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<document id="2FBE64D6322233FBBCD6D7D8DF4B7935" ID-CLB-Dataset="32309" ID-DOI="10.11646/zootaxa.4311.3.9" ID-GBIF-Dataset="3bc96c5f-7556-4468-8370-b4c23237b757" ID-ISSN="1175-5326" ID-Zenodo-Dep="847968" ID-ZooBank="D8464F8A-9123-4Afe-9C30-B63A026C1D59" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1503550765495" checkinUser="plazi" docAuthor="Baňař, Petr" docDate="2017" docId="0399C531FF80FFE39FF7D5EC340BF933" docLanguage="en" docName="zootaxa.4311.3.9.pdf" docOrigin="Zootaxa 4311 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Madastiger Baňař 2017, gen. nov." docType="treatment" docVersion="6" lastPageNumber="436" masterDocId="FFA0BD49FF81FFE29F60D57B3645FFEC" masterDocTitle="The Pselaphinae (Coleoptera: Staphylinidae) of Madagascar. VII. Description of Madastiger robustus gen. and sp. nov. from Central Madagascar" masterLastPageNumber="440" masterPageNumber="435" pageNumber="436" updateTime="1698467240106" updateUser="ExternalLinkService">
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<mods:title id="C1207B02DF2BC2AEE779ED19513118CE">The Pselaphinae (Coleoptera: Staphylinidae) of Madagascar. VII. Description of Madastiger robustus gen. and sp. nov. from Central Madagascar</mods:title>
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<mods:name id="F2E45B523E0D0D6CCA76605B6B35144A" type="personal">
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<mods:namePart id="612B4DBB840A9F2E097218C59F3A50AB">Baňař, Petr</mods:namePart>
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<mods:title id="9733DC9EEBA49850327EE4C2387E8CFE">Zootaxa</mods:title>
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<mods:date id="53A5214FEA9C52A99493649DC86CD85A">2017</mods:date>
<mods:detail id="C6B1193953E55319872932ACF8710590" type="pubDate">
<mods:number id="89E4D6750A49B1E42BB5092DB562A7CC">2017-08-24</mods:number>
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<mods:number id="636A7669792360C279FBD83C0CC9DB74">4311</mods:number>
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<mods:classification id="8FB483DBC21C3CC10BC0542B03F829EF">journal article</mods:classification>
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<mods:identifier id="3BED8DB8D762593B8D20507E2A99BF33" type="DOI">10.11646/zootaxa.4311.3.9</mods:identifier>
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<mods:identifier id="FD109908321B254F95F891AB932F51BD" type="ISSN">1175-5326</mods:identifier>
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<treatment id="0399C531FF80FFE39FF7D5EC340BF933" ID-DOI="http://doi.org/10.5281/zenodo.5999834" ID-GBIF-Taxon="132923486" ID-Zenodo-Dep="5999834" LSID="urn:lsid:plazi:treatment:0399C531FF80FFE39FF7D5EC340BF933" httpUri="http://treatment.plazi.org/id/0399C531FF80FFE39FF7D5EC340BF933" lastPageNumber="436" pageId="1" pageNumber="436">
<subSubSection id="C32A27ACFF80FFE39FF7D5EC375AFF3F" pageId="1" pageNumber="436" type="nomenclature">
<paragraph id="8B8F7427FF80FFE39FF7D5EC342CFF5E" blockId="1.[151,617,151,211]" box="[151,617,151,178]" pageId="1" pageNumber="436">
<heading id="D0C7C34BFF80FFE39FF7D5EC342CFF5E" bold="true" box="[151,617,151,178]" fontSize="11" level="1" pageId="1" pageNumber="436" reason="1">
<emphasis id="B944A835FF80FFE39FF7D5EC342CFF5E" bold="true" box="[151,617,151,178]" pageId="1" pageNumber="436">
<taxonomicName id="4C300FA4FF80FFE39FF7D5EC37B1FF5E" ID-CoL="8XMKG" authority="Hlavac &amp; Banar" authorityName="Baňař" authorityYear="2017" box="[151,500,151,178]" class="Insecta" family="Staphylinidae" genus="Madastiger" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B944A835FF80FFE39FF7D5EC3765FF5D" bold="true" box="[151,288,151,177]" italics="true" pageId="1" pageNumber="436">Madastiger</emphasis>
Hlaváč &amp; Baňař
</taxonomicName>
<taxonomicNameLabel id="A277154EFF80FFE39E9BD5E3342CFF5E" box="[507,617,152,178]" pageId="1" pageNumber="436" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F7427FF80FFE39FF7D5C1375AFF3F" blockId="1.[151,617,151,211]" box="[151,287,186,211]" pageId="1" pageNumber="436">
(
<figureCitation id="130B68A2FF80FFE39FFFD5C13753FF3F" box="[159,278,186,211]" captionStart-0="FIGURES 1 – 5" captionStart-1="FIGURES 6 – 9" captionStart-2="FIGURES 10 – 11" captionStartId-0="2.[151,264,1968,1991]" captionStartId-1="3.[151,264,1796,1819]" captionStartId-2="4.[151,264,1972,1995]" captionTargetBox-0="[177,1415,195,1955]" captionTargetBox-1="[151,1413,210,1721]" captionTargetBox-2="[478,1107,210,1948]" captionTargetId-0="figure@2.[177,1415,195,1955]" captionTargetId-1="figure@3.[151,1427,210,1731]" captionTargetId-2="figure@4.[473,1119,193,1965]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1 – 5. Madastiger robustus gen. et sp. nov. Hlaváč &amp; Baňař, male holotype. 1, dorsal habitus; 2, head, dorsal view; 3, abdomen, dorsal view; 4, left elytron, dorsal view; 5, right antenna, latero-dorsal view. Scale bars in mm." captionText-1="FIGURES 6 – 9. Madastiger robustus gen. et sp. nov. Hlaváč &amp; Baňař, male holotype. 6, abdomen, dorsal view; 7, abdomen, ventral view; 8, inflated aedeagus, lateral view; 9, accessory lobes with long setae, apical view. Lettering: blf = basilateral fovea; pt I, II, III = paratergite I, II, III. Scale bars in mm, accessory lobes (Fig. 9) not measured in this view." captionText-2="FIGURES 10 – 11. Madastiger robustus gen. et sp. nov. Hlaváč &amp; Baňař, male holotype. 10, aedeagus, ventral view; 11, accessory lobes of aedeagus, dorsal view, not measured. Scale bar in mm." httpUri-0="https://zenodo.org/record/847970/files/figure.png" httpUri-1="https://zenodo.org/record/847972/files/figure.png" httpUri-2="https://zenodo.org/record/847974/files/figure.png" pageId="1" pageNumber="436">Figs. 1–11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32A27ACFF80FFE39FF7D4793529FEF7" box="[151,876,258,283]" pageId="1" pageNumber="436" type="materials_examined">
<paragraph id="8B8F7427FF80FFE39FF7D4793529FEF7" blockId="1.[151,1437,258,1759]" box="[151,876,258,283]" pageId="1" pageNumber="436">
<emphasis id="B944A835FF80FFE39FF7D4793771FEF7" bold="true" box="[151,308,258,283]" pageId="1" pageNumber="436">Type species:</emphasis>
<taxonomicName id="4C300FA4FF80FFE39E5BD4793467FEF7" authority="Baňař, 2017" authorityName="Baňař" authorityYear="2017" box="[315,546,258,283]" class="Insecta" family="Staphylinidae" genus="Madastiger" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
<emphasis id="B944A835FF80FFE39E5BD4793467FEF7" box="[315,546,258,283]" italics="true" pageId="1" pageNumber="436">Madastiger robustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A277154EFF80FFE39D4AD47934C6FEF7" box="[554,643,258,283]" pageId="1" pageNumber="436" rank="species">
<emphasis id="B944A835FF80FFE39D4AD47934C6FEF7" bold="true" box="[554,643,258,283]" pageId="1" pageNumber="436">sp. nov.</emphasis>
</taxonomicNameLabel>
Present designation.
