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<document id="EFD18DEBAA0265F7376B0355759C7217" ID-DOI="10.1515/aemnp-2017-0068" ID-ISSN="0374-1036" ID-Zenodo-Dep="5315588" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630122827440" checkinUser="marcus" docAuthor="Kaspřák, David, Borkent, Christopher J. &amp; Wahab, Rodzay Abdul" docDate="2017" docId="03BA878E5263FFB3FED4FEABCE38A374" docLanguage="en" docName="ActaEntMusNatPra.2017.57.1.195-203.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 57 (1)" docSource="https://www.aemnp.eu/acta-entomologica/volume-57-1/1673/leptomorphus-sevciki-sp-nov-a-remarkable-new-wasp-mimicking-fungus-gnat-from-brunei-diptera-mycetophilidae.html" docStyle="DocumentStyle:60E99616ED1F2F5EE637DA5E5309A4D2.7:ActaEntMusNatPra.2017.journal_article" docStyleId="60E99616ED1F2F5EE637DA5E5309A4D2" docStyleName="ActaEntMusNatPra.2017.journal_article" docStyleVersion="7" docTitle="Leptomorphus sevciki Kasprak &amp; Borkent 2017, sp. nov." docType="treatment" docVersion="3" lastPageNumber="200" masterDocId="FF83FFF65261FFB6FFB2FFBACD0BA667" masterDocTitle="Leptomorphus sevciki sp. nov., a remarkable new wasp-mimicking fungus gnat from Brunei (Diptera: Mycetophilidae)" masterLastPageNumber="203" masterPageNumber="195" pageNumber="197" updateTime="1699161169930" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="94262C58FD8D623011B6850CE78D85B8">Leptomorphus sevciki sp. nov., a remarkable new wasp-mimicking fungus gnat from Brunei (Diptera: Mycetophilidae)</mods:title>
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<mods:name id="806CF14D4D04FB7ADE5D9E3277CD3707" type="personal">
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<mods:namePart id="64E1939662759999E47C0EE5BA2EB783">Kaspřák, David</mods:namePart>
<mods:affiliation id="35010C0354135C5275401FA0635B7CE0">) &amp;) Department of Biology and Ecology &amp; Institute of Environmental Technologies, Faculty of Science, University of Ostrava, Chittussiho 10, CZ- 710 00 Ostrava, Czech Republic; e-mail: davidkasprak @ gmail. com</mods:affiliation>
</mods:name>
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<mods:role id="0E7A4C3FFF43DC0F810B479E62CDAA65">
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</mods:role>
<mods:namePart id="E063DECADC08C04AD94E3A01101C08AC">Borkent, Christopher J.</mods:namePart>
<mods:affiliation id="6E52BE720E6B302364216472D6BE8C6F">) &amp;) California State Collection of Arthropods, California Department of Food and Agriculture, 3294 Meadowview Rd. Sacramento, CA 95831, USA; e-mail: chris. borkent @ gmail. com</mods:affiliation>
</mods:name>
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<mods:namePart id="A7A2629E378BCEDAC4336513D93246D8">Wahab, Rodzay Abdul</mods:namePart>
<mods:affiliation id="B0990E0EEE6A7E8FF9F3EF04DC02C6A0">) &amp;) Universiti Brunei Darussalam, Institute for Biodiversity and Environmental Research, Tungku Highway, BE 1410, Brunei Darussalam; e-mails: rodzay. wahab @ ubd. edu. bn</mods:affiliation>
</mods:name>
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<mods:title id="9A71FC38C1AC406EBD23F872BF548C49">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="A93AB406E23DCC1BFDA37F1CA67A42DA">2017</mods:date>
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<mods:url id="4CD9128FA3BB4D76E69F4568DAD791C0">https://www.aemnp.eu/acta-entomologica/volume-57-1/1673/leptomorphus-sevciki-sp-nov-a-remarkable-new-wasp-mimicking-fungus-gnat-from-brunei-diptera-mycetophilidae.html</mods:url>
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<taxonomicName id="4C134D1B5263FFB4FED4FEABCE49A74E" ID-CoL="9HJF2" authority="Kasprak &amp; Borkent" authorityName="Kasprak &amp; Borkent" authorityYear="2017" box="[358,834,273,297]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="sevciki" status="sp. nov.">
<emphasis id="B967EA8A5263FFB4FED4FEABCF5CA74E" bold="true" box="[358,599,273,297]" italics="true" pageId="2" pageNumber="197">Leptomorphus sevciki</emphasis>
Kaspřák &amp; Borkent
</taxonomicName>
<taxonomicNameLabel id="A25457F15263FFB4FCFBFEA8CE95A74E" box="[841,926,274,297]" pageId="2" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C30965135263FFB4FF25FE85C96DA44F" pageId="2" pageNumber="197" type="materials_examined">
<paragraph id="8BAC36985263FFB4FF25FE85CCF1A795" blockId="2.[151,1128,314,552]" pageId="2" pageNumber="197">
<emphasis id="B967EA8A5263FFB4FF25FE85CC17A735" bold="true" box="[151,284,319,338]" pageId="2" pageNumber="197">Type material.</emphasis>
<materialsCitation id="3B7B3CC55263FFB4FE93FE85CCE3A7EF" collectingDate="2014-01-14" collectionCode="UBD" collectorName="J. Sevcik &amp; D. Kasprak &amp; Specimen" country="Brunei" county="Ulu" latitude="4.5520835" location="Kuala Belalong Field Studies Centre" longLatPrecision="1" longitude="115.15745" municipality="National Park" pageId="2" pageNumber="197" specimenCount="1" stateProvince="Temburong" typeStatus="holotype">
<typeStatus id="54A8883A5263FFB4FE93FE85CC7FA737" box="[289,372,319,338]" pageId="2" pageNumber="197" type="holotype">HOLOTYPE</typeStatus>
: J,
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<collectingCountry id="F30476085263FFB4FE2BFE85CCE4A735" box="[409,495,319,338]" name="Brunei" pageId="2" pageNumber="197">BRUNEI</collectingCountry>
:
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<collectingRegion id="49D7F87A5263FFB4FDAFFE85CF74A735" box="[541,639,319,338]" country="Brunei" name="Temburong" pageId="2" pageNumber="197">Temburong</collectingRegion>
<collectingMunicipality id="6BC8ACE25263FFB4FD37FE85CFF7A735" box="[645,764,319,338]" pageId="2" pageNumber="197">National Park</collectingMunicipality>
,
<location id="8ECC60435263FFB4FCB7FE85C934A735" LSID="urn:lsid:plazi:treatment:03BA878E5263FFB3FED4FEABCE38A374:8ECC60435263FFB4FCB7FE85C934A735" box="[773,1087,319,338]" country="Brunei" county="Ulu" latitude="4.5520835" longLatPrecision="1" longitude="115.15745" municipality="National Park" name="Kuala Belalong Field Studies Centre" pageId="2" pageNumber="197" stateProvince="Temburong">Kuala Belalong Field Studies Centre</location>
, primary forest,
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,
<date id="FFAD10585263FFB4FE4BFEE0CF40A70A" box="[505,587,346,365]" pageId="2" pageNumber="197" value="2014-01-14">
<collectingDate id="EFE9E9B05263FFB4FE4BFEE0CF40A70A" box="[505,587,346,365]" pageId="2" pageNumber="197" value="2014-01-14">14.i.2014</collectingDate>
</date>
,
<collectorName id="26E6534E5263FFB4FDE1FEE0CFABA70A" box="[595,672,346,365]" pageId="2" pageNumber="197">J. Ševčík</collectorName>
and
<collectorName id="26E6534E5263FFB4FD7AFEE0CE23A70A" box="[712,808,346,365]" pageId="2" pageNumber="197">D. Kaspřák</collectorName>
leg., ex pupa (coll.
<collectionCode id="ED02AE5D5263FFB4FC7DFEE0CEF5A70A" box="[975,1022,346,365]" pageId="2" pageNumber="197">UBD</collectionCode>
, in ethanol).
<collectorName id="26E6534E5263FFB4FF25FECFCDE0A7EF" box="[151,235,373,392]" pageId="2" pageNumber="197">Specimen</collectorName>
voucher number JSS25a-LS1
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.
<materialsCitation id="3B7B3CC55263FFB4FE5DFECFCC9CA7DA" collectingDate="2014-01-08" collectingDateMax="2014-01-20" collectingDateMin="2014-01-08" collectionCode="UBD" country="Brunei" county="Ulu" latitude="4.5520835" location="Kuala Belalong Field Studies Centre" longLatPrecision="1" longitude="115.15745" municipality="National Park" pageId="2" pageNumber="197" specimenCode="LS3" specimenCount="1" specimenCount-female="1" stateProvince="Temburong" typeStatus="paratype">
<typeStatus id="54A8883A5263FFB4FE5DFECFCF4EA7E1" box="[495,581,373,392]" pageId="2" pageNumber="197" type="paratype">PARATYPES</typeStatus>
:
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<collectingCountry id="F30476085263FFB4FDFDFECFCFAEA7EF" box="[591,677,373,392]" name="Brunei" pageId="2" pageNumber="197">BRUNEI</collectingCountry>
:
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<collectingCounty id="62CD4E145263FFB4FD1FFECFCFC6A7EF" box="[685,717,373,392]" pageId="2" pageNumber="197">Ulu</collectingCounty>
<collectingRegion id="49D7F87A5263FFB4FD63FECFCE38A7EF" box="[721,819,373,392]" country="Brunei" name="Temburong" pageId="2" pageNumber="197">Temburong</collectingRegion>
<collectingMunicipality id="6BC8ACE25263FFB4FC84FECFCEA7A7EF" box="[822,940,373,392]" pageId="2" pageNumber="197">National Park</collectingMunicipality>
,
<location id="8ECC60435263FFB4FC01FECFCC1DA7C5" LSID="urn:lsid:plazi:treatment:03BA878E5263FFB3FED4FEABCE38A374:8ECC60435263FFB4FC01FECFCC1DA7C5" country="Brunei" county="Ulu" latitude="4.5520835" longLatPrecision="1" longitude="115.15745" municipality="National Park" name="Kuala Belalong Field Studies Centre" pageId="2" pageNumber="197" stateProvince="Temburong">Kuala Belalong Field Studies Centre</location>
, primary forest,
<geoCoordinate id="EE27505F5263FFB4FE12FE35CF01A7C5" box="[416,522,399,418]" degrees="4" direction="north" minutes="33" orientation="latitude" pageId="2" pageNumber="197" precision="1" seconds="07.5" value="4.5520835">4°3307.5″N</geoCoordinate>
<geoCoordinate id="EE27505F5263FFB4FDBDFE35CF85A7C5" box="[527,654,399,418]" degrees="115" direction="east" minutes="09" orientation="longitude" pageId="2" pageNumber="197" precision="1" seconds="26.8" value="115.15745">115°0926.8″E</geoCoordinate>
,
<date id="FFAD10585263FFB4FD24FE35CFF5A7C5" box="[662,766,399,418]" pageId="2" pageNumber="197" value="2014-01-08" valueMax="2014-01-20" valueMin="2014-01-08">
<collectingDate id="EFE9E9B05263FFB4FD24FE35CFF5A7C5" box="[662,766,399,418]" pageId="2" pageNumber="197" value="2014-01-08" valueMax="2014-01-20" valueMin="2014-01-08">820.i.2014</collectingDate>
</date>
, ex pupa, J (JSS25c-LS2) and
<specimenCount id="9D15FD115263FFB4FBB6FE33C91FA7C5" box="[1028,1044,393,418]" pageId="2" pageNumber="197" type="female"></specimenCount>
(JSS25e- -
<specimenCode id="DBB59EE35263FFB4FF2FFE10CDC8A7DA" box="[157,195,426,445]" collectionCode="LS" country="United Kingdom" name="Linnean Society of London" pageId="2" pageNumber="197">LS3</specimenCode>
) (coll.
<collectionCode id="ED02AE5D5263FFB4FEB3FE10CC3BA7DA" box="[257,304,426,445]" pageId="2" pageNumber="197">UBD</collectionCode>
, in ethanol)
</materialsCitation>
;
<materialsCitation id="3B7B3CC55263FFB4FE12FE10CF2FA7BF" collectionCode="JSL-UOC" country="Brunei" pageId="2" pageNumber="197" specimenCode="LS4, LS5, LS6, LS7" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
3 JJ (pinned, JSS25-
<specimenCode id="DBB59EE35263FFB4FDECFE10CF88A7DA" box="[606,643,426,445]" collectionCode="LS" country="United Kingdom" name="Linnean Society of London" pageId="2" pageNumber="197">LS4</specimenCode>
, JSS25-
<specimenCode id="DBB59EE35263FFB4FD78FE10CFE5A7DA" box="[714,750,426,445]" collectionCode="LS" country="United Kingdom" name="Linnean Society of London" pageId="2" pageNumber="197">LS5</specimenCode>
, JSS25-
<specimenCode id="DBB59EE35263FFB4FC87FE10CE51A7DA" box="[821,858,426,445]" collectionCode="LS" country="United Kingdom" name="Linnean Society of London" pageId="2" pageNumber="197">LS6</specimenCode>
),
<specimenCount id="9D15FD115263FFB4FCDBFE10CE81A7DB" box="[873,906,419,445]" pageId="2" pageNumber="197" type="female">1 ♀</specimenCount>
(pinned, JSS25-
<specimenCode id="DBB59EE35263FFB4FBAAFE10C937A7DA" box="[1048,1084,426,445]" collectionCode="LS" country="United Kingdom" name="Linnean Society of London" pageId="2" pageNumber="197">LS7</specimenCode>
) and
<specimenCount id="9D15FD115263FFB4FF25FE7FCDBCA7B0" box="[151,183,446,472]" pageId="2" pageNumber="197" type="female">1 ♀</specimenCount>
(in ethanol, JSS25f-LS8) (coll.
<collectionCode id="ED02AE5D5263FFB4FE77FE7FCF15A7BF" box="[453,542,453,472]" pageId="2" pageNumber="197">JSL-UOC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B7B3CC55263FFB4FD9EFE7FC94BA7BF" box="[556,1088,446,472]" collectionCode="CSCA" country="Brunei" pageId="2" pageNumber="197" specimenCode="LS9, LS10" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
1 J (JSS25-
<specimenCode id="DBB59EE35263FFB4FD26FE7FCFB3A7BF" box="[660,696,453,472]" collectionCode="LS" country="United Kingdom" name="Linnean Society of London" pageId="2" pageNumber="197">LS9</specimenCode>
) and
<specimenCount id="9D15FD115263FFB4FD55FE7FCE0DA7B0" box="[743,774,446,472]" pageId="2" pageNumber="197" type="female">1 ♀</specimenCount>
(JSS25-
<specimenCode id="DBB59EE35263FFB4FCFCFE7FCE75A7BF" box="[846,894,453,472]" collectionCode="LS" country="United Kingdom" name="Linnean Society of London" pageId="2" pageNumber="197">LS10</specimenCode>
) (coll.
<collectionCode id="ED02AE5D5263FFB4FC08FE7FCEFFA7BF" box="[954,1012,453,472]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" lsid="urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="2" pageNumber="197" type="Museum">CSCA</collectionCode>
, pinned)
</materialsCitation>
;
<materialsCitation id="3B7B3CC55263FFB4FBFAFE7FCCFDA795" collectionCode="NMPC" country="Brunei" pageId="2" pageNumber="197" specimenCode="LS11" specimenCount="1" typeStatus="paratype">
and 1 J (JSS25-
<specimenCode id="DBB59EE35263FFB4FEB3FE65CC25A795" box="[257,302,479,498]" collectionCode="LS" country="United Kingdom" name="Linnean Society of London" pageId="2" pageNumber="197">LS11</specimenCode>
) (coll.
<collectionCode id="ED02AE5D5263FFB4FED9FE65CCA1A795" box="[363,426,479,498]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="197">NMPC</collectionCode>
, pinned)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAC36985263FFB4FF25FE40C96DA44F" blockId="2.[151,1128,314,552]" pageId="2" pageNumber="197">
<emphasis id="B967EA8A5263FFB4FF25FE40CF95A46A" bold="true" box="[151,670,506,525]" pageId="2" pageNumber="197">
Immature material not included in
<typeStatus id="54A8883A5263FFB4FE6AFE40CCF4A46A" box="[472,511,506,525]" pageId="2" pageNumber="197">type</typeStatus>
series:
<collectingCountry id="F30476085263FFB4FDF6FE40CF91A46A" box="[580,666,506,525]" name="Brunei" pageId="2" pageNumber="197">BRUNEI</collectingCountry>
:
</emphasis>
Ulu
<collectingRegion id="49D7F87A5263FFB4FD7AFE40CE21A46A" box="[712,810,506,525]" country="Brunei" name="Temburong" pageId="2" pageNumber="197">Temburong</collectingRegion>
National Park, Kuala Belalong Field Studies Centre, primary forest,
<geoCoordinate id="EE27505F5263FFB4FE11FDAFCF06A44F" box="[419,525,533,552]" degrees="4" direction="north" minutes="33" orientation="latitude" pageId="2" pageNumber="197" precision="1" seconds="07.5" value="4.5520835">4°3307.5″N</geoCoordinate>
<geoCoordinate id="EE27505F5263FFB4FDA1FDAFCF99A44F" box="[531,658,533,552]" degrees="115" direction="east" minutes="09" orientation="longitude" pageId="2" pageNumber="197" precision="1" seconds="26.8" value="115.15745">115°0926.8″E</geoCoordinate>
,
<date id="FFAD10585263FFB4FD29FDAFCE08A44F" box="[667,771,533,552]" pageId="2" pageNumber="197" value="2014-01-08" valueMax="2014-01-20" valueMin="2014-01-08">820.i.2014</date>
, 2 L and 2 P (coll.
<collectionCode id="ED02AE5D5263FFB4FC11FDAFCEF0A44F" box="[931,1019,533,552]" pageId="2" pageNumber="197">JSL-UOC</collectionCode>
, in ethanol).
</paragraph>
</subSubSection>
<subSubSection id="C30965135263FFB3FF25FDFACDE4A594" lastPageId="5" lastPageNumber="200" pageId="2" pageNumber="197" type="diagnosis">
<paragraph id="8BAC36985263FFB5FF25FDFACFB8A774" blockId="2.[151,1127,576,727]" lastBlockId="3.[113,1090,219,819]" lastPageId="3" lastPageNumber="198" pageId="2" pageNumber="197">
<emphasis id="B967EA8A5263FFB4FF25FDFACC07A430" bold="true" box="[151,268,576,599]" pageId="2" pageNumber="197">Diagnosis.</emphasis>
The only extant species of
<taxonomicName id="4C134D1B5263FFB4FD98FDFACFCFA430" authorityName="Curtis" authorityYear="1831" box="[554,708,576,599]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="B967EA8A5263FFB4FD98FDFACFCFA430" box="[554,708,576,599]" italics="true" pageId="2" pageNumber="197">Leptomorphus</emphasis>
</taxonomicName>
with the following combination of characters: male antennae covered in long setae (unique in the genus); yellow V-shaped marking on the dorsal side of dark brown scutum; katepisternum yellow and brown; metakatepisternum partially dark brown; abdominal segments swelling towards the apex of terminalia. Sternite 9 with medioventral lobe and a pair of lateroventral lobes bearing four dark megasetae apically.
<emphasis id="B967EA8A5262FFB5FFC3FF66CC4AA694" bold="true" box="[113,321,219,243]" pageId="3" pageNumber="198">
Description.
<emphasis id="B967EA8A5262FFB5FEB0FF61CC31A694" bold="true" box="[258,314,219,243]" italics="true" pageId="3" pageNumber="198">Male</emphasis>
.
</emphasis>
Body length
<quantity id="4CEB9B7D5262FFB5FE7CFF66CF3BA694" box="[462,560,220,243]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="3" pageNumber="198" unit="mm" value="14.3">14.3 mm</quantity>
. General colouration dark brown and dark yellow, yellow-orange and brown striped abdomen (
<figureCitation id="13282A1D5262FFB5FDF8FF46CFAEA774" box="[586,677,252,275]" captionStart="Figs 14" captionStartId="2.[151,188,1477,1496]" captionTargetBox="[151,1126,777,1456]" captionTargetId="figure-251@2.[151,1126,774,1456]" captionTargetPageId="2" captionText="Figs 14. Leptomorphus sevciki sp. nov.: 1 male (Photo: J. Ševčík); 2 female (Photo: J. Ševčík); 3 male, habitus; 4 female, habitus." figureDoi="http://doi.org/10.5281/zenodo.5315592" httpUri="https://zenodo.org/record/5315592/files/figure.png" pageId="3" pageNumber="198">Fig. 1, 3</figureCitation>
).
</paragraph>
<caption id="DF6C66105263FFB4FF25FA7FCC82A395" ID-DOI="http://doi.org/10.5281/zenodo.5315592" ID-Zenodo-Dep="5315592" httpUri="https://zenodo.org/record/5315592/files/figure.png" pageId="2" pageNumber="197" startId="2.[151,188,1477,1496]" targetBox="[151,1126,777,1456]" targetPageId="2">
<paragraph id="8BAC36985263FFB4FF25FA7FCC82A395" blockId="2.[151,1127,1477,1522]" pageId="2" pageNumber="197">
Figs 14.
<taxonomicName id="4C134D1B5263FFB4FF42FA7FCCA6A3BF" authority="Kaspřák &amp; Borkent &amp; Wahab, 2017" authorityName="Kaspřák &amp; Borkent &amp; Wahab" authorityYear="2017" box="[240,429,1477,1496]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="sevciki" status="sp. nov.">
<emphasis id="B967EA8A5263FFB4FF42FA7FCCA6A3BF" box="[240,429,1477,1496]" italics="true" pageId="2" pageNumber="197">Leptomorphus sevciki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25457F15263FFB4FE06FA7FCCF7A3BF" box="[436,508,1477,1496]" pageId="2" pageNumber="197" rank="species">sp. nov.</taxonomicNameLabel>
: 1 male (Photo: J. Ševčík); 2 female (Photo: J. Ševčík); 3 male, habitus; 4 female, habitus.
</paragraph>
</caption>
<paragraph id="8BAC36985262FFB5FF22FEA6CC4AA7D4" blockId="3.[113,1090,219,819]" pageId="3" pageNumber="198">
<emphasis id="B967EA8A5262FFB5FF22FEA6CDC1A754" box="[144,202,284,307]" italics="true" pageId="3" pageNumber="198">Head</emphasis>
. Dark brown, light brown laterally with many long setae. Ocelli yellow, in a straight line. Face yellow. Flagellomeres dark brown, densely covered with long, fine, dark brown setae (
<figureCitation id="13282A1D5262FFB5FF0AFEE6CDF0A714" box="[184,251,348,371]" captionStart="Figs 58. 57" captionStartId="3.[112,149,1541,1560]" captionTargetBox="[115,1083,871,1529]" captionTargetId="figure-253@3.[115,1083,871,1529]" captionTargetPageId="3" captionText="Figs 58. 57 Leptomorphus sevciki sp. nov., male: 5 antennae; 6 thorax, V-shape mark; 7 wing; 8 female, genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5315594" httpUri="https://zenodo.org/record/5315594/files/figure.png" pageId="3" pageNumber="198">Fig. 5</figureCitation>
). Basal part of first flagellomere yellow. Scape and pedicel yellow, setae on at least the anterior surface. Face and clypeus covered with many long setae. Clypeus and palpus light brown.
</paragraph>
<paragraph id="8BAC36985262FFB5FF22FE06CF61A4D4" blockId="3.[113,1090,219,819]" pageId="3" pageNumber="198">
<emphasis id="B967EA8A5262FFB5FF22FE06CDEFA7B4" box="[144,228,444,467]" italics="true" pageId="3" pageNumber="198">Thorax.</emphasis>
Mostly dark brown. Scutum dark brown with dorsal yellow V-shape marking, covered with fine hairs, several long light setae along margin (
<figureCitation id="13282A1D5262FFB5FD4EFE66CE37A794" box="[764,828,476,499]" captionStart="Figs 58. 57" captionStartId="3.[112,149,1541,1560]" captionTargetBox="[115,1083,871,1529]" captionTargetId="figure-253@3.[115,1083,871,1529]" captionTargetPageId="3" captionText="Figs 58. 57 Leptomorphus sevciki sp. nov., male: 5 antennae; 6 thorax, V-shape mark; 7 wing; 8 female, genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5315594" httpUri="https://zenodo.org/record/5315594/files/figure.png" pageId="3" pageNumber="198">Fig. 6</figureCitation>
). Scutellum light brown, with many short setae. Antepronotum, proepisternum, anepisternum, anepimeron and laterotergite dark brown. Proepisternum covered with light setae. Paratergite light to dark brown. Laterotergite with several long setae. Anepisternum and anepimeron bare. Katepisternum anterior dark brown and posterior yellow, bare. Metakatepisternum partially dark brown, bare. Mediotergite yellow, bare. Margin of anterior and posterior spiracles yellow with yellow trichia. Haltere: stem yellow, knob dark brown.
</paragraph>
<paragraph id="8BAC36985262FFB2FF22FD06CCF0A7F4" blockId="3.[113,1090,219,819]" lastBlockId="4.[151,1127,220,531]" lastPageId="4" lastPageNumber="199" pageId="3" pageNumber="198">
<emphasis id="B967EA8A5262FFB5FF22FD06CDCCA4B4" box="[144,199,700,723]" italics="true" pageId="3" pageNumber="198">Wing</emphasis>
(
<figureCitation id="13282A1D5262FFB5FF61FD06CC19A4B4" box="[211,274,700,723]" captionStart="Figs 58. 57" captionStartId="3.[112,149,1541,1560]" captionTargetBox="[115,1083,871,1529]" captionTargetId="figure-253@3.[115,1083,871,1529]" captionTargetPageId="3" captionText="Figs 58. 57 Leptomorphus sevciki sp. nov., male: 5 antennae; 6 thorax, V-shape mark; 7 wing; 8 female, genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5315594" httpUri="https://zenodo.org/record/5315594/files/figure.png" pageId="3" pageNumber="198">Fig. 7</figureCitation>
). Hyaline, membrane densely covered with irregularly arranged macrotrichia. Cell among base of R 1, crossvein r-m and M bare, sector between veins M and CuA bare except apically. Apical shading dark brown but fading towards apex and posterior margin (pale brown apical wing shading not reaching to wing tip), running from anterior to posterior wing margin, beginning halfway along R 5. Medial macula small, only around Rs. Costa ending at tip of R 5. Sc short, ending in costa beyond base of r-m. Vein sc-r ending beyond Rs. Vein R 4 absent. Vein R 5 slightly concave for entire length. Crossvein r-m relatively long (twice as long as radial sector). Apex of crossvein r-m lighter than base. Apices of M and CuA veins clearly reaching wing margin. Point of furcation of CuA and M 4 well before crossvein r-m. CuP thick, slightly shorter than A
</paragraph>
<caption id="DF6C66105262FFB5FFC2F9BFCC24A054" ID-DOI="http://doi.org/10.5281/zenodo.5315594" ID-Zenodo-Dep="5315594" httpUri="https://zenodo.org/record/5315594/files/figure.png" pageId="3" pageNumber="198" startId="3.[112,149,1541,1560]" targetBox="[115,1083,871,1529]" targetPageId="3">
<paragraph id="8BAC36985262FFB5FFC2F9BFCC24A054" blockId="3.[112,1089,1541,1587]" pageId="3" pageNumber="198">
Figs 58. 57
<taxonomicName id="4C134D1B5262FFB5FF48F9BFCCBFA07F" authority="Kaspřák &amp; Borkent &amp; Wahab, 2017" authorityName="Kaspřák &amp; Borkent &amp; Wahab" authorityYear="2017" box="[250,436,1541,1560]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="sevciki" status="sp. nov.">
<emphasis id="B967EA8A5262FFB5FF48F9BFCCBFA07F" box="[250,436,1541,1560]" italics="true" pageId="3" pageNumber="198">Leptomorphus sevciki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25457F15262FFB5FE0BF9BFCCF6A07F" box="[441,509,1541,1560]" pageId="3" pageNumber="198" rank="species">sp. nov.</taxonomicNameLabel>
, male: 5 antennae; 6 thorax, V-shape mark; 7 wing; 8 female, genitalia, lateral view.
</paragraph>
</caption>
<paragraph id="8BAC36985265FFB2FE49FE36CF08A7FD" blockId="4.[151,1127,220,531]" box="[507,515,396,410]" pageId="4" pageNumber="199">1</paragraph>
<paragraph id="8BAC36985265FFB2FDB1FEC6CF29A7F4" blockId="4.[151,1127,220,531]" box="[515,546,380,403]" pageId="4" pageNumber="199">. A</paragraph>
<paragraph id="8BAC36985265FFB2FD90FE36CF21A7FD" blockId="4.[151,1127,220,531]" box="[546,554,396,410]" pageId="4" pageNumber="199">1</paragraph>
<paragraph id="8BAC36985265FFB2FD83FEC6CF8CA7F4" blockId="4.[151,1127,220,531]" box="[561,647,380,403]" pageId="4" pageNumber="199">distinct.</paragraph>
<paragraph id="8BAC36985265FFB3FF07FE26CE97A754" blockId="4.[151,1127,220,531]" lastBlockId="5.[113,1089,220,1299]" lastPageId="5" lastPageNumber="200" pageId="4" pageNumber="199">
<emphasis id="B967EA8A5265FFB2FF07FE26CDE5A7D4" box="[181,238,412,435]" italics="true" pageId="4" pageNumber="199">Legs.</emphasis>
Coxae principally yellow, forecoxa dark brown and hind coxa with anterior dark brown mark. All trochanters brown. Femora yellow, dark long setae laterally, hind femur brownish, dense setulae and dark brown mark beginning one third of length of hind femur and reaching beyond midpoint. Fore tibia yellow with basal brown mark, dense row of anteroventral bristles absent. Mid tibia entirely dark brown with irregular rows of short dark bristles. Hind tibia yellow with basal dark brown mark longer than fore tibia basal mark. Tarsi dark brown. First and second hind tarsomeres yellowish to light brown with dark brown apex.
</paragraph>
<caption id="DF6C66105265FFB2FF24FA68C959A052" ID-DOI="http://doi.org/10.5281/zenodo.5315598" ID-Zenodo-Dep="5315598" httpUri="https://zenodo.org/record/5315598/files/figure.png" pageId="4" pageNumber="199" startId="4.[150,187,1490,1509]" targetBox="[303,970,561,1479]" targetPageId="4">
<paragraph id="8BAC36985265FFB2FF24FA68C959A052" blockId="4.[150,1129,1490,1589]" pageId="4" pageNumber="199">
Figs 912
<taxonomicName id="4C134D1B5265FFB2FF4FFA68CCBEA382" authority="Kaspřák &amp; Borkent &amp; Wahab, 2017" authorityName="Kaspřák &amp; Borkent &amp; Wahab" authorityYear="2017" box="[253,437,1490,1509]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="sevciki" status="sp. nov.">
<emphasis id="B967EA8A5265FFB2FF4FFA68CCBEA382" box="[253,437,1490,1509]" italics="true" pageId="4" pageNumber="199">Leptomorphus sevciki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25457F15265FFB2FE0BFA68CCF7A382" box="[441,508,1490,1509]" pageId="4" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
, male, terminalia: 9 ventral part, internal view; 10 ventral part, external view; 11 dorsal part, internal view; 12 dorsal part, external view. Abbreviations: cerc = cercus; ej ap = ejaculatory apodeme; epi = epiproct; gst = gonostylus; hyp = hypoproct; mvl st 9 = medioventral lobe of sternite 9; lvl st 9 = lateroventral lobe of sternite 9; par = paramere; par ap = parameral apodeme; st 9 = sternite 9; tg 9 = tergite 9.
</paragraph>
</caption>
<paragraph id="8BAC36985264FFB3FF22FE86CCEAA7F4" blockId="5.[113,1089,220,1299]" pageId="5" pageNumber="200">
<emphasis id="B967EA8A5264FFB3FF22FE86CDF7A734" box="[144,252,316,339]" italics="true" pageId="5" pageNumber="200">Abdomen.</emphasis>
Tergites principally dark brown, tergites 16 with anterior third light brown. Tergites 78 dark brown. Sternites brown. Posterior 4 segments of abdomen noticeably swollen relative to other segments (
<figureCitation id="13282A1D5264FFB3FE21FEC6CCDFA7F4" box="[403,468,380,403]" captionStart="Figs 14" captionStartId="2.[151,188,1477,1496]" captionTargetBox="[151,1126,777,1456]" captionTargetId="figure-251@2.[151,1126,774,1456]" captionTargetPageId="2" captionText="Figs 14. Leptomorphus sevciki sp. nov.: 1 male (Photo: J. Ševčík); 2 female (Photo: J. Ševčík); 3 male, habitus; 4 female, habitus." figureDoi="http://doi.org/10.5281/zenodo.5315592" httpUri="https://zenodo.org/record/5315592/files/figure.png" pageId="5" pageNumber="200">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BAC36985264FFB3FF22FE26C94AA4B4" blockId="5.[113,1089,220,1299]" pageId="5" pageNumber="200">
<emphasis id="B967EA8A5264FFB3FF22FE26CC0FA7D4" box="[144,260,412,435]" italics="true" pageId="5" pageNumber="200">Terminalia</emphasis>
(Figs 912). Light brown, apex of gonostyli and medioventral lobe of sternite 9 darker. Sternite 9 sclerotized, medially covered with long setae, with medioventral lobe and a pair of lateroventral lobes, dark brown suture mediobasally. Medioventral lobe bare, as long as sternite 9, covering cerci, with two rounded processes bent ventrally and apical dark blunt bristles. Lateroventral lobe with bumpy margin, with several long setae and four dark apical megasetae. Tergite 9 sclerotized, densely covered with short setae and V-shaped indentation on the apex, bare transparent tapering part of base. Gonostyli bare, except lateral margin with long setae, dark blunt bristles on medial margin of apex, lateral rounded process and medial hooked process. Apex of gonostylus dark, tapering towards apex. Cerci and hypoproct covered with short trichia and long apical setae. Parameres strongly hooked at apex.
</paragraph>
<paragraph id="8BAC36985264FFB3FF22FD61C91BA534" blockId="5.[113,1089,220,1299]" pageId="5" pageNumber="200">
<emphasis id="B967EA8A5264FFB3FF22FD61CDE8A494" bold="true" box="[144,227,731,755]" italics="true" pageId="5" pageNumber="200">Female</emphasis>
(
<figureCitation id="13282A1D5264FFB3FF42FD66CC5BA494" box="[240,336,732,755]" captionStart="Figs 14" captionStartId="2.[151,188,1477,1496]" captionTargetBox="[151,1126,777,1456]" captionTargetId="figure-251@2.[151,1126,774,1456]" captionTargetPageId="2" captionText="Figs 14. Leptomorphus sevciki sp. nov.: 1 male (Photo: J. Ševčík); 2 female (Photo: J. Ševčík); 3 male, habitus; 4 female, habitus." figureDoi="http://doi.org/10.5281/zenodo.5315592" httpUri="https://zenodo.org/record/5315592/files/figure.png" pageId="5" pageNumber="200">Figs 2, 4</figureCitation>
). Similar to male with following differences: antennal setae short, hind femur lighter with brown mark only anteriorly, genitalia lighter. Tergite 9 yellow, with long setae. Tergite 10 yellow, row of several long and shorter setae on posterior margin. Cerci brown, setose. Gonopore 9 brown. Gonocoxite 8 posteriorly brown and setose (
<figureCitation id="13282A1D5264FFB3FC73FC86C909A534" box="[961,1026,828,851]" captionStart="Figs 58. 57" captionStartId="3.[112,149,1541,1560]" captionTargetBox="[115,1083,871,1529]" captionTargetId="figure-253@3.[115,1083,871,1529]" captionTargetPageId="3" captionText="Figs 58. 57 Leptomorphus sevciki sp. nov., male: 5 antennae; 6 thorax, V-shape mark; 7 wing; 8 female, genitalia, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5315594" httpUri="https://zenodo.org/record/5315594/files/figure.png" pageId="5" pageNumber="200">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8BAC36985264FFB3FFC3FCE6CDE4A594" blockId="5.[113,1089,220,1299]" pageId="5" pageNumber="200">
<emphasis id="B967EA8A5264FFB3FFC3FCE6CC28A514" bold="true" box="[113,291,860,883]" pageId="5" pageNumber="200">DNA sequences.</emphasis>
<materialsCitation id="3B7B3CC55264FFB3FE95FCE6CED7A5D4" accessionNumber="MF045798, MF045800, MF045803, MF045805" pageId="5" pageNumber="200" specimenCount="3" specimenCount-male="1" typeStatus="holotype">
DNA sequences taken from the
<typeStatus id="54A8883A5264FFB3FDDEFCE6CFC3A514" box="[620,712,860,883]" pageId="5" pageNumber="200" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D15FD115264FFB3FD45FCE6CE5FA514" box="[759,852,860,883]" pageId="5" pageNumber="200" type="male">one male</specimenCount>
<typeStatus id="54A8883A5264FFB3FCEBFCE6CEB8A514" box="[857,947,860,883]" pageId="5" pageNumber="200" type="paratype">paratype</typeStatus>
are deposited in the GenBank database, with following GB accession numbers: JSS25a-LS1 (
<typeStatus id="54A8883A5264FFB3FC60FCC6C93DA5F4" box="[978,1078,892,915]" pageId="5" pageNumber="200" type="holotype">holotype</typeStatus>
): 12S:
<accessionNumber id="9440AB7B5264FFB3FF19FC26CC2EA5D4" box="[171,293,924,947]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF045798" pageId="5" pageNumber="200">MF045798</accessionNumber>
, 16S:
<accessionNumber id="9440AB7B5264FFB3FED9FC26CCEFA5D4" box="[363,484,924,947]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF045800" pageId="5" pageNumber="200">MF045800</accessionNumber>
, COI:
<accessionNumber id="9440AB7B5264FFB3FD9DFC26CFA2A5D4" box="[559,681,924,947]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF045803" pageId="5" pageNumber="200">MF045803</accessionNumber>
, cytB:
<accessionNumber id="9440AB7B5264FFB3FD4AFC26CE79A5D4" box="[760,882,924,947]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF045805" pageId="5" pageNumber="200">MF045805</accessionNumber>
, 28S: n/a
</materialsCitation>
;
<materialsCitation id="3B7B3CC55264FFB3FC5BFC26CDE7A594" accessionNumber="MF045799, MF045801, MF045804, MF045806, MF045802" collectionCode="LS" pageId="5" pageNumber="200" specimenCode="LS2" specimenCount="1" typeStatus="paratype">
JSS25c-
<specimenCode id="DBB59EE35264FFB3FFC3FC06CD95A5B4" box="[113,158,956,979]" collectionCode="LS" country="United Kingdom" name="Linnean Society of London" pageId="5" pageNumber="200">LS2</specimenCode>
(
<typeStatus id="54A8883A5264FFB3FF19FC06CC05A5B4" box="[171,270,956,979]" pageId="5" pageNumber="200" type="paratype">paratype</typeStatus>
): 12S:
<accessionNumber id="9440AB7B5264FFB3FEE6FC06CCC5A5B4" box="[340,462,956,979]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF045799" pageId="5" pageNumber="200">MF045799</accessionNumber>
, 16S:
