750 lines
104 KiB
XML
750 lines
104 KiB
XML
<document id="52D6F6CE883DF9E1C07064402849617E" ID-CLB-Dataset="24239" ID-DOI="10.11646/zootaxa.4729.1.1" ID-GBIF-Dataset="cf71c5fb-e106-440f-9f50-530e723f187f" ID-ISSN="1175-5326" ID-Zenodo-Dep="3628946" ID-ZooBank="A78BFE69-34BC-4D75-8862-CE789B7C0A2C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1580179969094" checkinUser="plazi" docAuthor="Quah, Evan S. H., Grismer, L. Lee, Lim, Kelvin K. P., Anuar, M. S. Shahrul & Chan, Kin Onn" docDate="2020" docId="BF3E5102FFF1FFA2BBB1FE79FCBDFF0D" docLanguage="en" docName="zootaxa.4729.1.1.pdf" docOrigin="Zootaxa 4729 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Asthenodipsas borneensis Quah & Grismer & Lim & Anuar & Chan 2020, sp. nov." docType="treatment" docUuid="51657C98-84D4-4B74-8AF3-7695E8D72FFB" docUuidSource="ZooBank" docVersion="5" lastPageNumber="17" masterDocId="4307297AFFFDFFB3BB26FFEBFFACFFF6" masterDocTitle="A taxonomic revision of Asthenodipsas malaccana Peters, 1864 (Squamata Pareidae) with a description of a new species from Borneo" masterLastPageNumber="24" masterPageNumber="1" pageNumber="13" updateTime="1698779413567" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="6439DED827A67A81C3DA1F804E71C438" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="A2A11E070965B057AF8463B6F2A599E8">
|
||
<mods:title id="4214C42EF505EEB0A4599D5EE8E7307B">A taxonomic revision of Asthenodipsas malaccana Peters, 1864 (Squamata Pareidae) with a description of a new species from Borneo</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="C511AFE390B5C1A834DE22809D44C628" type="personal">
|
||
<mods:role id="C2811DABA3B162FC7B8DABCD6517D7B7">
|
||
<mods:roleTerm id="F1A0BFF860E9FA5864CBC3B6C34993E5">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="6C6A2D766D25ADF24CA8004D1B48A4E6">Quah, Evan S. H.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="D26FD385CEF682A21DEA00BB3DCDDF49" type="personal">
|
||
<mods:role id="9721C2F19A3F0CD5951793A3D1B24BC0">
|
||
<mods:roleTerm id="4FFE9D2BF41EFDF727811FCBDA636F85">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="4824A0B4FF79AC3B538264C684BAAD71">Grismer, L. Lee</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="F83853665D87CCE5CC6AED92F24817F3" type="personal">
|
||
<mods:role id="42AE91CB310FBB7909DB1B58F25E1273">
|
||
<mods:roleTerm id="7DD1DAC9FE38D2613F8682C27752AEE4">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="66238731541F8C07A786729D1DDACA85">Lim, Kelvin K. P.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="0CF63D08E8955D7056D4DEAF4B828B0D" type="personal">
|
||
<mods:role id="B84B569BB1B87DBC889095FC407B03D0">
|
||
<mods:roleTerm id="725B25B69448183D6A129FECF8607897">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B863F37D70E3B8F0F702D7FCDCACEA2A">Anuar, M. S. Shahrul</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="518B5DD5B5A01D8B61F463F41AF656D8" type="personal">
|
||
<mods:role id="D8023B551C98F782DB3B6CCFE8335F7A">
|
||
<mods:roleTerm id="E7A47F23EB65959044F296CC198ECFAC">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="36D2AE42DAACDDC7886F115542A2DE4C">Chan, Kin Onn</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="174C582EC8217B2ECEA5BBB1F81AD972">text</mods:typeOfResource>
|
||
<mods:relatedItem id="FE1EACD178B6A457D227C7B0ED025777" type="host">
|
||
<mods:titleInfo id="BB14BA26FBD76A9ADEBF77C86059F4AF">
|
||
<mods:title id="F930141AB3E113C0D7E81252B7A81BFB">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="7B1AA97DA3389A5DCC8FE7D232A95655">
|
||
<mods:date id="4787511C60B5131D33E056925B4723F2">2020</mods:date>
|
||
<mods:detail id="A30D2A4401405FBEEC1C09E1348C6FAA" type="pubDate">
|
||
<mods:number id="45E3445B57D64A8F79A94FC8323043AF">2020-01-28</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="B57E21492853DF90D1DA4414B1EB84D9" type="volume">
|
||
<mods:number id="0B1A8ECB4CB69AA2322BBBEA06C81FFF">4729</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="B986030657C5CC6021E9515B26024325" type="issue">
|
||
<mods:number id="15D420EFF3386989C182AAABAC37F289">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="7FC3494512DA5310F7F01D35410387BB" unit="page">
|
||
<mods:start id="0804254E1949074BE4051825563E6F5C">1</mods:start>
|
||
<mods:end id="5BA4E890363A96C93486577ADF74C82F">24</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="8D499F209972A9C9AF58BBFEE5545C94">journal article</mods:classification>
|
||
<mods:identifier id="0E0D3E6490D85D5D3D33C927C834CD99" type="CLB-Dataset">24239</mods:identifier>
|
||
<mods:identifier id="C91956424B53C1B1449836189BE0AD32" type="DOI">10.11646/zootaxa.4729.1.1</mods:identifier>
|
||
<mods:identifier id="DE513129C54CCD06FC04B32B4643B357" type="GBIF-Dataset">cf71c5fb-e106-440f-9f50-530e723f187f</mods:identifier>
|
||
<mods:identifier id="E9BFE96F571BA950A09901C2F1BF3008" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="1E787AEA78A55C8D2BB5C041F32F5A75" type="Zenodo-Dep">3628946</mods:identifier>
|
||
<mods:identifier id="BD341FBF90F5EE473737D8717AEA23D8" type="ZooBank">A78BFE69-34BC-4D75-8862-CE789B7C0A2C</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="BF3E5102FFF1FFA2BBB1FE79FCBDFF0D" ID-GBIF-Taxon="162328480" LSID="urn:lsid:zoobank.org:act:51657C98-84D4-4B74-8AF3-7695E8D72FFB" httpUri="http://treatment.plazi.org/id/BF3E5102FFF1FFA2BBB1FE79FCBDFF0D" lastPageId="17" lastPageNumber="17" pageId="12" pageNumber="13">
|
||
<subSubSection id="7F8DB39FFFF1FFBFBBB1FE79FD98FE5B" box="[151,564,402,429]" pageId="12" pageNumber="13" type="nomenclature">
|
||
<paragraph id="3728E014FFF1FFBFBBB1FE79FD98FE5B" blockId="12.[151,957,402,464]" box="[151,564,402,429]" pageId="12" pageNumber="13">
|
||
<heading id="6C605778FFF1FFBFBBB1FE79FD98FE5B" bold="true" box="[151,564,402,429]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
|
||
<emphasis id="05E33C06FFF1FFBFBBB1FE79FD98FE5B" bold="true" box="[151,564,402,429]" pageId="12" pageNumber="13">
|
||
<taxonomicName id="F0979B97FFF1FFBFBBB1FE79FE63FE5B" authority="Quah & Grismer & Lim & Anuar & Chan, 2020" authorityName="Quah & Grismer & Lim & Anuar & Chan" authorityYear="2020" box="[151,463,402,429]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="05E33C06FFF1FFBFBBB1FE79FE63FE5B" bold="true" box="[151,463,402,429]" italics="true" pageId="12" pageNumber="13">Asthenodipsas borneensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1ED0817DFFF1FFBFBAF1FE78FD98FE5B" box="[471,564,403,429]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7F8DB39FFFF1FFBFBBB1FE5EFC11FE26" box="[151,957,437,464]" pageId="12" pageNumber="13" type="description">
|
||
<paragraph id="3728E014FFF1FFBFBBB1FE5EFC11FE26" blockId="12.[151,957,402,464]" box="[151,957,437,464]" pageId="12" pageNumber="13">
|
||
<uri id="4306EC16FFF1FFBFBBB1FE5EFC11FE26" box="[151,957,437,464]" pageId="12" pageNumber="13">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid id="4331DAC1FFF1FFBFBA8DFE5EFC11FE26" box="[427,957,437,464]" pageId="12" pageNumber="13">51657C98-84D4-4B74-8AF3-7695E8D72FFB</uuid>
|
||
</uri>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSection id="5E18F9FFFFF1FFA2BBB1FE14FCBDFF0D" lastPageId="17" lastPageNumber="18" pageId="12" pageNumber="13" type="multiple">
|
||
<subSubSection id="7F8DB39FFFF1FFBFBBB1FE14FEBEFDCA" pageId="12" pageNumber="13" type="vernacular_names">
|
||
<paragraph id="3728E014FFF1FFBFBBB1FE14FDECFDEE" blockId="12.[151,576,510,572]" box="[151,576,510,537]" pageId="12" pageNumber="13">
|
||
<heading id="6C605778FFF1FFBFBBB1FE14FDECFDEE" bold="true" box="[151,576,510,537]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
|
||
<emphasis id="05E33C06FFF1FFBFBBB1FE14FDECFDEE" bold="true" box="[151,576,510,537]" pageId="12" pageNumber="13">Bornean Dark-necked Slug Snake</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="3728E014FFF1FFBFBBB1FDC9FEBEFDCA" blockId="12.[151,576,510,572]" box="[151,274,546,572]" pageId="12" pageNumber="13">
|
||
<figureCitation id="AFACFC91FFF1FFBFBBB1FDC9FF75FDCA" box="[151,217,546,572]" captionStart="FIGURE 2" captionStartId="13.[151,250,1688,1713]" captionTargetBox="[151,1436,181,1662]" captionTargetId="figure@13.[151,1436,181,1662]" captionTargetPageId="13" captionText="FIGURE 2. A. Dorsum of holotype of Asthenodipsas borneensis sp. nov. (FMNH 243935). B. Venter of holotype of A. borneensis sp. nov. (FMNH 243935). C. Dorsum of paratype of A. borneensis sp. nov. (FMNH 243936). D. Venter of paratype of A. borneensis sp. nov. (FMNH 243936). E. Dorsum of paratype of A. borneensis sp. nov. (FMNH 249751). F. Venter of paratype of A. borneensis sp. nov. (FMNH 249751)." figureDoi="http://doi.org/10.5281/zenodo.3628950" httpUri="https://zenodo.org/record/3628950/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
|
||
&
|
||
<figureCitation id="AFACFC91FFF1FFBFBBDAFDC9FEA0FDCA" box="[252,268,546,572]" captionStart="FIGURE 3" captionStartId="14.[151,250,1908,1933]" captionTargetBox="[170,1416,453,1883]" captionTargetId="figure@14.[170,1416,453,1883]" captionTargetPageId="14" captionText="FIGURE 3. Asthenodipsas borneensis sp. nov. from Borneo. A: Specimen from Kuching, Sarawak (LSUDPC 10988). B: Specimen from Sayap, Mount Kinabalu, Sabah (LSUDPC 10982). C – F (LSUDPC 10983–10986): Adults from Poring, Sabah. G & H (LSUDPC 10987 & 10978): Juvenile from Poring, Sabah. (Photographs by Chien C. Lee [A & B], Zhou Hang [C & D], Anton Sorokin [E & F] & Jaroslav Karhánek [G & H])." figureDoi="http://doi.org/10.5281/zenodo.3628952" httpUri="https://zenodo.org/record/3628952/files/figure.png" pageId="12" pageNumber="13">3</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<paragraph id="3728E014FFF1FFBFBBB1FD8DFA30FD2D" blockId="12.[151,1437,614,884]" pageId="12" pageNumber="13">
|
||
<subSubSection id="7F8DB39FFFF1FFBFBBB1FD8DFA30FD6B" pageId="12" pageNumber="13" type="synonymic_list">
|
||
<taxonomicName id="F0979B97FFF1FFBFBBB1FD8DFE00FD88" authorityName="Peters" authorityYear="1864" box="[151,428,614,638]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="malaccanus">
|
||
<emphasis id="05E33C06FFF1FFBFBBB1FD8DFE00FD88" box="[151,428,614,638]" italics="true" pageId="12" pageNumber="13">Asthenodipsas malaccanus</emphasis>
|
||
</taxonomicName>
|
||
(in part): Chan-ard
|
||
<emphasis id="05E33C06FFF1FFBFB9A5FD8CFD1EFD88" box="[643,690,614,638]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
|
||
. 2015: 159;
|
||
<bibRefCitation id="53069DE5FFF1FFBFB81DFD8DFB8FFD89" author="Charlton, T." box="[827,1059,614,639]" pageId="12" pageNumber="13" refId="ref12952" refString="Charlton, T. (2020) A Guide to Snakes of Peninsular Malaysia and Singapore. Natural History Publications (Borneo), Kota Kinabalu, 299 pp." type="book" year="2020">Charlton 2020: 86–87</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="53069DE5FFF1FFBFBF16FD8CFB0FFD89" author="Das, I." box="[1072,1187,614,639]" pageId="12" pageNumber="13" pagination="3 - 11" refId="ref12982" refString="Das, I. (2006 a) Crocker Range National Park, Sabah, as a refuge for Borneo's montane herpetofauna. Amphibian and Reptile Conservation, 4 (1), 3 - 11." type="journal article" year="2006">Das 2006a</bibRefCitation>
|
||
: 007,
|
||
<bibRefCitation id="53069DE5FFF1FFBFBFC3FD8CFAD3FD89" author="Das, I." box="[1253,1407,614,639]" pageId="12" pageNumber="13" refId="ref13079" refString="Das, I. (2010) A Field Guide to the Reptiles of South-East Asia. New Holland Publishers, London, 376 pp." type="book" year="2010">Das 2010: 161</bibRefCitation>
|
||
& 344–345,
|
||
<bibRefCitation id="53069DE5FFF1FFBFBA0BFD6DFE23FD6B" author="Das, I." box="[301,399,645,669]" pageId="12" pageNumber="13" refId="ref13105" refString="Das, I. (2012) A Naturalist's Guide to the Snakes of South-east Asia including Malaysia, Singapore, Thailand, Myanmar, Borneo, Sumatra, Java and Bali. John Beaufoy Publishing, Oxford, 160 pp." type="book" year="2012">Das 2012</bibRefCitation>
|
||
&
|
||
<bibRefCitation id="53069DE5FFF1FFBFBA97FD6EFDB7FD6B" author="Das, I." box="[433,539,645,669]" pageId="12" pageNumber="13" refId="ref13147" refString="Das, I. (2018) A Naturalist's Guide to the Snakes of Thailand and South-east Asia. 2 nd edition. John Beaufoy Publishing, Oxford, 176 pp." type="book" year="2018">2018: 131</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="53069DE5FFF1FFBFB900FD6FFCE2FD6B" author="Iskandar, D. T. & Colijn, E." box="[550,846,644,669]" pageId="12" pageNumber="13" refId="ref14776" refString="Iskandar, D. T. & Colijn, E. (" 2001 " 2002) A Checklist of Southeast Asian and New Guinean Reptiles. Part I. Serpentes, Biodiversity Conservation Project, Jakarta, 195 pp." type="book" year="2001">Iskandar & Colijn 2001: 111</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="53069DE5FFF1FFBFB87CFD6FFB83FD6B" author="Marlon, E." box="[858,1071,644,669]" pageId="12" pageNumber="13" refId="ref15175" refString="Marlon, E. (2014) Panduan Visual dan Identifikasi Lapangan- 107 + Ular Indonesia. Indonesia Nature and Wildlife Publishing, Jakarta, 251 pp." type="book" year="2014">Marlon 2014: 27–28</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="53069DE5FFF1FFBFBF1CFD6FFA9FFD6B" author="Stuebing, R. B. & Inger, R. F. & Lardner, B." box="[1082,1331,644,669]" pageId="12" pageNumber="13" refId="ref15734" refString="Stuebing, R. B., Inger, R. F. & Lardner, B. (2014) A field guide to the snakes of Borneo. 2 nd Edition. Natural History Publications (Borneo), Kota Kinabalu, 310 pp." type="book" year="2014">
|
||
Stuebing
|
||
<emphasis id="05E33C06FFF1FFBFBFBDFD6DFB65FD6B" box="[1179,1225,645,669]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
|
||
. 2014: 87
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="53069DE5FFF1FFBFBE18FD6FFA30FD6B" author="Teynie, A. & David, P. & Ohler, A." box="[1342,1436,644,669]" pageId="12" pageNumber="13" pagination="1 - 43" refId="ref15813" refString="Teynie, A., David, P. & Ohler, A. (2010) Note on a collection of amphibians and reptiles from Western Sumatra (Indonesia), with the description of a new species of the genus Bufo. Zootaxa, 2416 (1), 1 - 43. https: // doi. org / 10.11646 / zootaxa. 2416.1.1" type="journal article" year="2010">
|
||
Teynié
|
||
<emphasis id="05E33C06FFF1FFBFBEACFD6DFA30FD6B" box="[1418,1436,646,669]" italics="true" pageId="12" pageNumber="13">et</emphasis>
|
||
</bibRefCitation>
|
||
</subSubSection>
|
||
<subSubSection id="7F8DB39FFFF1FFBFBBE1FD4FFA30FD2D" pageId="12" pageNumber="13" type="reference_group">
|
||
<emphasis id="05E33C06FFF1FFBFBBE1FD4FFF77FD4A" box="[199,219,676,700]" italics="true" pageId="12" pageNumber="13">al</emphasis>
|
||
. 2010: 39;
|
||
<bibRefCitation id="53069DE5FFF1FFBFBA68FD48FD90FD4A" author="Wallach, V. & Williams, K. L. & Boundy, J." box="[334,572,675,700]" pageId="12" pageNumber="13" refId="ref16458" refString="Wallach, V., Williams, K. L. & Boundy, J. (2014) Snakes of the World: A Catalogue of Living and Extinct Species. Taylor and Francis, CRC Press, Boca Raton, 1209 pp. [type catalogue] https: // doi. org / 10.1201 / b 16901" type="book" year="2014">
|
||
Wallach
|
||
<emphasis id="05E33C06FFF1FFBFBA81FD4EFE7FFD4A" box="[423,467,676,700]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
|
||
. 2014: 59
|
||
</bibRefCitation>
|
||
.
