209 lines
38 KiB
XML
209 lines
38 KiB
XML
<document id="726F75578673182694D3078252F9D1C8" ID-CLB-Dataset="7139" ID-DOI="10.11646/zootaxa.4963.2.1" ID-GBIF-Dataset="049b6936-13d8-42d6-9468-2019ddf78d14" ID-ISSN="1175-5326" ID-Zenodo-Dep="4700704" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618839122537" checkinUser="felipe" docAuthor="Roux, Michel, Martinez, Alain & Vizcaïno, Daniel" docDate="2021" docId="038187A7FF81FFC536A9FCCE3BFFFE8C" docLanguage="en" docName="zootaxa.4963.2.1.pdf" docOrigin="Zootaxa 4963 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudoconocrinus doncieuxi subsp. suboblongus Roux & Martinez & Vizcaïno 2021, n. subsp." docType="treatment" docVersion="7" lastPageNumber="219" masterDocId="FFB8FFDFFF90FFD7363EFFA83835FF91" masterDocTitle="A diverse crinoid fauna (Echinodermata, Crinoidea) from the Lower Eocene of the Gulf of Languedoc (Corbières, Aude, southern France)" masterLastPageNumber="242" masterPageNumber="201" pageNumber="218" updateTime="1698937187163" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="0BA932C33CE70BEDEBD954D63A846317">
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<mods:title id="317BA00910C6B3B7919D070D2BD38C8A">A diverse crinoid fauna (Echinodermata, Crinoidea) from the Lower Eocene of the Gulf of Languedoc (Corbières, Aude, southern France)</mods:title>
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<mods:roleTerm id="1802AE7D9571F0892E116CCEEFA2C1E3">Author</mods:roleTerm>
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<mods:namePart id="7B453EDFFF3C029F2364E1AEEB8824C5">Roux, Michel</mods:namePart>
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<mods:affiliation id="2C77E26891A436D1E948A9A554D0ECDB">Muséum national d’Histoire naturelle, UMR 7205 ISYEB MNHN-CNRS-UPMC-EPHE, Département Systématique et Évolution, CP 51, 57 rue Cuvier, 75231 Paris Cedex 05, France</mods:affiliation>
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<mods:roleTerm id="3B5D1D8003C77D08BA37E7796A1B7351">Author</mods:roleTerm>
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<mods:namePart id="A426B447A51AB76076E60DE35E63EA3A">Martinez, Alain</mods:namePart>
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<mods:affiliation id="AB669ECB2C7794BE6E7EDA94321AAFA7">Chemin de Saint Estève, 11200 Lézignan-Corbières. France</mods:affiliation>
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<mods:roleTerm id="86BCBC24622032F8D29086B1533A0D47">Author</mods:roleTerm>
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<mods:namePart id="365CE87F22E17D26BEFF4F4A64500E44">Vizcaïno, Daniel</mods:namePart>
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<mods:affiliation id="3667B4ED130A375925AF1A62E0DF52AA">7 rue J. - B. Chardin Maquens, 11000 Carcassonne, France</mods:affiliation>
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<mods:typeOfResource id="C5B4026D12F425064E87498C9B8A6974">text</mods:typeOfResource>
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<mods:title id="85521D2B524C769B8DBCD8941CA62C8F">Zootaxa</mods:title>
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<mods:part id="1B73BDFE1FD431FA8D9E4D69D34AF666">
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<mods:date id="5876C8546CECA0D02ABAEC5AC49B1596">2021</mods:date>
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<mods:number id="3D5F812067E82881082EEBA6B5BC59AE">2021-04-19</mods:number>
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<mods:number id="3D867A1758B5C529CEFA5FF7E667EE04">4963</mods:number>
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<treatment id="038187A7FF81FFC536A9FCCE3BFFFE8C" ID-DOI="http://doi.org/10.5281/zenodo.4730449" ID-GBIF-Taxon="181760282" ID-Zenodo-Dep="4730449" LSID="urn:lsid:plazi:treatment:038187A7FF81FFC536A9FCCE3BFFFE8C" httpUri="http://treatment.plazi.org/id/038187A7FF81FFC536A9FCCE3BFFFE8C" lastPageId="18" lastPageNumber="219" pageId="17" pageNumber="218">
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<subSubSection id="C332653AFF81FFC636A9FCCE3B3EFC11" box="[151,779,870,897]" pageId="17" pageNumber="218" type="nomenclature">
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<paragraph id="8B9736B1FF81FFC636A9FCCE3B3EFC11" blockId="17.[151,779,870,932]" box="[151,779,870,897]" pageId="17" pageNumber="218">
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<heading id="D0DF81DDFF81FFC636A9FCCE3B3EFC11" bold="true" box="[151,779,870,897]" fontSize="11" level="1" pageId="17" pageNumber="218" reason="1">
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<emphasis id="B95CEAA3FF81FFC636A9FCCE3B3EFC11" bold="true" box="[151,779,870,897]" pageId="17" pageNumber="218">
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<taxonomicName id="4C284D32FF81FFC636A9FCCE3AA0FC10" authority="Roux & Martinez & Vizcaïno, 2021" authorityName="Roux & Martinez & Vizcaïno" authorityYear="2021" box="[151,661,870,897]" class="Crinoidea" family="Bathycrinidae" genus="Pseudoconocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="17" pageNumber="218" phylum="Echinodermata" rank="subSpecies" species="doncieuxi" status="subsp. nov." subSpecies="suboblongus">
