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<document id="44F100A145A1E757861ACCDA11673283" ID-DOI="10.11646/phytotaxa.347.4.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13707821" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725591235838" checkinUser="felipe" docAuthor="Henderson, Andrew &amp; Dung, Nguyen Quoc" docDate="2018" docId="28428787FFA39767FF733CCBFBC5523D" docLanguage="en" docName="phytotaxa.347.4.1.pdf" docOrigin="Phytotaxa 347 (4)" docSource="http://dx.doi.org/10.11646/phytotaxa.347.4.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Calamus manglaensis Henderson &amp; Nguyen Quoc Dung 2018, sp. nov." docType="treatment" docVersion="3" lastPageNumber="257" masterDocId="D47BFFFFFFA59761FFFB3E71FF815568" masterDocTitle="New species and subspecies of Calamus (Arecaceae) from Vietnam" masterLastPageNumber="262" masterPageNumber="251" pageNumber="257" updateTime="1725602979930" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C4A50355260011B82C897288EB49730D">New species and subspecies of Calamus (Arecaceae) from Vietnam</mods:title>
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<mods:namePart id="D2DD032E1CFB4E6FA1DFEB313DC8DB1F">Henderson, Andrew</mods:namePart>
<mods:affiliation id="7F855575C134152B013C240EE96701F5">Institute of Systematic Botany, The New York Botanical Garden, Bronx, NY 10458 - 5126, U. S. A.</mods:affiliation>
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<mods:namePart id="2E53A113DB40ADE2BF054D85F83D58DD">Dung, Nguyen Quoc</mods:namePart>
<mods:affiliation id="ED40C3C8E163F17F107CE914A807148D">Forest Inventory and Planning Institute, Thanh Tri, Hanoi, Vietnam</mods:affiliation>
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<mods:title id="F59FABA08EA8EFD8D34E12F9F3ABE4AF">Phytotaxa</mods:title>
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<mods:date id="7C8C1CCCB1124A947A033407CA97A1F3">2018</mods:date>
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<treatment id="28428787FFA39767FF733CCBFBC5523D" ID-DOI="http://doi.org/10.5281/zenodo.13708011" ID-Zenodo-Dep="13708011" LSID="urn:lsid:plazi:treatment:28428787FFA39767FF733CCBFBC5523D" httpUri="http://treatment.plazi.org/id/28428787FFA39767FF733CCBFBC5523D" lastPageNumber="257" pageId="6" pageNumber="257">
<subSubSection id="E8F1651AFFA39767FF733CCBFCFF57BD" box="[136,894,698,725]" pageId="6" pageNumber="257" type="nomenclature">
<paragraph id="A0543691FFA39767FF733CCBFCFF57BD" blockId="6.[136,1452,698,831]" box="[136,894,698,725]" pageId="6" pageNumber="257">
<heading id="FB1C81FDFFA39767FF733CCBFCFF57BD" bold="true" box="[136,894,698,725]" fontSize="11" level="1" pageId="6" pageNumber="257" reason="1">
<emphasis id="929FEA83FFA39767FF733CCBFCFF57BD" bold="true" box="[136,894,698,725]" pageId="6" pageNumber="257">
<taxonomicName id="67EB4D12FFA39767FF733CCBFCA157BD" ID-CoL="8WS22" authority="Henderson &amp; Nguyen Quoc Dung" authorityName="Henderson &amp; Nguyen Quoc Dung" authorityYear="2018" box="[136,800,698,725]" class="Liliopsida" family="Arecaceae" genus="Calamus" kingdom="Plantae" order="Arecales" pageId="6" pageNumber="257" phylum="Tracheophyta" rank="species" species="manglaensis" status="sp. nov.">
<emphasis id="929FEA83FFA39767FF733CCBFE0657BD" bold="true" box="[136,391,698,725]" italics="true" pageId="6" pageNumber="257">Calamus manglaensis</emphasis>
Henderson &amp; Nguyen Quoc Dung
</taxonomicName>
<emphasis id="929FEA83FFA39767FCDC3CCAFCFF57BD" bold="true" box="[807,894,699,725]" italics="true" pageId="6" pageNumber="257">
<taxonomicNameLabel id="89AC57F8FFA39767FCDC3CCAFCFF57BD" box="[807,894,699,725]" pageId="6" pageNumber="257" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E8F1651AFFA39767FF733C93FCA15379" pageId="6" pageNumber="257" type="materials_examined">
<paragraph id="A0543691FFA39767FF733C93FE075657" blockId="6.[136,1452,698,831]" pageId="6" pageNumber="257">
Type:—
<collectingCountry id="D8FC7601FFA39767FF2D3C93FEC9579F" box="[214,328,738,759]" name="Vietnam" pageId="6" pageNumber="257">VIETNAM</collectingCountry>
.
<collectingRegion id="622FF873FFA39767FEA93C93FE33579F" box="[338,434,738,759]" country="Vietnam" name="Kon Tum" pageId="6" pageNumber="257">Kon Tum</collectingRegion>
: Kon Plong district, Dak Long commune,
<collectingRegion id="622FF873FFA39767FCB43C93FC50579F" box="[847,977,738,759]" country="Vietnam" name="Ho Chi Minh City" pageId="6" pageNumber="257">Ho Chi Minh</collectingRegion>
Highway, just north of intersection of road from Mang Den to Bato,
<geoCoordinate id="C5DF5056FFA39767FE7E3D77FE565673" box="[389,471,774,795]" degrees="14" direction="north" minutes="41" orientation="latitude" pageId="6" pageNumber="257" precision="925" value="14.683333">14°41N</geoCoordinate>
<geoCoordinate id="C5DF5056FFA39767FE1B3D77FDBF5673" box="[480,574,774,795]" degrees="108" direction="east" minutes="21" orientation="longitude" pageId="6" pageNumber="257" precision="925" value="108.35">108°21E</geoCoordinate>
,
<quantity id="67139B74FFA39767FDB13D77FD195673" box="[586,664,774,795]" metricMagnitude="3" metricUnit="m" metricValue="1.468" pageId="6" pageNumber="257" unit="m" value="1468.0">1468 m</quantity>
,
<date id="D4551051FFA39767FD5F3D77FCCD5673" box="[676,844,774,795]" pageId="6" pageNumber="257" value="2015-10-16">16 October 2015</date>
,
<emphasis id="929FEA83FFA39767FCA33D77FB7C5673" box="[856,1277,774,795]" italics="true" pageId="6" pageNumber="257">
<collectionCode id="C6FAAE54FFA39767FCA33D77FCEB5673" box="[856,874,774,795]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="6" pageNumber="257" type="Herbarium">A</collectionCode>
. Henderson &amp; Nguyen Quoc Dung 4031
</emphasis>
(
<typeStatus id="7F508833FFA39767FAF73D77FAE35673" box="[1292,1378,774,795]" pageId="6" pageNumber="257" type="holotype">holotype</typeStatus>
<collectionCode id="C6FAAE54FFA39767FA923D77FA255673" box="[1385,1444,774,795]" pageId="6" pageNumber="257">VFM</collectionCode>
!,
<typeStatus id="7F508833FFA39767FF463D5BFE835657" box="[189,258,810,831]" pageId="6" pageNumber="257" type="isotype">isotype</typeStatus>
<collectionCode id="C6FAAE54FFA39767FEF23D5BFEB25657" box="[265,307,810,831]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="6" pageNumber="257" type="Herbarium">NY</collectionCode>
!).
