180 lines
20 KiB
XML
180 lines
20 KiB
XML
<document id="F411619499849185BCDD73636A1D4CFE" ID-DOI="10.11646/zootaxa.1489.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5087337" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625848289847" checkinUser="felipe" docAuthor="PERKINS, PHILIP D." docDate="2007" docId="038187DBFFE1FF9CFF37F97DFBBF3498" docLanguage="en" docName="zt01489p140.pdf.imf" docOrigin="Zootaxa 1489 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.1489.1.1" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Hydraena cygnus Zwick 1977" docType="treatment" docVersion="4" lastPageNumber="92" masterDocId="FFB8FFA3FFBBFFC7FFA0FF90FFE83112" masterDocTitle="A revision of the Australian species of the water beetle genus Hydraena Kugelann (Coleoptera: Hydraenidae)" masterLastPageNumber="207" masterPageNumber="1" pageNumber="91" updateTime="1698997746102" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="555F30437CA1D6ED15FECBA983AE09E1">
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<mods:title id="61D9721F00B6749E569B1393082CD575">A revision of the Australian species of the water beetle genus Hydraena Kugelann (Coleoptera: Hydraenidae)</mods:title>
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<mods:namePart id="B285730D8ECC0D00A721789A9904C552">PERKINS, PHILIP D.</mods:namePart>
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<mods:title id="0942F756A98F1D19A840F39EFC9C20D2">Zootaxa</mods:title>
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<mods:part id="F39BABFF4D53BADA837EAA02AB83D021">
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<mods:date id="7E17D96F491F2D78B6DE082AB593F3C6">2007</mods:date>
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<mods:number id="D9E0769F9426CA44BACED5ACDC4886A8">2007-05-31</mods:number>
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<mods:number id="C2E2DF715ABB6F3C9B4AEAE5AA8ADEE5">1489</mods:number>
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<mods:classification id="F2A5B5D8EA7F864A0BD52A40C90948E8">journal article</mods:classification>
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<mods:identifier id="308FE5ECBA8F6F94EB06C2F59BB1FE65" type="DOI">10.11646/zootaxa.1489.1.1</mods:identifier>
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<mods:identifier id="CE8826DF09AD3FAB90FC8B12B918D931" type="ISSN">11755334</mods:identifier>
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<treatment id="038187DBFFE1FF9CFF37F97DFBBF3498" ID-DOI="http://doi.org/10.5281/zenodo.5087041" ID-Zenodo-Dep="5087041" LSID="urn:lsid:plazi:treatment:038187DBFFE1FF9CFF37F97DFBBF3498" httpUri="http://treatment.plazi.org/id/038187DBFFE1FF9CFF37F97DFBBF3498" lastPageId="91" lastPageNumber="92" pageId="90" pageNumber="91">
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<subSubSection id="C3326546FFE1FF9DFF37F97DFE2F3638" pageId="90" pageNumber="91" type="nomenclature">
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<paragraph id="8B9736CDFFE1FF9DFF37F97DFDE13615" blockId="90.[151,521,1773,1834]" box="[151,521,1773,1799]" pageId="90" pageNumber="91">
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<heading id="D0DF81A1FFE1FF9DFF37F97DFDE13615" bold="true" box="[151,521,1773,1799]" fontSize="11" level="1" pageId="90" pageNumber="91" reason="1">
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<taxonomicName id="4C284D4EFFE1FF9DFF37F97DFDE13615" ID-CoL="3N38N" authority="Zwick, 1977" authorityName="Zwick" authorityYear="1977" box="[151,521,1773,1799]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="cygnus">
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||
<emphasis id="B95CEADFFFE1FF9DFF37F97DFDE13615" bold="true" box="[151,521,1773,1799]" pageId="90" pageNumber="91">
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<emphasis id="B95CEADFFFE1FF9DFF37F97DFE833615" bold="true" box="[151,363,1773,1799]" italics="true" pageId="90" pageNumber="91">Hydraena cygnus</emphasis>
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||
<bibRefCitation id="EFB94B3CFFE1FF9DFED3F97DFDE13615" author="Zwick, P." box="[371,521,1773,1799]" pageId="90" pageNumber="91" pagination="147 - 184" refId="ref129398" refString="Zwick, P. (1977). Australian Hydraena (Coleoptera: Hydraenidae). Australian Journal of Zoology, 25, 147 - 184." type="journal article" year="1977">Zwick, 1977</bibRefCitation>
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</emphasis>
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</taxonomicName>
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||
</heading>
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</paragraph>
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<paragraph id="8B9736CDFFE1FF9DFF37F880FE2F3638" blockId="90.[151,521,1773,1834]" box="[151,455,1808,1834]" pageId="90" pageNumber="91">(Figs. 147, 151, 152, 240)</paragraph>
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</subSubSection>
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<subSubSection id="C3326546FFE1FF9DFF37F8C2FDCB367B" box="[151,547,1874,1897]" pageId="90" pageNumber="91" type="reference_group">
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||
<paragraph id="8B9736CDFFE1FF9DFF37F8C2FDCB367B" blockId="90.[151,547,1874,1897]" box="[151,547,1874,1897]" pageId="90" pageNumber="91">
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<treatmentCitationGroup id="AB3811E3FFE1FF9DFF37F8C2FDCB367B" box="[151,547,1874,1897]" pageId="90" pageNumber="91">
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<treatmentCitation id="0A8910DCFFE1FF9DFF37F8C2FDF7367B" author="Zwick, P." box="[151,543,1874,1897]" page="169" pageId="90" pageNumber="91" year="1977">
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<taxonomicName id="4C284D4EFFE1FF9DFF37F8C2FDF7367B" ID-CoL="3N38N" authority="Zwick, 1977: 169" authorityName="Zwick" authorityPageNumber="169" authorityYear="1977" box="[151,543,1874,1897]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="cygnus">
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<emphasis id="B95CEADFFFE1FF9DFF37F8C2FEBA367B" box="[151,338,1874,1897]" italics="true" pageId="90" pageNumber="91">Hydraena cygnus</emphasis>
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<bibRefCitation id="EFB94B3CFFE1FF9DFEF9F8C2FDF7367B" author="Zwick, P." box="[345,543,1874,1897]" pageId="90" pageNumber="91" pagination="147 - 184" refId="ref129398" refString="Zwick, P. (1977). Australian Hydraena (Coleoptera: Hydraenidae). Australian Journal of Zoology, 25, 147 - 184." type="journal article" year="1977">Zwick, 1977: 169</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3326546FFE1FF9DFF37F81AFDE336E6" pageId="90" pageNumber="91" type="materials_examined">
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<paragraph id="8B9736CDFFE1FF9DFF37F81AFDE336E6" blockId="90.[151,1436,1930,2036]" pageId="90" pageNumber="91">
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<emphasis id="B95CEADFFFE1FF9DFF37F81AFEA736B6" bold="true" box="[151,335,1930,1956]" pageId="90" pageNumber="91">Type Material.</emphasis>
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<materialsCitation id="3B403C90FFE1FF9DFEFDF81AFDE336E6" collectionCode="R, DIGITAL, IMAGE, SAMA" collectorName="A. M. Lea & P. D. Perkins & Deposited" location="Swan River" pageId="90" pageNumber="91" specimenCount="2" stateProvince="Western Australia" typeStatus="holotype">
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<typeStatus id="5493886FFFE1FF9DFEFDF81AFE2236B6" box="[349,458,1930,1956]" pageId="90" pageNumber="91" type="holotype">Holotype</typeStatus>
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(male):
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<emphasis id="B95CEADFFFE1FF9DFD94F81AFCF136B6" bold="true" box="[564,793,1930,1956]" pageId="90" pageNumber="91">
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<collectingRegion id="49ECF82FFFE1FF9DFD94F81AFCF136B6" box="[564,793,1930,1956]" country="Australia" name="Western Australia" pageId="90" pageNumber="91">Western Australia</collectingRegion>
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</emphasis>
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,
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<location id="8EF76016FFE1FF9DFC87F81AFC5836B6" LSID="urn:lsid:plazi:treatment:038187DBFFE1FF9CFF37F97DFBBF3498:8EF76016FFE1FF9DFC87F81AFC5836B6" box="[807,944,1930,1956]" name="Swan River" pageId="90" pageNumber="91" stateProvince="Western Australia">Swan River</location>
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[labels: Swan
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<collectionCode id="ED39AE08FFE1FF9DFBC3F81AFB9F36B6" box="[1123,1143,1930,1956]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="90" pageNumber="91">R</collectionCode>
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Lea // Griffith collection Id. by
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<collectorName id="26DD531BFFE1FF9DFF47F822FE8C36DE" box="[231,356,1970,1996]" pageId="90" pageNumber="91">A. M. Lea</collectorName>