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<emphasis id="B95CEAA3FF81FFC636A9FCCE3AA0FC10" bold="true" box="[151,661,870,897]" italics="true" pageId="17" pageNumber="218">Pseudoconocrinus doncieuxi suboblongus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26F57D8FF81FFC634A3FCCE3B3EFC11" box="[669,779,870,896]" pageId="17" pageNumber="218" rank="subSpecies">n. subsp.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C332653AFF81FFC636A9FC223994FBA4" pageId="17" pageNumber="218" type="description">
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<paragraph id="8B9736B1FF81FFC636A9FC22391AFC35" blockId="17.[151,779,870,932]" box="[151,303,906,932]" pageId="17" pageNumber="218">
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<figureCitation id="13132A34FF81FFC636A9FC223923FC35" box="[151,278,906,932]" captionStart="FIGURE 8" captionStartId="16.[151,250,1673,1697]" captionTargetBox="[210,1378,181,1649]" captionTargetId="figure-23@16.[210,1378,181,1649]" captionTargetPageId="16" captionText="FIGURE 8. Aboral cups and columnals of Pseudoconocrinus doncieuxi and aboral cup of?Democrinus cf. elongatus. A–F: P. doncieuxi suboblongus n. subsp.; A–C: holotype from Fontcouverte à l’Estagnol (MNHN.F.A81998), A: lateral view, B: distal face of radial circlet, C: aboral face with stalk insertion with proximalmost columnals preserved; D–F: paratypes, D–E: from Lavade (MNHN.F.A81999), F: from Réqui (MNHN.F.A82000), D: oblique lateral view showing stalk insertion, E: oblique lateral view of same specimen showing proximal face of radial circlet, F: distal face of radial circlet; G–I:?D. elongatus from Lavade (MNHN.F.A82010), G: lateral view, H: distal face of radial circlet, I: detail of H showing interradial crest and muscular synarthrial facet; J–Q: columnals of P. doncieuxi from Fontcouverte à l’Estagnol (MNHN.F.A69946), J–K: columnal of middle mesistele, K: detail of fulcral ridge; L–N: columnals of dististele, L: oblique proximal view, M: lateral view, N: distal facet, O–Q: columnal of distal mesistele, Q: detail of fulcral ridge. Scale bar equals 1 mm (A–B, D–F, G), 0.5 mm (C, H–I, L–P), 0.25 mm (J, Q) and 0.12 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.4700730" httpUri="https://zenodo.org/record/4700730/files/figure.png" pageId="17" pageNumber="218">Figs. 8A–F</figureCitation>
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,
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<figureCitation id="13132A34FF81FFC6371FFC22391AFC35" box="[289,303,906,932]" captionStart="FIGURE 9" captionStartId="18.[151,250,1653,1677]" captionTargetBox="[157,1424,923,1617]" captionTargetId="figure-282@18.[151,1436,921,1629]" captionTargetPageId="18" captionText="FIGURE 9. Variations in main quantitative characters related to columnal shape and aboral cups of P. doncieuxi suboblongus n. subsp. from Réqui. The coloured envelopes cover more than 90% of specimens; straight lines indicate values of H/D (left) and Hc/Dc (right) (in purple: specimens from Réqui, and, for comparison, in green: P. doncieuxi from the Fontcouverte sites, except for Lavade, in red: P. lavadensis n. sp. from Lavade). Dp and red bar: range of stalk insertion diameter (= proximalmost stalk diameter) measured on aboral cups from Réqui. Values in mm, except for ratios (black straight lines). See Fig. 2 for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4700735" httpUri="https://zenodo.org/record/4700735/files/figure.png" pageId="17" pageNumber="218">9</figureCitation>
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</paragraph>
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<paragraph id="8B9736B1FF81FFC636A9FC7A3994FBA4" blockId="17.[151,1437,978,2013]" pageId="17" pageNumber="218">
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<emphasis id="B95CEAA3FF81FFC636A9FC7A3972FC7D" bold="true" box="[151,327,978,1004]" pageId="17" pageNumber="218">Type-material.</emphasis>
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<materialsCitation id="3B403CECFF81FFC6376FFC7A3994FBA4" ID-GBIF-Occurrence="3095320301" accessionNumber="A81998, A81999" collectionCode="MNHN" pageId="17" pageNumber="218" specimenCount="3" typeStatus="holotype">
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<typeStatus id="54938813FF81FFC6376FFC7A398CFC7D" box="[337,441,978,1004]" pageId="17" pageNumber="218" type="holotype">Holotype</typeStatus>
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(
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<collectionCode id="ED39AE74FF81FFC637F5FC7A3A12FC7D" box="[459,551,978,1004]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="218">MNHN</collectionCode>
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.F.
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<accessionNumber id="947BAB52FF81FFC63400FC7A3AA9FC7D" box="[574,668,978,1004]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A81998" pageId="17" pageNumber="218" type="EnaNcbi">A81998</accessionNumber>
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;
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<figureCitation id="13132A34FF81FFC63494FC7A3B13FC7D" box="[682,806,978,1004]" captionStart="FIGURE 8" captionStartId="16.[151,250,1673,1697]" captionTargetBox="[210,1378,181,1649]" captionTargetId="figure-23@16.[210,1378,181,1649]" captionTargetPageId="16" captionText="FIGURE 8. Aboral cups and columnals of Pseudoconocrinus doncieuxi and aboral cup of?Democrinus cf. elongatus. A–F: P. doncieuxi suboblongus n. subsp.; A–C: holotype from Fontcouverte à l’Estagnol (MNHN.F.A81998), A: lateral view, B: distal face of radial circlet, C: aboral face with stalk insertion with proximalmost columnals preserved; D–F: paratypes, D–E: from Lavade (MNHN.F.A81999), F: from Réqui (MNHN.F.A82000), D: oblique lateral view showing stalk insertion, E: oblique lateral view of same specimen showing proximal face of radial circlet, F: distal face of radial circlet; G–I:?D. elongatus from Lavade (MNHN.F.A82010), G: lateral view, H: distal face of radial circlet, I: detail of H showing interradial crest and muscular synarthrial facet; J–Q: columnals of P. doncieuxi from Fontcouverte à l’Estagnol (MNHN.F.A69946), J–K: columnal of middle mesistele, K: detail of fulcral ridge; L–N: columnals of dististele, L: oblique proximal view, M: lateral view, N: distal facet, O–Q: columnal of distal mesistele, Q: detail of fulcral ridge. Scale bar equals 1 mm (A–B, D–F, G), 0.5 mm (C, H–I, L–P), 0.25 mm (J, Q) and 0.12 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.4700730" httpUri="https://zenodo.org/record/4700730/files/figure.png" pageId="17" pageNumber="218">Fig. 8A–C</figureCitation>
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, Plaziat Collection), from Fontcouverte (à l’Estagnol);
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<typeStatus id="54938813FF81FFC636A9FC5F38C2FB80" box="[151,247,1015,1041]" pageId="17" pageNumber="218" type="paratype">paratype</typeStatus>
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A (
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<collectionCode id="ED39AE74FF81FFC6371EFC5E394EFB81" box="[288,379,1014,1040]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="218">MNHN</collectionCode>
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.F.
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<accessionNumber id="947BAB52FF81FFC637ADFC5E39C4FB81" box="[403,497,1014,1040]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A81999" pageId="17" pageNumber="218" type="EnaNcbi">A81999</accessionNumber>
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;
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<figureCitation id="13132A34FF81FFC637C3FC5E3A43FB81" box="[509,630,1014,1040]" captionStart="FIGURE 8" captionStartId="16.[151,250,1673,1697]" captionTargetBox="[210,1378,181,1649]" captionTargetId="figure-23@16.[210,1378,181,1649]" captionTargetPageId="16" captionText="FIGURE 8. Aboral cups and columnals of Pseudoconocrinus doncieuxi and aboral cup of?Democrinus cf. elongatus. A–F: P. doncieuxi suboblongus n. subsp.; A–C: holotype from Fontcouverte à l’Estagnol (MNHN.F.A81998), A: lateral view, B: distal face of radial circlet, C: aboral face with stalk insertion with proximalmost columnals preserved; D–F: paratypes, D–E: from Lavade (MNHN.F.A81999), F: from Réqui (MNHN.F.A82000), D: oblique lateral view showing stalk insertion, E: oblique lateral view of same specimen showing proximal face of radial circlet, F: distal face of radial circlet; G–I:?D. elongatus from Lavade (MNHN.F.A82010), G: lateral view, H: distal face of radial circlet, I: detail of H showing interradial crest and muscular synarthrial facet; J–Q: columnals of P. doncieuxi from Fontcouverte à l’Estagnol (MNHN.F.A69946), J–K: columnal of middle mesistele, K: detail of fulcral ridge; L–N: columnals of dististele, L: oblique proximal view, M: lateral view, N: distal facet, O–Q: columnal of distal mesistele, Q: detail of fulcral ridge. Scale bar equals 1 mm (A–B, D–F, G), 0.5 mm (C, H–I, L–P), 0.25 mm (J, Q) and 0.12 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.4700730" httpUri="https://zenodo.org/record/4700730/files/figure.png" pageId="17" pageNumber="218">Fig. 8D–E</figureCitation>
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), from Fontcouverte (Lavade);
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||
<typeStatus id="54938813FF81FFC635E4FC5F3C0FFB80" box="[986,1082,1015,1041]" pageId="17" pageNumber="218" type="paratype">paratype</typeStatus>
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B (
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<collectionCode id="ED39AE74FF81FFC6325AFC5E3C8AFB81" box="[1124,1215,1014,1040]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="17" pageNumber="218">MNHN</collectionCode>
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.F.
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<accessionNumber id="947BAB52FF81FFC632E9FC5E3D00FB81" box="[1239,1333,1014,1040]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A81999" pageId="17" pageNumber="218" type="EnaNcbi">A81999</accessionNumber>
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;
|
||
<figureCitation id="13132A34FF81FFC6337FFC5E3DA1FB81" box="[1345,1428,1014,1040]" captionStart="FIGURE 8" captionStartId="16.[151,250,1673,1697]" captionTargetBox="[210,1378,181,1649]" captionTargetId="figure-23@16.[210,1378,181,1649]" captionTargetPageId="16" captionText="FIGURE 8. Aboral cups and columnals of Pseudoconocrinus doncieuxi and aboral cup of?Democrinus cf. elongatus. A–F: P. doncieuxi suboblongus n. subsp.; A–C: holotype from Fontcouverte à l’Estagnol (MNHN.F.A81998), A: lateral view, B: distal face of radial circlet, C: aboral face with stalk insertion with proximalmost columnals preserved; D–F: paratypes, D–E: from Lavade (MNHN.F.A81999), F: from Réqui (MNHN.F.A82000), D: oblique lateral view showing stalk insertion, E: oblique lateral view of same specimen showing proximal face of radial circlet, F: distal face of radial circlet; G–I:?D. elongatus from Lavade (MNHN.F.A82010), G: lateral view, H: distal face of radial circlet, I: detail of H showing interradial crest and muscular synarthrial facet; J–Q: columnals of P. doncieuxi from Fontcouverte à l’Estagnol (MNHN.F.A69946), J–K: columnal of middle mesistele, K: detail of fulcral ridge; L–N: columnals of dististele, L: oblique proximal view, M: lateral view, N: distal facet, O–Q: columnal of distal mesistele, Q: detail of fulcral ridge. Scale bar equals 1 mm (A–B, D–F, G), 0.5 mm (C, H–I, L–P), 0.25 mm (J, Q) and 0.12 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.4700730" httpUri="https://zenodo.org/record/4700730/files/figure.png" pageId="17" pageNumber="218">Fig. 8F</figureCitation>
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) from Montlaur (Réqui).
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C332653AFF81FFC636F9FB963BBDFBC8" box="[199,904,1086,1113]" pageId="17" pageNumber="218" type="etymology">
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<paragraph id="8B9736B1FF81FFC636F9FB963BBDFBC8" blockId="17.[151,1437,978,2013]" box="[199,904,1086,1113]" pageId="17" pageNumber="218">
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||
<emphasis id="B95CEAA3FF81FFC636F9FB96397BFBC9" bold="true" box="[199,334,1086,1112]" pageId="17" pageNumber="218">Etymology.</emphasis>
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In reference to the oblong shape of the aboral cup.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C332653AFF81FFC636F9FBCA3C1EFB55" pageId="17" pageNumber="218" type="materials_examined">
|
||
<paragraph id="8B9736B1FF81FFC636F9FBCA3C1EFB55" blockId="17.[151,1437,978,2013]" pageId="17" pageNumber="218">
|
||
<emphasis id="B95CEAA3FF81FFC636F9FBCA399FFBED" bold="true" box="[199,426,1122,1148]" pageId="17" pageNumber="218">Material examined</emphasis>
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||
.
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<materialsCitation id="3B403CECFF81FFC63786FBCB3C12FB55" ID-GBIF-Occurrence="3095320312" collectionCode="UCB-" collectorName="de Fontcouverte" location="Tuilerie de Fontcouverte" pageId="17" pageNumber="218" specimenCode="FSL 170248" specimenCount="1">
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In addition to the
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<typeStatus id="54938813FF81FFC634BEFBCB3A85FBEC" box="[640,688,1123,1149]" pageId="17" pageNumber="218">type</typeStatus>
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series, 10 aboral cups from the
|
||
<location id="8EF7606AFF81FFC63229FBCA3D19FBEC" LSID="urn:lsid:plazi:treatment:038187A7FF81FFC536A9FCCE3BFFFE8C:8EF7606AFF81FFC63229FBCA3D19FBEC" box="[1047,1324,1122,1149]" name="Tuilerie de Fontcouverte" pageId="17" pageNumber="218">
|
||
Tuilerie
|
||
<collectorName id="26DD5367FF81FFC63248FBCA3D19FBEC" box="[1142,1324,1122,1149]" pageId="17" pageNumber="218">de Fontcouverte</collectorName>
|
||
</location>
|
||
(in
|
||
<specimenCode id="DB8E9ECAFF81FFC63367FBCA3912FB31" collectionCode="FSL" country="France" httpUri="http://grbio.org/cool/p8j1-ms8j" name="Collections de la Faculte des Sciences de Lyon" pageId="17" pageNumber="218">
|
||
<collectionCode id="ED39AE74FF81FFC63367FBCA38F3FB31" pageId="17" pageNumber="218">UCB-FSL</collectionCode>
|
||
170248
|
||
</specimenCode>
|
||
b, and Martinez Collection); 125 aboral cups and 238 columnals from Réqui, some aboral cups and columnals from other sites (see
|
||
<tableCitation id="C6AA030AFF81FFC637C2FB023A7AFB55" box="[508,591,1194,1220]" captionStart="TABLE 2" captionStartId="7.[151,239,548,572]" captionText="TABLE 2. Sites with crinoids here studied with taxa, abundance and types of ossicles. For abbreviations, see Table 1." pageId="17" pageNumber="218">Table 2</tableCitation>
|
||
) in the Martinez and Vizcaïno collections
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C332653AFF81FFC636F9FB663ADCFAE9" pageId="17" pageNumber="218" type="diagnosis">
|
||
<paragraph id="8B9736B1FF81FFC636F9FB663ADCFAE9" blockId="17.[151,1437,978,2013]" pageId="17" pageNumber="218">
|
||
<emphasis id="B95CEAA3FF81FFC636F9FB663977FB79" bold="true" box="[199,322,1230,1256]" pageId="17" pageNumber="218">Diagnosis.</emphasis>
|
||
Variety with often very elongated aboral cup, Hc up to
|
||
<quantity id="4CD09B54FF81FFC63580FB663C2BFB78" box="[958,1054,1230,1257]" metricMagnitude="-1" metricUnit="m" metricValue="1.55" pageId="17" pageNumber="218" unit="mm" value="155.0">155 mm</quantity>
|
||
, H/D most often> 2.5 and sometimes up to 5.7, Dc maximum located in central or adoral third, aboral third in inverted truncated cone with base corresponding to stalk insertion, adoral third truncated conical or subcylindrical in shape, distal face of radial circlet almost flat with marked star-shaped central depression in largest specimens, stalk insertion wide (Dp/Dr>050), columnals usually without marked bulge at mid-height.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C332653AFF81FFC636F9FA2A398EF8B9" pageId="17" pageNumber="218" type="description">
|
||
<paragraph id="8B9736B1FF81FFC636F9FA2A3C07F909" blockId="17.[151,1437,978,2013]" pageId="17" pageNumber="218">
|
||
<emphasis id="B95CEAA3FF81FFC636F9FA2A396CFA0D" bold="true" box="[199,345,1410,1436]" pageId="17" pageNumber="218">Description.</emphasis>
|
||
Quantitative characters of the three aboral cups of the type series listed in
|
||
<tableCitation id="C6AA030AFF81FFC6329EFA2A3CC0FA0D" box="[1184,1269,1410,1436]" captionStart="TABLE 3" captionStartId="14.[151,239,368,392]" captionText="TABLE 3. Quantitative characters of aboral cups of Pseudoconocrinus doncieuxi here figured. Catalogue numbers are those of MNHN. * = P. doncieuxi suboblongus n. subsp., Dp: diameter of stalk insertion, Wr: width of distal articular facet of one radial; other abbreviations see Fig. 2. Values in mm, except for ratios." pageId="17" pageNumber="218">Table 3</tableCitation>
|
||
.
|
||
<typeStatus id="54938813FF81FFC6333FFA2A3D5BFA0D" box="[1281,1390,1410,1436]" pageId="17" pageNumber="218" type="holotype">Holotype</typeStatus>
|
||
: aboral cup with maximum Dc in central swollen third (
|
||
<figureCitation id="13132A34FF81FFC634D9FA0E3B74FA51" box="[743,833,1446,1472]" captionStart="FIGURE 8" captionStartId="16.[151,250,1673,1697]" captionTargetBox="[210,1378,181,1649]" captionTargetId="figure-23@16.[210,1378,181,1649]" captionTargetPageId="16" captionText="FIGURE 8. Aboral cups and columnals of Pseudoconocrinus doncieuxi and aboral cup of?Democrinus cf. elongatus. A–F: P. doncieuxi suboblongus n. subsp.; A–C: holotype from Fontcouverte à l’Estagnol (MNHN.F.A81998), A: lateral view, B: distal face of radial circlet, C: aboral face with stalk insertion with proximalmost columnals preserved; D–F: paratypes, D–E: from Lavade (MNHN.F.A81999), F: from Réqui (MNHN.F.A82000), D: oblique lateral view showing stalk insertion, E: oblique lateral view of same specimen showing proximal face of radial circlet, F: distal face of radial circlet; G–I:?D. elongatus from Lavade (MNHN.F.A82010), G: lateral view, H: distal face of radial circlet, I: detail of H showing interradial crest and muscular synarthrial facet; J–Q: columnals of P. doncieuxi from Fontcouverte à l’Estagnol (MNHN.F.A69946), J–K: columnal of middle mesistele, K: detail of fulcral ridge; L–N: columnals of dististele, L: oblique proximal view, M: lateral view, N: distal facet, O–Q: columnal of distal mesistele, Q: detail of fulcral ridge. Scale bar equals 1 mm (A–B, D–F, G), 0.5 mm (C, H–I, L–P), 0.25 mm (J, Q) and 0.12 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.4700730" httpUri="https://zenodo.org/record/4700730/files/figure.png" pageId="17" pageNumber="218">Fig. 8A</figureCitation>
|
||
); adoral third in truncated cone, distal surface flat entirely occupied by insertion of arms, strongly star-shaped central cavity (
|
||
<figureCitation id="13132A34FF81FFC635FEFA623C2DFA75" box="[960,1048,1482,1508]" captionStart="FIGURE 8" captionStartId="16.[151,250,1673,1697]" captionTargetBox="[210,1378,181,1649]" captionTargetId="figure-23@16.[210,1378,181,1649]" captionTargetPageId="16" captionText="FIGURE 8. Aboral cups and columnals of Pseudoconocrinus doncieuxi and aboral cup of?Democrinus cf. elongatus. A–F: P. doncieuxi suboblongus n. subsp.; A–C: holotype from Fontcouverte à l’Estagnol (MNHN.F.A81998), A: lateral view, B: distal face of radial circlet, C: aboral face with stalk insertion with proximalmost columnals preserved; D–F: paratypes, D–E: from Lavade (MNHN.F.A81999), F: from Réqui (MNHN.F.A82000), D: oblique lateral view showing stalk insertion, E: oblique lateral view of same specimen showing proximal face of radial circlet, F: distal face of radial circlet; G–I:?D. elongatus from Lavade (MNHN.F.A82010), G: lateral view, H: distal face of radial circlet, I: detail of H showing interradial crest and muscular synarthrial facet; J–Q: columnals of P. doncieuxi from Fontcouverte à l’Estagnol (MNHN.F.A69946), J–K: columnal of middle mesistele, K: detail of fulcral ridge; L–N: columnals of dististele, L: oblique proximal view, M: lateral view, N: distal facet, O–Q: columnal of distal mesistele, Q: detail of fulcral ridge. Scale bar equals 1 mm (A–B, D–F, G), 0.5 mm (C, H–I, L–P), 0.25 mm (J, Q) and 0.12 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.4700730" httpUri="https://zenodo.org/record/4700730/files/figure.png" pageId="17" pageNumber="218">Fig. 8B</figureCitation>
|
||
); aboral third in inverted truncated cone with flat base entirely occupied by stalk insertion housing proximalmost discoid columnals (
|
||
<figureCitation id="13132A34FF81FFC632EEFA463D12F999" box="[1232,1319,1518,1544]" captionStart="FIGURE 8" captionStartId="16.[151,250,1673,1697]" captionTargetBox="[210,1378,181,1649]" captionTargetId="figure-23@16.[210,1378,181,1649]" captionTargetPageId="16" captionText="FIGURE 8. Aboral cups and columnals of Pseudoconocrinus doncieuxi and aboral cup of?Democrinus cf. elongatus. A–F: P. doncieuxi suboblongus n. subsp.; A–C: holotype from Fontcouverte à l’Estagnol (MNHN.F.A81998), A: lateral view, B: distal face of radial circlet, C: aboral face with stalk insertion with proximalmost columnals preserved; D–F: paratypes, D–E: from Lavade (MNHN.F.A81999), F: from Réqui (MNHN.F.A82000), D: oblique lateral view showing stalk insertion, E: oblique lateral view of same specimen showing proximal face of radial circlet, F: distal face of radial circlet; G–I:?D. elongatus from Lavade (MNHN.F.A82010), G: lateral view, H: distal face of radial circlet, I: detail of H showing interradial crest and muscular synarthrial facet; J–Q: columnals of P. doncieuxi from Fontcouverte à l’Estagnol (MNHN.F.A69946), J–K: columnal of middle mesistele, K: detail of fulcral ridge; L–N: columnals of dististele, L: oblique proximal view, M: lateral view, N: distal facet, O–Q: columnal of distal mesistele, Q: detail of fulcral ridge. Scale bar equals 1 mm (A–B, D–F, G), 0.5 mm (C, H–I, L–P), 0.25 mm (J, Q) and 0.12 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.4700730" httpUri="https://zenodo.org/record/4700730/files/figure.png" pageId="17" pageNumber="218">Fig. 8C</figureCitation>
|
||
).
|
||
<typeStatus id="54938813FF81FFC63304FA473DA9F998" box="[1338,1436,1519,1545]" pageId="17" pageNumber="218" type="paratype">Paratype</typeStatus>
|
||
A: aboral cup similar in shape to
|
||
<typeStatus id="54938813FF81FFC63425F9BA3A48F9BD" box="[539,637,1554,1580]" pageId="17" pageNumber="218" type="holotype">holotype</typeStatus>
|
||
but more elongated (H/D 2.5) and illustrating general trend of aboral cups from the Lavade and Requi sites (
|
||
<figureCitation id="13132A34FF81FFC63468F99E3AE4F9C1" box="[598,721,1590,1616]" captionStart="FIGURE 8" captionStartId="16.[151,250,1673,1697]" captionTargetBox="[210,1378,181,1649]" captionTargetId="figure-23@16.[210,1378,181,1649]" captionTargetPageId="16" captionText="FIGURE 8. Aboral cups and columnals of Pseudoconocrinus doncieuxi and aboral cup of?Democrinus cf. elongatus. A–F: P. doncieuxi suboblongus n. subsp.; A–C: holotype from Fontcouverte à l’Estagnol (MNHN.F.A81998), A: lateral view, B: distal face of radial circlet, C: aboral face with stalk insertion with proximalmost columnals preserved; D–F: paratypes, D–E: from Lavade (MNHN.F.A81999), F: from Réqui (MNHN.F.A82000), D: oblique lateral view showing stalk insertion, E: oblique lateral view of same specimen showing proximal face of radial circlet, F: distal face of radial circlet; G–I:?D. elongatus from Lavade (MNHN.F.A82010), G: lateral view, H: distal face of radial circlet, I: detail of H showing interradial crest and muscular synarthrial facet; J–Q: columnals of P. doncieuxi from Fontcouverte à l’Estagnol (MNHN.F.A69946), J–K: columnal of middle mesistele, K: detail of fulcral ridge; L–N: columnals of dististele, L: oblique proximal view, M: lateral view, N: distal facet, O–Q: columnal of distal mesistele, Q: detail of fulcral ridge. Scale bar equals 1 mm (A–B, D–F, G), 0.5 mm (C, H–I, L–P), 0.25 mm (J, Q) and 0.12 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.4700730" httpUri="https://zenodo.org/record/4700730/files/figure.png" pageId="17" pageNumber="218">Fig. 8D–E</figureCitation>
|
||
).
|
||
<typeStatus id="54938813FF81FFC634DBF99F3B72F9C0" box="[741,839,1591,1617]" pageId="17" pageNumber="218" type="paratype">Paratype</typeStatus>
|
||
B: exceptionnally well-preserved, almost flat adoral face showing moderately protruding interradial ridges, with large, slightly star-shaped central cavity, and aboral and inner ligament areas of synarthries better developed than muscular areas (
|
||
<figureCitation id="13132A34FF81FFC635EEF9D63C10F909" box="[976,1061,1662,1688]" captionStart="FIGURE 8" captionStartId="16.[151,250,1673,1697]" captionTargetBox="[210,1378,181,1649]" captionTargetId="figure-23@16.[210,1378,181,1649]" captionTargetPageId="16" captionText="FIGURE 8. Aboral cups and columnals of Pseudoconocrinus doncieuxi and aboral cup of?Democrinus cf. elongatus. A–F: P. doncieuxi suboblongus n. subsp.; A–C: holotype from Fontcouverte à l’Estagnol (MNHN.F.A81998), A: lateral view, B: distal face of radial circlet, C: aboral face with stalk insertion with proximalmost columnals preserved; D–F: paratypes, D–E: from Lavade (MNHN.F.A81999), F: from Réqui (MNHN.F.A82000), D: oblique lateral view showing stalk insertion, E: oblique lateral view of same specimen showing proximal face of radial circlet, F: distal face of radial circlet; G–I:?D. elongatus from Lavade (MNHN.F.A82010), G: lateral view, H: distal face of radial circlet, I: detail of H showing interradial crest and muscular synarthrial facet; J–Q: columnals of P. doncieuxi from Fontcouverte à l’Estagnol (MNHN.F.A69946), J–K: columnal of middle mesistele, K: detail of fulcral ridge; L–N: columnals of dististele, L: oblique proximal view, M: lateral view, N: distal facet, O–Q: columnal of distal mesistele, Q: detail of fulcral ridge. Scale bar equals 1 mm (A–B, D–F, G), 0.5 mm (C, H–I, L–P), 0.25 mm (J, Q) and 0.12 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.4700730" httpUri="https://zenodo.org/record/4700730/files/figure.png" pageId="17" pageNumber="218">Fig. 8F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9736B1FF81FFC636F9F90A398EF8B9" blockId="17.[151,1437,978,2013]" pageId="17" pageNumber="218">
|
||
<tableCitation id="C6AA030AFF81FFC636F9F90A3910F92C" box="[199,293,1698,1725]" captionStart="TABLE 5" captionStartId="18.[151,239,332,356]" captionText="TABLE 5. Quantitative characters of a sample of 125 aboral cups of P. doncieuxi suboblongus n. subsp. from Réqui near Montlaur (Martinez and Vizcaïno collections). For abbreviations, see Fig. 2. Values in mm, except for ratios." pageId="17" pageNumber="218">Tables 5</tableCitation>
|
||
and
|
||
<tableCitation id="C6AA030AFF81FFC63763F90A395EF92D" box="[349,363,1698,1724]" captionStart="TABLE 6" captionStartId="18.[151,239,602,626]" captionText="TABLE 6. Quantitative characters of a sample of 238 rhizocrinid columnals from Réqui near Montlaur (Martinez and Vizcaïno collections) in a bed with abundant aboral cups of Pseudoconocrinus doncieuxi suboblongus n. subsp. For abbreviations, see Fig. 2. Values in mm, except for ratios." pageId="17" pageNumber="218">6</tableCitation>
|
||
list the general quantitative characters of aboral cups and columnals from the Réqui site where this
|
||
<taxonomicName id="4C284D32FF81FFC636F7F96E394EF971" box="[201,379,1734,1760]" pageId="17" pageNumber="218" rank="variety" variety="strongly">variety strongly</taxonomicName>
|
||
predominates; these specimens display a much smaller size than those of
|
||
<taxonomicName id="4C284D32FF81FFC632F1F96F3D68F971" box="[1231,1373,1734,1761]" class="Crinoidea" family="Bathycrinidae" genus="Pseudoconocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="17" pageNumber="218" phylum="Echinodermata" rank="species" species="doncieuxi">
|
||
<emphasis id="B95CEAA3FF81FFC632F1F96F3CD5F971" box="[1231,1248,1735,1760]" italics="true" pageId="17" pageNumber="218">P</emphasis>
|
||
.
|
||
<emphasis id="B95CEAA3FF81FFC632CEF96E3D68F971" box="[1264,1373,1734,1760]" italics="true" pageId="17" pageNumber="218">doncieuxi</emphasis>
|
||
</taxonomicName>
|
||
from the Tuilerie de Fontcouverte and a tendency towards greater and earlier elongation (Hc/Dc from 2.5 to 5.73 for 1.7<Dc<3.0 mm) (
|
||
<figureCitation id="13132A34FF81FFC63757F8A63998F8B9" box="[361,429,1806,1832]" captionStart="FIGURE 8" captionStartId="16.[151,250,1673,1697]" captionTargetBox="[210,1378,181,1649]" captionTargetId="figure-23@16.[210,1378,181,1649]" captionTargetPageId="16" captionText="FIGURE 8. Aboral cups and columnals of Pseudoconocrinus doncieuxi and aboral cup of?Democrinus cf. elongatus. A–F: P. doncieuxi suboblongus n. subsp.; A–C: holotype from Fontcouverte à l’Estagnol (MNHN.F.A81998), A: lateral view, B: distal face of radial circlet, C: aboral face with stalk insertion with proximalmost columnals preserved; D–F: paratypes, D–E: from Lavade (MNHN.F.A81999), F: from Réqui (MNHN.F.A82000), D: oblique lateral view showing stalk insertion, E: oblique lateral view of same specimen showing proximal face of radial circlet, F: distal face of radial circlet; G–I:?D. elongatus from Lavade (MNHN.F.A82010), G: lateral view, H: distal face of radial circlet, I: detail of H showing interradial crest and muscular synarthrial facet; J–Q: columnals of P. doncieuxi from Fontcouverte à l’Estagnol (MNHN.F.A69946), J–K: columnal of middle mesistele, K: detail of fulcral ridge; L–N: columnals of dististele, L: oblique proximal view, M: lateral view, N: distal facet, O–Q: columnal of distal mesistele, Q: detail of fulcral ridge. Scale bar equals 1 mm (A–B, D–F, G), 0.5 mm (C, H–I, L–P), 0.25 mm (J, Q) and 0.12 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.4700730" httpUri="https://zenodo.org/record/4700730/files/figure.png" pageId="17" pageNumber="218">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C332653AFF81FFC536F9F89A3BFFFE8C" lastPageId="18" lastPageNumber="219" pageId="17" pageNumber="218" type="discussion">
|
||
<paragraph id="8B9736B1FF81FFC536F9F89A3A15FF68" blockId="17.[151,1437,978,2013]" lastBlockId="18.[151,1436,151,285]" lastPageId="18" lastPageNumber="219" pageId="17" pageNumber="218">
|
||
<emphasis id="B95CEAA3FF81FFC636F9F89A390EF8DD" bold="true" box="[199,315,1842,1868]" pageId="17" pageNumber="218">Remarks.</emphasis>
|
||
<taxonomicName id="4C284D32FF81FFC6377AF89A3B15F8DD" authority="Roux & Martinez & Vizcaïno, 2021" authorityName="Roux & Martinez & Vizcaïno" authorityYear="2021" box="[324,800,1842,1868]" class="Crinoidea" family="Bathycrinidae" genus="Pseudoconocrinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Comatulida" pageId="17" pageNumber="218" phylum="Echinodermata" rank="subSpecies" species="doncieuxi" status="subsp. nov." subSpecies="suboblongus">
|
||
<emphasis id="B95CEAA3FF81FFC6377AF89A3B15F8DD" box="[324,800,1842,1868]" italics="true" pageId="17" pageNumber="218">Pseudoconocrinus doncieuxi suboblongus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95CEAA3FF81FFC63516F89A3BA6F8DD" bold="true" box="[808,915,1842,1868]" pageId="17" pageNumber="218">
|
||
<taxonomicNameLabel id="A26F57D8FF81FFC63516F89A3BA6F8DD" box="[808,915,1842,1868]" pageId="17" pageNumber="218" rank="subSpecies">n. subsp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is present at numerous sites, but with widely varying frequencies. In the particular environment of Réqui (see above), this
|
||
<taxonomicName id="4C284D32FF81FFC6321BF8FE3CEFF8E1" box="[1061,1242,1878,1904]" pageId="17" pageNumber="218" rank="variety" variety="strongly">variety strongly</taxonomicName>
|
||
predominates in several of the marl layers sampled. Aboral cups with Dc>
|
||
<quantity id="4CD09B54FF81FFC6351CF8D23B42F804" box="[802,887,1914,1941]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="17" pageNumber="218" unit="mm" value="1.7">1.7 mm</quantity>
|
||
have a much less variable shape than those with Dc>2.5 from the Tuilerie de Fontcouverte (
|
||
<figureCitation id="13132A34FF81FFC6344DF8363A83F829" box="[627,694,1950,1976]" captionStart="FIGURE 9" captionStartId="18.[151,250,1653,1677]" captionTargetBox="[157,1424,923,1617]" captionTargetId="figure-282@18.[151,1436,921,1629]" captionTargetPageId="18" captionText="FIGURE 9. Variations in main quantitative characters related to columnal shape and aboral cups of P. doncieuxi suboblongus n. subsp. from Réqui. The coloured envelopes cover more than 90% of specimens; straight lines indicate values of H/D (left) and Hc/Dc (right) (in purple: specimens from Réqui, and, for comparison, in green: P. doncieuxi from the Fontcouverte sites, except for Lavade, in red: P. lavadensis n. sp. from Lavade). Dp and red bar: range of stalk insertion diameter (= proximalmost stalk diameter) measured on aboral cups from Réqui. Values in mm, except for ratios (black straight lines). See Fig. 2 for abbreviations." figureDoi="http://doi.org/10.5281/zenodo.4700735" httpUri="https://zenodo.org/record/4700735/files/figure.png" pageId="17" pageNumber="218">Fig. 9</figureCitation>
|
||
). This suggests a tendency for elongation during ontogeny that was channeled by environmental factors early during growth. At Réqui, the absence of brachials and rhizoid segments, the abnormal relative abundance of aboral cups compared to columnals (about 3 columnals per 1 aboral cup), as well as the poor preservation due to attrition and biocorrosion indicate a displacement of the ossicles that does not allow us to estimate average stalk length.
|
||
</paragraph>
|
||
<paragraph id="8B9736B1FF82FFC536F9FEAA3BFFFE8C" blockId="18.[151,1436,151,285]" box="[199,970,258,285]" pageId="18" pageNumber="219">
|
||
<emphasis id="B95CEAA3FF82FFC536F9FEAA396FFE8D" bold="true" box="[199,346,258,284]" pageId="18" pageNumber="219">Occurrence.</emphasis>
|
||
Early Ypresian (middle Ilerdian, NP 10–11), Corbières.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |