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<document id="C282C2DD8475607D6EA8D76E23AC623E" ID="10.11646/zootaxa.4205.6.1" ID-CLB-Dataset="37562" ID-DOI="10.5281/zenodo.200269" ID-GBIF-Dataset="6c81cd0d-aeb6-4839-a588-bbdc59eac67f" ID-ISSN="1175-5326" ID-Zenodo-Dep="200269" ID-ZooBank="20C47092-7D67-4E65-ADE0-A73ED91DBB18" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1481526798141" checkinUser="plazi" docAuthor="Lohaj, Roman, Lakota, Ján, Quéinnec, Eric, Pavićević, Dragan &amp; Čeplík, Dávid" docDate="2016" docId="03C08791FFAAFF9F14990B27FEF0FE4A" docLanguage="en" docName="zootaxa.4205.6.1.pdf" docOrigin="Zootaxa 4205 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Adriaphaenops pretneri Scheibel 1935" docType="treatment" docVersion="8" lastPageNumber="515" masterDocId="FFF9FFE9FFA7FF91140E0E19FFF1FFBE" masterDocTitle="Studies on Adriaphaenops Noesske with the description of five new species from the Dinarides (Coleoptera: Carabidae: Trechini)" masterLastPageNumber="531" masterPageNumber="501" pageNumber="514" updateTime="1698433673803" updateUser="ExternalLinkService">
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<mods:title id="CFEE634893D4D9F959A25031441E0585">Studies on Adriaphaenops Noesske with the description of five new species from the Dinarides (Coleoptera: Carabidae: Trechini)</mods:title>
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<mods:namePart id="0C3C5E2DE8EA1A32ADD3D3ABE2577A43">Lohaj, Roman</mods:namePart>
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<mods:namePart id="7ECCC56BDB26C9E6805941FD8BCCF865">Lakota, Ján</mods:namePart>
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<treatment id="03C08791FFAAFF9F14990B27FEF0FE4A" ID-DOI="http://doi.org/10.5281/zenodo.6085111" ID-GBIF-Taxon="125104580" ID-Zenodo-Dep="6085111" LSID="urn:lsid:plazi:treatment:03C08791FFAAFF9F14990B27FEF0FE4A" httpUri="http://treatment.plazi.org/id/03C08791FFAAFF9F14990B27FEF0FE4A" lastPageId="14" lastPageNumber="515" pageId="13" pageNumber="514">
<subSubSection id="C373650CFFAAFF9C14990B27FD7DFAE7" box="[151,652,1342,1369]" pageId="13" pageNumber="514" type="nomenclature">
<paragraph id="8BD63687FFAAFF9C14990B27FD7DFAE7" blockId="13.[151,652,1342,1402]" box="[151,652,1342,1369]" pageId="13" pageNumber="514">
<heading id="D09E81EBFFAAFF9C14990B27FD7DFAE7" bold="true" box="[151,652,1342,1369]" fontSize="11" level="1" pageId="13" pageNumber="514" reason="1">
<emphasis id="B91DEA95FFAAFF9C14990B27FD7DFAE7" bold="true" box="[151,652,1342,1369]" pageId="13" pageNumber="514">
<emphasis id="B91DEA95FFAAFF9C14990B27FF5CFAE6" bold="true" box="[151,173,1342,1368]" italics="true" pageId="13" pageNumber="514">2.</emphasis>
<taxonomicName id="4C694D04FFAAFF9C14BA0B27FD7DFAE7" ID-CoL="64TN8" authority="Scheibel, 1935" authorityName="Scheibel" authorityYear="1935" box="[180,652,1342,1369]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="pretneri">
<emphasis id="B91DEA95FFAAFF9C14BA0B27FE25FAE6" bold="true" box="[180,468,1342,1368]" italics="true" pageId="13" pageNumber="514">Adriaphaenops pretneri</emphasis>
<bibRefCitation id="EFF84B76FFAAFF9C15D50B26FD7DFAE7" author="Scheibel" box="[475,652,1343,1369]" pageId="13" pageNumber="514" refString="Scheibel, O. (1935) Ein neuer Adriaphaenops und ein neuer Pterostichus aus Jugoslawien. Coleopterologische Rundschau, 21, 34 - 38." type="journal article" year="1935">Scheibel, 1935</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C373650CFFAAFF9C14990B78FE79FA60" pageId="13" pageNumber="514" type="description">
<paragraph id="8BD63687FFAAFF9C14990B78FED1FAC4" blockId="13.[151,652,1342,1402]" box="[151,288,1377,1402]" pageId="13" pageNumber="514">
(
<figureCitation id="13522A02FFAAFF9C14910B78FF1DFAC4" box="[159,236,1377,1402]" captionStart="FIGURES 1 4" captionStartId="5.[151,264,1952,1975]" captionTargetBox="[277,1300,210,1919]" captionTargetId="figure@5.[264,1323,195,1931]" captionTargetPageId="5" captionText="FIGURES 1 4. Adriaphaenops spp., head and pronotum, 1. A. pretneri, 2. A. jasminkoi sp. nov., 3. A. perreaui, 4. A. kevser." httpUri="https://zenodo.org/record/272639/files/figure.png" pageId="13" pageNumber="514">Figs. 1</figureCitation>
,
<figureCitation id="13522A02FFAAFF9C14F40B7BFEE6FAC4" box="[250,279,1378,1402]" captionStart="FIGURES 12 15" captionStartId="8.[151,264,1975,1998]" captionTargetBox="[287,1293,206,1937]" captionTargetId="figure@8.[276,1308,193,1953]" captionTargetPageId="8" captionText="FIGURES 12 15. Adriaphaenops spp., habitus: 12. A. antroherponomimus, Holotype male, 13. A. pretneri, Holotype female, 14. A. stirni, Lectotype male, 15. A. zupcense zupcense Topotype male." httpUri="https://zenodo.org/record/200273/files/figure.png" pageId="13" pageNumber="514">13</figureCitation>
)
</paragraph>
<paragraph id="8BD63687FFAAFF9C14990BB3FE79FA60" blockId="13.[151,1436,1450,1502]" pageId="13" pageNumber="514">
<materialsCitation id="3B013CDAFFAAFF9C14990BB3FE79FA60" ID-GBIF-Occurrence="1416605855" country="Bosnia and Herzegovina" location="Windhohle" municipality="Zavala" pageId="13" pageNumber="514" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="EFF84B76FFAAFF9C14990BB3FEC4FA7E" author="Scheibel" box="[151,309,1450,1472]" pageId="13" pageNumber="514" refString="Scheibel, O. (1935) Ein neuer Adriaphaenops und ein neuer Pterostichus aus Jugoslawien. Coleopterologische Rundschau, 21, 34 - 38." type="journal article" year="1935">Scheibel, 1935</bibRefCitation>
:35 (
<taxonomicName id="4C694D04FFAAFF9C15690BB3FDF3FA7E" box="[359,514,1450,1472]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="genus">
<emphasis id="B91DEA95FFAAFF9C15690BB3FDF3FA7E" box="[359,514,1450,1472]" italics="true" pageId="13" pageNumber="514">Adriaphaenops</emphasis>
</taxonomicName>
),
<typeStatus id="54D28825FFAAFF9C16120BB2FDBFFA7E" box="[540,590,1451,1472]" pageId="13" pageNumber="514">Type</typeStatus>
locality:
<location id="8EB6605CFFAAFF9C16B70BB3FCD9FA7E" LSID="urn:lsid:plazi:treatment:03C08791FFAAFF9F14990B27FEF0FE4A:8EB6605CFFAAFF9C16B70BB3FCD9FA7E" box="[697,808,1450,1472]" country="Bosnia and Herzegovina" municipality="Zavala" name="Windhohle" pageId="13" pageNumber="514">Windhöhle</location>
bei
<collectingMunicipality id="6BB2ACFDFFAAFF9C17510BB3FC57FA7E" box="[863,934,1450,1472]" pageId="13" pageNumber="514">Zavala</collectingMunicipality>
,
<collectingCountry id="F37E7617FFAAFF9C17BB0BB3FBCDFA7E" box="[949,1084,1450,1472]" name="Bosnia and Herzegovina" pageId="13" pageNumber="514">Herzegowina</collectingCountry>
[=Popovo polje, Ravno, Zavala, Vjetrenica pećina].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C373650CFFAAFF9C14990812FB30F931" pageId="13" pageNumber="514" type="materials_examined">
<paragraph id="8BD63687FFAAFF9C14990812FB30F931" blockId="13.[151,1437,1547,2004]" pageId="13" pageNumber="514">
<emphasis id="B91DEA95FFAAFF9C14990812FE70F99A" bold="true" box="[151,385,1547,1572]" pageId="13" pageNumber="514">Material examined.</emphasis>
Two females: Holotype female labelled: “Zavala
<date id="FFD71047FFAAFF9C17BF0815FBD2F99D" box="[945,1059,1547,1571]" pageId="13" pageNumber="514" value="1929-08">VIII 1929</date>
” (white label, hw)/“
<taxonomicName id="4C694D04FFAAFF9C110A0812FA91F99A" box="[1284,1376,1547,1572]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="pretneri">Pretneri</taxonomicName>
Typ! Scheibel” (white label, hw)/“Type” (red label, handwritten) (NMB), one female labelled: “Žira jama Turkovići” (white label, hw)/”Popovo polje
<date id="FFD71047FFAAFF9C1628084AFD41F9D2" box="[550,688,1619,1644]" pageId="13" pageNumber="514" value="1970-07-15">15.VII.1970</date>
” (white label, hw)/“leg. Bole Hercegov” (white label, hw,p)/ “Laboulbeniales n. 1096 Walter Rossi” (green label, hw,p)/“
<taxonomicName id="4C694D04FFAAFF9C173A086EFC7EF92E" box="[820,911,1655,1680]" class="Insecta" family="Carabidae" genus="Adriaphaenops" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="514" phylum="Arthropoda" rank="species" species="pretneri">pretneri</taxonomicName>
det. Pretner 1970” (NMP).
</paragraph>
</subSubSection>
<subSubSection id="C373650CFFAAFF9F14C90882FEF5FF6B" lastPageId="14" lastPageNumber="515" pageId="13" pageNumber="514" type="diagnosis">
<paragraph id="8BD63687FFAAFF9C14C90882FD57F811" blockId="13.[151,1437,1547,2004]" pageId="13" pageNumber="514">
<emphasis id="B91DEA95FFAAFF9C14C90882FEB0F90A" bold="true" box="[199,321,1691,1716]" pageId="13" pageNumber="514">Diagnosis.</emphasis>
L:
<quantity id="4C919B62FFAAFF9C156B0882FE00F90D" box="[357,497,1691,1716]" metricMagnitude="-3" metricUnit="m" metricValue="4.825" metricValueMax="5.0" metricValueMin="4.65" pageId="13" pageNumber="514" unit="mm" value="4.825" valueMax="5.0" valueMin="4.65">4.655 mm</quantity>
, TL:
<quantity id="4C919B62FFAAFF9C16250882FD3DF90D" box="[555,716,1691,1716]" metricMagnitude="-3" metricUnit="m" metricValue="4.425" metricValueMax="4.55" metricValueMin="4.3" pageId="13" pageNumber="514" unit="mm" value="4.425" valueMax="4.55" valueMin="4.3">4.34.55 mm</quantity>
. Head large, rounded (index HL/HW 1.25), frontal furrows moderately impressed, reaching the first third of head length, with two pairs of long supraorbital setae; neck constriction distinct; genae evidently curved. Clypeus with four pairs of setae, labrum with three pairs of setae, outer pairs longer. Antennae long (AL
<quantity id="4C919B62FFAAFF9C165C091EFD4AF8A1" box="[594,699,1799,1823]" metricMagnitude="-3" metricUnit="m" metricValue="3.28" pageId="13" pageNumber="514" unit="mm" value="3.28">3.28 mm</quantity>
). Pronotum elongate (index PL/PW 1.25), slightly narrower than head, with maximum width before the anterior quarter, basal part as wide as anterior, anterior and posterior angles small and acute. Elytra elongate oval, broader, distinctly longer than wide (index EL/EW 1.74), with maximum width at middle; elytral surface covered with sparse, short and erected pubescence; striae absent. Site of stria 3 with three (two discal and one preapical) setae.
</paragraph>
<paragraph id="8BD63687FFAAFF9C14C909A2FE86F86A" blockId="13.[151,1437,1547,2004]" box="[199,375,1979,2004]" pageId="13" pageNumber="514">Male unknown.</paragraph>
<paragraph id="8BD63687FFA9FF9F14C90E8EFEF5FF6B" blockId="14.[151,1436,151,501]" pageId="14" pageNumber="515">Female genitalia: examination not possible, in both female specimens available to the authors, missed or damaged.</paragraph>
</subSubSection>
<subSubSection id="C373650CFFA9FF9F14C90EC6FEF0FE4A" pageId="14" pageNumber="515" type="distribution">
<paragraph id="8BD63687FFA9FF9F14C90EC6FEF0FE4A" blockId="14.[151,1436,151,501]" pageId="14" pageNumber="515">
<emphasis id="B91DEA95FFA9FF9F14C90EC6FE91FF46" bold="true" box="[199,352,223,248]" pageId="14" pageNumber="515">Distribution.</emphasis>
<collectingCountry id="F37E7617FFA9FF9F15670EC6FE46FF46" box="[361,439,223,248]" name="Bosnia and Herzegovina" pageId="14" pageNumber="515">Bosnia</collectingCountry>
&amp; Hercegovina, Popovo polje, Vjetrenica and Žira caves. Vjetrenica cave is one of the best explored caves in the Balkan Peninsula and it is the
<typeStatus id="54D28825FFA9FF9F17300F1CFC9FFEA3" box="[830,878,261,285]" pageId="14" pageNumber="515">type</typeStatus>
locality also for other subterranean
<taxonomicName id="4C694D04FFA9FF9F11140F1DFA69FEA3" box="[1306,1432,260,285]" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
, namely
<taxonomicName id="4C694D04FFA9FF9F14F60F3EFD9BFEFE" authority="Muller, 1911" authorityName="Muller" authorityYear="1911" box="[248,618,295,320]" class="Insecta" family="Leiodidae" genus="Hadesia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="vasiceki">
<emphasis id="B91DEA95FFA9FF9F14F60F3EFE4EFEFE" box="[248,447,295,320]" italics="true" pageId="14" pageNumber="515">Hadesia vasiceki</emphasis>
Müller, 1911
</taxonomicName>
,
<taxonomicName id="4C694D04FFA9FF9F16730F3EFB29FEFE" authority="Muller, 1911" authorityName="Muller" authorityYear="1911" box="[637,1240,295,320]" class="Insecta" family="Leiodidae" genus="Anthroherpon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="subSpecies" species="apfelbecki" subSpecies="apfelbecki">
<emphasis id="B91DEA95FFA9FF9F16730F3EFBEFFEFE" box="[637,1054,295,320]" italics="true" pageId="14" pageNumber="515">Anthroherpon apfelbecki apfelbecki</emphasis>
(Müller, 1911)
</taxonomicName>
and
<taxonomicName id="4C694D04FFA9FF9F11110F3EFDC9FEDA" authority="Moravec &amp; Mlejnek, 2002" authorityName="Moravec &amp; Mlejnek" authorityYear="2002" class="Insecta" family="Leiodidae" genus="Nauticiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="stygivaga">
<emphasis id="B91DEA95FFA9FF9F11110F3EFEF2FEDA" italics="true" pageId="14" pageNumber="515">Nauticiella stygivaga</emphasis>
Moravec &amp; Mlejnek, 2002
</taxonomicName>
(
<taxonomicName id="4C694D04FFA9FF9F16460F55FCA8FEDB" authority="Leptodirini" authorityName="Leptodirini" box="[584,857,332,357]" class="Insecta" family="Cholevidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="family">Cholevidae, Leptodirini</taxonomicName>
) as well as
<taxonomicName id="4C694D04FFA9FF9F17EE0F55FF3EFE36" authority="Absolon, 1913" authorityName="Absolon" authorityYear="1913" class="Insecta" family="Carabidae" genus="Scotoplanetes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="arenstorffianus">
<emphasis id="B91DEA95FFA9FF9F17EE0F55FADFFEDA" box="[992,1326,332,357]" italics="true" pageId="14" pageNumber="515">Scotoplanetes arenstorffianus</emphasis>
Absolon, 1913
</taxonomicName>
(
<taxonomicName id="4C694D04FFA9FF9F14E00F76FE95FE36" box="[238,356,367,392]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
,
<taxonomicName id="4C694D04FFA9FF9F158E0F76FE12FE36" box="[384,483,367,392]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="tribe" tribe="Trechini">Trechini</taxonomicName>
).
<taxonomicName id="4C694D04FFA9FF9F16080F76FBDDFE36" authority="Winkler, 1925" authorityName="Winkler" authorityYear="1925" box="[518,1068,367,392]" class="Insecta" family="Staphylinidae" genus="Troglamaurops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="ganglbaueri">
<emphasis id="B91DEA95FFA9FF9F16080F76FCA0FE36" box="[518,849,367,392]" italics="true" pageId="14" pageNumber="515">Troglamaurops ganglbaueri</emphasis>
(Winkler, 1925)
</taxonomicName>
(
<taxonomicName id="4C694D04FFA9FF9F10420F76FB1BFE36" box="[1100,1258,367,392]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
,
<taxonomicName id="4C694D04FFA9FF9F11080F76FA61FE36" box="[1286,1424,367,392]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="subFamily" subFamily="Pselaphinae">Pselaphinae</taxonomicName>
),
<taxonomicName id="4C694D04FFA9FF9F14990F8CFC78FE12" authority="Apfelbeck, 1919" authorityName="Apfelbeck" authorityYear="1919" box="[151,905,404,429]" class="Insecta" family="Leiodidae" genus="Speonesiotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="subSpecies" species="naretinus" subGenus="Speonesiotes" subSpecies="latitarsis">
<emphasis id="B91DEA95FFA9FF9F14990F8CFD43FE13" box="[151,690,404,429]" italics="true" pageId="14" pageNumber="515">Speonesiotes (Speonesiotes) naretinus latitarsis</emphasis>
(Apfelbeck, 1919)
</taxonomicName>
(
<taxonomicName id="4C694D04FFA9FF9F17940F8DFB6FFE13" authority="Leptodirini" authorityName="Leptodirini" box="[922,1182,404,429]" class="Insecta" family="Leiodidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="family">Leiodidae, Leptodirini</taxonomicName>
),
<taxonomicName id="4C694D04FFA9FF9F10BD0F8DFE44FE6E" authority="Obenberger, 1917" authorityName="Obenberger" authorityYear="1917" class="Insecta" family="Carabidae" genus="Neotrechus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="subSpecies" species="suturalis" subSpecies="otiosus">
<emphasis id="B91DEA95FFA9FF9F10BD0F8DFF17FE6E" italics="true" pageId="14" pageNumber="515">Neotrechus suturalis otiosus</emphasis>
Obenberger, 1917
</taxonomicName>
and
<taxonomicName id="4C694D04FFA9FF9F15E20FAEFC63FE6E" authority="Miller, 1861" authorityName="Miller" authorityYear="1861" box="[492,914,439,464]" class="Insecta" family="Carabidae" genus="Neotrechus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="species" species="dalmatinus">
<emphasis id="B91DEA95FFA9FF9F15E20FAEFD1FFE6E" box="[492,750,439,464]" italics="true" pageId="14" pageNumber="515">Neotrechus dalmatinus</emphasis>
(Miller, 1861)
</taxonomicName>
(
<taxonomicName id="4C694D04FFA9FF9F17AF0FAEFBE6FE6E" box="[929,1047,439,464]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
,
<taxonomicName id="4C694D04FFA9FF9F102C0FAEFB73FE6E" box="[1058,1154,439,464]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="515" phylum="Arthropoda" rank="tribe" tribe="Trechini">Trechini</taxonomicName>
) were also reported from this cave.
</paragraph>
</subSubSection>
</treatment>
</document>