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//
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<taxonomicName id="4C284D4EFFE1FF9DFE27F823FD4336DE" authority="Zwick" authorityName="Zwick" box="[391,683,1970,1996]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="cygnus">
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<emphasis id="B95CEADFFFE1FF9DFE27F823FDBE36DE" box="[391,598,1971,1996]" italics="true" pageId="90" pageNumber="91">Hydraena cygnus</emphasis>
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Zwick
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</taxonomicName>
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male
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<typeStatus id="5493886FFFE1FF9DFD56F822FC8136DE" box="[758,873,1970,1996]" pageId="90" pageNumber="91" type="holotype">holotypus</typeStatus>
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//
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<collectionCode id="ED39AE08FFE1FF9DFC2DF822FBE936DE" box="[909,1025,1970,1996]" pageId="90" pageNumber="91">DIGITAL</collectionCode>
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<collectionCode id="ED39AE08FFE1FF9DFBABF822FB8336DE" box="[1035,1131,1970,1996]" country="USA" httpUri="http://grbio.org/cool/4pi7-qmy5" name="The I.M.A.G.E Consortium" pageId="90" pageNumber="91">IMAGE</collectionCode>
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captured 2004
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<collectorName id="26DD531BFFE1FF9DFA87F822FF2636E6" pageId="90" pageNumber="91">P. D. Perkins</collectorName>
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].
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<collectorName id="26DD531BFFE1FF9DFF47F84AFEB536E6" box="[231,349,2010,2036]" pageId="90" pageNumber="91">Deposited</collectorName>
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in the
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<collectionCode id="ED39AE08FFE1FF9DFE0EF84AFDEF36E6" box="[430,519,2010,2036]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="90" pageNumber="91" type="Museum">SAMA</collectionCode>
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||
.
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3326546FFE0FF9CFF66FF07FAE431C8" pageId="91" pageNumber="92" type="diagnosis">
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<paragraph id="8B9736CDFFE0FF9CFF66FF07FAE431C8" blockId="91.[151,1436,151,1418]" pageId="91" pageNumber="92">
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<emphasis id="B95CEADFFFE0FF9CFF66FF07FE0A31A3" bold="true" box="[198,482,151,177]" pageId="91" pageNumber="92">Differential Diagnosis.</emphasis>
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Externally very similar to other members of the
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<taxonomicName id="4C284D4EFFE0FF9CFBECFF08FB1F31A3" authorityName="Zwick Holo-Typus" authorityYear="1977" box="[1100,1271,152,177]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="ambiflagellata">
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<emphasis id="B95CEADFFFE0FF9CFBECFF08FB1F31A3" box="[1100,1271,152,177]" italics="true" pageId="91" pageNumber="92">ambiflagellata</emphasis>
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</taxonomicName>
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Group (Figs. 147–150); reliable determinations will require examination of the male genitalia (Figs. 151, 152).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3326546FFE0FF9CFF37FF77FDA035D0" pageId="91" pageNumber="92" type="description">
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<paragraph id="8B9736CDFFE0FF9CFF37FF77FCB53040" blockId="91.[151,1436,151,1418]" pageId="91" pageNumber="92">
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<emphasis id="B95CEADFFFE0FF9CFF37FF77FEC03013" bold="true" box="[151,296,231,257]" pageId="91" pageNumber="92">Description</emphasis>
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. Size:
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<typeStatus id="5493886FFFE0FF9CFE2CFF78FE1A3010" box="[396,498,232,258]" pageId="91" pageNumber="92" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 1.38/0.60; head 0.17/0.32; pronotum 0.36/0.47, PA 0.36, PB 0.46; elytra 0.84/0.60. Dorsum dark brown, anterior and posterior margins of pronotum slightly lighter, legs and maxillary palpi brown.