<figureCitation id="38D02A14FFA39767FEBC3D5BFE005657" box="[327,385,810,831]" captionStart="FIGURE 4" captionStartId="7.[136,229,1915,1936]" captionTargetBox="[151,1435,187,1887]" captionTargetId="figure-14@7.[151,1436,187,1888]" captionTargetPageId="7" captionText="FIGURE 4. Calamus manglaensis. A. Sheath with few, scattered, shortly recurved spines on a swollen base, not leaving a distal impression in the sheath (Henderson 4031). B. Leaf with irregularly arranged pinnae (Henderson 4031). C. Staminate partial inflorescence (image by M. Nuraliev). D. Infructescence with globose fruits (Henderson 4031)." figureDoi="http://doi.org/10.5281/zenodo.13707829" httpUri="https://zenodo.org/record/13707829/files/figure.png" pageId="6" pageNumber="257">Fig. 4</figureCitation>
.
</paragraph>
<paragraph id="A0543691FFA39767FF733D01FDAD56C5" blockId="6.[136,1452,880,1877]" pageId="6" pageNumber="257">
It differs from all other species of
<taxonomicName id="67EB4D12FFA39767FDFC3D01FDEA56E1" box="[519,619,880,905]" class="Liliopsida" family="Arecaceae" genus="Calamus" kingdom="Plantae" order="Arecales" pageId="6" pageNumber="257" phylum="Tracheophyta" rank="genus">
<emphasis id="929FEA83FFA39767FDFC3D01FDEA56E1" box="[519,619,880,905]" italics="true" pageId="6" pageNumber="257">Calamus</emphasis>
</taxonomicName>
by its pistillate inflorescences branched to 1 order and seeds covered with a non-tanniniferous, slimy sarcotesta.
</paragraph>
<paragraph id="A0543691FFA39767FF463DC9FCA15379" blockId="6.[136,1452,880,1877]" pageId="6" pageNumber="257">
<emphasis id="929FEA83FFA39767FF463DC9FE8456B9" bold="true" box="[189,261,952,977]" pageId="6" pageNumber="257">Stems</emphasis>
clustered, climbing, 20.0 m long, 0.8(0.61.0) cm diameter.
<emphasis id="929FEA83FFA39767FC303DC9FBE456B9" bold="true" box="[971,1125,952,977]" pageId="6" pageNumber="257">Leaf sheaths</emphasis>
tubular, closed opposite the petiole, with a knee below the petiole; leaf sheath spines few, scattered, shortly recurved on a swollen base, not leaving a distal impression in the sheath; ocreas short, membranous, usually non-spiny, with external and internal abscission zone, splitting and falling early; flagella well-developed; petioles 2.8(2.03.5) cm long; rachises 22.2(20.024.5) cm long, the apices a short stub between the distal pair of pinnae;
<emphasis id="929FEA83FFA39767FCB53A39FC1E5109" bold="true" box="[846,927,1096,1121]" pageId="6" pageNumber="257">pinnae</emphasis>
8(79) per side of rachis, irregularly arranged in groups, spreading in different planes, elliptic, without spinules on veins, proximalmost pinnae swept back across the sheath; middle pinnae 18.6(18.019.4) cm long, 1.6(1.21.8) cm wide; distal pair of pinnae joined at their bases for 1.0(0.71.4) cm.
<emphasis id="929FEA83FFA39767FE883AC5FD9D51A5" bold="true" box="[371,540,1204,1229]" pageId="6" pageNumber="257">Inflorescences</emphasis>
diverging from sheath near sheath apex; inflorescences erect to arching, slender, elongate, with recurved spines on rachis and rachis bracts abaxially, without a flagellum-like apex; rachis bracts tubular, narrow, persistent, closely sheathing the rachis, not swollen at the apex; rachillae sessile; staminate inflorescences branched to 2 orders, 88.0 cm long; staminate rachillae
<quantity id="67139B74FFA39767FD0C3B51FCC25051" box="[759,835,1312,1337]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="6" pageNumber="257" unit="cm" value="6.5">6.5 cm</quantity>
long; staminate flowers arranged in alternate, opposite rows; staminate sepals shorter than the petals, cupular, 3-lobed at the apex; stamens 6; filaments uniseriate, inflexed at the apex; pistillate inflorescences branched to 1 order, 110.0 cm long; pistillate rachillae
<quantity id="67139B74FFA39767FB903B19FB3650E9" box="[1131,1207,1384,1409]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" pageId="6" pageNumber="257" unit="cm" value="5.2">5.2 cm</quantity>
long, with dyads of 1 pistillate and 1 neuter flower;
<emphasis id="929FEA83FFA39767FE213BFDFD9A50CD" bold="true" box="[474,539,1420,1445]" pageId="6" pageNumber="257">fruits</emphasis>
globose,
<quantity id="67139B74FFA39767FD733BFDFD6D50CD" box="[648,748,1420,1445]" metricMagnitude="-2" metricUnit="m" metricValue="2.3899999999999997" pageId="6" pageNumber="257" unit="mm" value="23.9">23.9 mm</quantity>
long,
<quantity id="67139B74FFA39767FCC83BFDFC1650CD" box="[819,919,1420,1445]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="6" pageNumber="257" unit="mm" value="17.8">17.8 mm</quantity>
diameter, color not recorded; fruiting perianths tubular; fruit scales channeled vertically; seeds 1 per fruit, basally attached, large, globose, not reniform in longitudinal section, the dorsal surfaces pitted, covered with a non-tanniniferous, slimy sarcotesta; endosperm with numerous, deep, pit-like ruminations; embryos at or near base of seed.
</paragraph>
</subSubSection>
<subSubSection id="E8F1651AFFA39767FF46386DFBC5523D" pageId="6" pageNumber="257" type="distribution">
<paragraph id="A0543691FFA39767FF46386DFE625331" blockId="6.[136,1452,880,1877]" pageId="6" pageNumber="257">
<emphasis id="929FEA83FFA39767FF46386DFE64535D" bold="true" box="[189,485,1564,1589]" pageId="6" pageNumber="257">Distribution and habitat</emphasis>
:—Southern
<collectingCountry id="D8FC7601FFA39767FD8D386DFD59535D" box="[630,728,1564,1589]" name="Vietnam" pageId="6" pageNumber="257">Vietnam</collectingCountry>
, known only from
<collectingRegion id="622FF873FFA39767FC4F386DFB0F535D" box="[948,1166,1564,1589]" country="Vietnam" name="Kon Tum" pageId="6" pageNumber="257">Kon Tum province</collectingRegion>
, in montane rainforest at 1203(10001468) m elevation.
</paragraph>
<paragraph id="A0543691FFA39767FF463815FA1853AD" blockId="6.[136,1452,880,1877]" pageId="6" pageNumber="257">
<emphasis id="929FEA83FFA39767FF463815FE2A5315" bold="true" box="[189,427,1636,1661]" pageId="6" pageNumber="257">Taxonomic notes:—</emphasis>
In
<bibRefCitation id="C47A4B60FFA39767FE333815FC8A5315" author="Peters, C. &amp; Henderson, A." box="[456,779,1636,1661]" pageId="6" pageNumber="257" refId="ref6076" refString="Peters, C. &amp; Henderson, A. (2014) Systematics, ecology, and management of rattans in Cambodia, Laos, and Vietnam: The Biological Bases of Sustainable Use. World Wildlife Fund, Hanoi, 222 pp." type="book" year="2014">Peters and Henderson (2014)</bibRefCitation>
this species would not key. In fact, it is a remarkable species with a unique combination of character states in
<taxonomicName id="67EB4D12FFA39767FD6038F9FD7E53C9" box="[667,767,1672,1697]" class="Liliopsida" family="Arecaceae" genus="Calamus" kingdom="Plantae" order="Arecales" pageId="6" pageNumber="257" phylum="Tracheophyta" rank="genus">
<emphasis id="929FEA83FFA39767FD6038F9FD7E53C9" box="[667,767,1672,1697]" italics="true" pageId="6" pageNumber="257">Calamus</emphasis>
</taxonomicName>
. Pistillate inflorescences are branched to 1 order and seeds are covered with a non-tanniniferous, slimy sarcotesta and have deep, pit-like ruminations. Its relationships are unclear.
</paragraph>
<paragraph id="A0543691FFA39767FF4638A1FBC5523D" blockId="6.[136,1452,880,1877]" pageId="6" pageNumber="257">
<emphasis id="929FEA83FFA39767FF4638A1FDBE5381" bold="true" box="[189,575,1744,1769]" pageId="6" pageNumber="257">Additional specimens examined.</emphasis>
<materialsCitation id="10833CCCFFA39767FDBD38A1FE8D5259" collectingDate="2013-04-24" collectionCode="NY, VFM" collectorName="Henderson &amp; Nguyen Quoc Dung" country="Vietnam" elevation="1140" latitude="14.676" location="Hieu" longLatPrecision="76" longitude="108.397" municipality="Kon Plong district" pageId="6" pageNumber="257" specimenCount="1" stateProvince="Kon Tum">
<collectingCountry id="D8FC7601FFA39767FDBD38A1FD4A5381" box="[582,715,1744,1769]" name="Vietnam" pageId="6" pageNumber="257">VIETNAM</collectingCountry>
.
<collectingRegion id="622FF873FFA39767FD2E38A1FCC55381" box="[725,836,1744,1769]" country="Vietnam" name="Kon Tum" pageId="6" pageNumber="257">Kon Tum</collectingRegion>
:
<collectingMunicipality id="4030ACEBFFA39767FCB538A1FB9E5381" box="[846,1055,1744,1769]" pageId="6" pageNumber="257">Kon Plong district</collectingMunicipality>
,
<location id="A534604AFFA39767FBD138A1FBE15381" LSID="urn:lsid:plazi:treatment:28428787FFA39767FF733CCBFBC5523D:A534604AFFA39767FBD138A1FBE15381" box="[1066,1120,1744,1769]" country="Vietnam" latitude="14.676" longLatPrecision="76" longitude="108.397" municipality="Kon Plong district" name="Hieu" pageId="6" pageNumber="257" stateProvince="Kon Tum">Hieu</location>
commune,
<quantity id="67139B74FFA39767FB1F38A1FA9C5381" box="[1252,1309,1744,1769]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="257" unit="km" value="6.0">6 km</quantity>
north on
<collectingRegion id="622FF873FFA39767FA7238A1FF755265" country="Vietnam" name="Ho Chi Minh City" pageId="6" pageNumber="257">Ho Chi Minh</collectingRegion>
highway from road 24,
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<elevation id="2BC6D1A2FFA39767FDFB3885FDD75265" box="[512,598,1780,1805]" metricMagnitude="3" metricUnit="m" metricValue="1.14" pageId="6" pageNumber="257" unit="m" value="1140.0">1140 m</elevation>
</quantity>
,
<geoCoordinate id="C5DF5056FFA39767FD9A3885FD445265" box="[609,709,1780,1805]" degrees="14.676" direction="north" orientation="latitude" pageId="6" pageNumber="257" precision="55" value="14.676">14.676N</geoCoordinate>
,
<geoCoordinate id="C5DF5056FFA39767FD2B3885FCBE5265" box="[720,831,1780,1805]" degrees="108.397" direction="east" orientation="longitude" pageId="6" pageNumber="257" precision="55" value="108.397">108.397E</geoCoordinate>
,
<date id="D4551051FFA39767FCB23885FC665265" box="[841,999,1780,1805]" pageId="6" pageNumber="257" value="2013-04-24">
<collectingDate id="C411E9B9FFA39767FCB23885FC665265" box="[841,999,1780,1805]" pageId="6" pageNumber="257" value="2013-04-24">24 April 2013</collectingDate>
</date>
,
<emphasis id="929FEA83FFA39767FC093885FA2A5265" box="[1010,1451,1780,1805]" italics="true" pageId="6" pageNumber="257">
<collectorName id="0D1E5347FFA39767FC093885FBEC5265" box="[1010,1133,1780,1805]" pageId="6" pageNumber="257">Henderson</collectorName>
&amp;
<collectorName id="0D1E5347FFA39767FB743885FAEC5265" box="[1167,1389,1780,1805]" pageId="6" pageNumber="257">Nguyen Quoc Dung</collectorName>
3879
</emphasis>
(
<collectionCode id="C6FAAE54FFA39767FF6B3969FF385259" box="[144,185,1816,1841]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="6" pageNumber="257" type="Herbarium">NY</collectionCode>
,
<collectionCode id="C6FAAE54FFA39767FF393969FE855259" box="[194,260,1816,1841]" pageId="6" pageNumber="257">VFM</collectionCode>
)
</materialsCitation>
;
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<collectingCounty id="49354E1DFFA39767FEED3969FE655259" box="[278,484,1816,1841]" pageId="6" pageNumber="257">Kon Plong district</collectingCounty>
,
<collectingMunicipality id="4030ACEBFFA39767FE163969FDEE5259" box="[493,623,1816,1841]" pageId="6" pageNumber="257">Mang Canh</collectingMunicipality>
m.,
<location id="A534604AFFA39767FD663969FCAC5259" LSID="urn:lsid:plazi:treatment:28428787FFA39767FF733CCBFBC5523D:A534604AFFA39767FD663969FCAC5259" box="[669,813,1816,1841]" country="Vietnam" county="Kon Plong district" latitude="14.716666" longLatPrecision="1287" longitude="108.316666" municipality="Mang Canh" name="Thach Nham" pageId="6" pageNumber="257">Thach Nham</location>
protected forest,
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NNE from
<location id="A534604AFFA39767FB4F3969FAEB5259" LSID="urn:lsid:plazi:treatment:28428787FFA39767FF733CCBFBC5523D:A534604AFFA39767FB4F3969FAEB5259" box="[1204,1386,1816,1841]" country="Vietnam" county="Kon Plong district" latitude="14.716666" longLatPrecision="1287" longitude="108.316666" municipality="Mang Canh" name="Mang Den town" pageId="6" pageNumber="257">Mang Den town</location>
,
<quantity id="67139B74FFA39767FA883969FF21523D" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="257" unit="m" value="1000.0">
<elevation id="2BC6D1A2FFA39767FA883969FF21523D" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="257" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
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<geoCoordinate id="C5DF5056FFA39767FEEF394DFE00523D" box="[276,385,1852,1877]" degrees="108" direction="east" minutes="19" orientation="longitude" pageId="6" pageNumber="257" precision="925" value="108.316666">108°19E</geoCoordinate>
,
<date id="D4551051FFA39767FE76394DFDAA523D" box="[397,555,1852,1877]" pageId="6" pageNumber="257" value="2015-04-11">
<collectingDate id="C411E9B9FFA39767FE76394DFDAA523D" box="[397,555,1852,1877]" pageId="6" pageNumber="257" value="2015-04-11">11 April 2015</collectingDate>
</date>
,
<emphasis id="929FEA83FFA39767FDCD394DFC7A523D" box="[566,1019,1852,1877]" italics="true" pageId="6" pageNumber="257">
<collectorName id="0D1E5347FFA39767FDCD394DFD18523D" box="[566,665,1852,1877]" pageId="6" pageNumber="257">Nuraliev</collectorName>
,
<collectorName id="0D1E5347FFA39767FD5F394DFC96523D" box="[676,791,1852,1877]" pageId="6" pageNumber="257">Kuznetsov</collectorName>
&amp;
<collectorName id="0D1E5347FFA39767FCC0394DFC3D523D" box="[827,956,1852,1877]" pageId="6" pageNumber="257">Kuznetsova</collectorName>
1336
</emphasis>
(
<collectionCode id="C6FAAE54FFA39767FBF1394DFBB9523D" box="[1034,1080,1852,1877]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="6" pageNumber="257" type="Herbarium">NY</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
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