|
||
<taxonomicName id="F0979B97FFF1FFBFBBB1FD29FE1FFD2C" authorityName="Boulenger" authorityYear="1892" box="[151,435,706,730]" class="Squamata" family="Pareidae" genus="Amblycephalus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="malaccanus">
|
||
<emphasis id="05E33C06FFF1FFBFBBB1FD29FE1FFD2C" box="[151,435,706,730]" italics="true" pageId="12" pageNumber="13">Amblycephalus malaccanus</emphasis>
|
||
</taxonomicName>
|
||
(in part):
|
||
<bibRefCitation id="53069DE5FFF1FFBFB93BFD29FD5BFD2D" author="Boulenger, G. A." box="[541,759,706,731]" pageId="12" pageNumber="13" pagination="505 - 508" refId="ref12469" refString="Boulenger, G. A. (1892) An account of the reptiles and batrachians collected by Mr. C. Hose on Mt. Dulit, Borneo. Proceedings of the Zoological Society of London, 1892, 505 - 508." type="journal article" year="1892">Boulenger 1892: 507</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="53069DE5FFF1FFBFB825FD29FC0CFD2D" author="Boulenger, G. A." box="[771,928,706,731]" pageId="12" pageNumber="13" refId="ref12513" refString="Boulenger, G. A. (1896) Catalogue of the Snakes in the British Museum (Natural History). Vol. III. Trustees of the British Museum, London, xiv + 727 pp., 15 pls. https: // doi. org / 10.5962 / bhl. title. 8316" type="book" year="1896">1896: 442–443</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="53069DE5FFF1FFBFB88AFD28FBE5FD2D" author="Boulenger, G. A." box="[940,1097,707,731]" pageId="12" pageNumber="13" refId="ref12570" refString="Boulenger, G. A. (1912) A vertebrate fauna of the Malay Peninsula from the Isthmus of Kra to Singapore including the adjacent islands. Reptilia and Batrachia. Taylor & Francis, London, 294 pp. https: // doi. org / 10.5962 / bhl. title. 10813" type="book" year="1912">1912: 209–210</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="53069DE5FFF1FFBFBF72FD29FAA1FD2D" author="Flower, S. S." box="[1108,1293,706,731]" pageId="12" pageNumber="13" pagination="856 - 914" refId="ref13489" refString="Flower, S. S. (1896) Notes on a second collection of batrachians made in the Malay Peninsula 1895 - 96, with a list of the species recorded from that region. Proceedings of the Zoological Society of London, 1896, 856 - 914. https: // doi. org / 10.1111 / j. 1096 - 3642.1896. tb 03085. x" type="book chapter" year="1896">Flower 1896: 896</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="53069DE5FFF1FFBFBE3FFD28FA2DFD2D" author="Flower, S. S." box="[1305,1409,706,731]" pageId="12" pageNumber="13" pagination="600 - 696" refId="ref13558" refString="Flower, S. S. (1899) Notes on a second collection of reptiles made in the Malay Peninsula and Siam, from November 1896 - September 1898, with a list of the species recorded from those countries. Proceedings of the Zoological Society of London, 1899, 600 - 696. https: // doi. org / 10.1111 / j. 1469 - 7998.1899. tb 06880. x" type="book chapter" year="1899">1899: 607</bibRefCitation>
|
||
&
|
||
</subSubSection>
|
||
</paragraph>
|
||
<subSubSection id="7F8DB39FFFF1FFBFBBE1FD0BFA31FCC1" pageId="12" pageNumber="13" type="discussion">
|
||
<paragraph id="3728E014FFF1FFBFBBE1FD0BFA31FCC1" blockId="12.[151,1437,614,884]" pageId="12" pageNumber="13">
|
||
694;
|
||
<bibRefCitation id="53069DE5FFF1FFBFBBDDFD0BFE5FFD0F" author="de Rooij, N." box="[251,499,736,761]" pageId="12" pageNumber="13" refId="ref13443" refString="de Rooij, N. (1917) The Reptiles of the Indo-Australian Archipelago. II. Ophidia. E. J. Brill Ltd., Leiden, 334 pp. https: // doi. org / 10.5962 / bhl. title. 10610" type="book" year="1917">de Rooij 1917: 267–277</bibRefCitation>
|
||
;
|
||
<taxonomicName id="F0979B97FFF1FFBFBBB1FCEAFED3FCEE" authorityName="Rao & Yang" authorityYear="1992" box="[151,383,768,792]" class="Squamata" family="Pareidae" genus="Internatus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="malaccanus">
|
||
<emphasis id="05E33C06FFF1FFBFBBB1FCEAFED3FCEE" box="[151,383,768,792]" italics="true" pageId="12" pageNumber="13">Internatus malaccanus</emphasis>
|
||
</taxonomicName>
|
||
(in part):
|
||
<bibRefCitation id="53069DE5FFF1FFBFBAC3FD14FC8CFCEE" author="David, P. & Vogel, G." box="[485,800,767,792]" pageId="12" pageNumber="13" refId="ref13298" refString="David, P. & Vogel, G. (1996) The Snakes of Sumatra. An Annotated Checklist and Key with Natural History Notes. Edition Chimaira, Frankfurt am Main, 260 pp." type="book" year="1996">David & Vogel 1996: 137–138</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="53069DE5FFF1FFBFB80CFD14FB9EFCEE" author="Malkmus, R. & Manthey, U. & Vogel, G. & Hoffmann, P. & Kosuch, J." box="[810,1074,767,792]" pageId="12" pageNumber="13" refId="ref15123" refString="Malkmus, R., Manthey, U., Vogel, G., Hoffmann, P. & Kosuch, J. (2002) Amphibians and Reptiles of Mount Kinabalu (North Borneo). A. R. G. Ganter Verlag, Ruggel, 424 pp." type="book" year="2002">
|
||
Malkmus
|
||
<emphasis id="05E33C06FFF1FFBFB8B6FCEAFC10FCEE" box="[912,956,768,792]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
|
||
. 2002: 343
|
||
</bibRefCitation>
|
||
.
|
||
<taxonomicName id="F0979B97FFF1FFBFBBB1FCF4FEF2FCC0" authorityName="Robinson & Kloss" authorityYear="1920" box="[151,350,798,822]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="malaccanus">
|
||
<emphasis id="05E33C06FFF1FFBFBBB1FCF4FEF2FCC0" box="[151,350,798,822]" italics="true" pageId="12" pageNumber="13">Pareas malaccanus</emphasis>
|
||
</taxonomicName>
|
||
(in part): De Haas 1950: 529;
|
||
<bibRefCitation id="53069DE5FFF1FFBFB9BEFCF5FC93FCC1" author="Haile, N. S." box="[664,831,798,823]" pageId="12" pageNumber="13" pagination="743 - 771" refId="ref14666" refString="Haile, N. S. (1958) The snakes of Borneo, with a key to the species. Sarawak Museum Journal, 8 (12), 743 - 771." type="journal article" year="1958">Haile 1958: 766</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="53069DE5FFF1FFBFB86CFCF5FBBBFCC1" author="Grandison, A. G. C." box="[842,1047,798,823]" pageId="12" pageNumber="13" pagination="45 - 101" refId="ref13673" refString="Grandison, A. G. C. (1972) The Gunong Benom Expedition 1967. 5. Reptiles and amphibians of Gunong Benom with a description of a new species of Macrocalamus. Bulletin of the British Museum of Natural History, Zoology, 23, 45 - 101." type="journal article" year="1972">Grandison 1972: 92</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="53069DE5FFF1FFBFBF04FCF5FB5BFCC1" author="Marlon, E." box="[1058,1271,798,823]" pageId="12" pageNumber="13" refId="ref15175" refString="Marlon, E. (2014) Panduan Visual dan Identifikasi Lapangan- 107 + Ular Indonesia. Indonesia Nature and Wildlife Publishing, Jakarta, 251 pp." type="book" year="2014">Marlon 2014: 27–28</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="53069DE5FFF1FFBFBE24FCF5FA31FCC1" author="Stuebing, R. B." box="[1282,1437,798,823]" pageId="12" pageNumber="13" pagination="323 - 362" refId="ref15665" refString="Stuebing, R. B. (1991) A checklist of the snakes of Borneo. Raffles Bulletin of Zoology, 39 (2), 323 - 362." type="journal article" year="1991">Stuebing 1991:</bibRefCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7F8DB39FFFF1FFBFBBE1FCD6FDF3FC82" pageId="12" pageNumber="13" type="reference_group">
|
||
<paragraph id="3728E014FFF1FFBFBBE1FCD6FDF3FC82" blockId="12.[151,1437,614,884]" pageId="12" pageNumber="13">
|
||
331;
|
||
<bibRefCitation id="53069DE5FFF1FFBFBBDDFCD7FDA1FCA3" author="Stuebing, R. B. & Inger, R. F." box="[251,525,828,854]" pageId="12" pageNumber="13" refId="ref15696" refString="Stuebing, R. B. & Inger, R. F. (1999) A Field Guide to the Snakes of Borneo. Natural History Publications (Borneo), Kota Kinabalu, 254 pp." type="book" year="1999">Stuebing & Inger 1999: 88</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="53069DE5FFF1FFBFB931FCD7FD64FCA3" author="Taylor, E. H." box="[535,712,828,853]" pageId="12" pageNumber="13" pagination="609 - 1096" refId="ref15781" refString="Taylor, E. H. (1965) The serpents of Thailand and adjacent waters. University of Kansas Science Bulletin, 45 (9), 609 - 1096." type="journal article" year="1965">Taylor 1965: 686</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="53069DE5FFF1FFBFB9F4FCD7FCD9FCA3" author="Welch, K. R. G." box="[722,885,828,853]" pageId="12" pageNumber="13" refId="ref16518" refString="Welch, K. R. G. (1988) Snakes of the Orient: A Checklist. R. E. Krieger Publ. Co., Malabar Florida, 183 pp." type="book" year="1988">Welch 1988: 90</bibRefCitation>
|
||
.
|
||
<taxonomicName id="F0979B97FFF1FFBFBBB1FCB6FDF7FC82" authority=": Das 2006 b: 45" authorityName="Das" authorityPageNumber="45" authorityYear="2006" box="[151,603,859,884]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="carinatus">
|
||
<emphasis id="05E33C06FFF1FFBFBBB1FCB6FEE8FC82" box="[151,324,861,884]" italics="true" pageId="12" pageNumber="13">Pareas carinatus</emphasis>
|
||
:
|
||
<bibRefCitation id="53069DE5FFF1FFBFBA77FCB7FE44FC82" author="Das, I." box="[337,488,859,884]" pageId="12" pageNumber="13" refId="ref13019" refString="Das, I. (2006 b) A Photographic Guide to Snakes and Other Reptiles of Borneo. New Holland Publishers Ltd., London, 144 pp." type="book" year="2006">Das 2006b: 45</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="53069DE5FFF1FFBFBAD4FCB7FDF7FC82" author="Das, I." box="[498,603,859,884]" pageId="12" pageNumber="13" refId="ref13049" refString="Das, I. (2007) Amphibians and Reptiles of Brunei: A Pocket Guide. Natural History Publications (Borneo), Kota Kinabalu, 200 pp." type="book" year="2007">2007: 161</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7F8DB39FFFF1FFBFBBB1FC4BFE6FFAAC" pageId="12" pageNumber="13" type="materials_examined">
|
||
<paragraph id="3728E014FFF1FFBFBBB1FC4BFAA2FC2B" blockId="12.[151,1437,927,2030]" pageId="12" pageNumber="13">
|
||
<materialsCitation id="87FFEA49FFF1FFBFBBB1FC4BFAA6FC2B" ID-GBIF-Occurrence="2571579742" collectingDate="1990-08-25" collectionCode="FMNH" country="Malaysia" elevation="748" latitude="4.9" location="Sabah" longLatPrecision="1306" longitude="115.75" pageId="12" pageNumber="13" specimenCode="FMNH 243935" specimenCount="2" specimenCount-adult="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="holotype">
|
||
<emphasis id="05E33C06FFF1FFBFBBB1FC4BFEA7FC4C" bold="true" box="[151,267,928,954]" pageId="12" pageNumber="13">
|
||
<typeStatus id="E82C5EB6FFF1FFBFBBB1FC4BFEAAFC4C" box="[151,262,928,954]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCount id="21912B9DFFF1FFBFBA33FC74FEF9FC4F" box="[277,341,927,953]" pageId="12" pageNumber="13" type="adult">Adult</specimenCount>
|
||
<specimenCount id="21912B9DFFF1FFBFBA78FC74FE06FC4F" box="[350,426,927,953]" pageId="12" pageNumber="13" type="female">female</specimenCount>
|
||
<specimenCode id="6731486FFFF1FFBFBA92FC4BFDCEFC4C" box="[436,610,927,954]" pageId="12" pageNumber="13">FMNH 243935</specimenCode>
|
||
collected by Robert F. Inger and Tan Fu Lian on
|
||
<date id="4329C6D4FFF1FFBFBF80FC74FAF0FC4C" box="[1190,1372,927,954]" pageId="12" pageNumber="13" value="1990-08-25">
|
||
<collectingDate id="536D3F3CFFF1FFBFBF80FC74FAF0FC4C" box="[1190,1372,927,954]" pageId="12" pageNumber="13" value="1990-08-25">25 August 1990</collectingDate>
|
||
</date>
|
||
from Mendolong camp watershed, Sipitang District,
|
||
<collectingRegion id="F5532EF6FFF1FFBFB98FFC29FD5CFC2A" box="[681,752,962,988]" country="Malaysia" name="Sabah" pageId="12" pageNumber="13">Sabah</collectingRegion>
|
||
, East
|
||
<collectingCountry id="4F80A084FFF1FFBFB814FC29FC35FC2A" box="[818,921,962,988]" name="Malaysia" pageId="12" pageNumber="13">Malaysia</collectingCountry>
|
||
(
|
||
<geoCoordinate id="52A386D3FFF1FFBFB88FFC2AFC50FC2A" box="[937,1020,961,988]" degrees="4" direction="north" minutes="54" orientation="latitude" pageId="12" pageNumber="13" precision="925" value="4.9">4°54’N</geoCoordinate>
|
||
<geoCoordinate id="52A386D3FFF1FFBFBF25FC2AFBDCFC2A" box="[1027,1136,961,988]" degrees="115" direction="east" minutes="45" orientation="longitude" pageId="12" pageNumber="13" precision="925" value="115.75">115°45’E</geoCoordinate>
|
||
,
|
||
<quantity id="F06F4DF1FFF1FFBFBF5AFC29FB6FFC2B" box="[1148,1219,962,989]" metricMagnitude="2" metricUnit="m" metricValue="7.4799999999999995" pageId="12" pageNumber="13" unit="m" value="748.0">
|
||
<elevation id="BCBA0727FFF1FFBFBF5AFC29FB6FFC2B" box="[1148,1219,962,989]" metricMagnitude="2" metricUnit="m" metricValue="7.4799999999999995" pageId="12" pageNumber="13" unit="m" value="748.0">748 m</elevation>
|
||
</quantity>
|
||
a.s.l.)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="3728E014FFF1FFBFBBE1FC0EFA2AFB7C" blockId="12.[151,1437,927,2030]" pageId="12" pageNumber="13">
|
||
<emphasis id="05E33C06FFF1FFBFBBE1FC0EFEEAFC09" bold="true" box="[199,326,997,1023]" pageId="12" pageNumber="13">
|
||
<typeStatus id="E82C5EB6FFF1FFBFBBE1FC0EFEEDFC09" box="[199,321,997,1023]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult male FMNH 243936 collected by Robert F. Inger and Tan Fu Lian on
|
||
<date id="4329C6D4FFF1FFBFBFB0FC0EFACCFC09" box="[1174,1376,997,1023]" pageId="12" pageNumber="13" value="1990-09-09">9 September 1990</date>
|
||
from Purulon camp, Tenom District,
|
||
<collectingRegion id="F5532EF6FFF1FFBFBADCFBECFDEDFBD7" box="[506,577,1031,1057]" country="Malaysia" name="Sabah" pageId="12" pageNumber="13">Sabah</collectingRegion>
|
||
, East
|
||
<collectingCountry id="4F80A084FFF1FFBFB9A3FBECFD40FBD7" box="[645,748,1031,1057]" name="Malaysia" pageId="12" pageNumber="13">Malaysia</collectingCountry>
|
||
(
|
||
<geoCoordinate id="52A386D3FFF1FFBFB9DAFBECFCFCFBD4" box="[764,848,1031,1058]" degrees="5" direction="north" minutes="13" orientation="latitude" pageId="12" pageNumber="13" precision="925" value="5.2166667">5°13’N</geoCoordinate>
|
||
<geoCoordinate id="52A386D3FFF1FFBFB871FBECFC68FBD4" box="[855,964,1031,1058]" degrees="115" direction="east" minutes="57" orientation="longitude" pageId="12" pageNumber="13" precision="925" value="115.95">115°57’E</geoCoordinate>
|
||
,
|
||
<quantity id="F06F4DF1FFF1FFBFB8F6FBECFBB4FBD4" box="[976,1048,1031,1058]" metricMagnitude="2" metricUnit="m" metricValue="5.16" pageId="12" pageNumber="13" unit="m" value="516.0">516 m</quantity>
|
||
a.s.l.). Adult male FMNH 249751 collected by Robert F. Inger on
|
||
<date id="4329C6D4FFF1FFBFBAD2FBC1FD15FBB2" box="[500,697,1066,1092]" pageId="12" pageNumber="13" value="1991-12-06">6 December 1991</date>
|
||
from Poring station, Mount Kinabalu Park, Ranau District,
|
||
<collectingRegion id="F5532EF6FFF1FFBFBE77FBC1FA34FBB2" box="[1361,1432,1066,1092]" country="Malaysia" name="Sabah" pageId="12" pageNumber="13">Sabah</collectingRegion>
|
||
, East
|
||
<collectingCountry id="4F80A084FFF1FFBFBBF6FBA6FE9BFB91" box="[208,311,1101,1127]" name="Malaysia" pageId="12" pageNumber="13">Malaysia</collectingCountry>
|
||
(
|
||
<geoCoordinate id="52A386D3FFF1FFBFBA6EFBA6FE30FB91" box="[328,412,1101,1127]" degrees="6" direction="north" minutes="03" orientation="latitude" pageId="12" pageNumber="13" precision="925" value="6.05">6°03’N</geoCoordinate>
|
||
<geoCoordinate id="52A386D3FFF1FFBFBA82FBA6FDBDFB91" box="[420,529,1101,1127]" degrees="116" direction="east" minutes="42" orientation="longitude" pageId="12" pageNumber="13" precision="925" value="116.7">116°42’E</geoCoordinate>
|
||
,
|
||
<quantity id="F06F4DF1FFF1FFBFB938FBA6FDCBFB91" box="[542,615,1101,1127]" metricMagnitude="2" metricUnit="m" metricValue="6.06" pageId="12" pageNumber="13" unit="m" value="606.0">606 m</quantity>
|
||
a.s.l.). Adult female FMNH 71590 collected by T. H. Harrisson in
|
||
<date id="4329C6D4FFF1FFBFBE4FFBA6FF63FB7C" pageId="12" pageNumber="13" value="1952-05">May 1952</date>
|
||
from Pa Brayong, Lawas District,
|
||
<collectingCountry id="4F80A084FFF1FFBFB971FB84FD16FB7F" box="[599,698,1135,1161]" name="Malaysia" pageId="12" pageNumber="13">Sarawak</collectingCountry>
|
||
, East
|
||
<collectingCountry id="4F80A084FFF1FFBFB9DBFB84FCC8FB7F" box="[765,868,1135,1161]" name="Malaysia" pageId="12" pageNumber="13">Malaysia</collectingCountry>
|
||
(estimated:
|
||
<geoCoordinate id="52A386D3FFF1FFBFB8C8FB84FB92FB7C" box="[1006,1086,1135,1162]" degrees="4.45" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="555" value="4.45">4.45°N</geoCoordinate>
|
||
<geoCoordinate id="52A386D3FFF1FFBFBF63FB84FB77FB7C" box="[1093,1243,1135,1162]" degrees="115.51667" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="1" value="115.51667">115.51667°E</geoCoordinate>
|
||
,
|
||
<quantity id="F06F4DF1FFF1FFBFBFC3FB9BFA97FB7C" box="[1253,1339,1136,1162]" metricMagnitude="3" metricUnit="m" metricValue="1.077" pageId="12" pageNumber="13" unit="m" value="1077.0">1,077m</quantity>
|
||
a.s.l.).
|
||
</paragraph>
|
||
<paragraph id="3728E014FFF1FFBFBBE1FB7AFE6FFAAC" blockId="12.[151,1437,927,2030]" pageId="12" pageNumber="13">
|
||
<emphasis id="05E33C06FFF1FFBFBBE1FB7AFE1FFB5A" bold="true" box="[199,435,1169,1196]" pageId="12" pageNumber="13">Referred specimens</emphasis>
|
||
. Juvenile female MCZ R-178183 collected by L.H. Emmons on
|
||
<date id="4329C6D4FFF1FFBFBFB7FB78FAF2FB5A" box="[1169,1374,1170,1197]" pageId="12" pageNumber="13" value="1989-09-07">7 September 1989</date>
|
||
from Poring, Mount Kinabalu Park, Ranau District,
|
||
<collectingRegion id="F5532EF6FFF1FFBFB98CFB5EFD5DFB39" box="[682,753,1205,1231]" country="Malaysia" name="Sabah" pageId="12" pageNumber="13">Sabah</collectingRegion>
|
||
, East
|
||
<collectingCountry id="4F80A084FFF1FFBFB811FB5EFC32FB39" box="[823,926,1205,1231]" name="Malaysia" pageId="12" pageNumber="13">Malaysia</collectingCountry>
|
||
(
|
||
<geoCoordinate id="52A386D3FFF1FFBFB889FB5EFBEEFB39" box="[943,1090,1205,1231]" degrees="6" direction="north" minutes="03" orientation="latitude" pageId="12" pageNumber="13" precision="1" seconds="06.4" value="6.051778">6°03’06.4”N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="52A386D3FFF1FFBFBF69FB5FFB56FB39" box="[1103,1274,1204,1231]" degrees="116" direction="east" minutes="42" orientation="longitude" pageId="12" pageNumber="13" precision="1" seconds="03.5" value="116.700966">116°42’03.5”E</geoCoordinate>
|
||
,
|
||
<quantity id="F06F4DF1FFF1FFBFBE21FB5FFAE3FB39" box="[1287,1359,1204,1231]" metricMagnitude="2" metricUnit="m" metricValue="5.75" pageId="12" pageNumber="13" unit="m" value="575.0">575 m</quantity>
|
||
a.s.l.). Juvenile female FMNH248950/SP 04130 collected by Tan Fu Lian on
|
||
<date id="4329C6D4FFF1FFBFB883FB33FBD1FB04" box="[933,1149,1239,1266]" pageId="12" pageNumber="13" value="1991-09-14">14 September 1991</date>
|
||
from Tawau Hills, Tawau District,
|
||
<collectingRegion id="F5532EF6FFF1FFBFBBDEFB11FE93FAE2" box="[248,319,1274,1300]" country="Malaysia" name="Sabah" pageId="12" pageNumber="13">Sabah</collectingRegion>
|
||
, East
|
||
<collectingCountry id="4F80A084FFF1FFBFBAA7FB11FE44FAE2" box="[385,488,1274,1300]" name="Malaysia" pageId="12" pageNumber="13">Malaysia</collectingCountry>
|
||
(
|
||
<geoCoordinate id="52A386D3FFF1FFBFBAD0FB11FDE5FAE2" box="[502,585,1274,1300]" degrees="4" direction="north" minutes="37" orientation="latitude" pageId="12" pageNumber="13" precision="925" value="4.616667">4°37’N</geoCoordinate>
|
||
<geoCoordinate id="52A386D3FFF1FFBFB976FB12FD10FAE2" box="[592,700,1273,1300]" degrees="117" direction="east" minutes="54" orientation="longitude" pageId="12" pageNumber="13" precision="925" value="117.9">117°54’E</geoCoordinate>
|
||
,
|
||
<quantity id="F06F4DF1FFF1FFBFB9EEFB11FCA2FAE3" box="[712,782,1274,1301]" metricMagnitude="2" metricUnit="m" metricValue="2.66" pageId="12" pageNumber="13" unit="m" value="266.0">266 m</quantity>
|
||
a.s.l.). Adult female ZRC 2.7348 collected by an unknown collector in the 1990s from Wind & Fairy Caves, Bau district,
|
||
<collectingCountry id="4F80A084FFF1FFBFB840FAF6FC65FAC1" box="[870,969,1309,1335]" name="Malaysia" pageId="12" pageNumber="13">Sarawak</collectingCountry>
|
||
, East
|
||
<collectingCountry id="4F80A084FFF1FFBFBF29FAF6FBDAFAC1" box="[1039,1142,1309,1335]" name="Malaysia" pageId="12" pageNumber="13">Malaysia</collectingCountry>
|
||
(estimated:
|
||
<geoCoordinate id="52A386D3FFF1FFBFBE22FAF6FA34FAC1" box="[1284,1432,1309,1335]" degrees="01" direction="north" minutes="22.867" orientation="latitude" pageId="12" pageNumber="13" precision="1" value="1.3811166">01°22.867 N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="52A386D3FFF1FFBFBBB1FAD4FE9EFAAC" box="[151,306,1343,1370]" degrees="110" direction="east" minutes="07.164" orientation="longitude" pageId="12" pageNumber="13" precision="1" value="110.1194">110°07.164 E</geoCoordinate>
|
||
,
|
||
<quantity id="F06F4DF1FFF1FFBFBA18FAABFEDBFAAC" box="[318,375,1344,1370]" metricMagnitude="1" metricUnit="m" metricValue="7.3" pageId="12" pageNumber="13" unit="m" value="73.0">73 m</quantity>
|
||
a.s.l.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7F8DB39FFFF1FFBFBBE1FA8AFB56F851" pageId="12" pageNumber="13" type="diagnosis">
|
||
<paragraph id="3728E014FFF1FFBFBBE1FA8AFB56F851" blockId="12.[151,1437,927,2030]" pageId="12" pageNumber="13">
|
||
<emphasis id="05E33C06FFF1FFBFBBE1FA8AFEEEFA8A" bold="true" box="[199,322,1377,1404]" pageId="12" pageNumber="13">Diagnosis.</emphasis>
|
||
<taxonomicName id="F0979B97FFF1FFBFBA6FFA89FDC7FA8A" authority="Quah & Grismer & Lim & Anuar & Chan, 2020" authorityName="Quah & Grismer & Lim & Anuar & Chan" authorityYear="2020" box="[329,619,1378,1404]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="05E33C06FFF1FFBFBA6FFA89FDC7FA8A" box="[329,619,1378,1404]" italics="true" pageId="12" pageNumber="13">Asthenodipsas borneensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="05E33C06FFF1FFBFB955FA89FD62FA8A" bold="true" box="[627,718,1378,1404]" pageId="12" pageNumber="13">
|
||
<taxonomicNameLabel id="1ED0817DFFF1FFBFB955FA89FD62FA8A" box="[627,718,1378,1404]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is diagnosed as member of the genus
|
||
<taxonomicName id="F0979B97FFF1FFBFBF5BFA89FAB3FA8A" authorityName="Peters" authorityYear="1864" box="[1149,1311,1378,1404]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
||
<emphasis id="05E33C06FFF1FFBFBF5BFA89FAB3FA8A" box="[1149,1311,1378,1404]" italics="true" pageId="12" pageNumber="13">Asthenodipsas</emphasis>
|
||
</taxonomicName>
|
||
by its possession of smooth dorsals, absence of preoculars and suboculars, having supralabials in contact with the eyes, single anterior inframaxillary and two pairs of posterior inframaxillaries (
|
||
<bibRefCitation id="53069DE5FFF1FFBFB8A4FA43FB15FA34" author="Grossmann, W. & Tillack, F." box="[898,1209,1447,1474]" pageId="12" pageNumber="13" pagination="175 - 90" refId="ref14523" refString="Grossmann, W. & Tillack, F. (2003) On the Taxonomic Status of Asthenodipsas tropidonotus (Van Lidth De Jeude, 1923) and Pareas vertebralis (Boulenger, 1900) (Serpentes: Colubridae: Pareatinae). Russian Journal of Herpetology, 10 (3), 175 - 90." type="journal article" year="2003">Grossmann & Tillack 2003</bibRefCitation>
|
||
). It can be separated from
|
||
<taxonomicName id="F0979B97FFF1FFBFBBF2FA21FE75FA13" authority="(Boie)" baseAuthorityName="Boie" baseAuthorityYear="1828" box="[212,473,1482,1509]" class="Reptilia" family="Pareatidae" genus="Aplopeltura" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="boa">
|
||
<emphasis id="05E33C06FFF1FFBFBBF2FA21FE27FA12" box="[212,395,1482,1508]" italics="true" pageId="12" pageNumber="13">Aplopeltura boa</emphasis>
|
||
(Boie)
|
||
</taxonomicName>
|
||
by its higher number of mid-dorsal scale rows (15 vs. 13) and subcaudals (divided vs. undivided) (
|
||
<bibRefCitation id="53069DE5FFF1FFBFBA39FA06FE6CF9F1" author="de Rooij, N." box="[287,448,1517,1543]" pageId="12" pageNumber="13" refId="ref13443" refString="de Rooij, N. (1917) The Reptiles of the Indo-Australian Archipelago. II. Ophidia. E. J. Brill Ltd., Leiden, 334 pp. https: // doi. org / 10.5962 / bhl. title. 10610" type="book" year="1917">de Rooij 1917</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="53069DE5FFF1FFBFBAEDFA06FD57F9F1" author="Grossmann, W. & Tillack, F." box="[459,763,1517,1543]" pageId="12" pageNumber="13" pagination="175 - 90" refId="ref14523" refString="Grossmann, W. & Tillack, F. (2003) On the Taxonomic Status of Asthenodipsas tropidonotus (Van Lidth De Jeude, 1923) and Pareas vertebralis (Boulenger, 1900) (Serpentes: Colubridae: Pareatinae). Russian Journal of Herpetology, 10 (3), 175 - 90." type="journal article" year="2003">Grossmann & Tillack 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="53069DE5FFF1FFBFB820FA06FC45F9F1" author="Stuebing, R. B. & Inger, R. F. & Lardner, B." box="[774,1001,1517,1543]" pageId="12" pageNumber="13" refId="ref15734" refString="Stuebing, R. B., Inger, R. F. & Lardner, B. (2014) A field guide to the snakes of Borneo. 2 nd Edition. Natural History Publications (Borneo), Kota Kinabalu, 310 pp." type="book" year="2014">
|
||
Stuebing
|
||
<emphasis id="05E33C06FFF1FFBFB856FA05FC0CF9F1" box="[880,928,1517,1543]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
), and can be further differentiated from members of the genus
|
||
<taxonomicName id="F0979B97FFF1FFBFBAB0F9FBFE48F9DF" box="[406,484,1552,1577]" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
|
||
<emphasis id="05E33C06FFF1FFBFBAB0F9FBFE48F9DF" box="[406,484,1552,1577]" italics="true" pageId="12" pageNumber="13">Pareas</emphasis>
|
||
</taxonomicName>
|
||
by its possession of preocular and subocular scales (absent vs. present), supralabials in contact with orbit (3
|
||
<superScript id="C0E24D5CFFF1FFBFBABEF9DAFE0AF9C9" attach="left" box="[408,422,1585,1599]" fontSize="6" pageId="12" pageNumber="13">rd</superScript>
|
||
& 4
|
||
<superScript id="C0E24D5CFFF1FFBFBAF1F9DAFE48F9C9" attach="left" box="[471,484,1585,1599]" fontSize="6" pageId="12" pageNumber="13">th</superScript>
|
||
contact orbit vs. no supralabials in contact with orbit) and anterior single inframaxillary (present vs. absent) (
|
||
<bibRefCitation id="53069DE5FFF1FFBFBA88F9BEFD4FF999" author="Grossmann, W. & Tillack, F." box="[430,739,1621,1647]" pageId="12" pageNumber="13" pagination="175 - 90" refId="ref14523" refString="Grossmann, W. & Tillack, F. (2003) On the Taxonomic Status of Asthenodipsas tropidonotus (Van Lidth De Jeude, 1923) and Pareas vertebralis (Boulenger, 1900) (Serpentes: Colubridae: Pareatinae). Russian Journal of Herpetology, 10 (3), 175 - 90." type="journal article" year="2003">Grossmann & Tillack 2003</bibRefCitation>
|
||
).
|
||
<taxonomicName id="F0979B97FFF1FFBFB9D3F9BEFBBAF999" authority="Quah & Grismer & Lim & Anuar & Chan, 2020" authorityName="Quah & Grismer & Lim & Anuar & Chan" authorityYear="2020" box="[757,1046,1621,1647]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="05E33C06FFF1FFBFB9D3F9BEFBBAF999" box="[757,1046,1621,1647]" italics="true" pageId="12" pageNumber="13">Asthenodipsas borneensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="05E33C06FFF1FFBFBF3AF9BEFBD9F999" bold="true" box="[1052,1141,1621,1647]" pageId="12" pageNumber="13">
|
||
<taxonomicNameLabel id="1ED0817DFFF1FFBFBF3AF9BEFBD9F999" box="[1052,1141,1621,1647]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be differentiated from its congeners by the following combination of characters: a maximum SVL of
|
||
<quantity id="F06F4DF1FFF1FFBFBF27F993FBCCF964" box="[1025,1120,1656,1682]" metricMagnitude="-1" metricUnit="m" metricValue="4.41" pageId="12" pageNumber="13" unit="mm" value="441.0">441 mm</quantity>
|
||
; 15/15/15 dorsal scale rows; 166–179 ventrals; 35–48 subcaudal scales; two postoculars; 2(+0–1)+(1–3) temporals; seven or eight supralabials, 3
|
||
<superScript id="C0E24D5CFFF1FFBFBB83F957FF1FF93C" attach="left" box="[165,179,1724,1738]" fontSize="6" pageId="12" pageNumber="13">rd</superScript>
|
||
& 4
|
||
<superScript id="C0E24D5CFFF1FFBFBBC0F957FF5FF93C" attach="left" box="[230,243,1724,1738]" fontSize="6" pageId="12" pageNumber="13">th</superScript>
|
||
touching the eye; 4–7 infralabials, 2
|
||
<superScript id="C0E24D5CFFF1FFBFB9A9F957FD33F93C" attach="left" box="[655,671,1724,1738]" fontSize="6" pageId="12" pageNumber="13">nd</superScript>
|
||
or 3
|
||
<superScript id="C0E24D5CFFF1FFBFB9F2F957FD4EF93C" attach="left" box="[724,738,1724,1738]" fontSize="6" pageId="12" pageNumber="13">rd</superScript>
|
||
pair in contact; a sharp vertebral keel; dorsum of adults beige to orange-brown with a dark-brown to black patch on the neck followed by 23–40 irregularly-shaped, rhomboidal dark-brown bands that extend the length of body and tail and onto lateral edges of the ventral scales to form spots, but not encircling body; a narrow, light-coloured vertebral stripe starting from the nape or after the dark patch on the neck; throat and ventrals beige to yellow and mottled with small, dark spots except for neck region which is darkbrown to black, sometimes with a faint, light-coloured median stripe; head whitish to dark-grey with dark speckling on the snout, crown and labials; and iris reddish-brown and pupils black (
|
||
<tableCitation id="7A15D5AFFFF1FFBFB8EBF866FBB2F851" box="[973,1054,1933,1959]" captionStart="TABLE 1" captionStartId="2.[151,239,586,610]" captionTargetBox="[163,1422,662,1957]" captionText="TABLE 1. Scale counts and selected measurements (mm) of the type series of Asthenodipsas borneensis sp. nov. and A. malaccana. Abbreviations are listed in the Materials and methods." httpUri="http://table.plazi.org/id/63E8B09CFFFFFFB1BBB1FDA1FCF7FD71" pageId="12" pageNumber="13" tableUuid="63E8B09CFFFFFFB1BBB1FDA1FCF7FD71">Table 1</tableCitation>
|
||
,
|
||
<tableCitation id="7A15D5AFFFF1FFBFBF03F866FB9BF851" box="[1061,1079,1933,1959]" captionStart="TABLE 2" captionStartId="5.[151,244,484,511]" captionTargetBox="[162,1983,562,1664]" captionText="TABLE 2. Measurements and scale counts of Asthenodipsas borneensis sp. nov. and A. malaccana (*measurements from Grossmann & Tillack 2003; NA = measurement not available). Abbreviations are listed in the Materials and methods." httpUri="http://table.plazi.org/id/63E8B09CFFF8FFB6BBB1FE0FFC30FDD5" pageId="12" pageNumber="13" tableUuid="63E8B09CFFF8FFB6BBB1FE0FFC30FDD5">2</tableCitation>
|
||
&
|
||
<tableCitation id="7A15D5AFFFF1FFBFBF7CF866FBC6F851" box="[1114,1130,1933,1959]" captionStart="TABLE 7" captionStartId="17.[151,244,296,323]" captionTargetBox="[163,1380,375,1280]" captionText="TABLE 7. Summary statistics and diagnostic characters for Asthenodipsas borneensis sp. nov., A. malaccana, A. stuebingi and A. laevis. SD = standard deviation and n = sample size." httpUri="http://table.plazi.org/id/63E8B09CFFECFFA2BBB1FEC3FC32FE91" pageId="12" pageNumber="13" tableUuid="63E8B09CFFECFFA2BBB1FEC3FC32FE91">7</tableCitation>
|
||
;
|
||
<figureCitation id="AFACFC91FFF1FFBFBF51F866FB14F851" box="[1143,1208,1933,1959]" captionStart="FIGURE 2" captionStartId="13.[151,250,1688,1713]" captionTargetBox="[151,1436,181,1662]" captionTargetId="figure@13.[151,1436,181,1662]" captionTargetPageId="13" captionText="FIGURE 2. A. Dorsum of holotype of Asthenodipsas borneensis sp. nov. (FMNH 243935). B. Venter of holotype of A. borneensis sp. nov. (FMNH 243935). C. Dorsum of paratype of A. borneensis sp. nov. (FMNH 243936). D. Venter of paratype of A. borneensis sp. nov. (FMNH 243936). E. Dorsum of paratype of A. borneensis sp. nov. (FMNH 249751). F. Venter of paratype of A. borneensis sp. nov. (FMNH 249751)." figureDoi="http://doi.org/10.5281/zenodo.3628950" httpUri="https://zenodo.org/record/3628950/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
|
||
&
|
||
<figureCitation id="AFACFC91FFF1FFBFBFFAF866FB40F851" box="[1244,1260,1933,1959]" captionStart="FIGURE 3" captionStartId="14.[151,250,1908,1933]" captionTargetBox="[170,1416,453,1883]" captionTargetId="figure@14.[170,1416,453,1883]" captionTargetPageId="14" captionText="FIGURE 3. Asthenodipsas borneensis sp. nov. from Borneo. A: Specimen from Kuching, Sarawak (LSUDPC 10988). B: Specimen from Sayap, Mount Kinabalu, Sabah (LSUDPC 10982). C – F (LSUDPC 10983–10986): Adults from Poring, Sabah. G & H (LSUDPC 10987 & 10978): Juvenile from Poring, Sabah. (Photographs by Chien C. Lee [A & B], Zhou Hang [C & D], Anton Sorokin [E & F] & Jaroslav Karhánek [G & H])." figureDoi="http://doi.org/10.5281/zenodo.3628952" httpUri="https://zenodo.org/record/3628952/files/figure.png" pageId="12" pageNumber="13">3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7F8DB39FFFF1FFA3BBE1F844FC1AFE01" lastPageId="16" lastPageNumber="17" pageId="12" pageNumber="13" type="description">
|
||
<paragraph id="3728E014FFF1FFBDBBE1F844FE2FFE59" blockId="12.[151,1437,927,2030]" lastBlockId="14.[151,1436,153,431]" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
|
||
<emphasis id="05E33C06FFF1FFBFBBE1F844FE49F83C" bold="true" box="[199,485,1967,1994]" pageId="12" pageNumber="13">
|
||
Description of
|
||
<typeStatus id="E82C5EB6FFF1FFBFBA51F85BFE4CF83C" box="[375,480,1968,1994]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult female SVL
|
||
<quantity id="F06F4DF1FFF1FFBFB999F85BFCB7F83C" box="[703,795,1968,1994]" metricMagnitude="-1" metricUnit="m" metricValue="4.11" pageId="12" pageNumber="13" unit="mm" value="411.0">411 mm</quantity>
|
||
and Tal
|
||
<quantity id="F06F4DF1FFF1FFBFB85DF844FC61F83C" box="[891,973,1967,1994]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="12" pageNumber="13" unit="mm" value="55.0">55 mm</quantity>
|
||
; rostral slightly wider than tall; head bulbous, somewhat rectangular in shape, longer than wide (HL/HW 1.73); nasals undivided; internasals shorter than prefrontals; posterior margin of prefrontals contact eye; frontal hexagonal, slightly longer than wide; loreals present, longer than tall; supraoculars subpentagonal, approximately a quarter of both the length and width of frontal; preoculars absent; upper and lower postoculars present on both right and left sides of head, lower postocular extending to below orbit; suboculars absent; supralabials 7/7 with 3rd and 4th contacting orbit and 7th elongate; temporals 2(+1)+2/2+1, i.e. on left side of head there are two anterior temporals and two posterior temporals but between lower anterior and lower posterior temporal is an additional scale resulting in two upper temporals and three lower temporals, and on the right side of the head the upper anterior and posterior temporals are fused resulting in a single upper temporal and three lower temporals; mental triangular wider than long; anterior inframaxillary pentagonal, in contact with infralabials 1–3; two pairs of posterior inframaxillaries following anterior inframaxillary, both pairs of posterior inframaxillaries somewhat hexagonal, rhomboid-shaped, elongated; infralabials 5/5 with 3
|
||
<superScript id="C0E24D5CFFF3FFBDBFC9FEC3FB51FEC0" attach="left" box="[1263,1277,296,310]" fontSize="6" pageId="14" pageNumber="15">rd</superScript>
|
||
pair in medial contact. Body long, stout, laterally compressed, bearing a prominent keel-shaped vertebral region; dorsal scales smooth, in 15/15/15 rows, vertebrals slightly enlarged; 179 ventrals; 38 divided subcaudals; cloacal scute entire; tail tapering to point.
|
||
</paragraph>
|
||
<caption id="63E8B09CFFF0FFBEBBB1F973FDE2F8EB" ID-DOI="http://doi.org/10.5281/zenodo.3628950" ID-Zenodo-Dep="3628950" httpUri="https://zenodo.org/record/3628950/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,250,1688,1713]" targetBox="[151,1436,181,1662]" targetPageId="13">
|
||
<paragraph id="3728E014FFF0FFBEBBB1F973FDE2F8EB" blockId="13.[151,1437,1688,1821]" pageId="13" pageNumber="14">
|
||
<emphasis id="05E33C06FFF0FFBEBBB1F973FEA0F947" bold="true" box="[151,268,1688,1713]" pageId="13" pageNumber="14">FIGURE 2</emphasis>
|
||
. A. Dorsum of holotype of
|
||
<taxonomicName id="F0979B97FFF0FFBEB93AF972FC8DF947" authority="Quah & Grismer & Lim & Anuar & Chan, 2020" authorityName="Quah & Grismer & Lim & Anuar & Chan" authorityYear="2020" box="[540,801,1689,1713]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="05E33C06FFF0FFBEB93AF972FC8DF947" box="[540,801,1689,1713]" italics="true" pageId="13" pageNumber="14">Asthenodipsas borneensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="05E33C06FFF0FFBEB800F972FCD9F947" bold="true" box="[806,885,1689,1713]" pageId="13" pageNumber="14">
|
||
<taxonomicNameLabel id="1ED0817DFFF0FFBEB800F972FCD9F947" box="[806,885,1689,1713]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(FMNH 243935). B. Venter of holotype of
|
||
<taxonomicName id="F0979B97FFF0FFBEBE00F972FF1EF922" authority="Quah & Grismer & Lim & Anuar & Chan, 2020" authorityName="Quah & Grismer & Lim & Anuar & Chan" authorityYear="2020" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="05E33C06FFF0FFBEBE00F972FF1EF922" italics="true" pageId="13" pageNumber="14">A. borneensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="05E33C06FFF0FFBEBB9FF956FEA7F923" bold="true" box="[185,267,1725,1749]" pageId="13" pageNumber="14">
|
||
<taxonomicNameLabel id="1ED0817DFFF0FFBEBB9FF956FEA7F923" box="[185,267,1725,1749]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(FMNH 243935). C. Dorsum of paratype of
|
||
<taxonomicName id="F0979B97FFF0FFBEB9F8F956FCC5F923" authority="Quah & Grismer & Lim & Anuar & Chan, 2020" authorityName="Quah & Grismer & Lim & Anuar & Chan" authorityYear="2020" box="[734,873,1725,1749]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="05E33C06FFF0FFBEB9F8F956FCC5F923" box="[734,873,1725,1749]" italics="true" pageId="13" pageNumber="14">A. borneensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="05E33C06FFF0FFBEB856F956FC6EF923" bold="true" box="[880,962,1725,1749]" pageId="13" pageNumber="14">
|
||
<taxonomicNameLabel id="1ED0817DFFF0FFBEB856F956FC6EF923" box="[880,962,1725,1749]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(FMNH 243936). D. Venter of paratype of
|
||
<taxonomicName id="F0979B97FFF0FFBEBEA0F956FEA9F90F" authority="Quah & Grismer & Lim & Anuar & Chan, 2020" authorityName="Quah & Grismer & Lim & Anuar & Chan" authorityYear="2020" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="05E33C06FFF0FFBEBEA0F956FEA9F90F" italics="true" pageId="13" pageNumber="14">A. borneensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="05E33C06FFF0FFBEBA2DF90AFEF0F90F" bold="true" box="[267,348,1761,1785]" pageId="13" pageNumber="14">
|
||
<taxonomicNameLabel id="1ED0817DFFF0FFBEBA2DF90AFEF0F90F" box="[267,348,1761,1785]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(FMNH 243936). E. Dorsum of paratype of
|
||
<taxonomicName id="F0979B97FFF0FFBEB800F90AFC1DF90F" authority="Quah & Grismer & Lim & Anuar & Chan, 2020" authorityName="Quah & Grismer & Lim & Anuar & Chan" authorityYear="2020" box="[806,945,1761,1785]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="05E33C06FFF0FFBEB800F90AFC1DF90F" box="[806,945,1761,1785]" italics="true" pageId="13" pageNumber="14">A. borneensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="05E33C06FFF0FFBEB891F90AFBA4F90F" bold="true" box="[951,1032,1761,1785]" pageId="13" pageNumber="14">
|
||
<taxonomicNameLabel id="1ED0817DFFF0FFBEB891F90AFBA4F90F" box="[951,1032,1761,1785]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(FMNH 249751). F. Venter of paratype of
|
||
<taxonomicName id="F0979B97FFF0FFBEBB95F8EEFE91F8EB" authority="Quah & Grismer & Lim & Anuar & Chan, 2020" authorityName="Quah & Grismer & Lim & Anuar & Chan" authorityYear="2020" box="[179,317,1797,1821]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="05E33C06FFF0FFBEBB95F8EEFE91F8EB" box="[179,317,1797,1821]" italics="true" pageId="13" pageNumber="14">A. borneensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="05E33C06FFF0FFBEBA65F8EEFE3FF8EB" bold="true" box="[323,403,1797,1821]" pageId="13" pageNumber="14">
|
||
<taxonomicNameLabel id="1ED0817DFFF0FFBEBA65F8EEFE3FF8EB" box="[323,403,1797,1821]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(FMNH 249751).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="63E8B09CFFF3FFBDBBB1F89FFD66F80F" ID-DOI="http://doi.org/10.5281/zenodo.3628952" ID-Zenodo-Dep="3628952" httpUri="https://zenodo.org/record/3628952/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,250,1908,1933]" targetBox="[170,1416,453,1883]" targetPageId="14">
|
||
<paragraph id="3728E014FFF3FFBDBBB1F89FFD66F80F" blockId="14.[151,1437,1908,2041]" pageId="14" pageNumber="15">
|
||
<emphasis id="05E33C06FFF3FFBDBBB1F89FFEBCF87B" bold="true" box="[151,272,1908,1933]" pageId="14" pageNumber="15">FIGURE 3</emphasis>
|
||
.
|
||
<taxonomicName id="F0979B97FFF3FFBDBA06F89EFD86F87B" authority="Quah & Grismer & Lim & Anuar & Chan, 2020" authorityName="Quah & Grismer & Lim & Anuar & Chan" authorityYear="2020" box="[288,554,1909,1933]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="05E33C06FFF3FFBDBA06F89EFD86F87B" box="[288,554,1909,1933]" italics="true" pageId="14" pageNumber="15">Asthenodipsas borneensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1ED0817DFFF3FFBDB915F89EFD24F87A" box="[563,648,1909,1933]" pageId="14" pageNumber="15" rank="species">
|
||
<emphasis id="05E33C06FFF3FFBDB915F89EFD2EF87B" bold="true" box="[563,642,1909,1933]" pageId="14" pageNumber="15">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
from Borneo. A: Specimen from Kuching, Sarawak (LSUDPC 10988). B: Specimen from Sayap, Mount Kinabalu, Sabah (LSUDPC 10982). C – F (LSUDPC 10983–10986): Adults from Poring, Sabah. G & H (LSUDPC 10987 & 10978): Juvenile from Poring, Sabah. (Photographs by Chien C. Lee [A & B], Zhou Hang [C & D], Anton Sorokin [E & F] & Jaroslav Karhánek [G & H]).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="63E8B09CFFF2FFBCBBB1F95EFEFDF874" ID-DOI="http://doi.org/10.5281/zenodo.3628954" ID-Zenodo-Dep="3628954" httpUri="https://zenodo.org/record/3628954/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,250,1717,1742]" targetBox="[151,1437,181,1691]" targetPageId="15">
|
||
<paragraph id="3728E014FFF2FFBCBBB1F95EFEFDF874" blockId="15.[151,1437,1717,1922]" pageId="15" pageNumber="16">
|
||
<emphasis id="05E33C06FFF2FFBCBBB1F95EFEA1F938" bold="true" box="[151,269,1717,1742]" pageId="15" pageNumber="16">FIGURE 4</emphasis>
|
||
.
|
||
<taxonomicName id="F0979B97FFF2FFBCBA3CF95DFD8EF938" box="[282,546,1718,1742]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="malaccana">
|
||
<emphasis id="05E33C06FFF2FFBCBA3CF95DFD8EF938" box="[282,546,1718,1742]" italics="true" pageId="15" pageNumber="16">Asthenodipsas malaccana</emphasis>
|
||
</taxonomicName>
|
||
from Peninsular Malaysia, Thailand and Sumatra. A: Adult male (USMHC 1949) from Puncak Janing, Kedah, Peninsular Malaysia. B: Adult female (LSUHC 12740) from Gunung Besar Hantu, Negeri Sembilan, Peninsular Malaysia. C: Specimen from Betong district, Yala province, Thailand (LSUDPC 10979). D: Specimen from Jambi, Sumatra, Indonesia (LSUDPC 10980). E: Specimen from near Bukit Barisan Selatan National Park, Lampung, Sumatra, Indonesia (LSUDPC 10981). (Photographs by Evan Quah [A], Chan Kin Onn [B], Parinya Pawangkhanant [C], Mediyansyah [D] & Raúl León [E]).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="3728E014FFF2FFA3BBE1F844FC1AFE01" blockId="15.[151,1436,1967,2030]" lastBlockId="16.[151,1437,152,2015]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
|
||
<emphasis id="05E33C06FFF2FFBCBBE1F844FD6FF83C" bold="true" box="[199,707,1967,1994]" pageId="15" pageNumber="16">
|
||
Colouration in preservation (
|
||
<figureCitation id="AFACFC91FFF2FFBCB905F844FD1FF83C" box="[547,691,1967,1994]" captionStart="FIGURE 2" captionStartId="13.[151,250,1688,1713]" captionTargetBox="[151,1436,181,1662]" captionTargetId="figure@13.[151,1436,181,1662]" captionTargetPageId="13" captionText="FIGURE 2. A. Dorsum of holotype of Asthenodipsas borneensis sp. nov. (FMNH 243935). B. Venter of holotype of A. borneensis sp. nov. (FMNH 243935). C. Dorsum of paratype of A. borneensis sp. nov. (FMNH 243936). D. Venter of paratype of A. borneensis sp. nov. (FMNH 243936). E. Dorsum of paratype of A. borneensis sp. nov. (FMNH 249751). F. Venter of paratype of A. borneensis sp. nov. (FMNH 249751)." figureDoi="http://doi.org/10.5281/zenodo.3628950" httpUri="https://zenodo.org/record/3628950/files/figure.png" pageId="15" pageNumber="16">Fig. 2A & B</figureCitation>
|
||
).
|
||
</emphasis>
|
||
The ground colour of the head, body and tail are beige with very fine speckling. The speckling is lighter on the flanks and becomes heavier on the back closer to the vertebral scale rows. The top and underside of the head is lightly speckled and there are dark spots on the rostal, 1
|
||
<superScript id="C0E24D5CFFEDFFA3BFFBFF73FB44FF50" attach="left" box="[1245,1256,152,166]" fontSize="6" pageId="16" pageNumber="17">st</superScript>
|
||
supralabials, 7
|
||
<superScript id="C0E24D5CFFEDFFA3BEB6FF73FA31FF50" attach="none" box="[1424,1437,152,166]" fontSize="6" pageId="16" pageNumber="17">th</superScript>
|
||
supralabials along opening of the mouth, mental, inframaxillaries, and infralabials 1–3 along their medial sutures. On the neck is a dark patch starting from the nape, extending to approximately the position of the 20
|
||
<superScript id="C0E24D5CFFEDFFA3BE27FF0BFAA2FF18" attach="left" box="[1281,1294,224,238]" fontSize="6" pageId="16" pageNumber="17">th</superScript>
|
||
ventral. The dark patch starts as a narrow stripe on the 5
|
||
<superScript id="C0E24D5CFFEDFFA3B951FEEFFD28FEE4" attach="left" box="[631,644,260,274]" fontSize="6" pageId="16" pageNumber="17">th</superScript>
|
||
& 6
|
||
<superScript id="C0E24D5CFFEDFFA3B993FEEFFD6EFEE4" attach="left" box="[693,706,260,274]" fontSize="6" pageId="16" pageNumber="17">th</superScript>
|
||
dorsal scale rows, gradually widens to dorsal scale rows 1–7, and does not extend cross the vertebral scales but completely encircles the ventrals. Starting behind the dark neck patch are 29 dark bands along the body that are somewhat rhomboidal to inverted-triangle in shape, and nine bands on the tail. The dark body bands range between 1–4 dorsal scales in length and are faint on the dorsum near the vertebral scale row and more prominent on the lower flanks and edges of the ventrals. The venter is beige and bears, dark lateral blotches from the extensions of the dorsal bands that meet the ventral scales as well as scattered dark spots the entire length of the body and tail. The thin vertebral stripe is cream.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7F8DB39FFFEDFFA3BBE1FDEBFE2BFCCD" pageId="16" pageNumber="17" type="materials_examined">
|
||
<paragraph id="3728E014FFEDFFA3BBE1FDEBFE2BFCCD" blockId="16.[151,1437,152,2015]" pageId="16" pageNumber="17">
|
||
<emphasis id="05E33C06FFEDFFA3BBE1FDEBFE62FDED" bold="true" box="[199,462,512,539]" pageId="16" pageNumber="17">
|
||
Variation (
|
||
<figureCitation id="AFACFC91FFEDFFA3BA60FDEBFE26FDED" box="[326,394,512,539]" captionStart="FIGURE 2" captionStartId="13.[151,250,1688,1713]" captionTargetBox="[151,1436,181,1662]" captionTargetId="figure@13.[151,1436,181,1662]" captionTargetPageId="13" captionText="FIGURE 2. A. Dorsum of holotype of Asthenodipsas borneensis sp. nov. (FMNH 243935). B. Venter of holotype of A. borneensis sp. nov. (FMNH 243935). C. Dorsum of paratype of A. borneensis sp. nov. (FMNH 243936). D. Venter of paratype of A. borneensis sp. nov. (FMNH 243936). E. Dorsum of paratype of A. borneensis sp. nov. (FMNH 249751). F. Venter of paratype of A. borneensis sp. nov. (FMNH 249751)." figureDoi="http://doi.org/10.5281/zenodo.3628950" httpUri="https://zenodo.org/record/3628950/files/figure.png" pageId="16" pageNumber="17">Fig. 2</figureCitation>
|
||
&
|
||
<figureCitation id="AFACFC91FFEDFFA3BA89FDEBFE13FDED" box="[431,447,512,539]" captionStart="FIGURE 3" captionStartId="14.[151,250,1908,1933]" captionTargetBox="[170,1416,453,1883]" captionTargetId="figure@14.[170,1416,453,1883]" captionTargetPageId="14" captionText="FIGURE 3. Asthenodipsas borneensis sp. nov. from Borneo. A: Specimen from Kuching, Sarawak (LSUDPC 10988). B: Specimen from Sayap, Mount Kinabalu, Sabah (LSUDPC 10982). C – F (LSUDPC 10983–10986): Adults from Poring, Sabah. G & H (LSUDPC 10987 & 10978): Juvenile from Poring, Sabah. (Photographs by Chien C. Lee [A & B], Zhou Hang [C & D], Anton Sorokin [E & F] & Jaroslav Karhánek [G & H])." figureDoi="http://doi.org/10.5281/zenodo.3628952" httpUri="https://zenodo.org/record/3628952/files/figure.png" pageId="16" pageNumber="17">3</figureCitation>
|
||
).
|
||
</emphasis>
|
||
The
|
||
<typeStatus id="E82C5EB6FFEDFFA3B920FDEAFDDDFDED" box="[518,625,513,539]" pageId="16" pageNumber="17" type="paratype">paratypes</typeStatus>
|
||
closely resemble the
|
||
<typeStatus id="E82C5EB6FFEDFFA3B845FDEAFC69FDED" box="[867,965,513,539]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
|
||
in overall colour pattern and morphology.
|
||
<typeStatus id="E82C5EB6FFEDFFA3BBB1FDCEFEA8FDC9" box="[151,260,549,575]" pageId="16" pageNumber="17" type="paratype">Paratypes</typeStatus>
|
||
FMNH 243936 and 249751 have more prominent banding on the body than the
|
||
<typeStatus id="E82C5EB6FFEDFFA3BF5FFDCEFB77FDC9" box="[1145,1243,549,575]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="AFACFC91FFEDFFA3BFCFFDCEFADFFDC9" box="[1257,1395,549,575]" captionStart="FIGURE 2" captionStartId="13.[151,250,1688,1713]" captionTargetBox="[151,1436,181,1662]" captionTargetId="figure@13.[151,1436,181,1662]" captionTargetPageId="13" captionText="FIGURE 2. A. Dorsum of holotype of Asthenodipsas borneensis sp. nov. (FMNH 243935). B. Venter of holotype of A. borneensis sp. nov. (FMNH 243935). C. Dorsum of paratype of A. borneensis sp. nov. (FMNH 243936). D. Venter of paratype of A. borneensis sp. nov. (FMNH 243936). E. Dorsum of paratype of A. borneensis sp. nov. (FMNH 249751). F. Venter of paratype of A. borneensis sp. nov. (FMNH 249751)." figureDoi="http://doi.org/10.5281/zenodo.3628950" httpUri="https://zenodo.org/record/3628950/files/figure.png" pageId="16" pageNumber="17">Fig. 2C & E</figureCitation>
|
||
). In FMNH 243936 the markings from the dark neck patch do not encircle the ventrals, leaving a narrow, light-coloured median stripe on the underside of the neck (
|
||
<figureCitation id="AFACFC91FFEDFFA3B95AFD86FD79FD71" box="[636,725,621,647]" captionStart="FIGURE 2" captionStartId="13.[151,250,1688,1713]" captionTargetBox="[151,1436,181,1662]" captionTargetId="figure@13.[151,1436,181,1662]" captionTargetPageId="13" captionText="FIGURE 2. A. Dorsum of holotype of Asthenodipsas borneensis sp. nov. (FMNH 243935). B. Venter of holotype of A. borneensis sp. nov. (FMNH 243935). C. Dorsum of paratype of A. borneensis sp. nov. (FMNH 243936). D. Venter of paratype of A. borneensis sp. nov. (FMNH 243936). E. Dorsum of paratype of A. borneensis sp. nov. (FMNH 249751). F. Venter of paratype of A. borneensis sp. nov. (FMNH 249751)." figureDoi="http://doi.org/10.5281/zenodo.3628950" httpUri="https://zenodo.org/record/3628950/files/figure.png" pageId="16" pageNumber="17">Fig. 2D</figureCitation>
|
||
).
|
||
<typeStatus id="E82C5EB6FFEDFFA3B9CEFD86FCF9FD71" box="[744,853,621,647]" pageId="16" pageNumber="17" type="paratype">Paratypes</typeStatus>
|
||
FMNH 71590 and 249751 also have chin and throat regions that are almost completely dark with only a few scattered light spots (
|
||
<figureCitation id="AFACFC91FFEDFFA3BF33FD7AFBCAFD5D" box="[1045,1126,657,683]" captionStart="FIGURE 2" captionStartId="13.[151,250,1688,1713]" captionTargetBox="[151,1436,181,1662]" captionTargetId="figure@13.[151,1436,181,1662]" captionTargetPageId="13" captionText="FIGURE 2. A. Dorsum of holotype of Asthenodipsas borneensis sp. nov. (FMNH 243935). B. Venter of holotype of A. borneensis sp. nov. (FMNH 243935). C. Dorsum of paratype of A. borneensis sp. nov. (FMNH 243936). D. Venter of paratype of A. borneensis sp. nov. (FMNH 243936). E. Dorsum of paratype of A. borneensis sp. nov. (FMNH 249751). F. Venter of paratype of A. borneensis sp. nov. (FMNH 249751)." figureDoi="http://doi.org/10.5281/zenodo.3628950" httpUri="https://zenodo.org/record/3628950/files/figure.png" pageId="16" pageNumber="17">Fig 2F</figureCitation>
|
||
). In life, the colouration of the head and body in this species is variable. The colouration of the head ranges from white with black mottling to dark-brown or grey, while the body ranges from beige to orange-brown (
|
||
<figureCitation id="AFACFC91FFEDFFA3B8E4FD32FBAAFD05" box="[962,1030,729,755]" captionStart="FIGURE 3" captionStartId="14.[151,250,1908,1933]" captionTargetBox="[170,1416,453,1883]" captionTargetId="figure@14.[170,1416,453,1883]" captionTargetPageId="14" captionText="FIGURE 3. Asthenodipsas borneensis sp. nov. from Borneo. A: Specimen from Kuching, Sarawak (LSUDPC 10988). B: Specimen from Sayap, Mount Kinabalu, Sabah (LSUDPC 10982). C – F (LSUDPC 10983–10986): Adults from Poring, Sabah. G & H (LSUDPC 10987 & 10978): Juvenile from Poring, Sabah. (Photographs by Chien C. Lee [A & B], Zhou Hang [C & D], Anton Sorokin [E & F] & Jaroslav Karhánek [G & H])." figureDoi="http://doi.org/10.5281/zenodo.3628952" httpUri="https://zenodo.org/record/3628952/files/figure.png" pageId="16" pageNumber="17">Fig. 3</figureCitation>
|
||
). Juveniles are lighter coloured than the adults (
|
||
<figureCitation id="AFACFC91FFEDFFA3BA34FD16FE0FFCE1" box="[274,419,765,791]" captionStart="FIGURE 3" captionStartId="14.[151,250,1908,1933]" captionTargetBox="[170,1416,453,1883]" captionTargetId="figure@14.[170,1416,453,1883]" captionTargetPageId="14" captionText="FIGURE 3. Asthenodipsas borneensis sp. nov. from Borneo. A: Specimen from Kuching, Sarawak (LSUDPC 10988). B: Specimen from Sayap, Mount Kinabalu, Sabah (LSUDPC 10982). C – F (LSUDPC 10983–10986): Adults from Poring, Sabah. G & H (LSUDPC 10987 & 10978): Juvenile from Poring, Sabah. (Photographs by Chien C. Lee [A & B], Zhou Hang [C & D], Anton Sorokin [E & F] & Jaroslav Karhánek [G & H])." figureDoi="http://doi.org/10.5281/zenodo.3628952" httpUri="https://zenodo.org/record/3628952/files/figure.png" pageId="16" pageNumber="17">Fig. 3G & H</figureCitation>
|
||
). Variation in scale counts and measurements within the type series and referred specimens are shown in
|
||
<tableCitation id="7A15D5AFFFEDFFA3BA0BFCCAFE2DFCCD" box="[301,385,801,827]" captionStart="TABLE 1" captionStartId="2.[151,239,586,610]" captionTargetBox="[163,1422,662,1957]" captionText="TABLE 1. Scale counts and selected measurements (mm) of the type series of Asthenodipsas borneensis sp. nov. and A. malaccana. Abbreviations are listed in the Materials and methods." httpUri="http://table.plazi.org/id/63E8B09CFFFFFFB1BBB1FDA1FCF7FD71" pageId="16" pageNumber="17" tableUuid="63E8B09CFFFFFFB1BBB1FDA1FCF7FD71">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7F8DB39FFFEDFFA3BBE1FCAFFB0CF9A5" pageId="16" pageNumber="17" type="nomenclature">
|
||
<paragraph id="3728E014FFEDFFA3BBE1FCAFFB0CF9A5" blockId="16.[151,1437,152,2015]" pageId="16" pageNumber="17">
|
||
<emphasis id="05E33C06FFEDFFA3BBE1FCAFFECEFCA9" bold="true" box="[199,354,836,863]" pageId="16" pageNumber="17">Comparison.</emphasis>
|
||
<taxonomicName id="F0979B97FFEDFFA3BA4DFCAEFD23FCA9" authority="Quah & Grismer & Lim & Anuar & Chan, 2020" authorityName="Quah & Grismer & Lim & Anuar & Chan" authorityYear="2020" box="[363,655,837,863]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="05E33C06FFEDFFA3BA4DFCAEFD23FCA9" box="[363,655,837,863]" italics="true" pageId="16" pageNumber="17">Asthenodipsas borneensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="05E33C06FFEDFFA3B9BEFCAEFD58FCA9" bold="true" box="[664,756,837,863]" pageId="16" pageNumber="17">
|
||
<taxonomicNameLabel id="1ED0817DFFEDFFA3B9BEFCAEFD58FCA9" box="[664,756,837,863]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be differentiated from
|
||
<taxonomicName id="F0979B97FFEDFFA3BF17FCAEFDD8FC75" authority="Loredo, Wood, Quah, Shahrul, Greer, Ahmad & Grismer" authorityName="Loredo, Wood, Quah, Shahrul, Greer, Ahmad & Grismer" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="lasgalenensis">
|
||
<emphasis id="05E33C06FFEDFFA3BF17FCAEFB45FCA9" box="[1073,1257,837,863]" italics="true" pageId="16" pageNumber="17">A. lasgalenensis</emphasis>
|
||
Loredo, Wood, Quah, Shahrul, Greer, Ahmad & Grismer
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F0979B97FFEDFFA3B9A7FC82FC47FC72" authority="(Lidth de Jude)" baseAuthorityName="Lidth de Jude" box="[641,1003,873,900]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="tropidonotus">
|
||
<emphasis id="05E33C06FFEDFFA3B9A7FC82FC9DFC75" box="[641,817,873,899]" italics="true" pageId="16" pageNumber="17">A. tropidonotus</emphasis>
|
||
(Lidth de Jude)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F0979B97FFEDFFA3BF00FC82FAFFFC72" authority="(Boulenger)" baseAuthorityName="Boulenger" baseAuthorityYear="1900" box="[1062,1363,873,900]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="vertebralis">
|
||
<emphasis id="05E33C06FFEDFFA3BF00FC82FB6DFC75" box="[1062,1217,873,899]" italics="true" pageId="16" pageNumber="17">A. vertebralis</emphasis>
|
||
(Boulenger)
|
||
</taxonomicName>
|
||
by its lower number subcaudals (35–48 vs. 54–77), pairs of infralabials in contact (3
|
||
<superScript id="C0E24D5CFFEDFFA3BF25FC67FBBDFC6C" attach="left" box="[1027,1041,908,922]" fontSize="6" pageId="16" pageNumber="17">rd</superScript>
|
||
vs. 1
|
||
<superScript id="C0E24D5CFFEDFFA3BF6AFC67FBFBFC6C" attach="none" box="[1100,1111,908,922]" fontSize="6" pageId="16" pageNumber="17">st</superScript>
|
||
) and fewer pairs of posterior inframaxillaries (two vs. three).
|
||
<taxonomicName id="F0979B97FFEDFFA3B92CFC5AFD08FC3D" authority="Quah & Grismer & Lim & Anuar & Chan, 2020" authorityName="Quah & Grismer & Lim & Anuar & Chan" authorityYear="2020" box="[522,676,945,971]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="05E33C06FFEDFFA3B92CFC5AFD08FC3D" box="[522,676,945,971]" italics="true" pageId="16" pageNumber="17">A. borneensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="05E33C06FFEDFFA3B988FC5AFCA6FC3D" bold="true" box="[686,778,945,971]" pageId="16" pageNumber="17">
|
||
<taxonomicNameLabel id="1ED0817DFFEDFFA3B988FC5AFCA6FC3D" box="[686,778,945,971]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is distinguished from
|
||
<taxonomicName id="F0979B97FFEDFFA3BF28FC5AFBDDFC3D" baseAuthorityName="Boie" baseAuthorityYear="1827" box="[1038,1137,945,971]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="laevis">
|
||
<emphasis id="05E33C06FFEDFFA3BF28FC5AFBDDFC3D" box="[1038,1137,945,971]" italics="true" pageId="16" pageNumber="17">A. laevis</emphasis>
|
||
</taxonomicName>
|
||
by its larger adult length, (Max SVL
|
||
<quantity id="F06F4DF1FFEDFFA3BA3EFC3EFEDAFC19" box="[280,374,981,1007]" metricMagnitude="-1" metricUnit="m" metricValue="4.41" pageId="16" pageNumber="17" unit="mm" value="441.0">441 mm</quantity>
|
||
vs.
|
||
<quantity id="F06F4DF1FFEDFFA3BA83FC3EFDABFC19" box="[421,519,981,1007]" metricMagnitude="-1" metricUnit="m" metricValue="3.7299999999999995" pageId="16" pageNumber="17" unit="mm" value="373.0">373 mm</quantity>
|
||
), dorsal scales rows (15/15/15 vs. 15/15/13) and sharp vertebral keel (present vs. absent). From
|
||
<taxonomicName id="F0979B97FFEDFFA3BA1FFC12FE7CFBE5" box="[313,464,1017,1043]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="jamilinaisi">
|
||
<emphasis id="05E33C06FFEDFFA3BA1FFC12FE7CFBE5" box="[313,464,1017,1043]" italics="true" pageId="16" pageNumber="17">A. jamilinaisi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F0979B97FFEDFFA3BAFBFC12FDD9FBE5" authority="Quah & Grismer & Lim & Anuar & Chan, 2020" authorityName="Quah & Grismer & Lim & Anuar & Chan" authorityYear="2020" box="[477,629,1017,1043]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="05E33C06FFEDFFA3BAFBFC12FDD9FBE5" box="[477,629,1017,1043]" italics="true" pageId="16" pageNumber="17">A. borneensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="05E33C06FFEDFFA3B95AFC12FD79FBE5" bold="true" box="[636,725,1017,1043]" pageId="16" pageNumber="17">
|
||
<taxonomicNameLabel id="1ED0817DFFEDFFA3B95AFC12FD79FBE5" box="[636,725,1017,1043]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be differentiated by its lower number of subcaudals (35–48 vs. 52–53), higher number of supralabials (seven or eight vs. six [rarely seven]), size of vertebral scales (slightly enlarged vs greatly enlarged), body form (robust and bulky vs. gracile and laterally compressed), and colour pattern (beige to orange-brown dorsum with a dark neck patch followed by distinct dark bands and a usually lighter coloured head vs. dark overall colouration of dorsum with muted banding). From
|
||
<taxonomicName id="F0979B97FFEDFFA3BF09FB62FB1AFB55" box="[1071,1206,1161,1187]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="stuebingi">
|
||
<emphasis id="05E33C06FFEDFFA3BF09FB62FB1AFB55" box="[1071,1206,1161,1187]" italics="true" pageId="16" pageNumber="17">A. stuebingi</emphasis>
|
||
</taxonomicName>
|
||
to which it is superficially similar to,
|
||
<taxonomicName id="F0979B97FFEDFFA3BA41FB46FDACFB31" authority="Quah & Grismer & Lim & Anuar & Chan, 2020" authorityName="Quah & Grismer & Lim & Anuar & Chan" authorityYear="2020" box="[359,512,1197,1223]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="05E33C06FFEDFFA3BA41FB46FDACFB31" box="[359,512,1197,1223]" italics="true" pageId="16" pageNumber="17">A. borneensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1ED0817DFFEDFFA3B92EFB46FDCFFB31" box="[520,611,1197,1223]" pageId="16" pageNumber="17" rank="species">
|
||
<emphasis id="05E33C06FFEDFFA3B92EFB46FDF1FB31" bold="true" box="[520,605,1197,1223]" pageId="16" pageNumber="17">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
can be distinguished by its higher number of supralabials (seven or eight vs. six), smaller adult length (Max SVL
|
||
<quantity id="F06F4DF1FFEDFFA3B97AFB3AFD16FB1D" box="[604,698,1233,1259]" metricMagnitude="-1" metricUnit="m" metricValue="4.41" pageId="16" pageNumber="17" unit="mm" value="441.0">441 mm</quantity>
|
||
vs.
|
||
<quantity id="F06F4DF1FFEDFFA3B9CEFB3BFCEAFB1D" box="[744,838,1232,1259]" metricMagnitude="-1" metricUnit="m" metricValue="5.57" pageId="16" pageNumber="17" unit="mm" value="557.0">557 mm</quantity>
|
||
) and colour of iris (red vs. black) (
|
||
<bibRefCitation id="53069DE5FFEDFFA3BFE8FB3AFA3DFB1D" author="Quah, E. S. H. & Grismer, L. L. & Lim, K. K. P. & Shahrul, A. M. S. & Imbun, P. Y." box="[1230,1425,1233,1259]" pageId="16" pageNumber="17" pagination="501 - 526" refId="ref15456" refString="Quah, E. S. H., Grismer, L. L., Lim, K. K. P., Shahrul, A. M. S. & Imbun, P. Y. (2019) A taxonomic reappraisal of the Smooth Slug Snake Asthenodipsas laevis (Boie, 1827) (Squamata: Pareidae) in Borneo with the description of two new species. Zootaxa, 4646 (3), 501 - 526. https: // doi. org / 10.11646 / zootaxa. 4646.3.4" type="journal article" year="2019">
|
||
Quah
|
||
<emphasis id="05E33C06FFEDFFA3BE32FB39FAE1FB1D" box="[1300,1357,1233,1259]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="F0979B97FFEDFFA3BBB1FB1EFEB3FAF9" box="[151,287,1269,1295]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="stuebingi">
|
||
<emphasis id="05E33C06FFEDFFA3BBB1FB1EFEB3FAF9" box="[151,287,1269,1295]" italics="true" pageId="16" pageNumber="17">A. stuebingi</emphasis>
|
||
</taxonomicName>
|
||
is also an upland species known only from the highlands of
|
||
<collectingRegion id="F5532EF6FFEDFFA3B8F2FB1EFBB7FAF9" box="[980,1051,1269,1295]" country="Malaysia" name="Sabah" pageId="16" pageNumber="17">Sabah</collectingRegion>
|
||
, East
|
||
<collectingCountry id="4F80A084FFEDFFA3BF46FB1EFB6BFAF9" box="[1120,1223,1269,1295]" name="Malaysia" pageId="16" pageNumber="17">Malaysia</collectingCountry>
|
||
from
|
||
<quantity id="F06F4DF1FFEDFFA3BE28FB1EFA31FAF9" box="[1294,1437,1269,1295]" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="2.0" metricValueMin="0.9" pageId="16" pageNumber="17" unit="m" value="1450.0" valueMax="2000.0" valueMin="900.0">900–2000 m</quantity>
|
||
a.s.l. in elevation (
|
||
<bibRefCitation id="53069DE5FFEDFFA3BA43FAF2FD88FAC5" author="Quah, E. S. H. & Grismer, L. L. & Lim, K. K. P. & Shahrul, A. M. S. & Imbun, P. Y." box="[357,548,1305,1331]" pageId="16" pageNumber="17" pagination="501 - 526" refId="ref15456" refString="Quah, E. S. H., Grismer, L. L., Lim, K. K. P., Shahrul, A. M. S. & Imbun, P. Y. (2019) A taxonomic reappraisal of the Smooth Slug Snake Asthenodipsas laevis (Boie, 1827) (Squamata: Pareidae) in Borneo with the description of two new species. Zootaxa, 4646 (3), 501 - 526. https: // doi. org / 10.11646 / zootaxa. 4646.3.4" type="journal article" year="2019">
|
||
Quah
|
||
<emphasis id="05E33C06FFEDFFA3BA8CFAF1FE70FAC5" box="[426,476,1305,1331]" italics="true" pageId="16" pageNumber="17">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
) whereas
|
||
<taxonomicName id="F0979B97FFEDFFA3B9B3FAF2FC81FAC5" authority="Quah & Grismer & Lim & Anuar & Chan, 2020" authorityName="Quah & Grismer & Lim & Anuar & Chan" authorityYear="2020" box="[661,813,1305,1331]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="05E33C06FFEDFFA3B9B3FAF2FC81FAC5" box="[661,813,1305,1331]" italics="true" pageId="16" pageNumber="17">A. borneensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1ED0817DFFEDFFA3B813FAF2FC3CFAC5" box="[821,912,1305,1331]" pageId="16" pageNumber="17" rank="species">
|
||
<emphasis id="05E33C06FFEDFFA3B813FAF2FC25FAC5" bold="true" box="[821,905,1305,1331]" pageId="16" pageNumber="17">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
ranges widely across Borneo (
|
||
<figureCitation id="AFACFC91FFEDFFA3BFCFFAF2FA80FAC5" box="[1257,1324,1304,1331]" captionStart="FIGURE 5" captionStartId="18.[151,250,1335,1360]" captionTargetBox="[151,1436,489,1310]" captionTargetId="figure@18.[151,1436,489,1310]" captionTargetPageId="18" captionText="FIGURE 5. Distribution of Asthenodipsas borneensis sp. nov. (red) and A. malaccana (blue). Star = type locality." figureDoi="http://doi.org/10.5281/zenodo.3628956" httpUri="https://zenodo.org/record/3628956/files/figure.png" pageId="16" pageNumber="17">Fig. 5</figureCitation>
|
||
) from the lowlands to approximately
|
||
<quantity id="F06F4DF1FFEDFFA3BAEEFAD6FDB0FAA1" box="[456,540,1341,1367]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="17" unit="m" value="1000.0">1000 m</quantity>
|
||
a.s.l. in elevation (
|
||
<bibRefCitation id="53069DE5FFEDFFA3B9C8FAD6FC79FAA1" author="Stuebing, R. B. & Inger, R. F. & Lardner, B." box="[750,981,1341,1367]" pageId="16" pageNumber="17" refId="ref15734" refString="Stuebing, R. B., Inger, R. F. & Lardner, B. (2014) A field guide to the snakes of Borneo. 2 nd Edition. Natural History Publications (Borneo), Kota Kinabalu, 310 pp." type="book" year="2014">
|
||
Stuebing
|
||
<emphasis id="05E33C06FFEDFFA3B87CFAD5FC27FAA1" box="[858,907,1341,1367]" italics="true" pageId="16" pageNumber="17">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
). Finally, from
|
||
<taxonomicName id="F0979B97FFEDFFA3BF59FAD6FAB5FAA1" box="[1151,1305,1341,1367]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="malaccana">
|
||
<emphasis id="05E33C06FFEDFFA3BF59FAD6FAB5FAA1" box="[1151,1305,1341,1367]" italics="true" pageId="16" pageNumber="17">A. malaccana</emphasis>
|
||
</taxonomicName>
|
||
with which it was considered synonymous,
|
||
<taxonomicName id="F0979B97FFEDFFA3B926FA8AFD35FA8D" authority="Quah & Grismer & Lim & Anuar & Chan, 2020" authorityName="Quah & Grismer & Lim & Anuar & Chan" authorityYear="2020" box="[512,665,1377,1403]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="05E33C06FFEDFFA3B926FA8AFD35FA8D" box="[512,665,1377,1403]" italics="true" pageId="16" pageNumber="17">A. borneensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="05E33C06FFEDFFA3B987FA8AFD57FA8D" bold="true" box="[673,763,1377,1403]" pageId="16" pageNumber="17">
|
||
<taxonomicNameLabel id="1ED0817DFFEDFFA3B987FA8AFD57FA8D" box="[673,763,1377,1403]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be differentiated by its larger adult length (max SVL =
|
||
<quantity id="F06F4DF1FFEDFFA3BBB1FA6EFF58FA69" box="[151,244,1413,1439]" metricMagnitude="-1" metricUnit="m" metricValue="4.41" pageId="16" pageNumber="17" unit="mm" value="441.0">441 mm</quantity>
|
||
vs.
|
||
<quantity id="F06F4DF1FFEDFFA3BA04FA6FFE28FA69" box="[290,388,1412,1439]" metricMagnitude="-1" metricUnit="m" metricValue="3.65" pageId="16" pageNumber="17" unit="mm" value="365.0">365 mm</quantity>
|
||
), a significantly higher number of ventrals (166–179 vs. 152–169) and subcaudals (35–48 vs. 26–42), a sharp vertebral keel (present vs. absent) and colour pattern (beige to orange-brown dorsum with a dark neck patch followed by distinct dark bands and a usually lighter coloured head vs. dark-brown to black dorsum with a series of silver-grey to dirty-white blotches that form irregular cross bands across the back that start behind a broad dark area on the neck, and the colouration of the head ranging from whitish with dark spots to solid black) (
|
||
<figureCitation id="AFACFC91FFEDFFA3BE42F9FEFF6EF9A5" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="13.[151,250,1688,1713]" captionStartId-1="14.[151,250,1908,1933]" captionStartId-2="15.[151,250,1717,1742]" captionTargetBox-0="[151,1436,181,1662]" captionTargetBox-1="[170,1416,453,1883]" captionTargetBox-2="[151,1437,181,1691]" captionTargetId-0="figure@13.[151,1436,181,1662]" captionTargetId-1="figure@14.[170,1416,453,1883]" captionTargetId-2="figure@15.[151,1437,181,1691]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionTargetPageId-2="15" captionText-0="FIGURE 2. A. Dorsum of holotype of Asthenodipsas borneensis sp. nov. (FMNH 243935). B. Venter of holotype of A. borneensis sp. nov. (FMNH 243935). C. Dorsum of paratype of A. borneensis sp. nov. (FMNH 243936). D. Venter of paratype of A. borneensis sp. nov. (FMNH 243936). E. Dorsum of paratype of A. borneensis sp. nov. (FMNH 249751). F. Venter of paratype of A. borneensis sp. nov. (FMNH 249751)." captionText-1="FIGURE 3. Asthenodipsas borneensis sp. nov. from Borneo. A: Specimen from Kuching, Sarawak (LSUDPC 10988). B: Specimen from Sayap, Mount Kinabalu, Sabah (LSUDPC 10982). C – F (LSUDPC 10983–10986): Adults from Poring, Sabah. G & H (LSUDPC 10987 & 10978): Juvenile from Poring, Sabah. (Photographs by Chien C. Lee [A & B], Zhou Hang [C & D], Anton Sorokin [E & F] & Jaroslav Karhánek [G & H])." captionText-2="FIGURE 4. Asthenodipsas malaccana from Peninsular Malaysia, Thailand and Sumatra. A: Adult male (USMHC 1949) from Puncak Janing, Kedah, Peninsular Malaysia. B: Adult female (LSUHC 12740) from Gunung Besar Hantu, Negeri Sembilan, Peninsular Malaysia. C: Specimen from Betong district, Yala province, Thailand (LSUDPC 10979). D: Specimen from Jambi, Sumatra, Indonesia (LSUDPC 10980). E: Specimen from near Bukit Barisan Selatan National Park, Lampung, Sumatra, Indonesia (LSUDPC 10981). (Photographs by Evan Quah [A], Chan Kin Onn [B], Parinya Pawangkhanant [C], Mediyansyah [D] & Raúl León [E])." figureDoi-0="http://doi.org/10.5281/zenodo.3628950" figureDoi-1="http://doi.org/10.5281/zenodo.3628952" figureDoi-2="http://doi.org/10.5281/zenodo.3628954" httpUri-0="https://zenodo.org/record/3628950/files/figure.png" httpUri-1="https://zenodo.org/record/3628952/files/figure.png" httpUri-2="https://zenodo.org/record/3628954/files/figure.png" pageId="16" pageNumber="17">Figs. 2–4</figureCitation>
|
||
) (
|
||
<tableCitation id="7A15D5AFFFEDFFA3BBFFF9D2FEF9F9A5" box="[217,341,1593,1619]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStart-3="TABLE 4" captionStartId-0="2.[151,239,586,610]" captionStartId-1="5.[151,244,484,511]" captionStartId-2="7.[151,244,152,179]" captionStartId-3="7.[151,244,627,654]" captionTargetBox-0="[163,1422,662,1957]" captionTargetBox-1="[162,1983,562,1664]" captionTargetBox-2="[163,1370,267,580]" captionTargetBox-3="[163,1370,267,580]" captionTargetPageId-2="7" captionTargetPageId-3="7" captionText-0="TABLE 1. Scale counts and selected measurements (mm) of the type series of Asthenodipsas borneensis sp. nov. and A. malaccana. Abbreviations are listed in the Materials and methods." captionText-1="TABLE 2. Measurements and scale counts of Asthenodipsas borneensis sp. nov. and A. malaccana (*measurements from Grossmann & Tillack 2003; NA = measurement not available). Abbreviations are listed in the Materials and methods." captionText-2="TABLE 3. Pairwise matrix of characters with statistically significant mean differences among Asthenodipsas borneensis sp. nov., A. malaccana, A. stuebingi, A. jamilinaisi and A. laevis. Abbreviations are in the Materials and methods." captionText-3="TABLE 4. p values for statistically significant mean differences of characters among the Asthenodipsas borneensis sp. nov., A. malaccana, A. stuebingi, A. jamilinaisi and A. laevis species pairs." httpUri-0="http://table.plazi.org/id/63E8B09CFFFFFFB1BBB1FDA1FCF7FD71" httpUri-1="http://table.plazi.org/id/63E8B09CFFF8FFB6BBB1FE0FFC30FDD5" httpUri-2="http://table.plazi.org/id/63E8B09CFFFAFFB4BBB1FF73FF52FF0D" httpUri-3="http://table.plazi.org/id/63E8B09CFFFAFFB4BBB1FD98FBAAFD44" pageId="16" pageNumber="17" tableUuid-0="63E8B09CFFFFFFB1BBB1FDA1FCF7FD71" tableUuid-1="63E8B09CFFF8FFB6BBB1FE0FFC30FDD5" tableUuid-2="63E8B09CFFFAFFB4BBB1FF73FF52FF0D" tableUuid-3="63E8B09CFFFAFFB4BBB1FD98FBAAFD44">Tables 1–4</tableCitation>
|
||
&
|
||
<tableCitation id="7A15D5AFFFEDFFA3BA5EF9D2FE24F9A5" box="[376,392,1593,1619]" captionStart="TABLE 7" captionStartId="17.[151,244,296,323]" captionTargetBox="[163,1380,375,1280]" captionText="TABLE 7. Summary statistics and diagnostic characters for Asthenodipsas borneensis sp. nov., A. malaccana, A. stuebingi and A. laevis. SD = standard deviation and n = sample size." httpUri="http://table.plazi.org/id/63E8B09CFFECFFA2BBB1FEC3FC32FE91" pageId="16" pageNumber="17" tableUuid="63E8B09CFFECFFA2BBB1FEC3FC32FE91">7</tableCitation>
|
||
). An updated key to the family
|
||
<taxonomicName id="F0979B97FFEDFFA3B9CCF9D2FCE6F9A5" authorityName="Romer" authorityYear="1956" box="[746,842,1593,1619]" class="Reptilia" family="Pareidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="family">Pareidae</taxonomicName>
|
||
of Borneo is presented below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7F8DB39FFFEDFFA3BBE1F9B7FB47F8F1" pageId="16" pageNumber="17" type="distribution">
|
||
<paragraph id="3728E014FFEDFFA3BBE1F9B7FB47F8F1" blockId="16.[151,1437,152,2015]" pageId="16" pageNumber="17">
|
||
<emphasis id="05E33C06FFEDFFA3BBE1F9B7FECDF981" bold="true" box="[199,353,1628,1655]" pageId="16" pageNumber="17">Distribution.</emphasis>
|
||
<taxonomicName id="F0979B97FFEDFFA3BA41F9B6FD25F981" authority="Quah & Grismer & Lim & Anuar & Chan, 2020" authorityName="Quah & Grismer & Lim & Anuar & Chan" authorityYear="2020" box="[359,649,1629,1655]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="05E33C06FFEDFFA3BA41F9B6FD25F981" box="[359,649,1629,1655]" italics="true" pageId="16" pageNumber="17">Asthenodipsas borneensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1ED0817DFFEDFFA3B9B6F9B6FD47F981" box="[656,747,1629,1655]" pageId="16" pageNumber="17" rank="species">
|
||
<emphasis id="05E33C06FFEDFFA3B9B6F9B6FD49F981" bold="true" box="[656,741,1629,1655]" pageId="16" pageNumber="17">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
ranges from as far west as Kuching,
|
||
<collectingCountry id="4F80A084FFEDFFA3BFACF9B6FB46F981" box="[1162,1258,1629,1655]" name="Malaysia" pageId="16" pageNumber="17">Sarawak</collectingCountry>
|
||
to Tawau Hills,
|
||
<collectingRegion id="F5532EF6FFEDFFA3BBB1F96AFF77F96D" box="[151,219,1665,1691]" country="Malaysia" name="Sabah" pageId="16" pageNumber="17">Sabah</collectingRegion>
|
||
in the east (
|
||
<tableCitation id="7A15D5AFFFEDFFA3BA48F96AFE6BF96D" box="[366,455,1665,1691]" captionStart="TABLE 1" captionStartId="2.[151,239,586,610]" captionTargetBox="[163,1422,662,1957]" captionText="TABLE 1. Scale counts and selected measurements (mm) of the type series of Asthenodipsas borneensis sp. nov. and A. malaccana. Abbreviations are listed in the Materials and methods." httpUri="http://table.plazi.org/id/63E8B09CFFFFFFB1BBB1FDA1FCF7FD71" pageId="16" pageNumber="17" tableUuid="63E8B09CFFFFFFB1BBB1FDA1FCF7FD71">Table 1</tableCitation>
|
||
;
|
||
<figureCitation id="AFACFC91FFEDFFA3BAFEF96AFDB0F96D" box="[472,540,1665,1691]" captionStart="FIGURE 3" captionStartId="14.[151,250,1908,1933]" captionTargetBox="[170,1416,453,1883]" captionTargetId="figure@14.[170,1416,453,1883]" captionTargetPageId="14" captionText="FIGURE 3. Asthenodipsas borneensis sp. nov. from Borneo. A: Specimen from Kuching, Sarawak (LSUDPC 10988). B: Specimen from Sayap, Mount Kinabalu, Sabah (LSUDPC 10982). C – F (LSUDPC 10983–10986): Adults from Poring, Sabah. G & H (LSUDPC 10987 & 10978): Juvenile from Poring, Sabah. (Photographs by Chien C. Lee [A & B], Zhou Hang [C & D], Anton Sorokin [E & F] & Jaroslav Karhánek [G & H])." figureDoi="http://doi.org/10.5281/zenodo.3628952" httpUri="https://zenodo.org/record/3628952/files/figure.png" pageId="16" pageNumber="17">Fig. 3</figureCitation>
|
||
&
|
||
<figureCitation id="AFACFC91FFEDFFA3B963F96BFDF9F96D" box="[581,597,1664,1691]" captionStart="FIGURE 5" captionStartId="18.[151,250,1335,1360]" captionTargetBox="[151,1436,489,1310]" captionTargetId="figure@18.[151,1436,489,1310]" captionTargetPageId="18" captionText="FIGURE 5. Distribution of Asthenodipsas borneensis sp. nov. (red) and A. malaccana (blue). Star = type locality." figureDoi="http://doi.org/10.5281/zenodo.3628956" httpUri="https://zenodo.org/record/3628956/files/figure.png" pageId="16" pageNumber="17">5</figureCitation>
|
||
). Other reported localities for this species in East Malaysian Borneo are Crocker Range,
|
||
<collectingRegion id="F5532EF6FFEDFFA3BA69F94EFE3FF949" box="[335,403,1701,1727]" country="Malaysia" name="Sabah" pageId="16" pageNumber="17">Sabah</collectingRegion>
|
||
(
|
||
<bibRefCitation id="53069DE5FFEDFFA3BA85F94EFD8CF936" author="Das, I." box="[419,544,1701,1728]" pageId="16" pageNumber="17" pagination="3 - 11" refId="ref12982" refString="Das, I. (2006 a) Crocker Range National Park, Sabah, as a refuge for Borneo's montane herpetofauna. Amphibian and Reptile Conservation, 4 (1), 3 - 11." type="journal article" year="2006">Das 2006a</bibRefCitation>
|
||
), Bongon,
|
||
<collectingRegion id="F5532EF6FFEDFFA3B9BDF94EFD73F949" box="[667,735,1701,1727]" country="Malaysia" name="Sabah" pageId="16" pageNumber="17">Sabah</collectingRegion>
|
||
and Mount Dulit,
|
||
<collectingCountry id="4F80A084FFEDFFA3B897F94EFBBDF949" box="[945,1041,1701,1727]" name="Malaysia" pageId="16" pageNumber="17">Sarawak</collectingCountry>
|
||
(De Roij 1917;
|
||
<bibRefCitation id="53069DE5FFEDFFA3BFECF94EFADDF949" author="Stuebing, R. B." box="[1226,1393,1701,1727]" pageId="16" pageNumber="17" pagination="323 - 362" refId="ref15665" refString="Stuebing, R. B. (1991) A checklist of the snakes of Borneo. Raffles Bulletin of Zoology, 39 (2), 323 - 362." type="journal article" year="1991">Stuebing 1991</bibRefCitation>
|
||
). In Kalimantan, it has been reported from Samarinda (De Roij 1917) and Bulungan Research Forest, Malinau (
|
||
<bibRefCitation id="53069DE5FFEDFFA3BE1CF922FF78F8F1" author="Iskandar, D. T." pageId="16" pageNumber="17" refId="ref14700" refString="Iskandar, D. T. (2004) The Amphibians and Reptiles of Malinau Region, Bulungan Research Forest, East Kalimantan: Annotated checklist with notes on ecological preferences of the species and local utilization. Center for International Forestry Research, Jakarta, 27 pp. [Sheil, D. & Wan, M. (Ed.)] https: // doi. org / 10.17528 / cifor / 001503" type="book" year="2004">Iskandar 2004</bibRefCitation>
|
||
). The species is expected to be more wide ranging in Borneo in the appropriate forest habitat.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7F8DB39FFFEDFFA2BBE1F8FAFCBDFF0D" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="etymology">
|
||
<paragraph id="3728E014FFEDFFA3BBE1F8FAFAAFF885" blockId="16.[151,1437,152,2015]" pageId="16" pageNumber="17">
|
||
<emphasis id="05E33C06FFEDFFA3BBE1F8FAFEE2F8DD" bold="true" box="[199,334,1809,1835]" pageId="16" pageNumber="17">Etymology.</emphasis>
|
||
The specific epithet
|
||
<taxonomicName id="F0979B97FFEDFFA3B91FF8FAFD1EF8DD" authorityName="Quah & Grismer & Lim & Anuar & Chan" authorityYear="2020" box="[569,690,1809,1835]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="borneensis">
|
||
<emphasis id="05E33C06FFEDFFA3B91FF8FAFD1EF8DD" box="[569,690,1809,1835]" italics="true" pageId="16" pageNumber="17">borneensis</emphasis>
|
||
</taxonomicName>
|
||
is in reference to its restriction to the island of Borneo. The suffix
|
||
<emphasis id="05E33C06FFEDFFA3BBB1F8DDFF63F8B9" box="[151,207,1846,1871]" italics="true" pageId="16" pageNumber="17">ensis</emphasis>
|
||
is a Latin derivation meaning “from” or “inhabiting.” It renders the specific epithet an adjective that must be in grammatical accord with the gender of
|
||
<taxonomicName id="F0979B97FFEDFFA3B94AF8B2FCA2F885" authorityName="Peters" authorityYear="1864" box="[620,782,1881,1907]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
|
||
<emphasis id="05E33C06FFEDFFA3B94AF8B2FCA2F885" box="[620,782,1881,1907]" italics="true" pageId="16" pageNumber="17">Asthenodipsas</emphasis>
|
||
</taxonomicName>
|
||
that is feminine (Frank Tillack
|
||
<emphasis id="05E33C06FFEDFFA3BF56F8B1FB00F885" box="[1136,1196,1881,1907]" italics="true" pageId="16" pageNumber="17">in litt</emphasis>
|
||
. 2019).
|
||
</paragraph>
|
||
<paragraph id="3728E014FFEDFFA2BBE1F896FCBDFF0D" blockId="16.[151,1437,152,2015]" lastBlockId="17.[151,1437,152,251]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
|
||
<emphasis id="05E33C06FFEDFFA3BBE1F896FE2AF861" bold="true" box="[199,390,1916,1943]" pageId="16" pageNumber="17">Natural History</emphasis>
|
||
.
|
||
<taxonomicName id="F0979B97FFEDFFA3BAB2F896FD1AF861" authority="Quah & Grismer & Lim & Anuar & Chan, 2020" authorityName="Quah & Grismer & Lim & Anuar & Chan" authorityYear="2020" box="[404,694,1917,1943]" class="Reptilia" family="Pareatidae" genus="Asthenodipsas" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="borneensis" status="sp. nov.">
|
||
<emphasis id="05E33C06FFEDFFA3BAB2F896FD1AF861" box="[404,694,1917,1943]" italics="true" pageId="16" pageNumber="17">Asthenodipsas borneensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1ED0817DFFEDFFA3B99BF896FCB4F861" box="[701,792,1917,1943]" pageId="16" pageNumber="17" rank="species">
|
||
<emphasis id="05E33C06FFEDFFA3B99BF896FCBEF861" bold="true" box="[701,786,1917,1943]" pageId="16" pageNumber="17">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
has been recorded from lowland rainforests and montane regions up to elevations of
|
||
<quantity id="F06F4DF1FFEDFFA3BAF2F84AFD80F84D" box="[468,556,1953,1979]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="17" unit="m" value="1000.0">1000 m</quantity>
|
||
(
|
||
<bibRefCitation id="53069DE5FFEDFFA3B91BF84AFC9EF84D" author="Malkmus, R. & Manthey, U. & Vogel, G. & Hoffmann, P. & Kosuch, J." box="[573,818,1953,1979]" pageId="16" pageNumber="17" refId="ref15123" refString="Malkmus, R., Manthey, U., Vogel, G., Hoffmann, P. & Kosuch, J. (2002) Amphibians and Reptiles of Mount Kinabalu (North Borneo). A. R. G. Ganter Verlag, Ruggel, 424 pp." type="book" year="2002">
|
||
Malkmus
|
||
<emphasis id="05E33C06FFEDFFA3B994F849FD4AF84D" box="[690,742,1953,1979]" italics="true" pageId="16" pageNumber="17">et al</emphasis>
|
||
. 2002
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="53069DE5FFEDFFA3B866F84AFB82F84D" author="Stuebing, R. B. & Inger, R. F. & Lardner, B." box="[832,1070,1953,1979]" pageId="16" pageNumber="17" refId="ref15734" refString="Stuebing, R. B., Inger, R. F. & Lardner, B. (2014) A field guide to the snakes of Borneo. 2 nd Edition. Natural History Publications (Borneo), Kota Kinabalu, 310 pp." type="book" year="2014">
|
||
Stuebing
|
||
<emphasis id="05E33C06FFEDFFA3B88BF849FC4DF84D" box="[941,993,1953,1979]" italics="true" pageId="16" pageNumber="17">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
). It is nocturnal and forages for snails on low vegetation at night. During the day it reportedly shelters beneath leaf litter (
|
||
<bibRefCitation id="53069DE5FFEDFFA3BF84F82EFA3DF829" author="Stuebing, R. B. & Inger, R. F. & Lardner, B." box="[1186,1425,1989,2015]" pageId="16" pageNumber="17" refId="ref15734" refString="Stuebing, R. B., Inger, R. F. & Lardner, B. (2014) A field guide to the snakes of Borneo. 2 nd Edition. Natural History Publications (Borneo), Kota Kinabalu, 310 pp." type="book" year="2014">
|
||
Stuebing
|
||
<emphasis id="05E33C06FFEDFFA3BE36F82DFAE8F829" box="[1296,1348,1989,2015]" italics="true" pageId="16" pageNumber="17">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
). The referred juvenile specimens MCZ R-178183 and FMNH248950/ SP 04130 were collected during the month of September but other juveniles have been photographed in the months of May (LSUDPC 10977) and November (
|
||
<figureCitation id="AFACFC91FFECFFA2BE49FF56FF5EFF0D" captionStart="FIGURE 3" captionStartId="14.[151,250,1908,1933]" captionTargetBox="[170,1416,453,1883]" captionTargetId="figure@14.[170,1416,453,1883]" captionTargetPageId="14" captionText="FIGURE 3. Asthenodipsas borneensis sp. nov. from Borneo. A: Specimen from Kuching, Sarawak (LSUDPC 10988). B: Specimen from Sayap, Mount Kinabalu, Sabah (LSUDPC 10982). C – F (LSUDPC 10983–10986): Adults from Poring, Sabah. G & H (LSUDPC 10987 & 10978): Juvenile from Poring, Sabah. (Photographs by Chien C. Lee [A & B], Zhou Hang [C & D], Anton Sorokin [E & F] & Jaroslav Karhánek [G & H])." figureDoi="http://doi.org/10.5281/zenodo.3628952" httpUri="https://zenodo.org/record/3628952/files/figure.png" pageId="17" pageNumber="18">Fig. 3G & H</figureCitation>
|
||
) indicating breeding may take place year round.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</subSection>
|
||
</treatment>
|
||
</document> |