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</paragraph>
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<paragraph id="8B9736CDFFE0FF9CFF66FEF0FF213308" blockId="91.[151,1436,151,1418]" pageId="91" pageNumber="92">Frons punctures ca. 1xef; interstices shining, 0.5–3xpd. Clypeus microreticulate laterally, finely sparsely punctate medially. Mentum shining, very finely sparsely punctate; postmentum microreticulate. Genae raised, lacking posterior ridge. Pronotal punctures on disc ca. 1xpd those of frons, interstices shining, 2–4xpd, punctures denser at anterior, similar to frons; PF1 and PF4 absent; PF2 very shallow, nearly absent; PF3 shallow.</paragraph>
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<paragraph id="8B9736CDFFE0FF9CFF66FDB8FCE93380" blockId="91.[151,1436,151,1418]" pageId="91" pageNumber="92">Elytral punctures about equal size of largest pronotal punctures. Intervals not raised, shining, width ca. 2xpd, as are interstices between punctures of a row. Apices in dorsal aspect gradually rounded, in posterior aspect margins form shallow angle with one another.</paragraph>
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<paragraph id="8B9736CDFFE0FF9CFF66FD30FC6532C0" blockId="91.[151,1436,151,1418]" pageId="91" pageNumber="92">Plaque/intercoxal process ratios (P2/w/l/s) ca. 1/1/4/2. P1 laminate; median carina posterior extreme slightly more elevated than remainder. P2 narrow, l/w ca. 2/1, sides parallel, apex blunt. Plaques located in posterior 7/10 of metaventrite on sides of deep median subtriangular depression, plaques anteriorly very narrowly separated and lateral margins arcuate. Low midlongitudinal carina between mesoventral intercoxal process and plaques. Cuticle asperite under hydrofuge pubescence. AIS flat, width at arcuate posterior margin 2–3x P2. Protibia gradually widened to about distal 4/5, then markedly excavate, a prominent spine at base of excavation; metatibia with brush of setae on medial surface over distal 3/5. Last sternite symmetrical or nearly so; last tergite with small, slightly off-center concavity.</paragraph>
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<paragraph id="8B9736CDFFE0FF9CFF66FC70FDA035D0" blockId="91.[151,1436,151,1418]" pageId="91" pageNumber="92">Aedeagus (Figs. 151, 152) main-piece extending as wide process on right side, as wide as distal piece, apex blunt; distal piece irregularly shaped, with an apical process visible in lateral view, gonopore flagellum very short, base of distal piece with large, striated lobe, visible in lateral view; left paramere wide, almost trilobed on margin, two small distal lobes each with two setae, ventral margin with row of long setae; right paramere about same length as left paramere, ventral margin curling around main-piece and bearing setae in two groups. Females not yet known.</paragraph>
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||
</subSubSection>
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<subSubSection id="C3326546FFE0FF9CFF66FB5FFAD235F8" box="[198,1338,1231,1258]" pageId="91" pageNumber="92" type="distribution">
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||
<paragraph id="8B9736CDFFE0FF9CFF66FB5FFAD235F8" blockId="91.[151,1436,151,1418]" box="[198,1338,1231,1258]" pageId="91" pageNumber="92">
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||
<emphasis id="B95CEADFFFE0FF9CFF66FB5FFE8F35FB" bold="true" box="[198,359,1231,1257]" pageId="91" pageNumber="92">Distribution.</emphasis>
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||
Currently known only from two localities in western
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||
<collectingRegion id="49ECF82FFFE0FF9CFC44FB40FB5F35F8" box="[996,1207,1232,1258]" country="Australia" name="Western Australia" pageId="91" pageNumber="92">Western Australia</collectingRegion>
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(Fig. 240).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3326546FFE0FF9CFF66FB67FED73428" pageId="91" pageNumber="92" type="discussion">
|
||
<paragraph id="8B9736CDFFE0FF9CFF66FB67FED73428" blockId="91.[151,1436,151,1418]" pageId="91" pageNumber="92">
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||
<emphasis id="B95CEADFFFE0FF9CFF66FB67FED03403" bold="true" box="[198,312,1271,1297]" pageId="91" pageNumber="92">Remarks</emphasis>
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||
. The aedeagus has the left paramere complete, not fused to the main-piece as illustrated by
|
||
<bibRefCitation id="EFB94B3CFFE0FF9CFF37FAB0FED33428" author="Zwick, P." box="[151,315,1312,1338]" pageId="91" pageNumber="92" pagination="147 - 184" refId="ref129398" refString="Zwick, P. (1977). Australian Hydraena (Coleoptera: Hydraenidae). Australian Journal of Zoology, 25, 147 - 184." type="journal article" year="1977">Zwick (1977)</bibRefCitation>
|
||
.
|
||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C3326546FFE0FF9CFF66FAD7FBBF3498" pageId="91" pageNumber="92" type="materials_examined">
|
||
<paragraph id="8B9736CDFFE0FF9CFF66FAD7FBBF3498" blockId="91.[151,1436,151,1418]" pageId="91" pageNumber="92">
|
||
<emphasis id="B95CEADFFFE0FF9CFF66FAD7FE043473" bold="true" box="[198,492,1351,1377]" pageId="91" pageNumber="92">Material Examined (4):</emphasis>
|
||
<materialsCitation id="3B403C90FFE0FF9CFE5BFAD7FBBB3498" collectingDate="2002-10-06" collectionCode="SAMA" collectorName="C. H. S. Watts & Lea" latitude="-31.983334" location="Swan River" longLatPrecision="1213" longitude="115.833336" municipality="Lake Nalyerin" pageId="91" pageNumber="92" specimenCount="1" stateProvince="Western Australia">
|
||
<emphasis id="B95CEADFFFE0FF9CFE5BFAD7FD083473" bold="true" box="[507,736,1351,1377]" pageId="91" pageNumber="92">
|
||
<collectingRegion id="49ECF82FFFE0FF9CFE5BFAD7FD083473" box="[507,736,1351,1377]" country="Australia" name="Western Australia" pageId="91" pageNumber="92">Western Australia</collectingRegion>
|
||
</emphasis>
|
||
,
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||
<collectingMunicipality id="6BF3ACB7FFE0FF9CFD4FFAD8FC723470" box="[751,922,1352,1378]" pageId="91" pageNumber="92">Lake Nalyerin</collectingMunicipality>
|
||
,
|
||
<geoCoordinate id="EE1C500AFFE0FF9CFC06FAD8FBEB3470" box="[934,1027,1352,1378]" degrees="33" direction="south" minutes="8" orientation="latitude" pageId="91" pageNumber="92" precision="925" value="-33.133335">33° 8' S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1C500AFFE0FF9CFBB0FAD8FB643470" box="[1040,1164,1352,1378]" degrees="116" direction="east" minutes="22" orientation="longitude" pageId="91" pageNumber="92" precision="925" value="116.36667">116° 22' E</geoCoordinate>
|
||
,
|
||
<date id="FF96100DFFE0FF9CFB38FAD8FAB93470" box="[1176,1361,1352,1378]" pageId="91" pageNumber="92" value="2002-10-06">
|
||
<collectingDate id="EFD2E9E5FFE0FF9CFB38FAD8FAB93470" box="[1176,1361,1352,1378]" pageId="91" pageNumber="92" value="2002-10-06">6 October 2002</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26DD531BFFE0FF9CFAFDFAD8FF103498" pageId="91" pageNumber="92">C. H. S. Watts</collectorName>
|
||
(3
|
||
<collectionCode id="ED39AE08FFE0FF9CFEBFFAE0FE913498" box="[287,377,1392,1418]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="91" pageNumber="92" type="Museum">SAMA</collectionCode>
|
||
);
|
||
<location id="8EF76016FFE0FF9CFE2DFAE0FDFE3498" LSID="urn:lsid:plazi:treatment:038187DBFFE1FF9CFF37F97DFBBF3498:8EF76016FFE0FF9CFE2DFAE0FDFE3498" box="[397,534,1392,1418]" latitude="-31.983334" longLatPrecision="1213" longitude="115.833336" municipality="Lake Nalyerin" name="Swan River" pageId="91" pageNumber="92" stateProvince="Western Australia">Swan River</location>
|
||
[no date],
|
||
<geoCoordinate id="EE1C500AFFE0FF9CFD35FAE0FCE93498" box="[661,769,1392,1418]" degrees="31" direction="south" minutes="59" orientation="latitude" pageId="91" pageNumber="92" precision="925" value="-31.983334">31° 59' S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE1C500AFFE0FF9CFCAEFAE0FC623498" box="[782,906,1392,1418]" degrees="115" direction="east" minutes="50" orientation="longitude" pageId="91" pageNumber="92" precision="925" value="115.833336">115° 50' E</geoCoordinate>
|
||
,,
|
||
<collectorName id="26DD531BFFE0FF9CFC3EFAE0FC223498" box="[926,970,1392,1418]" pageId="91" pageNumber="92">Lea</collectorName>
|
||
(1
|
||
<collectionCode id="ED39AE08FFE0FF9CFC51FAE0FBA33498" box="[1009,1099,1392,1418]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="91" pageNumber="92" type="Museum">SAMA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |