146 lines
17 KiB
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146 lines
17 KiB
XML
<document id="B4DEE260B31E2C601540DA8ED6608228" ID-DOI="http://dx.doi.org/10.3897/zookeys.542.5975" ID-GBIF-Dataset="b53aa92a-b9ac-43bd-a9b9-f468d62f0bdc" ID-PMC="PMC4714360" ID-Pensoft-Pub="1313-2970-542-1" ID-PubMed="26798285" ID-ZooBank="026407877355425BAB10BF1674510F12" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-542-1" ModsDocOrigin="ZooKeys 542" ModsDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" checkinTime="1451243652205" checkinUser="pensoft" docAuthor="Bistroem, Olof, Nilsson, Anders N. & Bergsten, Johannes" docDate="2015" docId="FC3151A6FACE6075B23D21B555275BD5" docLanguage="en" docName="ZooKeys 542: 1-379" docOrigin="ZooKeys 542" docSource="http://dx.doi.org/10.3897/zookeys.542.5975" docTitle="Laccophilus canthydroides Omer-Cooper 1957" docType="treatment" docVersion="5" lastPageNumber="231" masterDocId="E3118410B56DFF839B32FF89FFCCFFB0" masterDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" masterLastPageNumber="379" masterPageNumber="1" pageNumber="230" updateTime="1701314941532" updateUser="plazi">
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<mods:titleInfo id="708563B4C9532C31B93E0A8041252754">
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<mods:title id="967221B372D8ACF3B6B2BF394809373F">Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
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<mods:namePart id="408D9B30D93A7BC8E1E75E5139334EC1">Bistroem, Olof</mods:namePart>
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<mods:namePart id="8C7856E016ED514EBE9B510048CB05F8">Nilsson, Anders N.</mods:namePart>
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<mods:namePart id="AD67277BE207C08A6AEEB01560999852">Bergsten, Johannes</mods:namePart>
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<mods:typeOfResource id="C0F58AC65680D760B02DCD6F9AD839B8">text</mods:typeOfResource>
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<mods:part id="E78B01A841C607497A004B51AC00E26F">
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<mods:date id="B54A31CB5C77E897950C14D3C754F8E1">2015</mods:date>
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<mods:detail id="596BA400CCFB40E36E94872F56520896" type="volume">
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<mods:number id="B5974F74E015BA6D103F75485A1D4494">542</mods:number>
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<mods:url id="C3CF992132E3EBBF303E0D238B1EE425">http://dx.doi.org/10.3897/zookeys.542.5975</mods:url>
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<mods:classification id="40D59A1075B888197A13985EF031B67E">journal article</mods:classification>
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<mods:identifier id="A5C45556D863689B7BE114FE1FCD9CE8" type="DOI">http://dx.doi.org/10.3897/zookeys.542.5975</mods:identifier>
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<treatment id="FC3151A6FACE6075B23D21B555275BD5" ID-GBIF-Taxon="127862696" LSID="urn:lsid:plazi:treatment:FC3151A6FACE6075B23D21B555275BD5" httpUri="http://treatment.plazi.org/id/FC3151A6FACE6075B23D21B555275BD5" lastPageId="230" lastPageNumber="231" pageId="229" pageNumber="230">
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<subSubSection id="C2E6E1F9F8E4A28A8C8DB3CA045EBEB5" pageId="229" pageNumber="230" type="multiple">
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<paragraph id="717A1B470D92177E400A43E3C2A877E2" pageId="229" pageNumber="230">
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<pageBreakToken id="76B03D3B3873D788D67682848B515285" pageId="229" pageNumber="230" start="start">Taxon</pageBreakToken>
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classification Animalia Coleoptera Dytiscidae
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</paragraph>
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</subSubSection>
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<subSubSection id="93F85E0EE7F53036BC18A1B76AFBF4A4" pageId="229" pageNumber="230" type="nomenclature">
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<paragraph id="0EE0BC44876EBB12A84E6AFDCD7A83A2" pageId="229" pageNumber="230">
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<taxonomicName id="64DF8699DA9569BC9991892EFC777CA3" ID-CoL="3RM5T" authority="Omer-Cooper, 1957" authorityName="Omer-Cooper" authorityYear="1957" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus canthydroides" order="Coleoptera" pageId="229" pageNumber="230" phylum="Arthropoda" rank="species" species="canthydroides">Laccophilus canthydroides Omer-Cooper, 1957</taxonomicName>
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Figs 184-185, 362, 512, 513, 575
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</paragraph>
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</subSubSection>
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<subSubSection id="D77CD76C3912E6CCB194046294C78D97" pageId="230" pageNumber="231" type="reference_group">
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<paragraph id="070FE3E4EA60581491E80B541011003F" pageId="230" pageNumber="231">
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<taxonomicName id="0B6113D0B93CD9C66EE6885BC28B97F9" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus canthydroides" order="Coleoptera" pageId="230" pageNumber="231" phylum="Arthropoda" rank="species" species="canthydroides">
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<pageBreakToken id="FB57A000EA340BD5FA3A0EA82954F2A7" pageId="230" pageNumber="231" start="start">Laccophilus</pageBreakToken>
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canthydroides
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</taxonomicName>
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<bibRefCitation id="4FE030AB0B38C24B37A8870287E86916" author="Omer-Cooper, Joyce" journalOrPublisher="Memoirs of the Entomological Society of Southern Africa" pageId="260" pageNumber="261" pagination="1 - 90" title="Dytiscidae (Coleoptera) collected in the Transvaal (Rhodes University Expedition 1948)." volume="5" year="1957">Omer-Cooper 1957</bibRefCitation>
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: 13, 90 (original description, faunistics);
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<bibRefCitation id="B76E9CBFF82FBC16A1C0D5AB894E5364" author="Omer-Cooper, Joyce" journalOrPublisher="South African Animal Life" pageId="261" pageNumber="262" pagination="59 - 214" title="Chapter 2. Coleoptera: Dytiscidae." volume="11" year="1965">Omer-Cooper 1965</bibRefCitation>
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: 76, 81 (description, faunistics);
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<bibRefCitation id="319896FB717FA4D9ABFC3C57B68788FE" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
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: 241 (catalogue, faunistics);
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<bibRefCitation id="E0A7DA6E9984EB2F246F895BF4507A03" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
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: 210 (catalogue, faunistics).
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</paragraph>
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</subSubSection>
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<subSubSection id="3D723061461B1FFCFE2DB3D986072804" pageId="230" pageNumber="231" type="type locality">
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<paragraph id="FA0AF54613F73456D31A4CEB48CB99C0" pageId="230" pageNumber="231">Type locality.</paragraph>
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<paragraph id="1DEF036338CA4464BDD6D90633ED0664" pageId="230" pageNumber="231">South Africa: Transvaal, Barberton, (trib. of Koop River).</paragraph>
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</subSubSection>
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<subSubSection id="E8E7D82E7142142DB8AF86921BB67D32" pageId="230" pageNumber="231" type="type material">
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<paragraph id="495B095EA7D9A360EEB4F6B2F3850E9E" pageId="230" pageNumber="231">Type material studied</paragraph>
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<paragraph id="F3545EAE31797D0BABD70D430C3CBBE6" pageId="230" pageNumber="231">
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(2 exs.). Holotype: female: "Type / Transvaal Trib. of Koop R. nr Barberton / Brit. Mus. 1957-660 /
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<taxonomicName id="65042558BCF7A849D91EFD100C02B219" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus" order="Coleoptera" pageId="230" pageNumber="231" phylum="Arthropoda" rank="genus">Laccophilus</taxonomicName>
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? sp. n. J. Balfour-Browwne det. /
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<taxonomicName id="CD7447C4FF11EEEA9F7E94BD0B90E2D3" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus canthydroides" order="Coleoptera" pageId="230" pageNumber="231" phylum="Arthropoda" rank="species" species="canthydroides">Laccophilus canthydroides</taxonomicName>
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" (BMNH; habitus in Fig. 512). - Paratype: female: "Paratype / Transvaal, Trib. of Koop R. nr. Barberton 1.12. 1948 /
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<taxonomicName id="FBAEF694C0D59BEA6AFB624CF7F80F02" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus canthydroides" order="Coleoptera" pageId="230" pageNumber="231" phylum="Arthropoda" rank="species" species="canthydroides">Laccophilus canthydroides</taxonomicName>
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/
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<taxonomicName id="6647DF595228FC7208AC23CEA864681E" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus" order="Coleoptera" pageId="230" pageNumber="231" phylum="Arthropoda" rank="genus">Laccophilus</taxonomicName>
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? sp. n. J. Balfour-Browne det." (1 ex. AMGS; according to the original description to be deposited in TMSA).
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</paragraph>
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</subSubSection>
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<subSubSection id="A805496A5F34515328407185F256C7AB" pageId="230" pageNumber="231" type="materials_examined">
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<paragraph id="680924EEC9A5F58E401BCA0DE6F4C03C" pageId="230" pageNumber="231">Additional material studied</paragraph>
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<paragraph id="7FCC6857A15DC818E4B31231CDA1076A" pageId="230" pageNumber="231">
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(56 exs.). Ethiopia: "Bahar Dar 8.10. 1968 Harde" (2 exs. NHMB). - Cameroon: "Emana Obala 16.5. 1970" (1 ex. NHMB). - Zaire: "Parc National Garamba 10.3. 1952 De Saeger" (3 exs. MRAC, 2 exs. NHMB, 1 ex. MZH); "Elisabethville,
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<normalizedToken id="93A586F3F2462D59A34A5B374ECEEC12" originalValue="à">a</normalizedToken>
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la
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<normalizedToken id="98B135694C1A7A4A29092631AE8629D8" originalValue="lumière">lumiere</normalizedToken>
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1957-1958 Seydel" (1 ex. AMGS, 10 exs. MRAC, 2 exs. MZH; habitus in Fig. 513); same but "1. 1956-1. 1957" (1 ex. MRAC); same but "11. 1951-2. 1952" (1 ex. MNHN); same but "11. 1950-6. 1951" (1 ex. MNHN); same but
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<normalizedToken id="6CBD77431C580DFA653781563323BFA4" originalValue="“1953-1955”">"1953-1955"</normalizedToken>
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(1 ex. AMGS, 2 exs. NHMB); "Elisabethville 2. 1940 H.J.
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<normalizedToken id="421486129033061FF6284B1CCB5BD184" originalValue="Brédo”">Bredo"</normalizedToken>
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(2 exs. OLML); "Elisabethville,
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<normalizedToken id="5B256D159D2AB6BB04689CF598D470A0" originalValue="lumière">lumiere</normalizedToken>
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11. 1951-2. 1952 Seydel" (2 exs. IRSNB); "Kivu, Kavimvira (Uvira),
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<normalizedToken id="FC6BCC1A1D6FE1B0071B85DDB0E9C523" originalValue="à">a</normalizedToken>
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la
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<normalizedToken id="5B896C7408367F2051EEC7A103F25359" originalValue="lumière">lumiere</normalizedToken>
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1. 1956 Marlier" (1 ex. NHMB). - Kenya: "Meru distr., Matiri (Mitunguu) mt. 800, 18.10. 1982 Mourglia" (3 exs. NHMB); same but "8.11. 1983" (1 ex. NHMB). - Tanzania: "Tang. Terr. Ukerewe 7. 1933" (3 exs. OLML); "Mufundi Mafinga m. 1900 21.11.-4.12. 1989 leg. R. Mouglia" (1 ex. CSR); "NP Udzungwa, Ifakara, at street from entrance, S08.02.55, E36.43.59, 251 m, trunk under bark 18.7. 2004 Sprecher" (1 ex. NHMB); "Zanzibar Mangapwani rd 13.9. 1955 JOC."(6 exs. AMGS). - Zambia: "Mbesuma Ranch (Isoka) 9-10.12. 2004 Werner & Smrz" (5 exs. NHRS); "Luapula Pr., Lake Bangweulu, Chilubi 11.2. 1982 Selander" (1 ex. MZH). - Malawi: "Nkhata Bay env. rainforest 1.1. 2002 Kantner" (1 ex. NHMB). - South Africa: "Wellington, Witte River 1500 ft." (1 ex. SAMC).
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</paragraph>
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</subSubSection>
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<subSubSection id="DA5944C45DE98DF0D7E868BF3B83EFCB" pageId="230" pageNumber="231" type="diagnosis">
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<paragraph id="0406B3FFBBF60325B06161A369087CE3" pageId="230" pageNumber="231">Diagnosis.</paragraph>
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<paragraph id="213DDB4800560960182C81C935EEC979" pageId="230" pageNumber="231">
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<taxonomicName id="A2BA6362EF0F38C483B7B596BD602DD3" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus canthydroides" order="Coleoptera" pageId="230" pageNumber="231" phylum="Arthropoda" rank="species" species="canthydroides">Laccophilus canthydroides</taxonomicName>
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seems to be closest related to
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<taxonomicName id="50C9E07FB3E2CD55FCFFAD6A4B3D5B88" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus inornatus" order="Coleoptera" pageId="230" pageNumber="231" phylum="Arthropoda" rank="species" species="inornatus">Laccophilus inornatus</taxonomicName>
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- for separation of the two species, vide diagnosis of
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<taxonomicName id="9BA919ACC9BF8288F5DF266EC8577A39" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus inornatus" order="Coleoptera" pageId="230" pageNumber="231" phylum="Arthropoda" rank="species" species="inornatus">Laccophilus inornatus</taxonomicName>
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(p. 241).
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</paragraph>
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</subSubSection>
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<subSubSection id="CDEA8295A637852DD4D756813DC6C286" pageId="230" pageNumber="231" type="description">
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<paragraph id="92EA9A9D048320DF70261E9FAD67B9B7" pageId="230" pageNumber="231">Description.</paragraph>
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<paragraph id="88807F3FE7DE74B926F29A9898299E9C" pageId="230" pageNumber="231">Body length 3.2-3.6 mm, width 1.8-1.9 mm. Head and pronotum extensively pale while elytra often somewhat darker. Often with two, pale spots on base of elytra. Sometimes pale spots absent or reduced to one humeral spot (Figs 512-513).</paragraph>
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<paragraph id="6EDB79D7A954001E68809491FF1013F7" pageId="230" pageNumber="231">Head: Pale ferrugineous, posteriorly slightly darker but darker area anteriorly with vague delimitation. Submat, finely microsculptured; reticulation double but large meshes only slightly stronger than fine meshes. By large, 3-7 fine meshes included in one large mesh. At eyes with irregular cluster of fine, rather indistinct punctures.</paragraph>
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<paragraph id="C289D14BBF132E021FD1CDF0C2CDC0EA" pageId="230" pageNumber="231">Pronotum: Pale ferrugineous, basally in middle often with rather narrow darker area. Rather shiny, although finely microsculptured; reticulation double but large meshes only slightly stronger than fine meshes. By large, 3-7 fine meshes included in one large mesh. Laterally and frontally on pronotum with scattered, fine punctures.</paragraph>
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<paragraph id="B5EFCA5083967325563F05E33CAE5F08" pageId="230" pageNumber="231">Elytra: Dark ferrugineous to ferrugineous, at humeral region and posteriorly often with vague, small, pale spots (Figs 512-513). Rather shiny, although finely microsculptured; reticulation double but in part (laterally and posteriorly) division of meshes in two size-classes indistinct. In general large meshes (when discernible), contain 2-5 fine meshes. Impunctate, except for discal, dorsolateral and lateral rows of fine, somewhat irregular punctures.</paragraph>
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<paragraph id="B85881644F9DAD3B0237DC0C30E2B3DD" pageId="230" pageNumber="231">Ventral aspect: Pale ferrugineous, except ventrites often somewhat darker; ferrugineous. Rather shiny, although with fine reticulation (densely located fine lines). Metacoxal plates with some irregular and indistinct, laterally projecting striae. Basal, abdominal ventrites laterally with a few, distinct striae. Almost impunctate. Apex of prosternal process slender and pointed. Apical ventrite on one side with a minute knob (Fig. 184).</paragraph>
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<paragraph id="1790737443F7E8132F00046B170E3AEA" pageId="230" pageNumber="231">Legs: Pale ferrugineous. Pro- and mesotarsus slightly enlarged, provided with suckers.</paragraph>
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<paragraph id="891DCA456025EDAAEA9B8178F6398515" pageId="230" pageNumber="231">Male genitalia: External, extreme, apex of penis not distinctly extended, nor sharp (Fig. 362).</paragraph>
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<paragraph id="A803C182C2CE4631A6B3D96F5109928A" pageId="230" pageNumber="231">Female: Pro- and mesotarsus slender. Apical ventrite (Fig. 185).</paragraph>
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</subSubSection>
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<subSubSection id="B421B98431098D12F9A27BD5DF57AC14" pageId="230" pageNumber="231" type="distribution">
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<paragraph id="2022CCD5417A3FD8EC90CC2D121980E7" pageId="230" pageNumber="231">Distribution.</paragraph>
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<paragraph id="BAFEFD59C83E6C1C439A86E29E948DE3" pageId="230" pageNumber="231">Ethiopia, Cameroon, Zaire, Kenya, Tanzania, Zambia, Malawi, South Africa (Fig. 575).</paragraph>
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</subSubSection>
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<subSubSection id="82154E3560B8E15B6D1875668DF71340" pageId="230" pageNumber="231" type="collecting circumstances">
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<paragraph id="68D85F683222CBC304F1A6858E331DBF" pageId="230" pageNumber="231">Collecting circumstances.</paragraph>
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<paragraph id="A30C60194BFFDCF519079FE98136683B" pageId="230" pageNumber="231">Largely unknown. Collected in a light trap.</paragraph>
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</subSubSection>
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</treatment>
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</document> |