</paragraph>
</subSubSection>
<subSubSection id="C32A27ACFF80FFE39FA7D45D346BFE1F" pageId="1" pageNumber="436" type="diagnosis">
<paragraph id="8B8F7427FF80FFE39FA7D45D346BFE1F" blockId="1.[151,1437,258,1759]" pageId="1" pageNumber="436">
<emphasis id="B944A835FF80FFE39FA7D45D3704FED3" bold="true" box="[199,321,294,319]" pageId="1" pageNumber="436">Diagnosis.</emphasis>
Body length
<quantity id="4CC8D9C2FF80FFE39EBCD45D3401FED2" box="[476,580,294,319]" metricMagnitude="-3" metricUnit="m" metricValue="1.58" pageId="1" pageNumber="436" unit="mm" value="1.58">1.58 mm</quantity>
; head pentagonal, slightly tapered anteriad, rostrum triangular, head clearly narrower than pronotum, head with laterally located vertexal foveae, frontal fovea absent, pronotum with antebasal foveae, elongate median sulcus, and lateral foveae; antennae with 4 antennomeres, terminal antennomere cylindrical, truncate at apex; elytra with 2–3 evanescent foveae, with very weakly defined elytral striae, lacking trichomes; abdomen slightly shorter than elytra, composite tergite (IV–VI) with transverse impression and trichomes, both hidden under elytra.
</paragraph>
</subSubSection>
<subSubSection id="C32A27ACFF80FFE39FA7D48532BEFAE7" pageId="1" pageNumber="436" type="description">
<paragraph id="8B8F7427FF80FFE39FA7D4853795FBE3" blockId="1.[151,1437,258,1759]" pageId="1" pageNumber="436">
<emphasis id="B944A835FF80FFE39FA7D485371CFDFB" bold="true" box="[199,345,510,535]" pageId="1" pageNumber="436">Description.</emphasis>
Body (
<figureCitation id="130B68A2FF80FFE39ECDD48537AAFDFB" box="[429,495,510,535]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1968,1991]" captionTargetBox="[177,1415,195,1955]" captionTargetId="figure@2.[177,1415,195,1955]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Madastiger robustus gen. et sp. nov. Hlaváč &amp; Baňař, male holotype. 1, dorsal habitus; 2, head, dorsal view; 3, abdomen, dorsal view; 4, left elytron, dorsal view; 5, right antenna, latero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/847970/files/figure.png" pageId="1" pageNumber="436">Fig. 1</figureCitation>
) uniformly reddish-brown, sparsely covered with very short appressed, posteriorly directed setae, head coarsely punctured, pronotum and elytra with some discal coarse punctures, body length
<quantity id="4CC8D9C2FF80FFE39A0BD7593680FDB2" metricMagnitude="-3" metricUnit="m" metricValue="1.58" pageId="1" pageNumber="436" unit="mm" value="1.58">1.58 mm</quantity>
, maximum width at posterior corners of elytra
<quantity id="4CC8D9C2FF80FFE39DB0D73C3572FDB2" box="[720,823,583,607]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="436" unit="mm" value="0.8">0.80 mm</quantity>
. Head (
<figureCitation id="130B68A2FF80FFE39CEBD73D358BFDB3" box="[907,974,582,607]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1968,1991]" captionTargetBox="[177,1415,195,1955]" captionTargetId="figure@2.[177,1415,195,1955]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Madastiger robustus gen. et sp. nov. Hlaváč &amp; Baňař, male holotype. 1, dorsal habitus; 2, head, dorsal view; 3, abdomen, dorsal view; 4, left elytron, dorsal view; 5, right antenna, latero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/847970/files/figure.png" pageId="1" pageNumber="436">Fig. 2</figureCitation>
) slightly shorter than wide, widest across eyes, evenly tapered anteriad, rostrum anteriorly triangular, with well-defined lateral vertexal foveae. Antennae (