<accessionNumber id="9440AB7B5264FFB3FDBFFC06CF8DA5B4" box="[525,646,956,979]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF045801" pageId="5" pageNumber="200">MF045801</accessionNumber>
, COI:
<accessionNumber id="9440AB7B5264FFB3FD79FC06CE4FA5B4" box="[715,836,956,979]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF045804" pageId="5" pageNumber="200">MF045804</accessionNumber>
, cytB:
<accessionNumber id="9440AB7B5264FFB3FC3FFC06C90DA5B4" box="[909,1030,956,979]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF045806" pageId="5" pageNumber="200">MF045806</accessionNumber>
, 28S:
<accessionNumber id="9440AB7B5264FFB3FFC3FC66CDE7A594" box="[113,236,988,1011]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MF045802" pageId="5" pageNumber="200">MF045802</accessionNumber>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30965135264FFB3FFC3FC46CFA9A234" pageId="5" pageNumber="200" type="etymology">
<paragraph id="8BAC36985264FFB3FFC3FC46CFA9A234" blockId="5.[113,1089,220,1299]" pageId="5" pageNumber="200">
<emphasis id="B967EA8A5264FFB3FFC3FC46CDF9A274" bold="true" box="[113,242,1020,1043]" pageId="5" pageNumber="200">Etymology.</emphasis>
This species is named after Jan Ševčík, a specialist on
<taxonomicName id="4C134D1B5264FFB3FCF7FC46CE9EA274" box="[837,917,1020,1043]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Diptera" pageId="5" pageNumber="200" phylum="Tracheophyta" rank="order">Diptera</taxonomicName>
associated with fungi and supervisor of David Kaspřáks Ph.D. study, who was the first to spot the specimens of the new species in Ulu
<collectingRegion id="49D7F87A5264FFB3FE34FB86CF0AA234" box="[390,513,1084,1107]" country="Brunei" name="Temburong" pageId="5" pageNumber="200">Temburong</collectingRegion>
National Park.
</paragraph>
</subSubSection>
<subSubSection id="C30965135264FFB3FFC3FBE6CE38A374" pageId="5" pageNumber="200" type="biology_ecology">
<paragraph id="8BAC36985264FFB3FFC3FBE6CE38A374" blockId="5.[113,1089,220,1299]" pageId="5" pageNumber="200">
<emphasis id="B967EA8A5264FFB3FFC3FBE6CDC6A214" bold="true" box="[113,205,1116,1139]" pageId="5" pageNumber="200">Biology.</emphasis>
The larvae and pupae of the new species were found on the underside of a log over a small brook. The larvae spun a silk sheet over the surface of an unidentified wood encrusting fungus (
<figureCitation id="13282A1D5264FFB3FF79FB26CC17A2D4" box="[203,284,1180,1203]" captionStart="Figs 1315. 13" captionStartId="6.[150,187,949,968]" captionTargetBox="[151,1126,219,930]" captionTargetId="figure-228@6.[148,1129,219,930]" captionTargetPageId="6" captionText="Figs 1315. 13 The larvae of Leptomorphus sevciki sp. nov. on the surface of an unidentified wood encrusting fungus growing on a log in a rainforest in Brunei. 14 The pupae of L. sevciki sp. nov., on the underside of a log in a rainforest in Brunei. 15 The fallen tree over a pathway with hanging pupae in Ulu Temburong National Park in Brunei. (Photos: J. Ševčík)." figureDoi="http://doi.org/10.5281/zenodo.5315600" httpUri="https://zenodo.org/record/5315600/files/figure.png" pageId="5" pageNumber="200">Fig. 13</figureCitation>
, Video: https://youtu.be/89HN2255ZOs). The pupae were hanging from a singly anchored
<quantity id="4CEB9B7D5264FFB3FE91FB06CC7FA2B4" box="[291,372,1212,1235]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="5" pageNumber="200" unit="cm" value="4.0" valueMax="5.0" valueMin="3.0">35 cm</quantity>
long yellowish thread, bearing a drop of brownish liquid (
<figureCitation id="13282A1D5264FFB3FC5EFB06C937A2B4" box="[1004,1084,1212,1235]" captionStart="Figs 1315. 13" captionStartId="6.[150,187,949,968]" captionTargetBox="[151,1126,219,930]" captionTargetId="figure-228@6.[148,1129,219,930]" captionTargetPageId="6" captionText="Figs 1315. 13 The larvae of Leptomorphus sevciki sp. nov. on the surface of an unidentified wood encrusting fungus growing on a log in a rainforest in Brunei. 14 The pupae of L. sevciki sp. nov., on the underside of a log in a rainforest in Brunei. 15 The fallen tree over a pathway with hanging pupae in Ulu Temburong National Park in Brunei. (Photos: J. Ševčík)." figureDoi="http://doi.org/10.5281/zenodo.5315600" httpUri="https://zenodo.org/record/5315600/files/figure.png" pageId="5" pageNumber="200">Fig. 14</figureCitation>
, Video: https://youtu.be/89HN2255ZOs). The behaviour of the flying adults and their colouration were reminiscent of wasps or ichneumonids (Hymenoptera).
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