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<document id="05F0B3C793B60BC1AA34D987F6C61095" ID-CLB-Dataset="296642" ID-DOI="10.11646/zootaxa.5453.1.3" ID-GBIF-Dataset="040fe0d3-0f53-4778-a814-bef399078de2" ID-ISSN="1175-5326" ID-Zenodo-Dep="11233494" ID-ZooBank="721FB81B-B372-41C6-9661-685AE7F834ED" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1716281936411" checkinUser="plazi" docAuthor="Pham, Duy Long, Chi, Nguyen Minh, Grehan, John R., Khai, Tran Quang, Ignatev, Nikolai, Jones, Loren D., Mielke, Carlos G. C., Quang, Dao Ngoc &amp; Yakovlev, Roman" docDate="2024" docId="03818796DE5DFFF69ACC4792FE2A6524" docLanguage="en" docName="zootaxa.5453.1.3.pdf" docOrigin="Zootaxa 5453 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5453.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Endoclita phuthoensis Long, Chi &amp; Grehan 2024, sp. n." docType="treatment" docVersion="4" lastPageNumber="66" masterDocId="FFB8FFEEDE5FFFF59A5B474DFFA7671B" masterDocTitle="Notes on Endoclita C. &amp; R. Felder (Lepidoptera: Hepialidae) feeding on Eucalyptus in Vietnam, with new records and a new species" masterLastPageNumber="85" masterPageNumber="63" pageNumber="65" updateTime="1716383097457" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C0D07625185E0985B5A5F02BC0898F30">Notes on Endoclita C. &amp; R. Felder (Lepidoptera: Hepialidae) feeding on Eucalyptus in Vietnam, with new records and a new species</mods:title>
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<mods:namePart id="959FCEEF17457FCA4509453EB6D14AAB">Pham, Duy Long</mods:namePart>
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<mods:affiliation id="C093D1AFA3C6A7C1F2B6A0F5C0640D27">Forest Protection Research Centre, Vietnamese Academy of Forest Sciences, 46 Duc Thang Street, Bac Tu Liem District, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="0C2CFDBFE63C587BFB532FEBE77C98A7">Chi, Nguyen Minh</mods:namePart>
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<mods:namePart id="29061DED83A3B1D4108DF437B484AF6D">Grehan, John R.</mods:namePart>
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<mods:affiliation id="20042B4AF6BEE89C1D0039BE52BB57AD">Research Associate, McGuire Center for Lepidoptera and Biodiversity, Gainesville, Florida, USA</mods:affiliation>
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<mods:affiliation id="B4EE8EEEF666A5586C935926434F0EC6">Tay Bac University, Son La Province, Vietnam</mods:affiliation>
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<mods:affiliation id="0C553EE7D603346F5DCA229FC798FE75">Institute of Entomology, Biology Centre of the Czech Academy of Science, České Budějovice, Czech Republic</mods:affiliation>
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<mods:affiliation id="29D9126EBEDF2C50FDC381365292AF17">Department of Entomology, Comstock Hall, Cornell University, Ithaca, NY 14853 - 2601 USA</mods:affiliation>
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<mods:namePart id="69B7BE1927DD968601EBCAD155572B57">Mielke, Carlos G. C.</mods:namePart>
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<mods:affiliation id="F73B09E6EEE5D2DC6044FC28CCDFF62F">Research Associate, McGuire Center for Lepidoptera and Biodiversity, Gainesville, Florida, USA</mods:affiliation>
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<mods:affiliation id="9B1BC8E814F674EF80EA05040D3D3706">Forest Protection Research Centre, Vietnamese Academy of Forest Sciences, 46 Duc Thang Street, Bac Tu Liem District, Hanoi, Vietnam</mods:affiliation>
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<mods:affiliation id="B24A85C9E7E6753E90202752462321AE">Altai State University, pr. Lenina 61, RUS- 656049 Barnaul, Russia &amp; Tomsk State University, Laboratory of Biodiversity and Ecology, Lenina pr. 36, RUS- 634050 Tomsk, Russia &amp; Western Caspian University, Istiglaliyyat Street 31, Baku 1001, Azerbaijan</mods:affiliation>
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<mods:date id="2B3AEDA88ED7DC2C4D00CD535EB1E214">2024</mods:date>
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<treatment id="03818796DE5DFFF69ACC4792FE2A6524" ID-DOI="http://doi.org/10.5281/zenodo.11233391" ID-Zenodo-Dep="11233391" LSID="urn:lsid:plazi:treatment:03818796DE5DFFF69ACC4792FE2A6524" httpUri="http://treatment.plazi.org/id/03818796DE5DFFF69ACC4792FE2A6524" lastPageId="3" lastPageNumber="66" pageId="2" pageNumber="65">