<figureCitation id="130B68A2FF80FFE39FFFD7F536A1FD4B" box="[159,228,654,679]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1968,1991]" captionTargetBox="[177,1415,195,1955]" captionTargetId="figure@2.[177,1415,195,1955]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Madastiger robustus gen. et sp. nov. Hlaváč &amp; Baňař, male holotype. 1, dorsal habitus; 2, head, dorsal view; 3, abdomen, dorsal view; 4, left elytron, dorsal view; 5, right antenna, latero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/847970/files/figure.png" pageId="1" pageNumber="436">Fig. 5</figureCitation>
) with four antennomeres, about as long as head, partly hidden in antennal cavity; scapes minuscule, not visible dorsally, pedicels much larger than scapes and larger than antennomeres III, terminal antenomeres short, at apex truncate, with stout dense setae, more than 4 times as long as antennomere III. Pronotum slightly wider than long and slightly longer than head, widest at level of projected posterior corners; antebasal fovea, elongate median sulcus, and lateral foveae present. Elytra (
<figureCitation id="130B68A2FF80FFE39D0CD66534EAFCDB" box="[620,687,798,823]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1968,1991]" captionTargetBox="[177,1415,195,1955]" captionTargetId="figure@2.[177,1415,195,1955]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Madastiger robustus gen. et sp. nov. Hlaváč &amp; Baňař, male holotype. 1, dorsal habitus; 2, head, dorsal view; 3, abdomen, dorsal view; 4, left elytron, dorsal view; 5, right antenna, latero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/847970/files/figure.png" pageId="1" pageNumber="436">Fig. 4</figureCitation>
) robust, wider than long, with 2–3 evanescent basal foveae, with 4 rows of uneven coarse punctures, lacking trichomes. Abdomen (
<figureCitation id="130B68A2FF80FFE39C11D6393584FCB7" box="[881,961,834,859]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1968,1991]" captionTargetBox="[177,1415,195,1955]" captionTargetId="figure@2.[177,1415,195,1955]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Madastiger robustus gen. et sp. nov. Hlaváč &amp; Baňař, male holotype. 1, dorsal habitus; 2, head, dorsal view; 3, abdomen, dorsal view; 4, left elytron, dorsal view; 5, right antenna, latero-dorsal view. Scale bars in mm." httpUri="https://zenodo.org/record/847970/files/figure.png" pageId="1" pageNumber="436">Figs. 3</figureCitation>
,
<figureCitation id="130B68A2FF80FFE39CAFD63935BEFCB7" box="[975,1019,834,859]" captionStart="FIGURES 6 – 9" captionStartId="3.[151,264,1796,1819]" captionTargetBox="[151,1413,210,1721]" captionTargetId="figure@3.[151,1427,210,1731]" captionTargetPageId="3" captionText="FIGURES 6 – 9. Madastiger robustus gen. et sp. nov. Hlaváč &amp; Baňař, male holotype. 6, abdomen, dorsal view; 7, abdomen, ventral view; 8, inflated aedeagus, lateral view; 9, accessory lobes with long setae, apical view. Lettering: blf = basilateral fovea; pt I, II, III = paratergite I, II, III. Scale bars in mm, accessory lobes (Fig. 9) not measured in this view." httpUri="https://zenodo.org/record/847972/files/figure.png" pageId="1" pageNumber="436">6, 7</figureCitation>
) short, slightly narrower than elytra, composite tergite with basal, heavily sclerotized impression and robust lateral trichomes (
<figureCitation id="130B68A2FF80FFE39BE4D61D328CFC93" box="[1156,1225,870,895]" captionStart="FIGURES 6 – 9" captionStartId="3.[151,264,1796,1819]" captionTargetBox="[151,1413,210,1721]" captionTargetId="figure@3.[151,1427,210,1731]" captionTargetPageId="3" captionText="FIGURES 6 – 9. Madastiger robustus gen. et sp. nov. Hlaváč &amp; Baňař, male holotype. 6, abdomen, dorsal view; 7, abdomen, ventral view; 8, inflated aedeagus, lateral view; 9, accessory lobes with long setae, apical view. Lettering: blf = basilateral fovea; pt I, II, III = paratergite I, II, III. Scale bars in mm, accessory lobes (Fig. 9) not measured in this view." httpUri="https://zenodo.org/record/847972/files/figure.png" pageId="1" pageNumber="436">Fig. 6</figureCitation>
), with both hidden under elytra in undissected specimen, first three visible paratergites well-developed, paratergites I with minute trichomes (
<figureCitation id="130B68A2FF80FFE39E7BD6D53720FC2B" box="[283,357,942,967]" captionStart="FIGURES 6 – 9" captionStartId="3.[151,264,1796,1819]" captionTargetBox="[151,1413,210,1721]" captionTargetId="figure@3.[151,1427,210,1731]" captionTargetPageId="3" captionText="FIGURES 6 – 9. Madastiger robustus gen. et sp. nov. Hlaváč &amp; Baňař, male holotype. 6, abdomen, dorsal view; 7, abdomen, ventral view; 8, inflated aedeagus, lateral view; 9, accessory lobes with long setae, apical view. Lettering: blf = basilateral fovea; pt I, II, III = paratergite I, II, III. Scale bars in mm, accessory lobes (Fig. 9) not measured in this view." httpUri="https://zenodo.org/record/847972/files/figure.png" pageId="1" pageNumber="436">Fig. 6</figureCitation>
). First visible sternite (
<figureCitation id="130B68A2FF80FFE39D1FD6D5348CFC2B" box="[639,713,942,967]" captionStart="FIGURES 6 – 9" captionStartId="3.[151,264,1796,1819]" captionTargetBox="[151,1413,210,1721]" captionTargetId="figure@3.[151,1427,210,1731]" captionTargetPageId="3" captionText="FIGURES 6 – 9. Madastiger robustus gen. et sp. nov. Hlaváč &amp; Baňař, male holotype. 6, abdomen, dorsal view; 7, abdomen, ventral view; 8, inflated aedeagus, lateral view; 9, accessory lobes with long setae, apical view. Lettering: blf = basilateral fovea; pt I, II, III = paratergite I, II, III. Scale bars in mm, accessory lobes (Fig. 9) not measured in this view." httpUri="https://zenodo.org/record/847972/files/figure.png" pageId="1" pageNumber="436">Fig. 7</figureCitation>
) slightly shorter than second (IV), second with well-defined basolateral foveae (
<figureCitation id="130B68A2FF80FFE39E19D6A93784FC06" box="[377,449,978,1003]" captionStart="FIGURES 6 – 9" captionStartId="3.[151,264,1796,1819]" captionTargetBox="[151,1413,210,1721]" captionTargetId="figure@3.[151,1427,210,1731]" captionTargetPageId="3" captionText="FIGURES 6 – 9. Madastiger robustus gen. et sp. nov. Hlaváč &amp; Baňař, male holotype. 6, abdomen, dorsal view; 7, abdomen, ventral view; 8, inflated aedeagus, lateral view; 9, accessory lobes with long setae, apical view. Lettering: blf = basilateral fovea; pt I, II, III = paratergite I, II, III. Scale bars in mm, accessory lobes (Fig. 9) not measured in this view." httpUri="https://zenodo.org/record/847972/files/figure.png" pageId="1" pageNumber="436">Fig. 7</figureCitation>
,
<emphasis id="B944A835FF80FFE39EB1D6A937ABFC07" box="[465,494,978,1003]" italics="true" pageId="1" pageNumber="436">blf</emphasis>
), slightly longer than third visible sternite (V), third (V) and fourth (VI) visible sternites subequal in length.
</paragraph>
<paragraph id="8B8F7427FF80FFE39FA7D1603696FBBA" blockId="1.[151,1437,258,1759]" pageId="1" pageNumber="436">Legs short and stout, tibiae about as long as femora, mesotibiae with preapical spurs, metatibiae curved before apex.</paragraph>
<paragraph id="8B8F7427FF80FFE39FA7D11937EAFB0B" blockId="1.[151,1437,258,1759]" pageId="1" pageNumber="436">
Aedeagus (
<figureCitation id="130B68A2FF80FFE39E25D11937F8FB97" box="[325,445,1122,1147]" captionStart-0="FIGURES 6 – 9" captionStart-1="FIGURES 10 – 11" captionStartId-0="3.[151,264,1796,1819]" captionStartId-1="4.[151,264,1972,1995]" captionTargetBox-0="[151,1413,210,1721]" captionTargetBox-1="[478,1107,210,1948]" captionTargetId-0="figure@3.[151,1427,210,1731]" captionTargetId-1="figure@4.[473,1119,193,1965]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 6 – 9. Madastiger robustus gen. et sp. nov. Hlaváč &amp; Baňař, male holotype. 6, abdomen, dorsal view; 7, abdomen, ventral view; 8, inflated aedeagus, lateral view; 9, accessory lobes with long setae, apical view. Lettering: blf = basilateral fovea; pt I, II, III = paratergite I, II, III. Scale bars in mm, accessory lobes (Fig. 9) not measured in this view." captionText-1="FIGURES 10 – 11. Madastiger robustus gen. et sp. nov. Hlaváč &amp; Baňař, male holotype. 10, aedeagus, ventral view; 11, accessory lobes of aedeagus, dorsal view, not measured. Scale bar in mm." httpUri-0="https://zenodo.org/record/847972/files/figure.png" httpUri-1="https://zenodo.org/record/847974/files/figure.png" pageId="1" pageNumber="436">Figs. 8–11</figureCitation>
) symmetrical, parameres fused to median lobe, basal bulb about as long as apical lobe, which is tapered to apex, diaphragm opening large, oval. Apex of apical lobe with pair of accessory lobes bearing dense fringe of long setae (
<figureCitation id="130B68A2FF80FFE39EADD1D13456FB2F" box="[461,531,1194,1219]" captionStart="FIGURES 6 – 9" captionStartId="3.[151,264,1796,1819]" captionTargetBox="[151,1413,210,1721]" captionTargetId="figure@3.[151,1427,210,1731]" captionTargetPageId="3" captionText="FIGURES 6 – 9. Madastiger robustus gen. et sp. nov. Hlaváč &amp; Baňař, male holotype. 6, abdomen, dorsal view; 7, abdomen, ventral view; 8, inflated aedeagus, lateral view; 9, accessory lobes with long setae, apical view. Lettering: blf = basilateral fovea; pt I, II, III = paratergite I, II, III. Scale bars in mm, accessory lobes (Fig. 9) not measured in this view." httpUri="https://zenodo.org/record/847972/files/figure.png" pageId="1" pageNumber="436">Fig. 9</figureCitation>
,
<figureCitation id="130B68A2FF80FFE39D41D1D0347BFB2F" box="[545,574,1195,1219]" captionStart="FIGURES 10 – 11" captionStartId="4.[151,264,1972,1995]" captionTargetBox="[478,1107,210,1948]" captionTargetId="figure@4.[473,1119,193,1965]" captionTargetPageId="4" captionText="FIGURES 10 – 11. Madastiger robustus gen. et sp. nov. Hlaváč &amp; Baňař, male holotype. 10, aedeagus, ventral view; 11, accessory lobes of aedeagus, dorsal view, not measured. Scale bar in mm." httpUri="https://zenodo.org/record/847974/files/figure.png" pageId="1" pageNumber="436">11</figureCitation>
). Proximal two-thirds of inflated endophallus bare, distal third with heavily sclerotized teeth (
<figureCitation id="130B68A2FF80FFE39E3DD1B537E5FB0B" box="[349,416,1230,1255]" captionStart="FIGURES 6 – 9" captionStartId="3.[151,264,1796,1819]" captionTargetBox="[151,1413,210,1721]" captionTargetId="figure@3.[151,1427,210,1731]" captionTargetPageId="3" captionText="FIGURES 6 – 9. Madastiger robustus gen. et sp. nov. Hlaváč &amp; Baňař, male holotype. 6, abdomen, dorsal view; 7, abdomen, ventral view; 8, inflated aedeagus, lateral view; 9, accessory lobes with long setae, apical view. Lettering: blf = basilateral fovea; pt I, II, III = paratergite I, II, III. Scale bars in mm, accessory lobes (Fig. 9) not measured in this view." httpUri="https://zenodo.org/record/847972/files/figure.png" pageId="1" pageNumber="436">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8B8F7427FF80FFE39FA7D18932BEFAE7" blockId="1.[151,1437,258,1759]" box="[199,1275,1266,1291]" pageId="1" pageNumber="436">
<emphasis id="B944A835FF80FFE39FA7D18937F2FAE7" bold="true" box="[199,439,1266,1291]" pageId="1" pageNumber="436">Sexual dimorphism.</emphasis>
Female unknown but most probably lacking preapical spur on mesotibiae.
</paragraph>
</subSubSection>
<subSubSection id="C32A27ACFF80FFE39FA7D06D3758F99F" pageId="1" pageNumber="436" type="diagnosis">
<paragraph id="8B8F7427FF80FFE39FA7D06D3758F99F" blockId="1.[151,1437,258,1759]" pageId="1" pageNumber="436">
<emphasis id="B944A835FF80FFE39FA7D06D3797FAC3" bold="true" box="[199,466,1302,1327]" pageId="1" pageNumber="436">Differential diagnosis:</emphasis>
<taxonomicName id="4C300FA4FF80FFE39EBBD06D3419FAC3" authority="Baňař, 2017" authorityName="Baňař" authorityYear="2017" box="[475,604,1302,1327]" class="Insecta" family="Staphylinidae" genus="Madastiger" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="435" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B944A835FF80FFE39EBBD06D3419FAC3" box="[475,604,1302,1327]" italics="true" pageId="1" pageNumber="436">Madastiger</emphasis>
</taxonomicName>
is tentatively assigned to the subtribe Mastigerina, although it disagrees with certain character states that have been used to define Mastigerina (
<bibRefCitation id="EFA109D6FF80FFE39CB7D0413231FABF" author="Hlavac" box="[983,1140,1338,1363]" pageId="1" pageNumber="436" refString="Hlavac, P. (2011) Contribution to the knowledge of the tribe [sic] Mastigerina (Coleoptera: Staphylinidae: Pselaphinae, Clavigeritae), with a description of a new genus from Borneo. Zootaxa, 3070, 51 - 59." type="journal article" year="2011">Hlaváč, 2011</bibRefCitation>
). The following character states should be withdraw from the subtribal diagnosis of Mastigerina: the lateral antennal cavities cover only small, basal part of the antennae (also found in some undescribed
<taxonomicName id="4C300FA4FF80FFE39C13D0FF359DFA77" authorityName="Motschulsky" authorityYear="1851" box="[883,984,1412,1435]" class="Insecta" family="Staphylinidae" genus="Mastiger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="genus">
<emphasis id="B944A835FF80FFE39C13D0FF359DFA77" box="[883,984,1412,1435]" italics="true" pageId="1" pageNumber="436">Mastiger</emphasis>
</taxonomicName>
) thus antennae are well-visible dorsally; humeri are not carinate, and mesotibiae are not flat.
<taxonomicName id="4C300FA4FF80FFE39D84D0DD3520FA53" authority="Baňař, 2017" authorityName="Baňař" authorityYear="2017" box="[740,869,1446,1471]" class="Insecta" family="Staphylinidae" genus="Madastiger" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="435" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B944A835FF80FFE39D84D0DD3520FA53" box="[740,869,1446,1471]" italics="true" pageId="1" pageNumber="436">Madastiger</emphasis>
</taxonomicName>
is placed near
<emphasis id="B944A835FF80FFE39B6FD0DC3231FA52" box="[1039,1140,1447,1470]" italics="true" pageId="1" pageNumber="436">Tasmiger</emphasis>
and
<emphasis id="B944A835FF80FFE39BCDD0DC3352FA52" box="[1197,1303,1447,1470]" italics="true" pageId="1" pageNumber="436">
<taxonomicName id="4C300FA4FF80FFE39BCDD0DC3357FA52" authorityName="Motschulsky" authorityYear="1851" box="[1197,1298,1447,1470]" class="Insecta" family="Staphylinidae" genus="Mastiger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="genus">Mastiger</taxonomicName>
,
</emphasis>
which both have the terminal antennomeres truncate at their apices. The new genus is readily separated from
<emphasis id="B944A835FF80FFE39B87D0B73315FA0F" box="[1255,1360,1484,1507]" italics="true" pageId="1" pageNumber="436">
<taxonomicName id="4C300FA4FF80FFE39B87D0B73309FA0F" authorityName="Motschulsky" authorityYear="1851" box="[1255,1356,1484,1507]" class="Insecta" family="Staphylinidae" genus="Mastiger" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="genus">Mastiger</taxonomicName>
,
</emphasis>
which has antennae with 3 antennomeres, and from
<emphasis id="B944A835FF80FFE39DF8D09434B8F9EA" box="[664,765,1519,1542]" italics="true" pageId="1" pageNumber="436">Tasmiger</emphasis>
by the absence of trichomes on the posterior margin of the elytra as well as the foveation pattern of head and as well as the foveation pattern of pronotum, and by the very remarkable structure of the aedeagus (presence of accessory lobes on its apex), which is unique in the tribe
<taxonomicName id="4C300FA4FF80FFE39FF7D321375CF99F" authorityName="Leach" authorityYear="1815" box="[151,281,1626,1651]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="tribe" tribe="Clavigerini">Clavigerini</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C32A27ACFF80FFE39FA7D30535E5F957" pageId="1" pageNumber="436" type="etymology">
<paragraph id="8B8F7427FF80FFE39FA7D30535E5F957" blockId="1.[151,1437,258,1759]" pageId="1" pageNumber="436">
<emphasis id="B944A835FF80FFE39FA7D305370BF97B" bold="true" box="[199,334,1662,1687]" pageId="1" pageNumber="436">Etymology.</emphasis>
The new generic name,
<taxonomicName id="4C300FA4FF80FFE39DF4D3053550F97B" authority="Baňař, 2017" authorityName="Baňař" authorityYear="2017" box="[660,789,1662,1687]" class="Insecta" family="Staphylinidae" genus="Madastiger" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="435" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B944A835FF80FFE39DF4D3053550F97B" box="[660,789,1662,1687]" italics="true" pageId="1" pageNumber="436">Madastiger</emphasis>
</taxonomicName>
is a combination composed from “
<taxonomicName id="4C300FA4FF80FFE39BBCD3053358F97B" box="[1244,1309,1662,1687]" class="Insecta" family="Coccinellidae" genus="Mada" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="genus">
<emphasis id="B944A835FF80FFE39BBCD3053358F97B" box="[1244,1309,1662,1687]" italics="true" pageId="1" pageNumber="436">Mada</emphasis>
</taxonomicName>
” meaning
<collectingCountry id="F32734B7FF80FFE39FF7D3D9375BF957" box="[151,286,1698,1723]" name="Madagascar" pageId="1" pageNumber="436">Madagascar</collectingCountry>
and suffix “
<emphasis id="B944A835FF80FFE39EC9D3DF37ACF957" box="[425,489,1700,1723]" italics="true" pageId="1" pageNumber="436">stiger</emphasis>
” from Mastigerina. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="C32A27ACFF80FFE39FA7D3BD340BF933" box="[199,590,1734,1759]" pageId="1" pageNumber="436" type="distribution">
<paragraph id="8B8F7427FF80FFE39FA7D3BD340BF933" blockId="1.[151,1437,258,1759]" box="[199,590,1734,1759]" pageId="1" pageNumber="436">
<emphasis id="B944A835FF80FFE39FA7D3BD3725F933" bold="true" box="[199,352,1734,1759]" pageId="1" pageNumber="436">Distribution.</emphasis>
Central
<collectingCountry id="F32734B7FF80FFE39EA0D3BD340FF933" box="[448,586,1734,1759]" name="Madagascar" pageId="1" pageNumber="436">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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