian Rift (
<bibRefCitation id="EF854B50FFED222DE2B1FAD8BD3E3432" author="Beyene" box="[960,1306,1318,1344]" pageId="8" pageNumber="51" refString="Beyene, A. &amp; Abdelsalam, M. G. (2005) Tectonics of the Afar depression: A review and synthesis. Journal of African Earth Sciences, 41, 41 - 59." type="journal article" year="2005">Beyene &amp; Abdelsalam 2005</bibRefCitation>
). It was an intermittent marine basin (at least in the northern region) during the late Pleistocene, and with the formation of volcanic structures at the northern end it is thought to have separated from the Red Sea sometime around 32,000–25,000 years ago (
<bibRefCitation id="EF854B50FFED222DE0ADFA68BAA434C2" author="Bonatti" box="[476,640,1430,1456]" pageId="8" pageNumber="51" refString="Bonatti, E., Emiliani, C., Ostlund, G. &amp; Rydell, H. (1971) Final desiccation of the Afar Rift. Science, 172, 468 - 469." type="journal article" year="1971">Bonatti 1971</bibRefCitation>
,
<bibRefCitation id="EF854B50FFED222DE3FCFA68BB5E34C2" author="Barberi" box="[653,890,1430,1456]" pageId="8" pageNumber="51" refString="Barberi, F., Borsi, S., Ferrara, G., Marinelli, G., Santacroce, R., Tazieff, H. &amp; Varet, J. (1972) Evolution of the Danakil Depression (Afar, Ethiopia) in Light of Radiometric Age. Journal of Geology, 80, 720 - 729." type="journal article" year="1972">
Barberi
<emphasis id="B960EAB3FFED222DE380FA68BB0B34DD" box="[753,815,1430,1455]" italics="true" pageId="8" pageNumber="51">et al.</emphasis>
1972
</bibRefCitation>
). Separation of the two species of
<taxonomicName id="4C144D22FFED222DE452FA68BDB934DD" box="[1315,1437,1430,1455]" class="Actinopterygii" family="Cichlidae" genus="Danakilia" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="51" phylum="Chordata" rank="genus">
<emphasis id="B960EAB3FFED222DE452FA68BDB934DD" box="[1315,1437,1430,1455]" italics="true" pageId="8" pageNumber="51">Danakilia</emphasis>
</taxonomicName>
might therefore date back to the Pleistocene to the time of the formation of the Alayta e Tat'Ale shield volcanoes that surround Lake Afdera (
<bibRefCitation id="EF854B50FFED222DE324FA1FBB113489" author="Barberi" box="[597,821,1505,1531]" pageId="8" pageNumber="51" refString="Barberi, F., Borsi, S., Ferrara, G., Marinelli, G., Santacroce, R., Tazieff, H. &amp; Varet, J. (1972) Evolution of the Danakil Depression (Afar, Ethiopia) in Light of Radiometric Age. Journal of Geology, 80, 720 - 729." type="journal article" year="1972">
Barberi
<emphasis id="B960EAB3FFED222DE3C4FA1FBAD43488" box="[693,752,1505,1530]" italics="true" pageId="8" pageNumber="51">et al.</emphasis>
1972
</bibRefCitation>
). Alternatively, their separation may be considerably younger having occurred after the drying of extensive saline lakes that occupied the bottom of the Danakil Depression during a major arid phase that started about 9,000–5,000 years ago (
<bibRefCitation id="EF854B50FFED222DE53EF9D5BCF93737" author="Gasse" box="[1103,1245,1579,1605]" pageId="8" pageNumber="51" refString="Gasse, F. (1974) Les Diatomees des Sediments Holocenes du Bassin du Lac Afrera (Giulietti) (Afar Septentrional, Ethiopie). Essai de Reconstitution de l'Evolution du Milieu. Internationale Revue der gesamten Hydrobiologie und Hydrographie, 59, 95 - 122." type="journal article" year="1974">Gasse 1974</bibRefCitation>
,
<bibRefCitation id="EF854B50FFED222DE59BF9D5B8F33719" author="Gasse" pageId="8" pageNumber="51" refString="Gasse, F. &amp; Street, F. A. (1978). Late Quaternary lake level fluctuations and environments of the Northern Rift Valley and Afar Region (Ethiopia and Djibouti). Paleogeography, Paleoclimatology, Paleoecology, 24, 279 - 325." type="journal article" year="1978">Gasse &amp; Street 1978</bibRefCitation>
). Unfortunately, despite considerable effort all attempts to extract DNA from either species have so far failed, and molecular divergence time estimates cannot be made until additional material becomes available.
</paragraph>
</subSubSection>
</treatment>
</document>
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