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<taxonomicName id="4C284D03DE5DFFF79ACC4792FD1367E2" authority="Long, Chi &amp; Grehan" authorityName="Long, Chi &amp; Grehan" authorityYear="2024" box="[151,692,223,250]" class="Insecta" family="Hepialidae" genus="Endoclita" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="phuthoensis" status="sp. nov.">
<emphasis id="B95CEA92DE5DFFF79ACC4792FE0067E1" bold="true" box="[151,423,223,250]" italics="true" pageId="2" pageNumber="65">Endoclita phuthoensis</emphasis>
Long, Chi &amp; Grehan
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(
<figureCitation id="13132A05DE5DFFF79AC4464EFEB66606" box="[159,273,259,285]" captionStart="FIGURE 1" captionStartId="2.[151,250,1053,1077]" captionTargetBox="[324,1263,309,1029]" captionTargetId="figure-48@2.[324,1263,309,1029]" captionTargetPageId="2" captionText="FIGURE 1. Endoclita phuthoensis sp. n.: holotype female (1a dorsal, 1b ventral), and two paratype females (1c–d). Photos by Nguyen Minh Chi." figureDoi="http://doi.org/10.5281/zenodo.11233496" httpUri="https://zenodo.org/record/11233496/files/figure.png" pageId="2" pageNumber="65">Figs 1a–d</figureCitation>
,
<figureCitation id="13132A05DE5DFFF79B47464EFE126606" box="[284,437,259,286]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1871,1895]" captionTargetBox="[283,1303,525,1847]" captionTargetId="figure-196@6.[283,1303,525,1847]" captionTargetPageId="6" captionText="FIGURES 5–8. Endoclita spp. head, thorax, and appendages. Endoclita phuthoensis sp. n.: (5a) head and thorax, (6a) antenna, (7b) labial palpus and (8b) legs (foreleg left, midleg centre, hindleg right); Endoclita coomani: (5b–c) head and thorax, (6b) antenna, (7a) labial palpus, and legs (8a, c–d respectively)—8a (top foreleg, centre midleg, bottom hindleg), 8c (foreleg left, midleg centre, hindleg right), 8d (top foreleg, centre midleg, bottom hindleg). Unscaled. Photos 6–6 by Duy Long Pham, 7–8 by John Grehan." figureDoi="http://doi.org/10.5281/zenodo.11233504" httpUri="https://zenodo.org/record/11233504/files/figure.png" pageId="2" pageNumber="65">5a, 6a, 7b, 8b</figureCitation>
,
<figureCitation id="13132A05DE5DFFF79B9A464EFDB76606" box="[449,528,259,286]" captionStart="FIGURE 9" captionStartId="7.[151,250,1391,1415]" captionTargetBox="[215,1372,181,1367]" captionTargetId="figure-18@7.[215,1372,181,1367]" captionTargetPageId="7" captionText="FIGURE 9. Endoclita spp. wing venation and forewing patterns. Endoclita phuthoensis sp. n.: (9a) venation, (9c) forewing basal portion, (9f) forewing ornamentation; Endoclita coomani: (9b) venation, (9d–e) forewing basal portion, (9g) forewing ornamentation." figureDoi="http://doi.org/10.5281/zenodo.11233506" httpUri="https://zenodo.org/record/11233506/files/figure.png" pageId="2" pageNumber="65">9a, c, f</figureCitation>
,
<figureCitation id="13132A05DE5DFFF79840464EFDC66606" box="[539,609,259,285]" captionStart="FIGURES 10–11" captionStartId="8.[151,264,1689,1713]" captionTargetBox="[151,1436,181,1665]" captionTargetId="figure-23@8.[151,1436,181,1665]" captionTargetPageId="8" captionText="FIGURES 10–11. Endoclita spp. abdominal sclerites. Endoclita phuthoensis sp. n. (FPRC#68): (10a) habitus lateral view of tergum I and tergosternal bar, (10b) tergosternal sclerite, (10c) anterior abdominal segments, (10d) posterior abdominal segments (SVIII broken away from SVII), (10e) ventral habitus view of lamella antevaginalis. Endoclita coomani: (11a) ventral habitus view of lamella antevaginalis, (11b) ventral habitus view of SVIII and pseudotegumen, (11c) anterior abdominal segments, (11d) posterior abdominal segments, (11e) anterior abdominal segments, (11f) tergosternal sclerite, (11g) posterior abdominal segments. Unscaled. Photos by John Grehan (10a), Duy Long Pham (10e, 11a–b), and Loren Jones (10b–d, 11c–g)." figureDoi="http://doi.org/10.5281/zenodo.11233508" httpUri="https://zenodo.org/record/11233508/files/figure.png" pageId="2" pageNumber="65">10a–e</figureCitation>
,
<figureCitation id="13132A05DE5DFFF79837464EFD6C6605" box="[620,715,259,286]" captionStart="FIGURE 12" captionStartId="9.[151,250,1838,1862]" captionTargetBox="[174,1413,345,1814]" captionTargetId="figure-82@9.[174,1413,345,1814]" captionTargetPageId="9" captionText="FIGURE 12. Endoclita spp. female external and internal genitalia. Endoclita phuthoensis sp. n.: (12a) posterior view, (12c) ductus and corpus bursae, (12e) spermatheca; E. coomani: (12b) posterior view, (12d) ductus and corpus bursae. Unscaled. Photos by Loren Jones." figureDoi="http://doi.org/10.5281/zenodo.11233510" httpUri="https://zenodo.org/record/11233510/files/figure.png" pageId="2" pageNumber="65">12a, c, e</figureCitation>
,
<figureCitation id="13132A05DE5DFFF7988C464EFCBB6606" box="[727,796,259,285]" captionStart="FIGURE 20" captionStartId="17.[151,250,1200,1224]" captionTargetBox="[151,1436,181,1176]" captionTargetId="figure-18@17.[151,1436,181,1176]" captionTargetPageId="17" captionText="FIGURE 20. Live adults of Endoclita phuthoensis sp. n. (20b, e) and E. coomani (20a, c–d, f) at rest. Photos by Nguyen Minh Chi." figureDoi="http://doi.org/10.5281/zenodo.11233537" httpUri="https://zenodo.org/record/11233537/files/figure.png" pageId="2" pageNumber="65">20b, e</figureCitation>
)
</paragraph>
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<caption id="DF576608DE5DFFF79ACC4350FEF06342" ID-DOI="http://doi.org/10.5281/zenodo.11233496" ID-Zenodo-Dep="11233496" httpUri="https://zenodo.org/record/11233496/files/figure.png" pageId="2" pageNumber="65" startId="2.[151,250,1053,1077]" targetBox="[324,1263,309,1029]" targetPageId="2" targetType="figure">
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<emphasis id="B95CEA92DE5DFFF79ACC4350FEB4632E" bold="true" box="[151,275,1053,1077]" pageId="2" pageNumber="65">FIGURE 1.</emphasis>
<taxonomicName id="4C284D03DE5DFFF79B424350FE5C632E" authority="Pham &amp; Chi &amp; Grehan &amp; Khai &amp; Ignatev &amp; Jones &amp; Mielke &amp; Quang &amp; Yakovlev, 2024" authorityName="Pham &amp; Chi &amp; Grehan &amp; Khai &amp; Ignatev &amp; Jones &amp; Mielke &amp; Quang &amp; Yakovlev" authorityYear="2024" box="[281,507,1053,1077]" class="Insecta" family="Hepialidae" genus="Endoclita" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="phuthoensis" status="sp. nov.">
<emphasis id="B95CEA92DE5DFFF79B424350FE5C632E" box="[281,507,1053,1077]" italics="true" pageId="2" pageNumber="65">Endoclita phuthoensis</emphasis>
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<taxonomicNameLabel id="A26F57E9DE5DFFF798594350FD9C632E" box="[514,571,1053,1077]" pageId="2" pageNumber="65" rank="species">sp. n.</taxonomicNameLabel>
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: holotype female (1a dorsal, 1b ventral), and two paratype females (1c–d). Photos by Nguyen Minh Chi.
</paragraph>
</caption>
<caption id="DF576608DE5DFFF79ACC4030FDAD60C6" ID-DOI="http://doi.org/10.5281/zenodo.11233498" ID-Zenodo-Dep="11233498" httpUri="https://zenodo.org/record/11233498/files/figure.png" pageId="2" pageNumber="65" startId="2.[151,250,1917,1941]" targetBox="[325,1262,1137,1893]" targetPageId="2" targetType="figure">
<paragraph id="8B973680DE5DFFF79ACC4030FDAD60C6" blockId="2.[151,1437,1917,2014]" pageId="2" pageNumber="65">
<emphasis id="B95CEA92DE5DFFF79ACC4030FEB4608E" bold="true" box="[151,275,1917,1941]" pageId="2" pageNumber="65">FIGURE 2.</emphasis>
<taxonomicName id="4C284D03DE5DFFF79B424030FE7E608E" baseAuthorityName="Viette" baseAuthorityYear="1949" box="[281,473,1917,1941]" class="Insecta" family="Hepialidae" genus="Endoclita" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="coomani">
<emphasis id="B95CEA92DE5DFFF79B424030FE7E608E" box="[281,473,1917,1941]" italics="true" pageId="2" pageNumber="65">Endoclita coomani</emphasis>
</taxonomicName>
(a–f): specimens from a
<taxonomicName id="4C284D03DE5DFFF7988D4030FCE0608E" box="[726,839,1917,1941]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="65" phylum="Tracheophyta" rank="genus">
<emphasis id="B95CEA92DE5DFFF7988D4030FCE0608E" box="[726,839,1917,1941]" italics="true" pageId="2" pageNumber="65">Eucalyptus</emphasis>
</taxonomicName>
plantation in Yen Lap district, Phu Tho province, Vietnam. Note size range of females, with smallest matching the two male specimens, including the holotype male. Photos 2a–e by Nguyen Minh Chi, 2f by Jöel Minet.
</paragraph>
</caption>
<subSubSection id="C332650BDE5CFFF69ACC47DAFE31667C" pageId="3" pageNumber="66" type="materials_examined">
<paragraph id="8B973680DE5CFFF69ACC47DAFE356610" blockId="3.[151,1437,150,359]" pageId="3" pageNumber="66">
<materialsCitation id="3B403CDDDE5CFFF69ACC47DAFDCA67F7" collectingDate="2022-10-16" collectingDateMax="2022-11-11" collectingDateMin="2022-10-16" collectionCode="FPRC" collectorName="Nguyen Minh Chi &amp; Duy Long Pham" country="Vietnam" county="Adult" location="Host" municipality="Yen Lap district" pageId="3" pageNumber="66" specimenCode="FPRC#51" specimenCount="2" specimenCount-female="2" stateProvince="Phu Tho" typeStatus="holotype">
<emphasis id="B95CEA92DE5CFFF69ACC47DAFEB267AA" bold="true" box="[151,277,151,177]" pageId="3" pageNumber="66">
<typeStatus id="54938822DE5CFFF69ACC47DAFF5D67B4" box="[151,250,151,175]" pageId="3" pageNumber="66" type="holotype">Holotype</typeStatus>
<specimenCount id="9D2EFD09DE5CFFF69B5947D5FEB267AA" box="[258,277,152,177]" count="1" pageId="3" pageNumber="66" type="female">♀</specimenCount>
</emphasis>
(with the following labels separated by forward slashes): / “
<specimenCode id="DB8E9EFBDE5CFFF699D347DBFC5667B5" box="[904,1009,150,174]" collectionCode="FPRC" pageId="3" pageNumber="66">FPRC#51</specimenCode>
,
<location id="8EF7605BDE5CFFF699A747DAFB9767B5" LSID="urn:lsid:plazi:treatment:03818796DE5DFFF69ACC4792FE2A6524:8EF7605BDE5CFFF699A747DAFB9767B5" box="[1020,1072,151,174]" country="Vietnam" county="Adult" municipality="Yen Lap district" name="Host" pageId="3" pageNumber="66" stateProvince="Phu Tho">Host</location>
:
<taxonomicName id="4C284D03DE5CFFF69E6047DAFB0B67B4" box="[1083,1196,151,175]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="66" phylum="Tracheophyta" rank="genus">
<emphasis id="B95CEA92DE5CFFF69E6047DAFB0B67B4" box="[1083,1196,151,175]" italics="true" pageId="3" pageNumber="66">Eucalyptus</emphasis>
</taxonomicName>
hybrid (
<taxonomicName id="4C284D03DE5CFFF69F5D47DAFA2167B4" box="[1286,1414,151,175]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="66" phylum="Tracheophyta" rank="species" species="urophylla">
<emphasis id="B95CEA92DE5CFFF69F5D47DAFA2167B4" box="[1286,1414,151,175]" italics="true" pageId="3" pageNumber="66">E. urophylla</emphasis>
</taxonomicName>
×
<taxonomicName id="4C284D03DE5CFFF69A9C47FBFE9767D6" box="[199,304,181,205]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="66" phylum="Tracheophyta" rank="species" species="grandis">
<emphasis id="B95CEA92DE5CFFF69A9C47FBFE9767D6" box="[199,304,181,205]" italics="true" pageId="3" pageNumber="66">E. grandis</emphasis>
</taxonomicName>
),
<location id="8EF7605BDE5CFFF69B1D47F8FE0367D6" LSID="urn:lsid:plazi:treatment:03818796DE5DFFF69ACC4792FE2A6524:8EF7605BDE5CFFF69B1D47F8FE0367D6" box="[326,420,181,205]" country="Vietnam" county="Adult" municipality="Yen Lap district" name="Location" pageId="3" pageNumber="66" stateProvince="Phu Tho">Location</location>
:
<collectingMunicipality id="6BF3ACFADE5CFFF69BF747FBFDF667D6" box="[428,593,181,205]" pageId="3" pageNumber="66">Yen Lap district</collectingMunicipality>
,
<collectingRegion id="49ECF862DE5CFFF6980047F8FCB467D6" box="[603,787,181,205]" country="Vietnam" name="Phu Tho" pageId="3" pageNumber="66">Phu Tho province</collectingRegion>
,
<collectingCountry id="F33F7610DE5CFFF6994647F8FCD267D6" box="[797,885,181,205]" name="Vietnam" pageId="3" pageNumber="66">Vietnam</collectingCountry>
,
<collectingCounty id="62F64E0CDE5CFFF6992547F8FC1F67D6" box="[894,952,181,205]" pageId="3" pageNumber="66">Adult</collectingCounty>
emergence:
<date id="FF961040DE5CFFF69E6247F8FB0967D6" box="[1081,1198,181,205]" pageId="3" pageNumber="66" value="2022-10-16">
<collectingDate id="EFD2E9A8DE5CFFF69E6247F8FB0967D6" box="[1081,1198,181,205]" pageId="3" pageNumber="66" value="2022-10-16">16/10/2022</collectingDate>
</date>
–
<date id="FF961040DE5CFFF69EE747F8FA8A67D6" box="[1212,1325,181,205]" pageId="3" pageNumber="66" value="2022-11-11">
<collectingDate id="EFD2E9A8DE5CFFF69EE747F8FA8A67D6" box="[1212,1325,181,205]" pageId="3" pageNumber="66" value="2022-11-11">11/11/2022</collectingDate>
</date>
,
<collectorName id="26DD5356DE5CFFF69F6C47F8FA3E67D6" box="[1335,1433,181,205]" pageId="3" pageNumber="66">Collector</collectorName>
:
<collectorName id="26DD5356DE5CFFF69A9C4799FE2267F7" box="[199,389,212,236]" pageId="3" pageNumber="66">Nguyen Minh Chi</collectorName>
and Duy Long Pham/
</materialsCitation>
<materialsCitation id="3B403CDDDE5CFFF6982D4799FE356610" collectingDate="2022-10-16" collectingDateMax="2022-11-11" collectingDateMin="2022-10-16" collectionCode="FPRC" country="Vietnam" county="Adult" location="Host" municipality="Yen Lap district" pageId="3" pageNumber="66" specimenCode="FPRC#51" specimenCount="2" specimenCount-female="2" stateProvince="Phu Tho" typeStatus="holotype">
<typeStatus id="54938822DE5CFFF6982D4799FD4567F7" box="[630,738,212,236]" pageId="3" pageNumber="66" type="holotype">Holotypus</typeStatus>
,
<taxonomicName id="4C284D03DE5CFFF698B44799FB4767F7" authority="♀, Long, Chi &amp; Grehan." authorityName="Long, Chi &amp; Grehan." box="[751,1248,212,238]" class="Insecta" family="Hepialidae" genus="Endoclita" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="phuthoensis">
<emphasis id="B95CEA92DE5CFFF698B44799FC7467F7" box="[751,979,212,236]" italics="true" pageId="3" pageNumber="66">Endoclita phuthoensis</emphasis>
<emphasis id="B95CEA92DE5CFFF699874798FC4867F5" bold="true" box="[988,1007,213,238]" pageId="3" pageNumber="66">
<specimenCount id="9D2EFD09DE5CFFF699874798FC4867F5" box="[988,1007,213,238]" count="1" pageId="3" pageNumber="66" type="female">♀</specimenCount>
</emphasis>
, Long, Chi &amp; Grehan.
</taxonomicName>
”
<collectorName id="26DD5356DE5CFFF69EAA4799FAFF67F7" box="[1265,1368,212,236]" pageId="3" pageNumber="66">Deposited</collectorName>
in the collection of
<collectionCode id="ED39AE45DE5CFFF69B1647BEFE296610" box="[333,398,243,267]" pageId="3" pageNumber="66">FPRC</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B973680DE5CFFF69ACC465CFE31667C" blockId="3.[151,1437,150,359]" pageId="3" pageNumber="66">
<materialsCitation id="3B403CDDDE5CFFF69ACC465CFCB16632" box="[151,790,273,300]" collectionCode="FPRC" pageId="3" pageNumber="66" specimenCode="FPRC#53, 54, 60-62, 65-68, FPRC#72-73" specimenCount="11" specimenCount-female="11" typeStatus="paratype">
<emphasis id="B95CEA92DE5CFFF69ACC465CFEA46632" bold="true" box="[151,259,273,297]" pageId="3" pageNumber="66">
<typeStatus id="54938822DE5CFFF69ACC465CFEA46632" box="[151,259,273,297]" pageId="3" pageNumber="66" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D2EFD09DE5CFFF69B48465CFEE16637" box="[275,326,273,300]" count="11" pageId="3" pageNumber="66" type="female">
11
<emphasis id="B95CEA92DE5CFFF69B68465EFEE16637" bold="true" box="[307,326,275,300]" pageId="3" pageNumber="66">♀</emphasis>
</specimenCount>
:
<specimenCode id="DB8E9EFBDE5CFFF69B0E465CFE1C6632" box="[341,443,273,297]" collectionCode="FPRC" pageId="3" pageNumber="66">FPRC#53</specimenCode>
,
<specimenCode id="DB8E9EFBDE5CFFF69B9C465CFE446632" box="[455,483,273,297]" collectionCode="FPRC" pageId="3" pageNumber="66">54</specimenCode>
,
<specimenCode id="DB8E9EFBDE5CFFF69BB4465CFD976632" box="[495,560,273,297]" collectionCode="FPRC" pageId="3" pageNumber="66">60–62</specimenCode>
,
<specimenCode id="DB8E9EFBDE5CFFF69867465CFDD96632" box="[572,638,273,297]" collectionCode="FPRC" pageId="3" pageNumber="66">65–68</specimenCode>
,
<specimenCode id="DB8E9EFBDE5CFFF698D1465CFCB16632" box="[650,790,273,297]" collectionCode="FPRC" pageId="3" pageNumber="66">FPRC#72–73</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B403CDDDE5CFFF69979465FFEF06653" collectingDate="2022-10-16" collectingDateMax="2022-11-11" collectingDateMin="2022-10-16" collectionCode="CUIC, FPRC" collectorName="Nguyen Minh Chi and Duy Long Pham" country="Vietnam" county="Adult" location="Host" municipality="Yen Lap district" pageId="3" pageNumber="66" specimenCode="FRPC#53, 68" specimenCount="1" stateProvince="Phu Tho" typeStatus="paratype">
same site data as holotype (
<specimenCode id="DB8E9EFBDE5CFFF69E1B465CFB006632" box="[1088,1191,273,297]" collectionCode="FRPC" pageId="3" pageNumber="66">FRPC#53</specimenCode>
,
<specimenCode id="DB8E9EFBDE5CFFF69EE8465CFB6B6632" box="[1203,1228,273,297]" collectionCode="FRPC" pageId="3" pageNumber="66">68</specimenCode>
deposited in
<collectionCode id="ED39AE45DE5CFFF69F02465CFA3F6632" box="[1369,1432,273,297]" country="USA" httpUri="http://grbio.org/cool/sxrc-6x8s" name="Cornell University Insect Collection" pageId="3" pageNumber="66" type="University or college">CUIC</collectionCode>
, rest in
<collectionCode id="ED39AE45DE5CFFF69B56467DFEF76653" box="[269,336,304,328]" pageId="3" pageNumber="66">FPRC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B403CDDDE5CFFF69B3A467DFC8C6653" box="[353,811,304,330]" collectionCode="FPRC" country="Vietnam" pageId="3" pageNumber="66" specimenCode="FPRC#54, 60-62, 65-67, FPRC#72-73" specimenCount="9" specimenCount-female="9" stateProvince="Phu Tho" typeStatus="paratype">
<specimenCount id="9D2EFD09DE5CFFF69B3A467DFE206651" box="[353,391,304,330]" count="9" pageId="3" pageNumber="66" type="female">
9
<emphasis id="B95CEA92DE5CFFF69B2F467CFE206651" bold="true" box="[372,391,305,330]" pageId="3" pageNumber="66">♀</emphasis>
</specimenCount>
:
<specimenCode id="DB8E9EFBDE5CFFF69BCE467DFE5B6653" box="[405,508,304,328]" collectionCode="FPRC" pageId="3" pageNumber="66">FPRC#54</specimenCode>
,
<specimenCode id="DB8E9EFBDE5CFFF6985D467DFDEF6653" box="[518,584,304,328]" collectionCode="FPRC" pageId="3" pageNumber="66">60–62</specimenCode>
,
<specimenCode id="DB8E9EFBDE5CFFF69808467DFD336653" box="[595,660,304,328]" collectionCode="FPRC" pageId="3" pageNumber="66">65–67</specimenCode>
,
<specimenCode id="DB8E9EFBDE5CFFF698C4467DFC8C6653" box="[671,811,304,328]" collectionCode="FPRC" pageId="3" pageNumber="66">FPRC#72–73</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B403CDDDE5CFFF6996D467DFBCF6653" box="[822,1128,304,328]" collectionCode="FPRC" country="Vietnam" pageId="3" pageNumber="66" specimenCount="1" stateProvince="Phu Tho" typeStatus="paratype">
same site as
<typeStatus id="54938822DE5CFFF699EC467DFBB76653" box="[951,1040,304,328]" pageId="3" pageNumber="66" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED39AE45DE5CFFF69E46467DFBC66653" box="[1053,1121,304,328]" pageId="3" pageNumber="66">FPRC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B403CDDDE5CFFF69E29467DFE35667C" collectionCode="CUIC" country="Vietnam" pageId="3" pageNumber="66" specimenCount="2" specimenCount-female="2" stateProvince="Phu Tho" typeStatus="paratype">
<specimenCount id="9D2EFD09DE5CFFF69E29467DFB3E6651" box="[1138,1177,304,330]" count="2" pageId="3" pageNumber="66" type="female">
2
<emphasis id="B95CEA92DE5CFFF69EDD467CFB3E6651" bold="true" box="[1158,1177,305,330]" pageId="3" pageNumber="66">♀</emphasis>
</specimenCount>
: FRPC#53, 68, same site as
<typeStatus id="54938822DE5CFFF69AB84602FE9B667C" box="[227,316,335,359]" pageId="3" pageNumber="66" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED39AE45DE5CFFF69B124602FE2C667C" box="[329,395,335,359]" country="USA" httpUri="http://grbio.org/cool/sxrc-6x8s" name="Cornell University Insect Collection" pageId="3" pageNumber="66" type="University or college">CUIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C332650BDE5CFFF69ACC46D8FE2A6524" pageId="3" pageNumber="66" type="diagnosis">
<paragraph id="8B973680DE5CFFF69ACC46D8FE2A6524" blockId="3.[151,1437,405,575]" pageId="3" pageNumber="66">
<emphasis id="B95CEA92DE5CFFF69ACC46D8FEB566B4" bold="true" box="[151,274,405,431]" pageId="3" pageNumber="66">Diagnosis.</emphasis>
The forewing costal margin is almost liner, and this feature separates this species from all
<taxonomicName id="4C284D03DE5CFFF69EA346D8FAC266B4" authorityName="Felder" authorityYear="1874" box="[1272,1381,405,431]" class="Insecta" family="Hepialidae" genus="Endoclita" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B95CEA92DE5CFFF69EA346D8FAC266B4" box="[1272,1381,405,431]" italics="true" pageId="3" pageNumber="66">Endoclita</emphasis>
</taxonomicName>
with a lobe-like expansion near Sc1 (
<figureCitation id="13132A05DE5CFFF69BA046F4FDEB66C8" box="[507,588,441,467]" captionStart="FIGURE 4" captionStartId="3.[151,250,1925,1949]" captionTargetBox="[283,1304,1522,1901]" captionTargetId="figure-261@3.[283,1304,1522,1901]" captionTargetPageId="3" captionText="FIGURE 4. Endoclita spp. Characteristics of the forewing costal margin and discal cells that are not present in E. phuthoesnsis sp.n. Photos by Nguyen Minh Chi." figureDoi="http://doi.org/10.5281/zenodo.11233502" httpUri="https://zenodo.org/record/11233502/files/figure.png" pageId="3" pageNumber="66">Fig. 4a</figureCitation>
). Distinguished from linear costal margin species in eastern Asia by absence of a sub-triangular dark shaded patch over outer forewing discal region (
<figureCitation id="13132A05DE5CFFF6999B4690FBB366EC" box="[960,1044,477,503]" captionStart="FIGURE 4" captionStartId="3.[151,250,1925,1949]" captionTargetBox="[283,1304,1522,1901]" captionTargetId="figure-261@3.[283,1304,1522,1901]" captionTargetPageId="3" captionText="FIGURE 4. Endoclita spp. Characteristics of the forewing costal margin and discal cells that are not present in E. phuthoesnsis sp.n. Photos by Nguyen Minh Chi." figureDoi="http://doi.org/10.5281/zenodo.11233502" httpUri="https://zenodo.org/record/11233502/files/figure.png" pageId="3" pageNumber="66">Fig. 4b</figureCitation>
), or by absence of forewing stigma in comparison to eastern Asian linear costal margin species
<taxonomicName id="4C284D03DE5CFFF69974454FFC046500" baseAuthorityName="Hampson" baseAuthorityYear="1893" box="[815,931,514,539]" class="Insecta" family="Hepialidae" genus="Endoclita" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="auratus">
<emphasis id="B95CEA92DE5CFFF69974454FFC046500" box="[815,931,514,539]" italics="true" pageId="3" pageNumber="66">E. auratus</emphasis>
</taxonomicName>
(
<figureCitation id="13132A05DE5CFFF699EA454CFBA66500" box="[945,1025,513,539]" captionStart="FIGURE 3" captionStartId="3.[151,250,1366,1390]" captionTargetBox="[280,1308,599,1342]" captionTargetId="figure-196@3.[280,1308,599,1342]" captionTargetPageId="3" captionText="FIGURE 3. Endoclita spp. Endoclita marginenotatus: (a) Mt. Fansipan, Vietnam, ZSM (photo by Nikolai Ignatev), (b) holotype male, Sichuan, China, NHMUK (photo by David Lees: copyright David C. Lees/Trustees of NHMUK); E. auratus (c) Bernardmyo, Myanmar, 5500–7000', May 1890, NHMUK (copyright Alessandro Giusti/Trustees of the Natural History Museum UK); E. undulifer (d), Khasia Hills, October 1896, SAMA (photo by Peter Hudson)." figureDoi="http://doi.org/10.5281/zenodo.11233500" httpUri="https://zenodo.org/record/11233500/files/figure.png" pageId="3" pageNumber="66">Fig. 3c</figureCitation>
),
<taxonomicName id="4C284D03DE5CFFF69E4D454FFB4B6500" baseAuthorityName="Leech" baseAuthorityYear="1898" box="[1046,1260,514,539]" class="Insecta" family="Hepialidae" genus="Endoclita" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="marginentotatus">
<emphasis id="B95CEA92DE5CFFF69E4D454FFB4B6500" box="[1046,1260,514,539]" italics="true" pageId="3" pageNumber="66">E. marginentotatus</emphasis>
</taxonomicName>
(
<figureCitation id="13132A05DE5CFFF69EA1454CFAC16500" box="[1274,1382,513,539]" captionStart="FIGURE 3" captionStartId="3.[151,250,1366,1390]" captionTargetBox="[280,1308,599,1342]" captionTargetId="figure-196@3.[280,1308,599,1342]" captionTargetPageId="3" captionText="FIGURE 3. Endoclita spp. Endoclita marginenotatus: (a) Mt. Fansipan, Vietnam, ZSM (photo by Nikolai Ignatev), (b) holotype male, Sichuan, China, NHMUK (photo by David Lees: copyright David C. Lees/Trustees of NHMUK); E. auratus (c) Bernardmyo, Myanmar, 5500–7000', May 1890, NHMUK (copyright Alessandro Giusti/Trustees of the Natural History Museum UK); E. undulifer (d), Khasia Hills, October 1896, SAMA (photo by Peter Hudson)." figureDoi="http://doi.org/10.5281/zenodo.11233500" httpUri="https://zenodo.org/record/11233500/files/figure.png" pageId="3" pageNumber="66">Fig. 3a–b</figureCitation>
) and
<taxonomicName id="4C284D03DE5CFFF69ACC456BFEBA6524" baseAuthorityName="Walker" baseAuthorityYear="1869" box="[151,285,549,575]" class="Insecta" family="Hepialidae" genus="Endoclita" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="undulifer">
<emphasis id="B95CEA92DE5CFFF69ACC456BFEBA6524" box="[151,285,549,575]" italics="true" pageId="3" pageNumber="66">E. undulifer</emphasis>
</taxonomicName>
(
<figureCitation id="13132A05DE5CFFF69B774568FE276524" box="[300,384,549,575]" captionStart="FIGURE 3" captionStartId="3.[151,250,1366,1390]" captionTargetBox="[280,1308,599,1342]" captionTargetId="figure-196@3.[280,1308,599,1342]" captionTargetPageId="3" captionText="FIGURE 3. Endoclita spp. Endoclita marginenotatus: (a) Mt. Fansipan, Vietnam, ZSM (photo by Nikolai Ignatev), (b) holotype male, Sichuan, China, NHMUK (photo by David Lees: copyright David C. Lees/Trustees of NHMUK); E. auratus (c) Bernardmyo, Myanmar, 5500–7000', May 1890, NHMUK (copyright Alessandro Giusti/Trustees of the Natural History Museum UK); E. undulifer (d), Khasia Hills, October 1896, SAMA (photo by Peter Hudson)." figureDoi="http://doi.org/10.5281/zenodo.11233500" httpUri="https://zenodo.org/record/11233500/files/figure.png" pageId="3" pageNumber="66">Fig. 3d</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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